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<document id="7008390EAF788C2C916A5547ABC94D0A" ID-DOI="10.5281/zenodo.186351" ID-GBIF-Dataset="bb17e039-5346-4e01-a9ba-59b87e9e6a4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="186351" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050352768" checkinUser="plazi" docAuthor="Maus, Mariana Chani-Posse De" docDate="2009" docId="03D787C9FFAFFF9FFF1BCF17CE9AE62B" docLanguage="en" docName="zt02034p042.pdf" docOrigin="Zootaxa 2034" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gabrius nidicola Bernhauer 1921, comb. nov." docType="treatment" docVersion="8" lastPageNumber="34" masterDocId="FFEEFFB1FFACFF9CFF8CCD70CA68E133" masterDocTitle="Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes" masterLastPageNumber="42" masterPageNumber="31" pageNumber="34" updateTime="1698578662429" updateUser="plazi">
<mods:mods id="2A5BF2967CB088E608C5AB953F2EF7E7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4437257D34FF4182B87B5A8D8B04E5DC">Revision of some types of Philonthus Stephens and Gabrius Stephens from southern South America (Coleoptera: Staphylinidae), with nomenclatural notes</mods:title>
</mods:titleInfo>
<mods:name id="C3EF29F15A6792D6CBF4C06859B9BEFD" type="personal">
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<mods:namePart id="A09BE38E294A0FA5EFDF042BB55D4D6D">Maus, Mariana Chani-Posse De</mods:namePart>
</mods:name>
<mods:typeOfResource id="B4A879FF15226A7DCBDCD8D5ABADAFEC">text</mods:typeOfResource>
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<mods:title id="E488D1575B76CAD730E1EB2111C383CA">Zootaxa</mods:title>
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<mods:date id="33D3AAA29637B7944EF315305BC796A3">2009</mods:date>
<mods:detail id="9F1962FEBBEACA7E0C96EA0799C0C257" type="volume">
<mods:number id="9320ECA87D087F152C91FD7168AC410C">2034</mods:number>
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<mods:classification id="FD8A2E9A6413BECB920B1AFCA298C5DB">journal article</mods:classification>
<mods:identifier id="728C68505FC43C46B1B311B4C1C8CC14" type="DOI">10.5281/zenodo.186351</mods:identifier>
<mods:identifier id="C8F8031816B461EEF04CF6D9DBD0B0F8" type="GBIF-Dataset">bb17e039-5346-4e01-a9ba-59b87e9e6a4c</mods:identifier>
<mods:identifier id="C8E91C57780D610F1B52D3BDA9955D32" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="32D9EE3496316B45D2FFC049C8A678FF" type="Zenodo-Dep">186351</mods:identifier>
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<treatment id="03D787C9FFAFFF9FFF1BCF17CE9AE62B" ID-DOI="http://doi.org/10.5281/zenodo.6216656" ID-GBIF-Taxon="119611789" ID-Zenodo-Dep="6216656" LSID="urn:lsid:plazi:treatment:03D787C9FFAFFF9FFF1BCF17CE9AE62B" httpUri="http://treatment.plazi.org/id/03D787C9FFAFFF9FFF1BCF17CE9AE62B" lastPageNumber="34" pageId="3" pageNumber="34">
<subSubSection id="C3646554FFAFFF9FFF1BCF17CE7EE3FB" pageId="3" pageNumber="34" type="nomenclature">
<paragraph id="8BC136DFFFAFFF9FFF1BCF17C8B7E3B2" blockId="3.[151,735,615,641]" box="[151,735,615,641]" pageId="3" pageNumber="34">
<heading id="D08981B3FFAFFF9FFF1BCF17C8B7E3B2" bold="true" box="[151,735,615,641]" fontSize="11" level="1" pageId="3" pageNumber="34" reason="1">
<emphasis id="B90AEACDFFAFFF9FFF1BCF17C8B7E3B2" bold="true" box="[151,735,615,641]" pageId="3" pageNumber="34">
<taxonomicName id="4C7E4D5CFFAFFF9FFF1BCF17C823E3B2" ID-CoL="6JV6B" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" box="[151,587,615,641]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola" status="comb. nov.">
<emphasis id="B90AEACDFFAFFF9FFF1BCF17CB0AE3B2" bold="true" box="[151,354,615,641]" italics="true" pageId="3" pageNumber="34">Gabrius nidicola</emphasis>
(
<bibRefCitation id="EFEF4B2EFFAFFF9FFEFCCF17C82AE3B2" author="Bernhauer" box="[368,578,615,641]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A23957B6FFAFFF9FFDD5CF17C8B7E3B2" box="[601,735,615,641]" pageId="3" pageNumber="34">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC136DFFFAFFF9FFF1BCFC1CE7EE3FB" blockId="3.[151,1436,689,776]" box="[151,1046,689,712]" pageId="3" pageNumber="34">
<treatmentCitationGroup id="AB6E11F1FFAFFF9FFF1BCFC1CE7EE3FB" box="[151,1046,689,712]" pageId="3" pageNumber="34">
<taxonomicName id="4C7E4D5CFFAFFF9FFF1BCFC1CB81E3FB" ID-CoL="4GBC9" authority="Bernhauer" authorityName="Bernhauer" box="[151,489,689,712]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFF1BCFC1CB00E3FB" box="[151,360,689,712]" italics="true" pageId="3" pageNumber="34">Philonthus nidicola</emphasis>
&quot;Bernhauer
</taxonomicName>
in litt.&quot;;
<treatmentCitation id="0ADF10CEFFAFFF9FFDC4CFC1C96AE3FB" author="Bruch" box="[584,770,689,712]" page="445" pageId="3" pageNumber="34" year="1915">
<bibRefCitation id="EFEF4B2EFFAFFF9FFDC4CFC1C8B8E3FB" author="Bruch" box="[584,720,689,712]" pageId="3" pageNumber="41" refString="Bruch, C. (1915) Catalogo Sistematico de los coleopteros de la Republica Argentina. Pars VI. Revista del Museo de La Plata, 19 (Segunda Serie) (6), 235 - 302." type="journal article" year="1915">Bruch, 1915</bibRefCitation>
: 445
</treatmentCitation>
(catalog).
<emphasis id="B90AEACDFFAFFF9FFCF9CFC1CE67E3FB" box="[885,1039,689,712]" italics="true" pageId="3" pageNumber="34">Nomen nudum</emphasis>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAFFF9FFF1BCFA1C899E23B" pageId="3" pageNumber="34" type="reference_group">
<paragraph id="8BC136DFFFAFFF9FFF1BCFA1C899E23B" blockId="3.[151,1436,689,776]" pageId="3" pageNumber="34">
<treatmentCitationGroup id="AB6E11F1FFAFFF9FFF1BCFA1C899E23B" pageId="3" pageNumber="34">
<taxonomicName id="4C7E4D5CFFAFFF9FFF1BCFA1C80BE3DB" ID-CoL="4GBC9" authority="Bernhauer, 1921: 108" authorityName="Bernhauer" authorityPageNumber="108" authorityYear="1921" box="[151,611,721,744]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFF1BCFA1CB02E3DB" box="[151,362,721,744]" italics="true" pageId="3" pageNumber="34">Philonthus nidicola</emphasis>
<treatmentCitation id="0ADF10CEFFAFFF9FFEF8CFA1C80BE3DB" author="Bernhauer" box="[372,611,721,744]" page="108" pageId="3" pageNumber="34" year="1921">
<bibRefCitation id="EFEF4B2EFFAFFF9FFEF8CFA1C843E3DB" author="Bernhauer" box="[372,555,721,744]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
: 108
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0ADF10CEFFAFFF9FFDFCCFA1C91FE3DB" author="Scheerpeltz" box="[624,887,721,744]" page="1352" pageId="3" pageNumber="34" year="1933">
<bibRefCitation id="EFEF4B2EFFAFFF9FFDFCCFA1C95DE3DB" author="Scheerpeltz" box="[624,821,721,744]" pageId="3" pageNumber="42" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 129. W. Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1352
</treatmentCitation>
(catalog);
<treatmentCitation id="0ADF10CEFFAFFF9FFC7CCFA1CE9BE3DB" author="Blackwelder" box="[1008,1267,721,744]" page="134" pageId="3" pageNumber="34" year="1944">
<bibRefCitation id="EFEF4B2EFFAFFF9FFC7CCFA1CED7E3DB" author="Blackwelder" box="[1008,1215,721,744]" pageId="3" pageNumber="41" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous insects of Mexico, Central America, the West Indies, and South America. Part I. United States National Museum Bulletin, 185, 1 - 188." type="journal article" year="1944">Blackwelder, 1944</bibRefCitation>
: 134
</treatmentCitation>
(list);
<treatmentCitation id="0ADF10CEFFAFFF9FFAB3CFA1CB57E23B" author="Herman" page="2889" pageId="3" pageNumber="34" year="2001">
<bibRefCitation id="EFEF4B2EFFAFFF9FFAB3CFA1CB68E23B" author="Herman" pageId="3" pageNumber="42" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. Part V. Bulletin of the American Museum of Natural History, 265, 2441 - 3020." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 2889
</treatmentCitation>
(catalog); Chani-Posse, 2004: 231 (list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAFFF9FFF1BCE47CBFAE5C8" pageId="3" pageNumber="34" type="materials_examined">
<paragraph id="8BC136DFFFAFFF9FFF1BCE47CBA9E509" blockId="3.[151,1437,823,1816]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF1BCE47CBB5E262" bold="true" box="[151,477,823,849]" pageId="3" pageNumber="34">
<typeStatus id="54C5887DFFAFFF9FFF1BCE47CABEE262" box="[151,214,823,849]" pageId="3" pageNumber="34">Type</typeStatus>
material.
<typeStatus id="54C5887DFFAFFF9FFED1CE47CBB5E262" box="[349,477,823,849]" pageId="3" pageNumber="34" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated;
<figureCitation id="13452A5AFFAFFF9FFD4CCE48C963E261" box="[704,779,824,850]" captionStart="FIGURES 1 – 12" captionStartId="6.[151,275,1634,1658]" captionText="FIGURES 1 – 12. Type specimens: habitus and labels; labels with lectotype designation are not included. (1 a – b) Philonthus argentinus Bernhauer (now in Gabrius), (lectotype with labels *). (2 a – b) Philonthus jujuyensis Bernhauer (now in Gabrius), (lectotype with labels *). (3 a – b) Philonthus tucumanensis Bernhauer (now in Gabrius), (lectotype with labels *). (4) Philonthus nidicola Bernhauer (now in Gabrius), (only labels are shown *). (5 a – b) Philonthus quadraticeps (lectotype with labels *). (6 a – b) P. tenebrosus (= P. quadraticeps), (lectotype with labels *). (7 a – b) Philonthus bonariensis (lectotype with labels *). (8 a – b) Philonthus weiseri (= P. bonariensis), (lectotype with labels *). (9 a – b) Philonthus catamarcanus (= P. bonariensis), (lectotype with labels *). (10 a – b) Philonthus cribriventris (lectotype with labels *). (11) Philonthus catamarcanus var. densior (= P. cribriventris), (lectotype, labels are not shown). (12) G. chiliensis (= G. nigritulus), (paratype, labels are not shown). * Labels with lectotype designation are not included (pictures were taken prior to any designation)." pageId="3" pageNumber="34">Fig. 4</figureCitation>
), 3, with labels: “Rep.
<collectingCountry id="F369764FFFAFFF9FFBBCCE48CEB7E261" box="[1072,1247,824,850]" name="Argentina" pageId="3" pageNumber="34">ARGENTINA</collectingCountry>
/ Prov. Buenos Aires /
<date id="FFC0101FFFAFFF9FFF7DCE2ECB18E24B" box="[241,368,862,888]" pageId="3" pageNumber="34" value="1919-10-17">17.X.1919</date>
/ C. Bruch”, “Monte Veloz /
<date id="FFC0101FFFAFFF9FFD59CE2EC93CE24B" box="[725,852,862,888]" pageId="3" pageNumber="34" value="1919-10-17">17.X.1919</date>
/ ex nido
<taxonomicName id="4C7E4D5CFFAFFF9FFC5CCE2FCE2EE24B" box="[976,1094,863,888]" class="Mammalia" family="Ctenomyidae" genus="Ctenomys" kingdom="Animalia" order="Rodentia" pageId="3" pageNumber="34" phylum="Chordata" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFC5CCE2FCE2EE24B" box="[976,1094,863,888]" italics="true" pageId="3" pageNumber="34">Ctenomys</emphasis>
</taxonomicName>
”, “
<taxonomicName id="4C7E4D5CFFAFFF9FFBFDCE2FCF45E24B" authority="Bernh." authorityName="Bernh." box="[1137,1325,862,888]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFBFDCE2FCEBCE24B" box="[1137,1236,863,888]" italics="true" pageId="3" pageNumber="34">nidicola</emphasis>
Bernh.
</taxonomicName>
/
<typeStatus id="54C5887DFFAFFF9FFACACE2ECFFAE24B" box="[1350,1426,862,888]" pageId="3" pageNumber="34">Typus</typeStatus>
” (yellow label), “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFAFFF9FFC13CEF5CE72E2AC" box="[927,1050,901,927]" pageId="3" pageNumber="34" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFAFFF9FFBAFCEF5CABCE2F5" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFBAFCEF5CF79E2AD" box="[1059,1297,901,926]" italics="true" pageId="3" pageNumber="34">Philonthus nidicola</emphasis>
<bibRefCitation id="EFEF4B2EFFAFFF9FFA95CEF5CABCE2F5" author="Bernhauer" pageId="3" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFAFFF9FFD14CEDCC89EE2F5" LSID="urn:lsid:biocol.org:col:34795" box="[664,758,940,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="34">FMNH</collectionCode>
).
<emphasis id="B90AEACDFFAFFF9FFC87CEDBC9AFE2F6" bold="true" box="[779,967,939,965]" pageId="3" pageNumber="34">
<typeStatus id="54C5887DFFAFFF9FFC87CEDBC9AAE2F6" box="[779,962,939,965]" pageId="3" pageNumber="34" type="paralectotype">Paralectotype</typeStatus>
:
</emphasis>
Ƥ, with labels: “Rep.
<collectingCountry id="F369764FFFAFFF9FFB50CEDCCFE3E2F5" box="[1244,1419,940,966]" name="Argentina" pageId="3" pageNumber="34">ARGENTINA</collectingCountry>
/ Prov. Buenos Aires /
<date id="FFC0101FFFAFFF9FFE16CEA2C871E2DF" box="[410,537,978,1004]" pageId="3" pageNumber="34" value="1919-10-17">17.X.1919</date>
/ C. Bruch”, “189”, “
<taxonomicName id="4C7E4D5CFFAFFF9FFCAECEA2C9B7E2DF" authority="Bernh." authorityName="Bernh." box="[802,991,978,1004]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFCAECEA2C9EDE2D8" box="[802,901,978,1003]" italics="true" pageId="3" pageNumber="34">nidicola</emphasis>
Bernh.
</taxonomicName>
/ Cotypus” (yellow label), “Chicago NHMus. / M. Bernhauer Collection”, “
<typeStatus id="54C5887DFFAFFF9FFDD1CE89C968E520" box="[605,768,1017,1043]" pageId="3" pageNumber="34" type="paralectotype">Paralectotype</typeStatus>
<taxonomicName id="4C7E4D5CFFAFFF9FFC8BCE89CEDFE520" authority="Bernhauer, 1921" authorityName="Bernhauer" authorityYear="1921" box="[775,1207,1017,1043]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFC8BCE89C985E521" box="[775,1005,1017,1042]" italics="true" pageId="3" pageNumber="34">Philonthus nidicola</emphasis>
<bibRefCitation id="EFEF4B2EFFAFFF9FFC78CE89CEDFE520" author="Bernhauer" box="[1012,1207,1017,1043]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer, 1921</bibRefCitation>
</taxonomicName>
/ Des. Chani-Posse de Maus 2008” (
<collectionCode id="ED6FAE1AFFAFFF9FFED5C950CBDDE509" LSID="urn:lsid:biocol.org:col:34795" box="[345,437,1056,1082]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="3" pageNumber="34">FMNH</collectionCode>
).
</paragraph>
<paragraph id="8BC136DFFFAFFF9FFF4AC936CBFAE5C8" blockId="3.[151,1437,823,1816]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF4AC936CB7CE553" bold="true" box="[198,276,1094,1120]" pageId="3" pageNumber="34">Notes.</emphasis>
Although references about the
<typeStatus id="54C5887DFFAFFF9FFD25C936C8B4E553" box="[681,732,1094,1120]" pageId="3" pageNumber="34">type</typeStatus>
locality and a male character are given in the original description, there is no indication about how many specimens were examined. Of them, the male individual is the
<typeStatus id="54C5887DFFAFFF9FFF4DC9E4CB45E59D" box="[193,301,1172,1198]" pageId="3" pageNumber="34" type="lectotype">lectotype</typeStatus>
hereby designated because Bernhauer explicitly referred to the locality (Monte Veloz) where this specimen was collected in the original description. The two specimens here studied agree with the description by
<bibRefCitation id="EFEF4B2EFFAFFF9FFF30C991CBE6E5C8" author="Bernhauer" box="[188,398,1249,1275]" pageId="3" pageNumber="41" refString="Bernhauer, M. (1921) Neue Staphyliniden aus Sudamerika, besonders aus Argentinien. Wiener Entomologische Zeitung, 38, 101 - 108, 169 - 179." type="journal article" year="1921">Bernhauer (1921)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646554FFAFFF9FFF4AC877CE9AE62B" pageId="3" pageNumber="34" type="discussion">
<paragraph id="8BC136DFFFAFFF9FFF4AC877CE9AE62B" blockId="3.[151,1437,823,1816]" pageId="3" pageNumber="34">
<emphasis id="B90AEACDFFAFFF9FFF4AC877CB3AE412" bold="true" box="[198,338,1287,1313]" pageId="3" pageNumber="34">Discussion.</emphasis>
My examination of the
<typeStatus id="54C5887DFFAFFF9FFD0AC878C8D1E411" box="[646,697,1288,1314]" pageId="3" pageNumber="34">type</typeStatus>
material of
<taxonomicName id="4C7E4D5CFFAFFF9FFCDBC878C99DE412" box="[855,1013,1288,1313]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="B90AEACDFFAFFF9FFCDBC878C99DE412" box="[855,1013,1288,1313]" italics="true" pageId="3" pageNumber="34">P. argentinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E4D5CFFAFFF9FFB85C878CEC8E412" box="[1033,1184,1288,1313]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="jujuyensis">
<emphasis id="B90AEACDFFAFFF9FFB85C878CEC8E412" box="[1033,1184,1288,1313]" italics="true" pageId="3" pageNumber="34">P. jujuyensis</emphasis>
</taxonomicName>
,
<emphasis id="B90AEACDFFAFFF9FFB38C878CA92E47B" italics="true" pageId="3" pageNumber="34">
<taxonomicName id="4C7E4D5CFFAFFF9FFB38C878CF1EE412" box="[1204,1398,1288,1313]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="hornaditanus">P. hornaditanus</taxonomicName>
,
<taxonomicName id="4C7E4D5CFFAFFF9FFA0AC878CA92E47B" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">P. nidicola</taxonomicName>
</emphasis>
and
<taxonomicName id="4C7E4D5CFFAFFF9FFEB9C85FCB90E47B" box="[309,504,1327,1352]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAFFF9FFEB9C85FCB90E47B" box="[309,504,1327,1352]" italics="true" pageId="3" pageNumber="34">P. tucumanensis</emphasis>
</taxonomicName>
established that these five species do not belong to the genus
<taxonomicName id="4C7E4D5CFFAFFF9FFB67C85FCF05E47B" box="[1259,1389,1327,1352]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFB67C85FCF05E47B" box="[1259,1389,1327,1352]" italics="true" pageId="3" pageNumber="34">Philonthus</emphasis>
</taxonomicName>
but belong to the genus
<taxonomicName id="4C7E4D5CFFAFFF9FFE0CC825CB88E45D" box="[384,480,1365,1390]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFE0CC825CB88E45D" box="[384,480,1365,1390]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
. In Smetana’s (1995) key to Holarctic genera of
<taxonomicName id="4C7E4D5CFFAFFF9FFB9BC825CEC9E45C" box="[1047,1185,1365,1391]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="subTribe" subTribe="Philonthina">Philonthina</taxonomicName>
, these species will go to couplet 13, where they share with
<taxonomicName id="4C7E4D5CFFAFFF9FFDC4C80CC8C0E4A6" box="[584,680,1404,1429]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFDC4C80CC8C0E4A6" box="[584,680,1404,1429]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
the “first four segments of front tarsus with ventral face bearing only regular, unmodified marginal setae; each simple, not dilated”, “lateral puncture of pronotum, bearing long seta, situated close to superior line of pronotal hypomeron, separated from it by distance about equal to diameter of puncture” and the “last segment of labial palpus slender, distinctly narrower than preceding segment”. Although
<taxonomicName id="4C7E4D5CFFAFFF9FFE01CB67C840E703" box="[397,552,1559,1584]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="B90AEACDFFAFFF9FFE01CB67C840E703" box="[397,552,1559,1584]" italics="true" pageId="3" pageNumber="34">P. argentinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E4D5CFFAFFF9FFDB4CB67C8A3E703" box="[568,715,1559,1584]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="jujuyensis">
<emphasis id="B90AEACDFFAFFF9FFDB4CB67C8A3E703" box="[568,715,1559,1584]" italics="true" pageId="3" pageNumber="34">P. jujuyensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7E4D5CFFAFFF9FFC82CB67C9E5E703" box="[782,909,1559,1584]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">
<emphasis id="B90AEACDFFAFFF9FFC82CB67C9E5E703" box="[782,909,1559,1584]" italics="true" pageId="3" pageNumber="34">P. nidicola</emphasis>
</taxonomicName>
do not share with
<taxonomicName id="4C7E4D5CFFAFFF9FFBE7CB67CEA4E703" box="[1131,1228,1559,1584]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFBE7CB67CEA4E703" box="[1131,1228,1559,1584]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
“the sternite 9 of male genital segment with proximal portion not reduced, symmetrical”, an exception for this character state (
<taxonomicName id="4C7E4D5CFFAFFF9FFF2DCB14CBB0E74D" authority="Horn, 1884" authorityName="Horn" authorityYear="1884" box="[161,472,1636,1662]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="brevipennis">
<emphasis id="B90AEACDFFAFFF9FFF2DCB14CB21E74E" box="[161,329,1636,1661]" italics="true" pageId="3" pageNumber="34">G. brevipennis</emphasis>
<bibRefCitation id="EFEF4B2EFFAFFF9FFEDCCB14CBB0E74D" author="Horn" box="[336,472,1636,1662]" pageId="3" pageNumber="42" refString="Horn, G. (1884) Synopsis of the Philonthini of Boreal America. Transactions of the American Entomological Society, 11, 177 - 244." type="journal article" year="1884">Horn, 1884</bibRefCitation>
</taxonomicName>
) is observed in that same key. Concerning
<taxonomicName id="4C7E4D5CFFAFFF9FFC5FCB14CEE4E74E" box="[979,1164,1636,1661]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="hornaditanus">
<emphasis id="B90AEACDFFAFFF9FFC5FCB14CEE4E74E" box="[979,1164,1636,1661]" italics="true" pageId="3" pageNumber="34">P. hornaditanus</emphasis>
</taxonomicName>
and
<emphasis id="B90AEACDFFAFFF9FFB4ACB14CFE8E74E" box="[1222,1408,1636,1661]" italics="true" pageId="3" pageNumber="34">P. t u c u m a n e n s i s</emphasis>
, I was able to confirm (in the case of
<taxonomicName id="4C7E4D5CFFAFFF9FFDA3CBFBC883E797" box="[559,747,1675,1700]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAFFF9FFDA3CBFBC883E797" box="[559,747,1675,1700]" italics="true" pageId="3" pageNumber="34">P. tucumanensis</emphasis>
</taxonomicName>
from additional non-type material) that these species share with
<taxonomicName id="4C7E4D5CFFAFFF9FFF54CBC1CB28E7F9" box="[216,320,1713,1738]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFF54CBC1CB28E7F9" box="[216,320,1713,1738]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
the sternite 9 with proximal portion symmetrical. In addition, all these five species,
<taxonomicName id="4C7E4D5CFFAFFF9FFA09CBC1CB71E7C2" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="argentinus">
<emphasis id="B90AEACDFFAFFF9FFA09CBC1CB71E7C2" italics="true" pageId="3" pageNumber="34">P. argentinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7E4D5CFFAFFF9FFEA6CBA8CBACE7C2" box="[298,452,1752,1777]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="jujuyensis">
<emphasis id="B90AEACDFFAFFF9FFEA6CBA8CBACE7C2" box="[298,452,1752,1777]" italics="true" pageId="3" pageNumber="34">P. jujuyensis</emphasis>
</taxonomicName>
,
<emphasis id="B90AEACDFFAFFF9FFE58CBA8C944E7C2" box="[468,812,1752,1777]" italics="true" pageId="3" pageNumber="34">
<taxonomicName id="4C7E4D5CFFAFFF9FFE58CBA8C8F3E7C2" box="[468,667,1752,1777]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="hornaditanus">P. hornaditanus</taxonomicName>
,
<taxonomicName id="4C7E4D5CFFAFFF9FFD24CBA8C944E7C2" box="[680,812,1752,1777]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="nidicola">P. nidicola</taxonomicName>
</emphasis>
and
<taxonomicName id="4C7E4D5CFFAFFF9FFCE6CBA8CE58E7C2" box="[874,1072,1752,1777]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="tucumanensis">
<emphasis id="B90AEACDFFAFFF9FFCE6CBA8CE58E7C2" box="[874,1072,1752,1777]" italics="true" pageId="3" pageNumber="34">P. tucumanensis</emphasis>
</taxonomicName>
, share with
<taxonomicName id="4C7E4D5CFFAFFF9FFB45CBA8CF45E7C2" box="[1225,1325,1752,1777]" class="Insecta" family="Staphylinidae" genus="Gabrius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACDFFAFFF9FFB45CBA8CF45E7C2" box="[1225,1325,1752,1777]" italics="true" pageId="3" pageNumber="34">Gabrius</emphasis>
</taxonomicName>
all other characters summarized by
<bibRefCitation id="EFEF4B2EFFAFFF9FFE5CCB8EC8AEE62B" author="Smetana" box="[464,710,1790,1816]" pageId="3" pageNumber="42" refString="Smetana, A. (1995) Rove beetles of the subtribe Philonthina of America north of Mexico (Coleoptera: Staphylinidae): Classification, phylogeny and taxonomic revision. Memoirs on Entomology, International, 3, 1 - 946." type="journal article" year="1995">Smetana (1995: 606)</bibRefCitation>
for the recognition of the species of this genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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