152 lines
21 KiB
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152 lines
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<document id="C17DBDA78784D7E3346750464CF4FF37" ID-DOI="10.11646/zootaxa.2457.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5313709" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629674756648" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="2B7E87E4FFFBFFCFF5A6FEA2FB89FA8B" docLanguage="en" docName="zt02457p128.pdf" docOrigin="Zootaxa 2457 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2457.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Marilia mendolonga Oláh & Johanson 2010, new species" docType="treatment" docVersion="3" lastPageNumber="89" masterDocId="D747FF9CFFADFF97F531FFFDFFD9FFE9" masterDocTitle="Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457" masterLastPageNumber="128" masterPageNumber="1" pageNumber="87" updateTime="1699094080944" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="29353CEA8FD0D402C15B36819B102593">Description of 33 new species of Calamoceratidae, Molannidae, Odontoceridae and Philorheithridae (Trichoptera), with detailed presentation of their cephalic setal warts and grooves 2457</mods:title>
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<mods:namePart id="2787F62034055C0969593D6323D712F0">Oláh, János</mods:namePart>
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<mods:name id="4FC154306F614DFC5D904FE18570AA3E" type="personal">
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<mods:namePart id="82FB2981AA28E49E8C64E877F102675C">Johanson, Kjell Arne</mods:namePart>
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<mods:typeOfResource id="FD621CF97B63BD00A871DD7852110A9D">text</mods:typeOfResource>
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<mods:title id="F30B9111D673C92686305AB77D30591A">Zootaxa</mods:title>
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<mods:part id="5743172297F635A4BDBDA0ABB81F5CC3">
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<mods:date id="A7014E674ED2FA67A96C00C3C31CA387">2010</mods:date>
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<mods:detail id="76D857D6908ED5A095EE5CDC3D704CC8" type="pubDate">
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<mods:number id="541A7F974A98397CF9DCED4338CE56B0">2010-05-14</mods:number>
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<mods:number id="C89F63457CB0A94997F15A2F8B8C355A">2457</mods:number>
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<mods:number id="94392EA048994CC2815C1B83A08D34D9">1</mods:number>
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<mods:url id="D1E61D050970E7F2B84C169DAB2BB2ED">https://biotaxa.org/Zootaxa/article/view/zootaxa.2457.1.1</mods:url>
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<mods:classification id="1E83B56E80444CDE041E1FB893052B46">journal article</mods:classification>
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<mods:identifier id="0BE9CBD1B84FD854E4DC8A3A2D912DAD" type="DOI">10.11646/zootaxa.2457.1.1</mods:identifier>
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<mods:identifier id="0A2AA95F1FE1E4F68D4EFBF80E6F9F71" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="000A0445ED94B52E0527C396BD89263C" type="Zenodo-Dep">5313709</mods:identifier>
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<treatment id="2B7E87E4FFFBFFCFF5A6FEA2FB89FA8B" ID-DOI="http://doi.org/10.5281/zenodo.5321918" ID-Zenodo-Dep="5321918" LSID="urn:lsid:plazi:treatment:2B7E87E4FFFBFFCFF5A6FEA2FB89FA8B" httpUri="http://treatment.plazi.org/id/2B7E87E4FFFBFFCFF5A6FEA2FB89FA8B" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87">
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<subSubSection id="EBCD6579FFFBFFC1F5A6FEA2FDF4FE90" box="[151,557,351,377]" pageId="86" pageNumber="87" type="nomenclature">
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<paragraph id="A36836F2FFFBFFC1F5A6FEA2FDF4FE90" blockId="86.[151,557,351,412]" box="[151,557,351,377]" pageId="86" pageNumber="87">
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<heading id="F820819EFFFBFFC1F5A6FEA2FDF4FE90" bold="true" box="[151,557,351,377]" fontSize="11" level="1" pageId="86" pageNumber="87" reason="1">
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<emphasis id="91A3EAE0FFFBFFC1F5A6FEA2FDF4FE90" bold="true" box="[151,557,351,377]" pageId="86" pageNumber="87">
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<emphasis id="91A3EAE0FFFBFFC1F5A6FEA2FE4FFE90" bold="true" box="[151,406,351,377]" italics="true" pageId="86" pageNumber="87">
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<taxonomicName id="64D74D71FFFBFFC1F5A6FEA2FE49FE90" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,400,351,377]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mendolonga" status="sp. nov.">Marilia mendolonga</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="8A90579BFFFBFFC1F4AFFEA2FDF4FE90" box="[414,557,351,377]" pageId="86" pageNumber="87" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBCD6579FFFBFFCFF5A6FE7FFDDEFB2B" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87" type="description">
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<paragraph id="A36836F2FFFBFFC1F5A6FE7FFEEEFE75" blockId="86.[151,557,351,412]" box="[151,311,386,412]" pageId="86" pageNumber="87">
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<figureCitation id="3BEC2A77FFFBFFC1F5A6FE7FFEEEFE75" box="[151,311,386,412]" captionStart="FIGURES 197–203" captionStartId="87.[151,269,1968,1992]" captionTargetBox="[156,1433,196,1937]" captionTargetId="figure-54@87.[151,1436,194,1944]" captionTargetPageId="87" captionText="FIGURES 197–203. Marilia mendolonga, new species, holotype. 197 —head, frontal; 198 —head, dorsal; 199 — genitalia, lateral; 200 — genitalia, dorsal; 201 — gonocoxite, ventral; 202 — phallus, lateral; 203 — phallus, ventral." figureDoi="http://doi.org/10.5281/zenodo.5313817" httpUri="https://zenodo.org/record/5313817/files/figure.png" pageId="86" pageNumber="87">Figs 197–203</figureCitation>
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</paragraph>
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<paragraph id="A36836F2FFFBFFC1F5A6FE2FFB1DFD65" blockId="86.[151,1437,466,2012]" pageId="86" pageNumber="87">
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Having a strongly constricted, narrow vertex, this species strongly resembles
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<taxonomicName id="64D74D71FFFBFFC1F166FE2FFB37FE02" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[1111,1262,466,491]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="katakaha" status="sp. nov.">
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<emphasis id="91A3EAE0FFFBFFC1F166FE2FFB37FE02" box="[1111,1262,466,491]" italics="true" pageId="86" pageNumber="87">M. katakaha</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="8A90579BFFFBFFC1F032FE2FFA41FE05" box="[1283,1432,466,492]" pageId="86" pageNumber="87" rank="species">new species</taxonomicNameLabel>
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.
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<taxonomicName id="64D74D71FFFBFFC1F5A6FE07FE50FDFA" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,393,506,531]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="mendolonga">
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<emphasis id="91A3EAE0FFFBFFC1F5A6FE07FE50FDFA" box="[151,393,506,531]" italics="true" pageId="86" pageNumber="87">Marilia mendolonga</emphasis>
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</taxonomicName>
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is separated from
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<taxonomicName id="64D74D71FFFBFFC1F753FE07FD2CFDFA" authorityName="Oláh & Johanson" authorityYear="2010" box="[610,757,506,531]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="katakaha">
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<emphasis id="91A3EAE0FFFBFFC1F753FE07FD2CFDFA" box="[610,757,506,531]" italics="true" pageId="86" pageNumber="87">M. katakaha</emphasis>
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</taxonomicName>
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in being smaller, and the vertexal lateroantennal compact setal warts are smaller and more elongate. The ridge pattern on segment X of
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<taxonomicName id="64D74D71FFFBFFC1F11DFDDFFB67FDD2" authorityName="Oláh & Johanson" authorityYear="2010" box="[1068,1214,546,571]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="86" pageNumber="87" phylum="Arthropoda" rank="species" species="katakaha">
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<emphasis id="91A3EAE0FFFBFFC1F11DFDDFFB67FDD2" box="[1068,1214,546,571]" italics="true" pageId="86" pageNumber="87">M. katakaha</emphasis>
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</taxonomicName>
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is weaker, like the almost indiscernible lateral longitudinal ridges. The apicolateral corners of tergum IX are short, narrow, and obliquely set, not forming transverse plates over the articulation of the preanal appendages.
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</paragraph>
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<paragraph id="A36836F2FFFBFFCFF5F6FD67FB32FE1B" blockId="86.[151,1437,466,2012]" lastBlockId="88.[151,1436,152,1378]" lastPageId="88" lastPageNumber="89" pageId="86" pageNumber="87">
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Male (in alcohol). Body medium-sized; dark brown; legs, antennae and palps light brown; wings uniformly light brown, without pattern. Eyes black, extremely large; vertex constricted to narrow strip; interocular distance about 0.1 times the width of each eye, almost 12 times longer than broad. Ocelli absent. Tentorium slender, with short, vestigial dorsal arm; posterior arms robust, ending in pair of large posterior tentorial pits; tentorial bridge slender, without protuberances; posterior half of anterior tentorial arms thin, anterior half more robust, with strongly developed median lamellate lobes; anterior tentorial arms with lateral widening on anterior half. Facial groove pattern modified, shortened frontal, laterally fragmented setose wart and large antennal sockets dominating on frons. Anterior remnant of frontal grooves branching from coronal groove invisible at ventral margin of antennal sockets. Frontogenal vertical grooves invisible between frontal lateral fragmented setose warts and vertically elongate, narrow, frontogenal compact setose warts. Clypeogenal vertical grooves located ventrally of anterior tentorial pits; long, running obliquely laterad, reaching subgenal process. Subantennal grooves running vertically in narrow stripe between ocellar groove and frontogenal compact setose wart. Subocular groove indiscernible, merging to anterior clypeogenal grooves. Vertex forming narrow strip; vertexal groove pattern simplified; coronal groove at posterior cleft invisible. Postoccipital groove encircling
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<emphasis id="91A3EAE0FFFBFFC1F720FB36FD38FB0D" box="[529,737,1227,1252]" italics="true" pageId="86" pageNumber="87">foramen magnum</emphasis>
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. Labrum pyriform, with short setae on freely hanging and movable anterior apical area. Mandibles almost indiscernible; lacinia broad. Pair of short, fragmented frontal lateral setal warts present on face beside frontogenal compact setal warts. Frontogenal compact setal warts running from antennal sockets along subantennal groove of pregenae. Narrow vertex with pair of elongate, posteriorly narrowing, fragmenting vertexal lateroantennal compact setal warts, having almost indiscernible posterior ending. Vertexal mediantennal compact setose warts separate. Occipital compact setose warts small, broad, elongated. Pair of
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<emphasis id="91A3EAE0FFFBFFC1F48FFA46FD72FA3D" box="[446,683,1467,1492]" italics="true" pageId="86" pageNumber="87">sensilla basiconicae</emphasis>
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present along end of coronial groove, tangential with anteromesal corner of occipital warts. Vertically elongate postgenal compact warts curving along posterior section of ocular grooves, narrow strip pressed to ocular groove. Postgenal surface glabrous, postgenal surface hidden by eyes. Maxillary palps paler than legs and antennae; maxillary palp formula V-IV-III-II-I; each basal maxillary palp segments significantly longer than each second maxillary palp segments, without setose subapicomesal nodule. Each scape slightly shorter than head. Each pedicel shorter than first segment of each antennal flagella. Single pair pronotal warts transversely elongated, located on posterior 1/3rd of segment, almost tangential mesally, narrowest laterally. Pair of diffuse mesoscutal setose warts present, represented by few pale alveoli on dark mesoscutal surface. Almost entire surface of mesoscutellar area covered by diffuse setal warts separated by median notal groove, represented by few setal alveoli. Each proepisternum with small-sized, ovoid, setose warts, much smaller than setose warts on precoxale. Large, compact, setal wart present anteriorly on cervical sclerites; apparently forming sclerotized surface on membranous part of neck, touching only anterior cervical sclerite. Lateral cervical sclerites forming narrow anterior arm articulating anteriorly to back of head, with occipital condyle above posterior tentorial pits, fused to posterior cervical sclerites. Posterior cervical sclerites forming posterad broadening plates reaching prothoracic episternum, articulating to weakly sclerotized anteromedian band of prothoracic eusternum by ventral intercervical sclerites; dark pattern of cervical sclerite complex, clearly visible on pale membranous neck. Legs claws symmetrical; spur formula 2, 4, 4; foreleg spurs almost equally long; each midleg posterior subapical spur half as long as foreleg spurs; posteroapical spur 1/4th as long as foreleg spurs; each hind leg posterior spurs half as long as anterior spurs. Forewings: length 8.0 mm; membrane brown, without visible pattern; termen straight; R1 confluent with R2 shortly before C; base of discoidal cell located proximally of mid-length of wing; fork I longer than fork II; forks III–V absent; fork I sessile, fork II short petiolate, almost sessile. Hind wings: R1 present, long, running to R2 near distal part of C; fork I slightly longer than fork II; long brush present.
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</paragraph>
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<caption id="F7A8667AFFFAFFC0F5A6F84DFA97F803" ID-DOI="http://doi.org/10.5281/zenodo.5313817" ID-Zenodo-Dep="5313817" httpUri="https://zenodo.org/record/5313817/files/figure.png" pageId="87" pageNumber="88" startId="87.[151,269,1968,1992]" targetBox="[156,1433,196,1937]" targetPageId="87">
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<paragraph id="A36836F2FFFAFFC0F5A6F84DFA97F803" blockId="87.[151,1436,1968,2026]" pageId="87" pageNumber="88">
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<emphasis id="91A3EAE0FFFAFFC0F5A6F84DFEA1F821" bold="true" box="[151,376,1968,1992]" pageId="87" pageNumber="88">FIGURES 197–203.</emphasis>
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<taxonomicName id="64D74D71FFFAFFC0F44FF84CFD83F821" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[382,602,1969,1992]" class="Insecta" family="Odontoceridae" genus="Marilia" kingdom="Animalia" order="Trichoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="mendolonga" status="sp. nov.">
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<emphasis id="91A3EAE0FFFAFFC0F44FF84CFD83F821" box="[382,602,1969,1992]" italics="true" pageId="87" pageNumber="88">Marilia mendolonga</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="8A90579BFFFAFFC0F759F84CFD32F821" box="[616,747,1969,1992]" pageId="87" pageNumber="88" rank="species">new species</taxonomicNameLabel>
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, holotype. 197 —head, frontal; 198 —head, dorsal; 199 — genitalia, lateral; 200 — genitalia, dorsal; 201 — gonocoxite, ventral; 202 — phallus, lateral; 203 — phallus, ventral.
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</paragraph>
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</caption>
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<paragraph id="A36836F2FFF5FFCFF5F6FDFDFDDEFB2B" blockId="88.[151,1436,152,1378]" pageId="88" pageNumber="89">Male genitalia. Abdominal segment IX fused annularly, ventral longitudinal grooves present; largest groove separating ventral and dorsal halfs in lateral view, running to end of apical lobes; tergum IX slightly longer than venter; anterior margins straight, vertical, slightly convex; apical lobes rounded, with deep excision near preanal appendages. Apicolateral corners of tergum IX forming short, narrow, oblique plates over articulation of preanal appendages. Antecosta on anterior margins of segment IX forming dark, marginal rims connected to broad longitudinal ventral grooves. Ventral longitudinal grooves turning ventrad, forming posterior grooves on ventral half of segment IX; spine rows absent on posterior margins of segment IX. Apicopleural and apicoventral setose areas absent. Intersegmental depression between segment IX and segment X sloping gently. Segment X weakly pigmented, forming quadrangular plate with slightly excised apex; longer than gonocoxites in lateral view; in dorsal view central ridge almost indiscernible on basal half, well developed on apical half; separating lateral concavities. Apicoventral setose lobes represented by few alveoli on lateral lobe before apex. Apicodorsal setose area not discernible. Apex of segment X filled; dorsal interlobular gap absent. Preanal appendages long, filiform in lateral and dorsal view; almost parallel-sided in dorsal view. Gonocoxites shorter than segment X; coxopodites curving, parallel-sided in lateral and ventral view; harpagones small, not clavate or narrowing apicad. Phallic apparatus with phallotheca being sclerotized at basal half and membranous at apical half. Phallicata difficult to separate. Sclerotized phallotheca curving basad, slightly constricted before apex. Small, elongate, poorly visible phallotremal sclerite present at phallicata base, U-shaped in ventral view.</paragraph>
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</subSubSection>
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<subSubSection id="EBCD6579FFF5FFCFF5F6FB32FE5BFAFB" pageId="88" pageNumber="89" type="materials_examined">
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<paragraph id="A36836F2FFF5FFCFF5F6FB32FE5BFAFB" blockId="88.[151,1436,152,1378]" pageId="88" pageNumber="89">
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<materialsCitation id="13BF3CAFFFF5FFCFF5F6FB32FE5BFAFB" collectingDate="1988-04-28" collectionCode="NRM" country="Malaysia" location="Mendolong Nursery" municipality="Sipitang" pageId="88" pageNumber="89" specimenCount="1" stateProvince="Borneo" typeStatus="holotype">
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<emphasis id="91A3EAE0FFF5FFCFF5F6FB32FCDCFB00" bold="true" box="[199,773,1231,1257]" pageId="88" pageNumber="89">
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<typeStatus id="7C6C8850FFF5FFCFF5F6FB32FEE0FB00" box="[199,313,1231,1257]" pageId="88" pageNumber="89" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="DBC07662FFF5FFCFF4A1FB32FDEBFB00" box="[400,562,1231,1257]" name="Malaysia" pageId="88" pageNumber="89">MALAYSIA</collectingCountry>
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:
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<collectingRegion id="6113F810FFF5FFCFF70EFB32FD47FB00" box="[575,670,1231,1257]" country="Malaysia" name="Borneo" pageId="88" pageNumber="89">Borneo</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="6113F810FFF5FFCFF79CFB32FD26FB00" box="[685,767,1231,1257]" country="Malaysia" name="Sabah" pageId="88" pageNumber="89">Sabah</collectingRegion>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
<collectingMunicipality id="430CAC88FFF5FFCFF63FFB2DFCABFB03" box="[782,882,1232,1258]" pageId="88" pageNumber="89">Sipitang</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="A6086029FFF5FFCFF64EFB2DFBB7FB03" LSID="urn:lsid:plazi:treatment:2B7E87E4FFFBFFCFF5A6FEA2FB89FA8B:A6086029FFF5FFCFF64EFB2DFBB7FB03" box="[895,1134,1232,1258]" country="Malaysia" municipality="Sipitang" name="Mendolong Nursery" pageId="88" pageNumber="89" stateProvince="Borneo">Mendolong Nursery</location>
|
|||
|
|
,
|
|||
|
|
<date id="D7691032FFF5FFCFF14DFB2DFB23FB03" box="[1148,1274,1232,1258]" pageId="88" pageNumber="89" value="1988-04-28">
|
|||
|
|
<collectingDate id="C72DE9DAFFF5FFCFF14DFB2DFB23FB03" box="[1148,1274,1232,1258]" pageId="88" pageNumber="89" value="1988-04-28">28.iv.1988</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
, at light [Stig Adebratt] – (
|
|||
|
|
<collectionCode id="C5C6AE37FFF5FFCFF41CFB05FEAFFAFB" box="[301,374,1272,1298]" country="Sweden" name="Swedish Museum of Natural History - Zoological Collections" pageId="88" pageNumber="89" type="Museum">NRM</collectionCode>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="EBCD6579FFF5FFCFF5F6FAE2FD9FFAD3" box="[199,582,1311,1338]" pageId="88" pageNumber="89" type="distribution">
|
|||
|
|
<paragraph id="A36836F2FFF5FFCFF5F6FAE2FD9FFAD3" blockId="88.[151,1436,152,1378]" box="[199,582,1311,1338]" pageId="88" pageNumber="89">
|
|||
|
|
<emphasis id="91A3EAE0FFF5FFCFF5F6FAE2FEB0FAD0" bold="true" box="[199,361,1311,1337]" pageId="88" pageNumber="89">Distribution:</emphasis>
|
|||
|
|
<collectingCountry id="DBC07662FFF5FFCFF440FADDFE05FAD3" box="[369,476,1312,1338]" name="Malaysia" pageId="88" pageNumber="89">Malaysia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<collectingRegion id="6113F810FFF5FFCFF4DDFADDFDE3FAD3" box="[492,570,1312,1338]" country="Malaysia" name="Sabah" pageId="88" pageNumber="89">Sabah</collectingRegion>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="EBCD6579FFF5FFCFF5F6FABAFB89FA8B" box="[199,1104,1351,1378]" pageId="88" pageNumber="89" type="etymology">
|
|||
|
|
<paragraph id="A36836F2FFF5FFCFF5F6FABAFB89FA8B" blockId="88.[151,1436,152,1378]" box="[199,1104,1351,1378]" pageId="88" pageNumber="89">
|
|||
|
|
<emphasis id="91A3EAE0FFF5FFCFF5F6FABAFE8EFA88" bold="true" box="[199,343,1351,1377]" pageId="88" pageNumber="89">Etymology:</emphasis>
|
|||
|
|
<emphasis id="91A3EAE0FFF5FFCFF46EFAB5FE2AFA88" box="[351,499,1352,1377]" italics="true" pageId="88" pageNumber="89">Mendolonga</emphasis>
|
|||
|
|
, named after the
|
|||
|
|
<typeStatus id="7C6C8850FFF5FFCFF78EFAB5FD28FA8B" box="[703,753,1352,1378]" pageId="88" pageNumber="89">type</typeStatus>
|
|||
|
|
locality, Mendolong Nursery.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|