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<document id="E6927F5CDE88624B86FE109AB00B7763" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638258267731" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2018" docId="03C887BEFFBAFFF186201611FBCCE3F9" docLanguage="en" docName="Euscorpius.260.1-11.pdf" docOrigin="Euscorpius 260" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Compsobuthus maidensis Kovařík 2018, sp. n." docType="treatment" docUuid="442AA0FF-2FFB-465A-9EE2-95404CFF2429" docUuidSource="ZooBank" docVersion="3" lastPageNumber="6" masterDocId="FFF1FFC6FFBFFFF687121750FF97E54A" masterDocTitle="Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVI. Compsobuthus maidensis sp. n. (Buthidae) from Somaliland" masterLastPageNumber="11" masterPageNumber="1" pageNumber="4" updateTime="1699436198362" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="D9B4F169AB2CD70EE8A024514B3AB79B">Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part XVI. Compsobuthus maidensis sp. n. (Buthidae) from Somaliland</mods:title>
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<mods:namePart id="99E4B98AB8EBC4FE68EF4B86C8952E07">Kovařík, František</mods:namePart>
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<mods:typeOfResource id="CCECB971F0DA31FB7BD85C94EAB1DBE9">text</mods:typeOfResource>
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<mods:title id="AEE09020FFB6B165D0F760457CE8243A">Euscorpius</mods:title>
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<mods:date id="F667AFC7053F9CE6C73BD9FB0808D911">2018</mods:date>
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<subSubSection id="C37B6523FFBAFFF386201611FD27E417" box="[306,688,321,349]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BDE36A8FFBAFFF386201611FD27E417" blockId="5.[194,799,321,440]" box="[306,688,321,349]" pageId="5" pageNumber="4">
<heading id="D09681C4FFBAFFF386201611FD27E417" box="[306,688,321,349]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="4" reason="2">
<emphasis id="B915EABAFFBAFFF386201611FD27E417" bold="true" box="[306,688,321,349]" pageId="5" pageNumber="4">
<taxonomicName id="4C614D2BFFBAFFF386201611FDF0E416" ID-CoL="XHFX" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[306,615,321,348]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFBAFFF386201611FDF0E416" bold="true" box="[306,615,321,348]" italics="true" pageId="5" pageNumber="4">Compsobuthus maidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657C1FFBAFFF3857C1613FD27E417" box="[622,688,323,349]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="C37B6523FFBAFFF386931633FD52E4F2" pageId="5" pageNumber="4" type="description">
<paragraph id="8BDE36A8FFBAFFF386931633FC88E4D3" blockId="5.[194,799,321,440]" pageId="5" pageNumber="4">
(
<figureCitation id="135A2A2DFFBAFFF3869A1633FE6BE430" box="[392,508,355,378]" captionStart-0="Figures 1–4" captionStart-1="Figures 5–12" captionStart-2="Figures 13–20" captionStart-3="Figures 21–37" captionStart-4="Figure 38" captionStart-5="Figure 39" captionStartId-0="4.[192,278,1662,1685]" captionStartId-1="6.[192,278,1654,1677]" captionStartId-2="7.[192,278,1398,1421]" captionStartId-3="8.[192,278,1733,1756]" captionStartId-4="9.[192,268,1305,1328]" captionStartId-5="11.[192,268,1262,1285]" captionTargetBox-0="[209,1415,217,1632]" captionTargetBox-1="[191,1440,191,1624]" captionTargetBox-2="[204,1440,179,1349]" captionTargetBox-3="[214,1420,232,1728]" captionTargetBox-4="[191,1921,268,1273]" captionTargetBox-5="[191,1440,191,1232]" captionTargetId-0="figure-12@4.[192,1440,206,1632]" captionTargetId-1="figure-12@6.[192,1440,192,1623]" captionTargetId-2="figure-6@7.[192,1440,168,1369]" captionTargetId-3="figure-12@8.[192,1440,223,1728]" captionTargetId-4="figure-9@9.[192,1921,311,796]" captionTargetId-5="figure-6@11.[192,1439,192,1231]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="11" captionText-0="Figures 1–4: Compsobuthus maidensis sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." captionText-1="Figures 5–12: Compsobuthus maidensis sp. n. Figures 5, 7. Male holotype, carapace and tergites I–IV (5), coxosternal area and sternites (70). Figures 6, 8–12. Female paratype, carapace and tergites I–IV (6), coxosternal area and sternites (8), left legs I– IV, retrolateral aspects (9–12)." captionText-2="Figures 13–20: Compsobuthus maidensis sp. n. Figures 13, 15–17. Male holotype, lateral view of telson (13), metasoma and telson lateral (15), ventral (16), and dorsal (17). Figures 14, 18–20. Female paratype, lateral view of telson (14), metasoma and telson lateral (18), ventral (19), and dorsal (20). Scale bar: 10 mm (15–20)." captionText-3="Figures 21–37: Compsobuthus maidensis sp. n. Figures 21–30. Male holotype, pedipalp chela dorsal (20), externodorsal (22) and ventrointernal (23), pedipalp patella dorsal (24), external (25) and ventral (26), trochanter and femur internal (27) dorsoexternal (28), and ventrointernal (29), movable finger dentition (30). Figures 31–37. Female paratype, pedipalp chela dorsal (31), externodorsal (32) and ventrointernal (33), pedipalp patella dorsal (34), external (35) and ventral (36), trochanter and femur dorsoexternal (37). Trichobothrial pattern is indicated in Figures 22–25 and 27–28." captionText-4="Figure 38: The type locality of Compsobuthus maidensis sp. n., Somaliland, Maid." captionText-5="Figure 39: Map showing confirmed distribution of Compsobuthus spp. Points indicate sites sampled mainly during 2011– 2017 expeditions. Scorpion in the inside map of the photos is the male paratype of C. somalilandus." pageId="5" pageNumber="4">Figs. 1–39</figureCitation>
,
<tableCitation id="C6E30313FFBAFFF385151633FDC0E430" box="[519,599,355,378]" captionStart="Table 1" captionStartId="10.[192,257,858,881]" captionText="Table 1: Comparative measurements of adults of Compsobuthus maidensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="5" pageNumber="4">Table 1</tableCitation>
)
<emphasis id="B915EABAFFBAFFF387D016D2FC88E4D3" bold="true" box="[194,799,386,409]" pageId="5" pageNumber="4">http://zoobank.org/urn:lsid:zoobank.org:act:442AA0</emphasis>
</paragraph>
<paragraph id="8BDE36A8FFBAFFF3860E16F1FD52E4F2" blockId="5.[194,799,321,440]" box="[284,709,417,440]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF3860E16F1FD52E4F2" bold="true" box="[284,709,417,440]" pageId="5" pageNumber="4">FF-2FFB-465A-9EE2-95404CFF2429</emphasis>
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</subSubSection>
<subSubSection id="C37B6523FFBAFFF387D2168EFE2EE7E7" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BDE36A8FFBAFFF387D2168EFD4EE75E" blockId="5.[192,802,478,532]" pageId="5" pageNumber="4">
<materialsCitation id="3B093CF5FFBAFFF387D2168EFE27E4BF" box="[192,432,478,501]" collectionCode="LOCALITY, AND" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DA880AFFBAFFF387D2168EFF6FE4BF" box="[192,248,478,501]" pageId="5" pageNumber="4">TYPE</typeStatus>
<collectionCode id="ED70AE6DFFBAFFF3861A16B2FE27E4BF" box="[264,432,482,501]" pageId="5" pageNumber="4">LOCALITY AND</collectionCode>
</materialsCitation>
<materialsCitation id="3B093CF5FFBAFFF386D216B2FD16E75E" collectionCode="DEPOSITORY" elevation="52" latitude="11.0008335" longLatPrecision="21" longitude="47.108334" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DA880AFFBAFFF386D216B2FE62E4BF" box="[448,501,482,501]" pageId="5" pageNumber="4">TYPE</typeStatus>
<collectionCode id="ED70AE6DFFBAFFF3851716B2FD1DE4BF" box="[517,650,482,501]" pageId="5" pageNumber="4">DEPOSITORY</collectionCode>
. Somaliland, Maid,
<geoCoordinate id="EE55506FFFBAFFF3861416ADFE14E75E" box="[262,387,509,532]" degrees="11" direction="north" minutes="00" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="03" value="11.0008335">11°00'03&quot;N</geoCoordinate>
<geoCoordinate id="EE55506FFFBAFFF3869816ADFD90E75E" box="[394,519,509,532]" degrees="47" direction="east" minutes="06" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="30" value="47.108334">47°06'30&quot;E</geoCoordinate>
,
<elevation id="004CD19BFFBAFFF3850016ADFD16E75E" box="[530,641,509,532]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="4" unit="m" value="52.0">
<quantity id="4C999B4DFFBAFFF3850016ADFDD0E75E" box="[530,583,509,532]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="4" unit="m" value="52.0">52 m</quantity>
a.s.l.
</elevation>
</materialsCitation>
;
<collectionCode id="ED70AE6DFFBAFFF3859C16ADFD43E75E" box="[654,724,509,532]" pageId="5" pageNumber="4">FKCP</collectionCode>
.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF387D2156AFE2EE7E7" blockId="5.[192,803,570,685]" pageId="5" pageNumber="4">
<materialsCitation id="3B093CF5FFBAFFF387D2156AFEF3E7E7" collectingDate="2017-09-03" collectingDateMax="2017-09-04" collectingDateMin="2017-09-03" collectionCode="TYPE, MATERIAL, EXAMINED" collectorName="Locality No. &amp; F. Kovarik" elevation="52" latitude="11.0008335" longLatPrecision="21" longitude="47.108334" pageId="5" pageNumber="4" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="ED70AE6DFFBAFFF387D2156AFE7AE71B" box="[192,493,570,593]" pageId="5" pageNumber="4">TYPE MATERIAL EXAMINED</collectionCode>
.
<emphasis id="B915EABAFFBAFFF386ED156AFD17E71B" bold="true" box="[511,640,570,593]" pageId="5" pageNumber="4">Somaliland</emphasis>
, Maid,
<geoCoordinate id="EE55506FFFBAFFF385CF156AFF6EE73A" degrees="11" direction="north" minutes="00" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="03" value="11.0008335">11°00' 03&quot;N</geoCoordinate>
<geoCoordinate id="EE55506FFFBAFFF386151509FE13E73A" box="[263,388,601,624]" degrees="47" direction="east" minutes="06" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="30" value="47.108334">47°06'30&quot;E</geoCoordinate>
,
<elevation id="004CD19BFFBAFFF386841509FD83E73A" box="[406,532,601,624]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="4" unit="m" value="52.0">
<quantity id="4C999B4DFFBAFFF386841509FE44E73A" box="[406,467,601,624]" metricMagnitude="1" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="4" unit="m" value="52.0">52 m</quantity>
a.s.l.
</elevation>
(
<figureCitation id="135A2A2DFFBAFFF385381509FD17E73A" box="[554,640,601,624]" captionStart="Figure 38" captionStartId="9.[192,268,1305,1328]" captionTargetBox="[191,1921,268,1273]" captionTargetId="figure-9@9.[192,1921,311,796]" captionTargetPageId="9" captionText="Figure 38: The type locality of Compsobuthus maidensis sp. n., Somaliland, Maid." pageId="5" pageNumber="4">Fig. 38</figureCitation>
,
<collectorName id="2694537EFFBAFFF385811509FCB5E73A" box="[659,802,601,624]" pageId="5" pageNumber="4">Locality No.</collectorName>
17SN),
<date id="FFDF1068FFBAFFF3860B1527FE33E7C4" box="[281,420,631,654]" pageId="5" pageNumber="4" value="2017-09-03" valueMax="2017-09-04" valueMin="2017-09-03">
<collectingDate id="EF9BE980FFBAFFF3860B1527FE33E7C4" box="[281,420,631,654]" pageId="5" pageNumber="4" value="2017-09-03" valueMax="2017-09-04" valueMin="2017-09-03">3.-4.IX.2017</collectingDate>
</date>
,
<specimenCount id="9D67FD21FFBAFFF386A41527FE40E7C4" box="[438,471,631,654]" pageId="5" pageNumber="4" type="male">1♂</specimenCount>
<specimenCount id="9D67FD21FFBAFFF386C51527FE6FE7C4" box="[471,504,631,654]" pageId="5" pageNumber="4" type="female">3♀</specimenCount>
(
<typeStatus id="54DA880AFFBAFFF3851F1527FDFBE7C4" box="[525,620,631,654]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
and
<typeStatus id="54DA880AFFBAFFF385BE1527FC81E7C4" box="[684,790,631,654]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
), leg.
<collectorName id="2694537EFFBAFFF387FC15C6FEF3E7E7" box="[238,356,662,685]" pageId="5" pageNumber="4">F. Kovařík</collectorName>
</materialsCitation>
,
<collectionCode id="ED70AE6DFFBAFFF3867D15C6FE22E7E7" box="[367,437,662,685]" pageId="5" pageNumber="4">FKCP</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C37B6523FFBAFFF387D21583FD70E7A0" box="[192,743,723,746]" pageId="5" pageNumber="4" type="etymology">
<paragraph id="8BDE36A8FFBAFFF387D21583FD70E7A0" blockId="5.[192,743,723,746]" box="[192,743,723,746]" pageId="5" pageNumber="4">ETYMOLOGY. Named after the village of collection.</paragraph>
</subSubSection>
<subSubSection id="C37B6523FFBAFFF387D21441FD8FE0C7" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BDE36A8FFBAFFF387D21441FD8FE0C7" blockId="5.[192,802,785,1421]" pageId="5" pageNumber="4">
DIAGNOSIS. Total length
<quantity id="4C999B4DFFBAFFF386F31441FDE4E662" box="[481,627,785,808]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" metricValueMax="3.35" metricValueMin="3.0" pageId="5" pageNumber="4" unit="mm" value="31.75" valueMax="33.5" valueMin="30.0">30–33.5 mm</quantity>
. Sexual dimorphism minor, fingers of pedipalps straight in both sexes; there is no difference in length and width of metasomal segments. Base color uniformly yellow to yellowish brown with dark spot on fifth and four metasomal segment. Movable finger of pedipalp bears 10 rows of granules, all without external and with internal accessory granules (
<taxonomicName id="4C614D2BFFBAFFF3862714B7FE43E6B4" authorityName=": Sissom" authorityYear="1994" box="[309,468,999,1022]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="B915EABAFFBAFFF3862714B7FE43E6B4" box="[309,468,999,1022]" italics="true" pageId="5" pageNumber="4">acutecarinatus</emphasis>
</taxonomicName>
group of Levy &amp; Amitati, 1980). Pedipalp chela length/width ratio
<quantity id="4C999B4DFFBAFFF385981356FD4FE157" box="[650,728,1030,1053]" metricMagnitude="-1" metricUnit="m" metricValue="1.14046" pageId="5" pageNumber="4" unit="in" value="4.49">4.49 in</quantity>
males and
<quantity id="4C999B4DFFBAFFF387E01375FEA4E176" box="[242,307,1061,1084]" metricMagnitude="-1" metricUnit="m" metricValue="1.2446" pageId="5" pageNumber="4" unit="in" value="4.9">4.9 in</quantity>
females. Manus of chela shorter than fixed finger. Pedipalp chela length/movable finger length ratio
<quantity id="4C999B4DFFBAFFF387D21332FED0E133" box="[192,327,1122,1145]" metricMagnitude="-2" metricUnit="m" metricValue="3.5686999999999998" metricValueMax="3.5814" metricValueMin="3.5559999999999996" pageId="5" pageNumber="4" unit="in" value="1.4049999999999998" valueMax="1.41" valueMin="1.4">1.40–1.41 in</quantity>
both sexes. Trochanter of pedipalps with 3– 7 spinules and 2 setae. Anterior margin of carapace bears 8 symmetrically distributed spinules. First to third metasomal segments bear 10 carinae, fourth bears 8 or 10 carinae. All metasomal segments longer than wide. Pectinal teeth number
<quantity id="4C999B4DFFBAFFF386D913ABFDA5E058" box="[459,562,1275,1298]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="5" pageNumber="4" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
male and
<quantity id="4C999B4DFFBAFFF385A913ABFCB6E058" box="[699,801,1275,1298]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="5" pageNumber="4" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
females. Sternites and ventral surface of metasoma granulated. Seventh sternite bears four crenulate carinae. Telson bulbous, aculeus shorter than vesicle. Subaculear tubercle present but not spinoid.
</paragraph>
</subSubSection>
<subSubSection id="C37B6523FFBAFFF187D212E2FBCCE3F9" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4" type="description">
<paragraph id="8BDE36A8FFBAFFF387D212E2FC9DE328" blockId="5.[192,802,1458,1910]" pageId="5" pageNumber="4">
DESCRIPTION. Total length
<quantity id="4C999B4DFFBAFFF386E312E2FDEBE083" box="[497,636,1458,1481]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" metricValueMax="3.35" metricValueMin="3.0" pageId="5" pageNumber="4" unit="mm" value="31.75" valueMax="33.5" valueMin="30.0">30–33.5 mm</quantity>
in both sexes. The habitus is shown in
<figureCitation id="135A2A2DFFBAFFF386F51280FDC3E0AD" box="[487,596,1488,1511]" captionStart="Figures 1–4" captionStartId="4.[192,278,1662,1685]" captionTargetBox="[209,1415,217,1632]" captionTargetId="figure-12@4.[192,1440,206,1632]" captionTargetPageId="4" captionText="Figures 1–4: Compsobuthus maidensis sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." pageId="5" pageNumber="4">Figs. 1–4</figureCitation>
. For position and distribution of trichobothria of pedipalps see
<figureCitation id="135A2A2DFFBAFFF385A812BFFF49E36F" captionStart-0="Figures 13–20" captionStart-1="Figures 21–37" captionStartId-0="7.[192,278,1398,1421]" captionStartId-1="8.[192,278,1733,1756]" captionTargetBox-0="[204,1440,179,1349]" captionTargetBox-1="[214,1420,232,1728]" captionTargetId-0="figure-6@7.[192,1440,168,1369]" captionTargetId-1="figure-12@8.[192,1440,223,1728]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 13–20: Compsobuthus maidensis sp. n. Figures 13, 15–17. Male holotype, lateral view of telson (13), metasoma and telson lateral (15), ventral (16), and dorsal (17). Figures 14, 18–20. Female paratype, lateral view of telson (14), metasoma and telson lateral (18), ventral (19), and dorsal (20). Scale bar: 10 mm (15–20)." captionText-1="Figures 21–37: Compsobuthus maidensis sp. n. Figures 21–30. Male holotype, pedipalp chela dorsal (20), externodorsal (22) and ventrointernal (23), pedipalp patella dorsal (24), external (25) and ventral (26), trochanter and femur internal (27) dorsoexternal (28), and ventrointernal (29), movable finger dentition (30). Figures 31–37. Female paratype, pedipalp chela dorsal (31), externodorsal (32) and ventrointernal (33), pedipalp patella dorsal (34), external (35) and ventral (36), trochanter and femur dorsoexternal (37). Trichobothrial pattern is indicated in Figures 22–25 and 27–28." pageId="5" pageNumber="4">Figs. 20– 28</figureCitation>
. Sexual dimorphism minor, fingers of pedipalps straight in both sexes (
<figureCitation id="135A2A2DFFBAFFF386DF117CFD1DE309" box="[461,650,1580,1603]" captionStart="Figures 21–37" captionStartId="8.[192,278,1733,1756]" captionTargetBox="[214,1420,232,1728]" captionTargetId="figure-12@8.[192,1440,223,1728]" captionTargetPageId="8" captionText="Figures 21–37: Compsobuthus maidensis sp. n. Figures 21–30. Male holotype, pedipalp chela dorsal (20), externodorsal (22) and ventrointernal (23), pedipalp patella dorsal (24), external (25) and ventral (26), trochanter and femur internal (27) dorsoexternal (28), and ventrointernal (29), movable finger dentition (30). Figures 31–37. Female paratype, pedipalp chela dorsal (31), externodorsal (32) and ventrointernal (33), pedipalp patella dorsal (34), external (35) and ventral (36), trochanter and femur dorsoexternal (37). Trichobothrial pattern is indicated in Figures 22–25 and 27–28." pageId="5" pageNumber="4">Figs. 22 and 32</figureCitation>
); there is no difference in length and width of metasomal segments.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF387D2113AFD84E397" blockId="5.[192,802,1458,1910]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF387D2113AFEACE3CB" bold="true" box="[192,315,1642,1665]" pageId="5" pageNumber="4">Coloration</emphasis>
(
<figureCitation id="135A2A2DFFBAFFF38642113AFE57E3CB" box="[336,448,1642,1665]" captionStart="Figures 1–4" captionStartId="4.[192,278,1662,1685]" captionTargetBox="[209,1415,217,1632]" captionTargetId="figure-12@4.[192,1440,206,1632]" captionTargetPageId="4" captionText="Figures 1–4: Compsobuthus maidensis sp. n. Figures 1–2. Male holotype in dorsal (1) and ventral (2) views. Figures 3–4: Female paratype in dorsal (3) and ventral (4) aspects. Scale bar: 10 mm." pageId="5" pageNumber="4">Figs. 1–4</figureCitation>
). The base color is uniformly yellow to yellowish brown, with dark spot on anterior half of the fifth and fourth metasomal segment; other spots missing or indicated only.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF387D211B4FB04E64C" blockId="5.[192,802,1458,1910]" lastBlockId="5.[830,1441,199,1725]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF387D211B4FE61E3B1" bold="true" box="[192,502,1764,1787]" pageId="5" pageNumber="4">Carapace and mesosoma</emphasis>
(
<figureCitation id="135A2A2DFFBAFFF3850011B4FD13E3B1" box="[530,644,1764,1787]" captionStart="Figures 5–12" captionStartId="6.[192,278,1654,1677]" captionTargetBox="[191,1440,191,1624]" captionTargetId="figure-12@6.[192,1440,192,1623]" captionTargetPageId="6" captionText="Figures 5–12: Compsobuthus maidensis sp. n. Figures 5, 7. Male holotype, carapace and tergites I–IV (5), coxosternal area and sternites (70). Figures 6, 8–12. Female paratype, carapace and tergites I–IV (6), coxosternal area and sternites (8), left legs I– IV, retrolateral aspects (9–12)." pageId="5" pageNumber="4">Figs. 5–8</figureCitation>
). The entire carapace is covered by granules of different sizes. The carinae are moderately to strongly developed and granular. The anterior margin of the carapace is medially weakly concave, and bears eight symmetrically distributed spinules. The tergites are granulated. Tergites I–VI bear very strong, denticulate lateral carinae. Each carina terminates in a spiniform process that extends well past the posterior margin of the tergite. Tergite VII is pentacarinate, with lateral pairs strong, serratocrenulate and the median carina moderate, crenulate and present only in the proximal half. The pectinal tooth count is
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male and 16–18 (2x16, 4x18) in females. The pectine marginal tips extend to one-third of the fourth sternite in the female and to half of the fifth sternite in the male. The pectines have three marginal lamellae and six to eight middle lamellae. The lamellae bear numerous dark setae, each fulcrum with two or three dark setae. All sternites are finely granulated. The glabrous wide zone on posterior part of fifth sternite developed medially and absent on other sternites in male.The sixth and seventh segments bear four ventral crenulate carinae, which are more strongly developed on the seventh segment. The other sternites bear two carinae.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF3842C145EFBF1E11D" blockId="5.[830,1441,199,1725]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF3842C145EFBA0E66F" bold="true" box="[830,1079,782,805]" pageId="5" pageNumber="4">Metasoma and telson</emphasis>
(
<figureCitation id="135A2A2DFFBAFFF3835B145EFB47E66F" box="[1097,1232,782,805]" captionStart="Figures 13–20" captionStartId="7.[192,278,1398,1421]" captionTargetBox="[204,1440,179,1349]" captionTargetId="figure-6@7.[192,1440,168,1369]" captionTargetPageId="7" captionText="Figures 13–20: Compsobuthus maidensis sp. n. Figures 13, 15–17. Male holotype, lateral view of telson (13), metasoma and telson lateral (15), ventral (16), and dorsal (17). Figures 14, 18–20. Female paratype, lateral view of telson (14), metasoma and telson lateral (18), ventral (19), and dorsal (20). Scale bar: 10 mm (15–20)." pageId="5" pageNumber="4">Figs. 13–20</figureCitation>
). The first to third segments bear 10 carinae, the fourth segment bears 8 or 10 carinae and the fifth segment bears five carinae. Intermediate carinae of the fourth segment are replaced by isolated granules that may also form carinae. All segments are sparsely setose and densely granulated. Accessory rows of granules are present on dorsal surfaces of segments as well as on the ventral surface of the fifth segment. The telson is bulbous, with the aculeus a little shorter than the vesicle. A subaculear tubercle is present and variously short.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF3842C130FFB8DE0C0" blockId="5.[830,1441,199,1725]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF3842C130FFC3AE13C" bold="true" box="[830,941,1119,1142]" pageId="5" pageNumber="4">Pedipalps</emphasis>
(
<figureCitation id="135A2A2DFFBAFFF384D3130FFBDFE13C" box="[961,1096,1119,1142]" captionStart="Figures 21–37" captionStartId="8.[192,278,1733,1756]" captionTargetBox="[214,1420,232,1728]" captionTargetId="figure-12@8.[192,1440,223,1728]" captionTargetPageId="8" captionText="Figures 21–37: Compsobuthus maidensis sp. n. Figures 21–30. Male holotype, pedipalp chela dorsal (20), externodorsal (22) and ventrointernal (23), pedipalp patella dorsal (24), external (25) and ventral (26), trochanter and femur internal (27) dorsoexternal (28), and ventrointernal (29), movable finger dentition (30). Figures 31–37. Female paratype, pedipalp chela dorsal (31), externodorsal (32) and ventrointernal (33), pedipalp patella dorsal (34), external (35) and ventral (36), trochanter and femur dorsoexternal (37). Trichobothrial pattern is indicated in Figures 22–25 and 27–28." pageId="5" pageNumber="4">Figs. 21–37</figureCitation>
). The pedipalps are granulated and hirsute. The femur bears five carinae. The patella bears seven granular carinae. The chela bears five carinae. The movable and fixed fingers bear 10 rows of granules, all without external and with internal granules. Pedipalp chela length/width ratio
<quantity id="4C999B4DFFBAFFF383A313A8FB69E045" box="[1201,1278,1272,1295]" metricMagnitude="-1" metricUnit="m" metricValue="1.14046" pageId="5" pageNumber="4" unit="in" value="4.49">4.49 in</quantity>
males and
<quantity id="4C999B4DFFBAFFF3826C13A8FCC4E064" metricMagnitude="-1" metricUnit="m" metricValue="1.2446" pageId="5" pageNumber="4" unit="in" value="4.9">4.9 in</quantity>
females. Manus of chela shorter than fixed finger. Pedipalp chela length/movable finger length ratio
<quantity id="4C999B4DFFBAFFF382761266FC1EE021" metricMagnitude="-2" metricUnit="m" metricValue="3.5686999999999998" metricValueMax="3.5814" metricValueMin="3.5559999999999996" pageId="5" pageNumber="4" unit="in" value="1.4049999999999998" valueMax="1.41" valueMin="1.4">1.40– 1.41 in</quantity>
both sexes. The trochanter of pedipalps bears 3–7 spinules and 2 setae.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF3842C12C2FC03E3D4" blockId="5.[830,1441,199,1725]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF3842C12C2FCE4E0E3" bold="true" box="[830,883,1426,1449]" pageId="5" pageNumber="4">Legs</emphasis>
(
<figureCitation id="135A2A2DFFBAFFF3849712C2FC6AE0E3" box="[901,1021,1426,1449]" captionStart="Figures 5–12" captionStartId="6.[192,278,1654,1677]" captionTargetBox="[191,1440,191,1624]" captionTargetId="figure-12@6.[192,1440,192,1623]" captionTargetPageId="6" captionText="Figures 5–12: Compsobuthus maidensis sp. n. Figures 5, 7. Male holotype, carapace and tergites I–IV (5), coxosternal area and sternites (70). Figures 6, 8–12. Female paratype, carapace and tergites I–IV (6), coxosternal area and sternites (8), left legs I– IV, retrolateral aspects (9–12)." pageId="5" pageNumber="4">Figs. 9–12</figureCitation>
). Legs III and IV bear moderate tibial spurs. Retrolateral and prolateral pedal spurs are present on all legs. The tarsomeres bear two rows of macrosetae on the ventral surface and several macrosetae on the other surfaces. Bristlecombs are absent. The femur bears four carinae and the patella bears four to six carinae. The femur and patella bear only solitary macrosetae and are granulated except for external lateral surfaces, which are smooth.
</paragraph>
<paragraph id="8BDE36A8FFBAFFF3842C11F6FBFBE3F7" blockId="5.[830,1441,199,1725]" box="[830,1132,1702,1725]" pageId="5" pageNumber="4">
<emphasis id="B915EABAFFBAFFF3842C11F6FC72E3F7" bold="true" box="[830,997,1702,1725]" pageId="5" pageNumber="4">Measurements</emphasis>
. See
<tableCitation id="C6E30313FFBAFFF3833211F6FBFBE3F7" box="[1056,1132,1702,1725]" captionStart="Table 1" captionStartId="10.[192,257,858,881]" captionText="Table 1: Comparative measurements of adults of Compsobuthus maidensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." pageId="5" pageNumber="4">Tab. 1.</tableCitation>
</paragraph>
<paragraph id="8BDE36A8FFBAFFF1842C11B3FEDFE398" blockId="5.[830,1441,1763,1909]" lastBlockId="7.[192,802,1508,1746]" lastPageId="7" lastPageNumber="6" pageId="5" pageNumber="4">
AFFINITIES. The described features distinguish
<taxonomicName id="4C614D2BFFBAFFF3825211B3FC6AE253" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFBAFFF3825211B3FC6AE253" italics="true" pageId="5" pageNumber="4">Compsobuthus maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFBAFFF3831B1052FBDCE253" bold="true" box="[1033,1099,1794,1817]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="A22657C1FFBAFFF3831B1052FBDCE253" box="[1033,1099,1794,1817]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other species of the genus.
<taxonomicName id="4C614D2BFFBAFFF3849F1070FB8DE27D" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[909,1050,1824,1847]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFBAFFF3849F1070FB8DE27D" box="[909,1050,1824,1847]" italics="true" pageId="5" pageNumber="4">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFBAFFF383361070FBF5E27D" bold="true" box="[1060,1122,1824,1847]" pageId="5" pageNumber="4">
<taxonomicNameLabel id="A22657C1FFBAFFF383361070FBF5E27D" box="[1060,1122,1824,1847]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is morphologically the most similar to
<taxonomicName id="4C614D2BFFBAFFF384B9106FFAB3E21C" authority="Kovarik et al., 2016" authorityName="Kovarik" authorityYear="2016" box="[939,1316,1855,1878]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFBAFFF384B9106FFBDFE21C" box="[939,1096,1855,1878]" italics="true" pageId="5" pageNumber="4">C. eritreaensis</emphasis>
<bibRefCitation id="EFF04B59FFBAFFF3835D106FFAB3E21C" author="KOVARIK" box="[1103,1316,1855,1878]" pageId="5" pageNumber="4" pagination="1 - 21" refId="ref6141" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae) with description of C. eritreaensis sp. n. Euscorpius, 226: 1 - 21." type="journal article" year="2016">Kovařík et al., 2016</bibRefCitation>
</taxonomicName>
(see key in
<bibRefCitation id="EFF04B59FFBAFFF3842C100EFBD4E23F" author="KOVARIK" box="[830,1091,1886,1909]" pageId="5" pageNumber="4" pagination="1 - 21" refId="ref6141" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae) with description of C. eritreaensis sp. n. Euscorpius, 226: 1 - 21." type="journal article" year="2016">Kovařík et al., 2016: 19</bibRefCitation>
) but these two species occur in remote areas (
<figureCitation id="135A2A2DFFB9FFF0867811B4FE55E3B1" box="[362,450,1764,1787]" captionStart="Figure 39" captionStartId="11.[192,268,1262,1285]" captionTargetBox="[191,1440,191,1232]" captionTargetId="figure-6@11.[192,1439,192,1231]" captionTargetPageId="11" captionText="Figure 39: Map showing confirmed distribution of Compsobuthus spp. Points indicate sites sampled mainly during 2011– 2017 expeditions. Scorpion in the inside map of the photos is the male paratype of C. somalilandus." pageId="6" pageNumber="5">Fig. 39</figureCitation>
) and can be morphologically unequivocally separated by:
<emphasis id="B915EABAFFB9FFF085151052FD83E253" bold="true" box="[519,532,1794,1817]" pageId="6" pageNumber="5">1</emphasis>
) Pectinal teeth number
<quantity id="4C999B4DFFB9FFF087D21071FEB7E272" box="[192,288,1825,1848]" metricMagnitude="-1" metricUnit="m" metricValue="4.953" metricValueMax="5.08" metricValueMin="4.826" pageId="6" pageNumber="5" unit="in" value="19.5" valueMax="20.0" valueMin="19.0">19–20 in</quantity>
male and
<quantity id="4C999B4DFFB9FFF086831071FE66E272" box="[401,497,1825,1848]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="6" pageNumber="5" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
females in
<taxonomicName id="4C614D2BFFB9FFF0857D1071FD6DE272" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[623,762,1825,1848]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB9FFF0857D1071FD6DE272" box="[623,762,1825,1848]" italics="true" pageId="6" pageNumber="5">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB9FFF084111071FF42E21D" bold="true" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A22657C1FFB9FFF084111071FF42E21D" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
vs.
<quantity id="4C999B4DFFB9FFF086011010FEEDE21D" box="[275,378,1856,1879]" metricMagnitude="-1" metricUnit="m" metricValue="6.096" metricValueMax="6.604" metricValueMin="5.588" pageId="6" pageNumber="5" unit="in" value="24.0" valueMax="26.0" valueMin="22.0">22–26 in</quantity>
males and
<quantity id="4C999B4DFFB9FFF0851E1010FDE3E21D" box="[524,628,1856,1879]" metricMagnitude="-1" metricUnit="m" metricValue="5.207" metricValueMax="5.842" metricValueMin="4.572" pageId="6" pageNumber="5" unit="in" value="20.5" valueMax="23.0" valueMin="18.0">18–23 in</quantity>
females in
<taxonomicName id="4C614D2BFFB9FFF0841B1010FEAAE23F" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFB9FFF0841B1010FEAAE23F" italics="true" pageId="6" pageNumber="5">C. eritreaensis</emphasis>
</taxonomicName>
;
<emphasis id="B915EABAFFB9FFF08646100EFEF6E23F" bold="true" box="[340,353,1886,1909]" pageId="6" pageNumber="5">2</emphasis>
) pedipalp chela length/movable finger length ratio is
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both sexes in
<taxonomicName id="4C614D2BFFB9FFF0820111B4FA37E3B1" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[1299,1440,1764,1787]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB9FFF0820111B4FA37E3B1" box="[1299,1440,1764,1787]" italics="true" pageId="6" pageNumber="5">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB9FFF0842C1052FCE9E253" bold="true" box="[830,894,1794,1817]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A22657C1FFB9FFF0842C1052FCE9E253" box="[830,894,1794,1817]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
vs.
<quantity id="4C999B4DFFB9FFF084A21052FBADE253" box="[944,1082,1794,1817]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="3.5052" metricValueMin="3.3528" pageId="6" pageNumber="5" unit="in" value="1.35" valueMax="1.38" valueMin="1.32">1.32–1.38 in</quantity>
<taxonomicName id="4C614D2BFFB9FFF083511052FB74E253" box="[1091,1251,1794,1817]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFB9FFF083511052FB74E253" box="[1091,1251,1794,1817]" italics="true" pageId="6" pageNumber="5">C. eritreaensis</emphasis>
</taxonomicName>
;
<emphasis id="B915EABAFFB9FFF083E61052FA96E253" bold="true" box="[1268,1281,1794,1817]" pageId="6" pageNumber="5">3</emphasis>
) trochanter of pedipalps with 3–7 spinules and 2 setae in
<taxonomicName id="4C614D2BFFB9FFF082071071FA37E272" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[1301,1440,1825,1848]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB9FFF082071071FA37E272" box="[1301,1440,1825,1848]" italics="true" pageId="6" pageNumber="5">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB9FFF084521010FCEAE21D" bold="true" box="[832,893,1856,1879]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A22657C1FFB9FFF084521010FCEAE21D" box="[832,893,1856,1879]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
vs. 10–12 spinules and 2 setae in
<taxonomicName id="4C614D2BFFB9FFF083E91010FA0EE21D" box="[1275,1433,1856,1879]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFB9FFF083E91010FA0EE21D" box="[1275,1433,1856,1879]" italics="true" pageId="6" pageNumber="5">C. eritreaensis</emphasis>
</taxonomicName>
;
<emphasis id="B915EABAFFB9FFF0842C100EFCDCE23F" bold="true" box="[830,843,1886,1909]" pageId="6" pageNumber="5">4</emphasis>
) fingers of pedipalps straight in both sexes in
<taxonomicName id="4C614D2BFFB9FFF18291100EFEBCE0B1" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" lastPageId="7" lastPageNumber="6" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB9FFF18291100EFEBCE0B1" italics="true" lastPageId="7" lastPageNumber="6" pageId="6" pageNumber="5">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB8FFF1862612B4FEE5E0B1" bold="true" box="[308,370,1508,1531]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A22657C1FFB8FFF1862612B4FEE5E0B1" box="[308,370,1508,1531]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135A2A2DFFB8FFF1869612B4FDA5E0B1" box="[388,562,1508,1531]" captionStart="Figures 21–37" captionStartId="8.[192,278,1733,1756]" captionTargetBox="[214,1420,232,1728]" captionTargetId="figure-12@8.[192,1440,223,1728]" captionTargetPageId="8" captionText="Figures 21–37: Compsobuthus maidensis sp. n. Figures 21–30. Male holotype, pedipalp chela dorsal (20), externodorsal (22) and ventrointernal (23), pedipalp patella dorsal (24), external (25) and ventral (26), trochanter and femur internal (27) dorsoexternal (28), and ventrointernal (29), movable finger dentition (30). Figures 31–37. Female paratype, pedipalp chela dorsal (31), externodorsal (32) and ventrointernal (33), pedipalp patella dorsal (34), external (35) and ventral (36), trochanter and femur dorsoexternal (37). Trichobothrial pattern is indicated in Figures 22–25 and 27–28." pageId="7" pageNumber="6">Figs. 28 and 30</figureCitation>
) vs. adult males with fingers of pedipalps slightly flexed proximally (figs. 28 and
<quantity id="4C999B4DFFB8FFF187E21172FEBEE373" box="[240,297,1570,1593]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="6" unit="in" value="30.0">30 in</quantity>
<bibRefCitation id="EFF04B59FFB8FFF186201172FDBAE373" author="KOVARIK" box="[306,557,1570,1593]" pageId="7" pageNumber="6" pagination="1 - 21" refId="ref6141" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae) with description of C. eritreaensis sp. n. Euscorpius, 226: 1 - 21." type="journal article" year="2016">Kovařík et al., 2016: 7</bibRefCitation>
) in
<taxonomicName id="4C614D2BFFB8FFF1854E1172FD6CE373" box="[604,763,1570,1593]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFB8FFF1854E1172FD6CE373" box="[604,763,1570,1593]" italics="true" pageId="7" pageNumber="6">C. eritreaensis</emphasis>
</taxonomicName>
;
<emphasis id="B915EABAFFB8FFF184191172FC8FE373" bold="true" box="[779,792,1570,1593]" pageId="7" pageNumber="6">5</emphasis>
) glabrous wide zone on posterior part of fifth sternite developed medially and absent on other sternites in male in
<taxonomicName id="4C614D2BFFB8FFF187CC112EFEFCE3DF" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[222,363,1662,1685]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB8FFF187CC112EFEFCE3DF" box="[222,363,1662,1685]" italics="true" pageId="7" pageNumber="6">C. maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB8FFF18666112EFE24E3DF" bold="true" box="[372,435,1662,1685]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A22657C1FFB8FFF18666112EFE24E3DF" box="[372,435,1662,1685]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135A2A2DFFB8FFF186D6112EFD90E3DF" box="[452,519,1662,1685]" captionStart="Figures 5–12" captionStartId="6.[192,278,1654,1677]" captionTargetBox="[191,1440,191,1624]" captionTargetId="figure-12@6.[192,1440,192,1623]" captionTargetPageId="6" captionText="Figures 5–12: Compsobuthus maidensis sp. n. Figures 5, 7. Male holotype, carapace and tergites I–IV (5), coxosternal area and sternites (70). Figures 6, 8–12. Female paratype, carapace and tergites I–IV (6), coxosternal area and sternites (8), left legs I– IV, retrolateral aspects (9–12)." pageId="7" pageNumber="6">Fig. 7</figureCitation>
) vs. reduced/indicated on sternites in male in
<taxonomicName id="4C614D2BFFB8FFF1868411CCFDA3E3F9" box="[406,564,1692,1715]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="B915EABAFFB8FFF1868411CCFDA3E3F9" box="[406,564,1692,1715]" italics="true" pageId="7" pageNumber="6">C. eritreaensis</emphasis>
</taxonomicName>
(fig.
<quantity id="4C999B4DFFB8FFF1856311CCFD30E3F9" box="[625,679,1692,1715]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="7" pageNumber="6" unit="in" value="11.0">11 in</quantity>
<bibRefCitation id="EFF04B59FFB8FFF185BD11CCFEACE398" author="KOVARIK" pageId="7" pageNumber="6" pagination="1 - 21" refId="ref6141" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae) with description of C. eritreaensis sp. n. Euscorpius, 226: 1 - 21." type="journal article" year="2016">Kovařík et al., 2016: 6</bibRefCitation>
).
</paragraph>
<caption id="DF1E6620FFB9FFF087D21126FE71E3F4" pageId="6" pageNumber="5" startId="6.[192,278,1654,1677]" targetBox="[191,1440,191,1624]" targetPageId="6">
<paragraph id="8BDE36A8FFB9FFF087D21126FE71E3F4" blockId="6.[192,1441,1654,1726]" pageId="6" pageNumber="5">
<emphasis id="B915EABAFFB9FFF087D21126FECAE3C7" bold="true" box="[192,349,1654,1677]" pageId="6" pageNumber="5">Figures 5–12:</emphasis>
<taxonomicName id="4C614D2BFFB9FFF086771127FDCBE3C6" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[357,604,1655,1676]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB9FFF086771127FDCBE3C6" box="[357,604,1655,1676]" italics="true" pageId="6" pageNumber="5">Compsobuthus maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB9FFF085771127FCB6E3C6" bold="true" box="[613,801,1655,1676]" pageId="6" pageNumber="5">
<taxonomicNameLabel id="A22657C1FFB9FFF085771127FD0AE3C6" box="[613,669,1655,1676]" pageId="6" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
Figures 5, 7
</emphasis>
. Male holotype, carapace and tergites I–IV (5), coxosternal area and sternites (70).
<emphasis id="B915EABAFFB9FFF0866111C0FD99E3EF" bold="true" box="[371,526,1680,1701]" pageId="6" pageNumber="5">Figures 6, 8–12</emphasis>
. Female paratype, carapace and tergites I–IV (6), coxosternal area and sternites (8), left legs I– IV, retrolateral aspects (9–12).
</paragraph>
</caption>
<caption id="DF1E6620FFB8FFF187D21226FC06E0F5" pageId="7" pageNumber="6" startId="7.[192,278,1398,1421]" targetBox="[204,1440,179,1349]" targetPageId="7">
<paragraph id="8BDE36A8FFB8FFF187D21226FC06E0F5" blockId="7.[192,1442,1398,1471]" pageId="7" pageNumber="6">
<emphasis id="B915EABAFFB8FFF187D21226FEFEE0C7" bold="true" box="[192,361,1398,1421]" pageId="7" pageNumber="6">Figures 13–20:</emphasis>
<taxonomicName id="4C614D2BFFB8FFF186631228FDF0E0C7" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[369,615,1400,1421]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="maidensis" status="sp. nov.">
<emphasis id="B915EABAFFB8FFF186631228FDF0E0C7" box="[369,615,1400,1421]" italics="true" pageId="7" pageNumber="6">Compsobuthus maidensis</emphasis>
</taxonomicName>
<emphasis id="B915EABAFFB8FFF1857C1228FCF4E0C7" bold="true" box="[622,867,1400,1421]" pageId="7" pageNumber="6">
<taxonomicNameLabel id="A22657C1FFB8FFF1857C1228FD31E0C7" box="[622,678,1400,1421]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
Figures 13, 15–17
</emphasis>
. Male holotype, lateral view of telson (13), metasoma and telson lateral (15), ventral (16), and dorsal (17).
<emphasis id="B915EABAFFB8FFF1858912C1FCD8E0EC" bold="true" box="[667,847,1425,1446]" pageId="7" pageNumber="6">Figures 14, 18–20</emphasis>
. Female paratype, lateral view of telson (14), metasoma and telson lateral (18), ventral (19), and dorsal (20). Scale bar: 10 mm (15–20).
</paragraph>
</caption>
<paragraph id="8BDE36A8FFB8FFF187D211ABFBCCE3F9" blockId="7.[192,802,1787,1902]" lastBlockId="7.[830,1441,1508,1715]" pageId="7" pageNumber="6">
COMMENTS ON LOCALITIES AND LIFE STRATEGY. The
<typeStatus id="54DA880AFFB8FFF187D2104AFF79E27B" box="[192,238,1818,1841]" pageId="7" pageNumber="6">type</typeStatus>
locality, 17SN is sandy semi-desert to desert (
<figureCitation id="135A2A2DFFB8FFF185E4104AFF4CE21A" captionStart="Figure 38" captionStartId="9.[192,268,1305,1328]" captionTargetBox="[191,1921,268,1273]" captionTargetId="figure-9@9.[192,1921,311,796]" captionTargetPageId="9" captionText="Figure 38: The type locality of Compsobuthus maidensis sp. n., Somaliland, Maid." pageId="7" pageNumber="6">Fig. 38</figureCitation>
). The
<typeStatus id="54DA880AFFB8FFF186231069FEFEE21A" box="[305,361,1849,1872]" pageId="7" pageNumber="6">types</typeStatus>
of
<taxonomicName id="4C614D2BFFB8FFF1868C1069FD58E21A" authority="Kovařík, 2018" authorityName="Kovařík" authorityYear="2018" box="[414,719,1849,1872]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="maidenensis" status="sp. nov.">
<emphasis id="B915EABAFFB8FFF1868C1069FD58E21A" box="[414,719,1849,1872]" italics="true" pageId="7" pageNumber="6">Compsobuthus maidenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22657C1FFB8FFF185CC1069FCB5E21A" box="[734,802,1849,1872]" pageId="7" pageNumber="6" rank="species">
<emphasis id="B915EABAFFB8FFF185CC1069FD60E21A" bold="true" box="[734,759,1849,1872]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="B915EABAFFB8FFF1841E1069FC8CE21A" bold="true" box="[780,795,1849,1872]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
were obtained at night during UV collecting together with
<taxonomicName id="4C614D2BFFB8FFF1846812B4FA81E0B1" authority="Kovarik et al., 2018" authorityName="Kovarik" authorityYear="2018" box="[890,1302,1508,1531]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="maidensis">
<emphasis id="B915EABAFFB8FFF1846812B4FBB6E0B1" box="[890,1057,1508,1531]" italics="true" pageId="7" pageNumber="6">Gint maidensis</emphasis>
Kovařík et al., 2018
</taxonomicName>
,
<taxonomicName id="4C614D2BFFB8FFF1823512B4FCC8E350" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B915EABAFFB8FFF1823512B4FA36E0B1" box="[1319,1441,1508,1531]" italics="true" pageId="7" pageNumber="6">Hottentotta</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C614D2BFFB8FFF1847E1153FC74E350" box="[876,995,1539,1562]" class="Arachnida" family="Buthidae" genus="Leiurus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B915EABAFFB8FFF1847E1153FC2AE350" box="[876,957,1539,1562]" italics="true" pageId="7" pageNumber="6">Leiurus</emphasis>
sp.
</taxonomicName>
, and
<taxonomicName id="4C614D2BFFB8FFF183311153FAAFE350" box="[1059,1336,1539,1562]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B915EABAFFB8FFF183311153FB04E350" box="[1059,1171,1539,1562]" italics="true" pageId="7" pageNumber="6">Neobuthus</emphasis>
sp. (Buthidae)
</taxonomicName>
. The first author arrived at the locality at night on 3th
<date id="FFDF1068FFB8FFF1823C1172FCE4E31D" pageId="7" pageNumber="6" value="2017-09">September 2017</date>
at 21.00. At this time the temperature was 38.6 ºC and humidity 52%. Minimum temperature of 31.9 ºC and humidity of 46% were recorded on
<date id="FFDF1068FFB8FFF18210112EFCE4E3F9" pageId="7" pageNumber="6" value="2017-09-04">
4th September
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</date>
in the early morning.
</paragraph>
</subSubSection>
</treatment>
</document>
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