treatments-xml/data/03/B4/87/03B487885B322A35FDC33C256674FB1F.xml

1015 lines
155 KiB
XML
Raw Normal View History

2024-06-21 12:22:17 +02:00
<document id="4AD0BB98A809ECEB9CAD43E90B54101B" ID-CLB-Dataset="168932" ID-DOI="10.5852/ejt.2022.844.1949" ID-GBIF-Dataset="3a08058c-14f8-4232-ad02-65ed5b28380f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222155" ID-ZooBank="193EDD91-B24D-455C-B8AA-8133586A00A1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126129383" checkinUser="felipe" docAuthor="Grzelak, Katarzyna &amp; Sørensen, Martin V." docDate="2022" docId="03B487885B322A35FDC33C256674FB1F" docLanguage="en" docName="ejt-844_1-108.pdf" docOrigin="European Journal of Taxonomy 844" docSource="http://dx.doi.org/10.5852/ejt.2022.844.1949" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes leduci Grzelak &amp; Sørensen 2022, sp. nov." docType="treatment" docUuid="B0BC5E3C-0F52-4854-B0BD-D80712B6EC30" docUuidSource="ZooBank" docVersion="12" lastPageNumber="38" masterDocId="FF8DFFF05B2C2A10FFB63A55654FFFB8" masterDocTitle="Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand" masterLastPageNumber="108" masterPageNumber="1" pageNumber="31" updateTime="1699407836451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="BC046515ACDAEF525EFE8B608E2118B3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E64874500A17277438367748CDCB90C">
<mods:title id="3537089A1A236F1D7BAB9DA00BB1604D">Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand</mods:title>
</mods:titleInfo>
<mods:name id="5CF9ADDF1FF4E5066C6C364FB789B214" type="personal">
<mods:role id="5EDF0B9597AD77517EBCD5E248294D84">
<mods:roleTerm id="FF82D6156ECA484325A1BB32DEFCB438">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1280E0ACDAFF17FFFF8666E761DFE506">Grzelak, Katarzyna</mods:namePart>
<mods:nameIdentifier id="2C4658C7822289CFBE46F06AF10EDEEA" type="LSID">ABFE9734-0BD8-4ED9-A780-469D0058FCED</mods:nameIdentifier>
<mods:affiliation id="3190937F89E27EF3CB742A5A8BB9BFA2">Polish Academy of Sciences, Institute of Oceanology, Sopot, Poland. &amp; Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
<mods:nameIdentifier id="3263752B27FCA914F0DA3DCB32E658B9" type="email">kgrzelak@iopan.gda.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="347413C90BA9D1D0A74450C85E91E515" type="personal">
<mods:role id="4F4DB6678161D8000FFC21CB65450055">
<mods:roleTerm id="C2AFBCCB82440EA6079B3F2B8A9B207A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DE64C13F8B87CB23313A6BCC54C99BB">Sørensen, Martin V.</mods:namePart>
<mods:nameIdentifier id="E52AD60F641177F96207270DF1D4A378" type="LSID">4143D650-12FC-4914-93F5-2C39339A7156</mods:nameIdentifier>
<mods:affiliation id="8037B27FAD56391C23E4D11512CAECA3">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
<mods:nameIdentifier id="A3A70A816F71513AF7592636746D0D27" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="14D947D82D5B5D990DFF73D2050D8914">text</mods:typeOfResource>
<mods:relatedItem id="1E5212D8F123F0B93870FAFAF844299E" type="host">
<mods:titleInfo id="88F6AB1B96CC6B72311717E0D7392735">
<mods:title id="747B105596666C0726629A117455FF40">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="17F701DE78F28C601740E3AAD520D74E">
<mods:date id="097254D543D663CD4F033309093CFEB6">2022</mods:date>
<mods:detail id="C9CF81EC0F0423FD100CAFAD11A37701" type="pubDate">
<mods:number id="F885953CAA17B41A46DF70466FB06A9F">2022-10-18</mods:number>
</mods:detail>
<mods:detail id="C0EDDBB9AD09E027F18DB528AF8D20B1" type="volume">
<mods:number id="C927C329913E0E1DC186FFA046094BF6">844</mods:number>
</mods:detail>
<mods:extent id="28D562A3780BED5A603E83E165FF8A22" unit="page">
<mods:start id="B4360B0B13FFC05FA94BB08905C78829">1</mods:start>
<mods:end id="3658100DC28233FE1ECF2BF32957874C">108</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0B6AB381439F3263D762B4CFAA1CEDE7">
<mods:url id="EA72BE4AA60AC3A726D0C8C0219D265F">http://dx.doi.org/10.5852/ejt.2022.844.1949</mods:url>
</mods:location>
<mods:classification id="93E0DB10EDAFD8C94D5AD0F2507F4D9E">journal article</mods:classification>
<mods:identifier id="88CC3861B360D14D8B49D9EB57761A01" type="CLB-Dataset">168932</mods:identifier>
<mods:identifier id="E4D3512F7C7FA6D7110A89085C3BF2B1" type="DOI">10.5852/ejt.2022.844.1949</mods:identifier>
<mods:identifier id="F2E3FBA0D198ED73662B1769C09866B8" type="GBIF-Dataset">3a08058c-14f8-4232-ad02-65ed5b28380f</mods:identifier>
<mods:identifier id="1B7007A82B64427F0BD5973BE60C125E" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="58D1C801A5A2574551B3ADA994C3888F" type="Zenodo-Dep">7222155</mods:identifier>
<mods:identifier id="74D5D94707B67FD68E5A38D980B6055E" type="ZooBank">193EDD91-B24D-455C-B8AA-8133586A00A1</mods:identifier>
</mods:mods>
<treatment id="03B487885B322A35FDC33C256674FB1F" ID-DOI="http://doi.org/10.5281/zenodo.7225521" ID-GBIF-Taxon="203115732" ID-Zenodo-Dep="7225521" LSID="urn:lsid:zoobank.org:act:B0BC5E3C-0F52-4854-B0BD-D80712B6EC30" httpUri="http://treatment.plazi.org/id/03B487885B322A35FDC33C256674FB1F" lastPageId="37" lastPageNumber="38" pageId="30" pageNumber="31">
<subSubSection id="C30765155B322A0EFDC33C2566F1F933" box="[629,958,1648,1675]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="8BA2369E5B322A0EFDC33C2566F1F933" blockId="30.[363,1225,1648,1752]" box="[629,958,1648,1675]" pageId="30" pageNumber="31">
<heading id="D0EA81F25B322A0EFDC33C2566F1F933" box="[629,958,1648,1675]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="4C1D4D1D5B322A0EFDC33C256612F933" ID-CoL="d8aa052c-c0f0-497b-b7f9-0a9033c5b9eb" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[629,861,1648,1675]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="30" pageNumber="31" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B322A0EFDC33C256612F933" bold="true" box="[629,861,1648,1675]" italics="true" pageId="30" pageNumber="31">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B322A0EFCD23C2466F1F933" box="[868,958,1649,1675]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30765155B322A0EFEDD3CC26186F90A" box="[363,1225,1687,1714]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BA2369E5B322A0EFEDD3CC26186F90A" blockId="30.[363,1225,1648,1752]" box="[363,1225,1687,1714]" pageId="30" pageNumber="31">
<uri id="FF8C3A9C5B322A0EFEDD3CC26186F90A" box="[363,1225,1687,1714]" pageId="30" pageNumber="31">
urn:lsid:zoobank.org:act:
<uuid id="FFBB0C4B5B322A0EFD3C3CC26186F90A" box="[650,1225,1687,1714]" pageId="30" pageNumber="31">B0BC5E3C-0F52-4854-B0BD-D80712B6EC30</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C30765155B322A0EFD273CE866EDF960" box="[657,930,1725,1752]" pageId="30" pageNumber="31" type="description">
<paragraph id="8BA2369E5B322A0EFD273CE866EDF960" blockId="30.[363,1225,1648,1752]" box="[657,930,1725,1752]" pageId="30" pageNumber="31">
<figureCitation id="13262A1B5B322A0EFD273CE86659F960" box="[657,790,1725,1752]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="32.[189,232,1927,1953]" captionStartId-1="34.[190,233,1857,1883]" captionStartId-2="35.[188,231,1855,1881]" captionTargetBox-0="[217,1368,271,1892]" captionTargetBox-1="[196,1394,587,1819]" captionTargetBox-2="[190,1396,857,1822]" captionTargetId-0="figure-39@32.[216,1371,267,1892]" captionTargetId-1="figure-92@34.[194,1404,586,1820]" captionTargetId-2="figure-0@35.[188,1398,856,1832]" captionTargetPageId-0="32" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." captionText-2="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222181" figureDoi-1="http://doi.org/10.5281/zenodo.7222183" figureDoi-2="http://doi.org/10.5281/zenodo.7222185" httpUri-0="https://zenodo.org/record/7222181/files/figure.png" httpUri-1="https://zenodo.org/record/7222183/files/figure.png" httpUri-2="https://zenodo.org/record/7222185/files/figure.png" pageId="30" pageNumber="31">Figs 1113</figureCitation>
;
<tableCitation id="C69F03255B322A0EFC953CE866EDF960" box="[803,930,1725,1752]" captionStart-0="Table 8" captionStart-1="Table 9" captionStartId-0="33.[189,258,265,291]" captionStartId-1="35.[189,258,265,291]" captionTargetPageId-0="33" captionTargetPageId-1="35" captionText-0="Table 8. Measurements from light microscopy ofEchinoderes leduci sp. nov. (in µm) from the Hikurangi Margin." captionText-1="Table 9. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes leduci sp. nov." pageId="30" pageNumber="31">Tables 89</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30765155B322A0EFF0B3D57602AF850" pageId="30" pageNumber="31" type="diagnosis">
<paragraph id="8BA2369E5B322A0EFF0B3D576479F8A4" blockId="30.[189,310,1794,1820]" box="[189,310,1794,1820]" pageId="30" pageNumber="31">
<heading id="D0EA81F25B322A0EFF0B3D576479F8A4" bold="true" box="[189,310,1794,1820]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B969EA8C5B322A0EFF0B3D576479F8A4" bold="true" box="[189,310,1794,1820]" pageId="30" pageNumber="31">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B322A0EFF0B3D61602AF850" blockId="30.[189,1400,1844,2025]" pageId="30" pageNumber="31">
<taxonomicName id="4C1D4D1D5B322A0EFF0B3D616400F8F6" authorityName="Claparede" authorityYear="1863" box="[189,335,1844,1870]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="30" pageNumber="31" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B322A0EFF0B3D616400F8F6" box="[189,335,1844,1870]" italics="true" pageId="30" pageNumber="31">Echinoderes</emphasis>
</taxonomicName>
with spines in middorsal position on segments 4, 6 and 8 and spines in lateroventral positions on segments 6 to 9. Tubes present in laterodorsal and ventrolateral positions on segment 2, lateroventral positions on segment 5, sublateral positions on segment 8, and laterodorsal positions on segment 9. Sexually dimorphic tubes furthermore present in laterodorsal positions on segment
<quantity id="4CE59B7B5B322A0EFA8E3DFD6038F87A" box="[1336,1399,1960,1986]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="30" pageNumber="31" unit="in" value="10.0">10 in</quantity>
males. Minute scales present on segments 2 to 10, but regular cuticular hairs absent throughout trunk.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B332A0FFF0B3B5C6070FECD" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8BA2369E5B332A0FFF0B3B5C640BFE9B" blockId="31.[189,324,265,291]" box="[189,324,265,291]" pageId="31" pageNumber="32">
<heading id="D0EA81F25B332A0FFF0B3B5C640BFE9B" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B969EA8C5B332A0FFF0B3B5C640BFE9B" bold="true" box="[189,324,265,291]" pageId="31" pageNumber="32">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3B626070FECD" blockId="31.[189,1399,311,373]" pageId="31" pageNumber="32">The species is named after Dr Daniel Leduc in recognition of his contribution in collecting Hikurangi Margin kinorhynchs and making them available for description to the authors of the present study.</paragraph>
</subSubSection>
<subSubSection id="C30765155B332A0FFF0B3BC9644CFC11" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BA2369E5B332A0FFF0B3BC964E5FE0E" blockId="31.[189,426,412,438]" box="[189,426,412,438]" pageId="31" pageNumber="32">
<heading id="D0EA81F25B332A0FFF0B3BC964E5FE0E" bold="true" box="[189,426,412,438]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B969EA8C5B332A0FFF0B3BC964E5FE0E" bold="true" box="[189,426,412,438]" pageId="31" pageNumber="32">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B753CC35B332A0FFF5C3B996465FDE8" ID-GBIF-Occurrence="3926367301" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="730" latitude="-41.4962" location="Pahaua Canyon" longLatPrecision="6" longitude="175.6828" pageId="31" pageNumber="32" specimenCode="NIWA-159405" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="8BA2369E5B332A0FFF5C3B996413FE5E" blockId="31.[189,1399,460,592]" box="[234,348,460,486]" pageId="31" pageNumber="32">
<emphasis id="B969EA8C5B332A0FFF5C3B996413FE5E" bold="true" box="[234,348,460,486]" pageId="31" pageNumber="32">
<typeStatus id="54A6883C5B332A0FFF5C3B996413FE5E" box="[234,348,460,486]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3BA56465FDE8" blockId="31.[189,1399,460,592]" pageId="31" pageNumber="32">
<collectingCountry id="F30A760E5B332A0FFF0B3BA564DDFDB1" box="[189,402,495,522]" name="New Zealand" pageId="31" pageNumber="32">NEW ZEALAND</collectingCountry>
<specimenCount id="9D1BFD175B332A0FFE1B3BA56489FDB2" box="[429,454,496,522]" pageId="31" pageNumber="32" type="male"></specimenCount>
;
<location id="8EC260455B332A0FFE623BBA67C0FDB1" LSID="urn:lsid:plazi:treatment:03B487885B322A35FDC33C256674FB1F:8EC260455B332A0FFE623BBA67C0FDB1" box="[468,655,495,521]" country="New Zealand" latitude="-41.4962" longLatPrecision="6" longitude="175.6828" name="Pahaua Canyon" pageId="31" pageNumber="32">Pahaua Canyon</location>
, stn TAN1004/31;
<geoCoordinate id="EE2950595B332A0FFCC63BBA66B9FDB1" box="[880,1014,495,521]" degrees="41.4962" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="5" value="-41.4962">41.4962° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B332A0FFBB23BBA61D4FDB2" box="[1028,1179,495,522]" degrees="175.6828" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="175.6828">175.6828° E</geoCoordinate>
;
<elevation id="0030D1AD5B332A0FFB1E3BBA6079FDB1" box="[1192,1334,495,522]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="31" pageNumber="32" unit="m" value="730.0">
<quantity id="4CE59B7B5B332A0FFB1E3BBA61BBFDB2" box="[1192,1268,495,522]" metricMagnitude="2" metricUnit="m" metricValue="7.3" pageId="31" pageNumber="32" unit="m" value="730.0">730 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B332A0FFAF53BBA65B4FD95" pageId="31" pageNumber="32" value="2010-04">
<collectingDate id="EFE7E9B65B332A0FFAF53BBA65B4FD95" pageId="31" pageNumber="32" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCode id="DBBB9EE55B332A0FFD5B384666EFFD96" box="[749,928,531,558]" pageId="31" pageNumber="32">NIWA-159405</specimenCode>
. Mounted for LM in Fluoromount G on HS slide.
</paragraph>
</materialsCitation>
<paragraph id="8BA2369E5B332A0FFF5C382D6413FD2A" blockId="31.[188,1398,632,730]" box="[234,348,632,658]" pageId="31" pageNumber="32">
<materialsCitation id="3B753CC35B332A0FFF5C382D6413FD2A" ID-GBIF-Occurrence="3926367302" box="[234,348,632,658]" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="670" latitude="-41.408" location="Honeycomb Canyon" longLatPrecision="6" longitude="175.8977" pageId="31" pageNumber="32" specimenCode="NIWA-159406" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B969EA8C5B332A0FFF5C382D6413FD2A" bold="true" box="[234,348,632,658]" pageId="31" pageNumber="32">
<typeStatus id="54A6883C5B332A0FFF5C382D6413FD2A" box="[234,348,632,658]" pageId="31" pageNumber="32" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B38C96190FD61" blockId="31.[188,1398,632,730]" pageId="31" pageNumber="32">
<collectingCountry id="F30A760E5B332A0FFF0B38C964DFFD0E" box="[189,400,668,694]" name="New Zealand" pageId="31" pageNumber="32">NEW ZEALAND</collectingCountry>
<specimenCount id="9D1BFD175B332A0FFE1E38C9649AFD0E" box="[424,469,668,694]" pageId="31" pageNumber="32" type="female">1 ♀</specimenCount>
;
<location id="8EC260455B332A0FFE5538C96798FD0E" LSID="urn:lsid:plazi:treatment:03B487885B322A35FDC33C256674FB1F:8EC260455B332A0FFE5538C96798FD0E" box="[483,727,668,694]" country="New Zealand" latitude="-41.408" longLatPrecision="6" longitude="175.8977" name="Honeycomb Canyon" pageId="31" pageNumber="32">Honeycomb Canyon</location>
, stn TAN1004/58;
<geoCoordinate id="EE2950595B332A0FFC0238C96177FD0E" box="[948,1080,668,694]" degrees="41.4080" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="5" value="-41.408">41.4080° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B332A0FFBF238CE6198FD0E" box="[1092,1239,667,694]" degrees="175.8977" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="175.8977">175.8977° E</geoCoordinate>
;
<elevation id="0030D1AD5B332A0FFB5038C9603EFD0E" box="[1254,1393,668,694]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="31" pageNumber="32" unit="m" value="670.0">
<quantity id="4CE59B7B5B332A0FFB5038C96060FD0E" box="[1254,1327,668,694]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="31" pageNumber="32" unit="m" value="670.0">670 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B332A0FFF0A38EA647AFD61" box="[188,309,703,729]" pageId="31" pageNumber="32" value="2010-04">
<collectingDate id="EFE7E9B65B332A0FFF0A38EA647AFD61" box="[188,309,703,729]" pageId="31" pageNumber="32" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCode id="DBBB9EE55B332A0FFC9438EA669BFD62" box="[802,980,703,730]" pageId="31" pageNumber="32">NIWA-159406</specimenCode>
. Mounted as holotype.
</paragraph>
<paragraph id="8BA2369E5B332A0FFF5C395464AEFCA3" blockId="31.[188,1398,769,937]" box="[234,481,769,795]" pageId="31" pageNumber="32">
<emphasis id="B969EA8C5B332A0FFF5C395464AEFCA3" bold="true" box="[234,481,769,795]" pageId="31" pageNumber="32">Additional material</emphasis>
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3970644CFC11" blockId="31.[188,1398,769,937]" pageId="31" pageNumber="32">
<materialsCitation id="3B753CC35B332A0FFF0B397065B7FC3E" ID-GBIF-Occurrence="3926364306" collectingDate="2010-04" collectionCode="NIWA, SEM" country="New Zealand" elevation="1013" latitude="-41.4983" location="Pahaua Canyon" longLatPrecision="6" longitude="175.7043" pageId="31" pageNumber="32" specimenCode="TAN1004" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F30A760E5B332A0FFF0B397064DCFC87" box="[189,403,805,831]" name="New Zealand" pageId="31" pageNumber="32">NEW ZEALAND</collectingCountry>
<specimenCount id="9D1BFD175B332A0FFE07397064ACFC87" box="[433,483,805,831]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
;
<location id="8EC260455B332A0FFE47397167E1FC87" LSID="urn:lsid:plazi:treatment:03B487885B322A35FDC33C256674FB1F:8EC260455B332A0FFE47397167E1FC87" box="[497,686,804,831]" country="New Zealand" latitude="-41.4983" longLatPrecision="6" longitude="175.7043" name="Pahaua Canyon" pageId="31" pageNumber="32">Pahaua Canyon</location>
, stn TAN1004/27;
<geoCoordinate id="EE2950595B332A0FFC2239706155FC87" box="[916,1050,805,831]" degrees="41.4983" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="5" value="-41.4983">41.4983° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B332A0FFB9C3971618DFC87" box="[1066,1218,804,831]" degrees="175.7043" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="175.7043">175.7043° E</geoCoordinate>
;
<elevation id="0030D1AD5B332A0FFB673970603EFC86" box="[1233,1393,804,831]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="31" pageNumber="32" unit="m" value="1013.0">
<quantity id="4CE59B7B5B332A0FFB6739706062FC87" box="[1233,1325,805,831]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="31" pageNumber="32" unit="m" value="1013.0">1013 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B332A0FFF0B391D647AFCDA" box="[189,309,840,866]" pageId="31" pageNumber="32" value="2010-04">
<collectingDate id="EFE7E9B65B332A0FFF0B391D647AFCDA" box="[189,309,840,866]" pageId="31" pageNumber="32" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment; personal reference collection of MVS. Mounted for SEM
</materialsCitation>
<materialsCitation id="3B753CC35B332A0FFEA3393965B0FC11" ID-GBIF-Occurrence="3926364329" collectingDate="2010-04" country="New Zealand" elevation="670" latitude="-41.408" location="Honeycomb Canyon" longLatPrecision="6" longitude="175.8977" pageId="31" pageNumber="32" specimenCount="1" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D1BFD175B332A0FFEA339396409FC3E" box="[277,326,876,902]" pageId="31" pageNumber="32" type="male">1 ♂</specimenCount>
; same collection data as for paratype; personal reference collection of MVS. Mounted for SEM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B332A34FF0B3984667AFB34" lastPageId="36" lastPageNumber="37" pageId="31" pageNumber="32" type="description">
<paragraph id="8BA2369E5B332A0FFF0B39846401FC53" blockId="31.[189,334,977,1003]" box="[189,334,977,1003]" pageId="31" pageNumber="32">
<heading id="D0EA81F25B332A0FFF0B39846401FC53" bold="true" box="[189,334,977,1003]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B969EA8C5B332A0FFF0B39846401FC53" bold="true" box="[189,334,977,1003]" pageId="31" pageNumber="32">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3E5564FEFB3D" blockId="31.[188,1399,1024,1157]" pageId="31" pageNumber="32">
GENERAL. Adults with head, neck and eleven trunk segments (
<figureCitation id="13262A1B5B332A0FFC233E556152FBA2" box="[917,1053,1024,1050]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="32.[189,232,1927,1953]" captionStartId-1="34.[190,233,1857,1883]" captionStartId-2="35.[188,231,1855,1881]" captionTargetBox-0="[217,1368,271,1892]" captionTargetBox-1="[196,1394,587,1819]" captionTargetBox-2="[190,1396,857,1822]" captionTargetId-0="figure-39@32.[216,1371,267,1892]" captionTargetId-1="figure-92@34.[194,1404,586,1820]" captionTargetId-2="figure-0@35.[188,1398,856,1832]" captionTargetPageId-0="32" captionTargetPageId-1="34" captionTargetPageId-2="35" captionText-0="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." captionText-2="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222181" figureDoi-1="http://doi.org/10.5281/zenodo.7222183" figureDoi-2="http://doi.org/10.5281/zenodo.7222185" httpUri-0="https://zenodo.org/record/7222181/files/figure.png" httpUri-1="https://zenodo.org/record/7222183/files/figure.png" httpUri-2="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">Figs 1113</figureCitation>
). Overview of measurements and dimensions in
<tableCitation id="C69F03255B332A0FFE2E3E7664BFFB86" box="[408,496,1059,1086]" captionStart="Table 8" captionStartId="33.[189,258,265,291]" captionTargetPageId="33" captionText="Table 8. Measurements from light microscopy ofEchinoderes leduci sp. nov. (in µm) from the Hikurangi Margin." pageId="31" pageNumber="32">Table 8</tableCitation>
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, summarized in
<tableCitation id="C69F03255B332A0FFDF03E1267EFFBD9" box="[582,672,1095,1121]" captionStart="Table 9" captionStartId="35.[189,258,265,291]" captionTargetPageId="35" captionText="Table 9. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes leduci sp. nov." pageId="31" pageNumber="32">Table 9</tableCitation>
. Head morphology could not be examined in detail in any of available specimens.
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3EF86038FB52" blockId="31.[189,1399,1196,1259]" pageId="31" pageNumber="32">
NECK. Consists of 16 placids. Midventral placid broadest, 12 µm in width and 14 µm in length, whereas all others narrower, measuring 7 µm in width at their bases (
<figureCitation id="13262A1B5B332A0FFCC53E856687FB52" box="[883,968,1232,1258]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="31" pageNumber="32">Fig. 11</figureCitation>
). Trichoscalid plates well developed.
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3F4761D9FA02" blockId="31.[188,1399,1298,1466]" pageId="31" pageNumber="32">
SEGMENT 1. Consists of complete cuticular ring. Sensory spots present in subdorsal, laterodorsal and ventromedial positions. Sensory spots relatively large with marginal hairs, located on anterior half of segment (
<figureCitation id="13262A1B5B332A0FFE9B3F0C64F2FACB" box="[301,445,1369,1395]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="31" pageNumber="32">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B332A0FFE7F3F0C6751FACB" box="[457,542,1369,1395]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="31" pageNumber="32">12BC</figureCitation>
,
<figureCitation id="13262A1B5B332A0FFD9F3F0C67CEFACB" box="[553,641,1369,1395]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">13BC</figureCitation>
). Glandular cell outlet
<typeStatus id="54A6883C5B332A0FFC3F3F0C66F3FACB" box="[905,956,1369,1395]" pageId="31" pageNumber="32">type</typeStatus>
1 present in middorsal position and in ventrolateral positions. Cuticular hairs or perforation sites not present. Posterior segment margin almost straight, forming pectinate fringe with short, sawtooth-like fringe tips (
<figureCitation id="13262A1B5B332A0FFC4D3FF561C6FA02" box="[1019,1161,1440,1466]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">Fig. 13BC</figureCitation>
).
</paragraph>
<paragraph id="8BA2369E5B332A0FFF0B3FB76685F884" blockId="31.[186,1400,1506,1852]" pageId="31" pageNumber="32">
SEGMENT 2. Consists of complete cuticular ring, with tubes located in laterodorsal and ventrolateral positions (
<figureCitation id="13262A1B5B332A0FFE823C50648CF9A7" box="[308,451,1541,1567]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="31" pageNumber="32">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B332A0FFE7B3C50676DF9A7" box="[461,546,1541,1567]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="31" pageNumber="32">12BC</figureCitation>
,
<figureCitation id="13262A1B5B332A0FFD9A3C5067CCF9A7" box="[556,643,1541,1567]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">13BC</figureCitation>
). Sensory spots of similar sizes as on preceding segment with long marginal hair, present in middorsal, laterodorsal and ventromedial positions. Glandular cell outlet
<typeStatus id="54A6883C5B332A0FFA9B3C7C602FF9FB" box="[1325,1376,1577,1603]" pageId="31" pageNumber="32">type</typeStatus>
1 present in middorsal position and in ventromedial positions. Pachycyclus of anterior segment margin of regular thickness, interrupted in middorsal position. Secondary pectinate fringe present near anterior segment margin of this and following segments, but mostly covered by preceding segment. This and following nine segments completely hairless. Cuticular hairs reduced to minute scales distributed around segment (
<figureCitation id="13262A1B5B332A0FFE303C8E6759F94D" box="[390,534,1755,1781]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">Fig. 13DG</figureCitation>
), emerging through perforation sites; perforation sites easily visible in LM (
<figureCitation id="13262A1B5B332A0FFF733CAB6419F8A0" box="[197,342,1790,1816]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="31" pageNumber="32">Fig. 12AH</figureCitation>
). Posterior segment margin almost straight (
<figureCitation id="13262A1B5B332A0FFCE33CAB66F0F8A0" box="[853,959,1790,1816]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="31" pageNumber="32">Fig. 13C</figureCitation>
); pectinate fringe tips as on preceding segment, except midventral area with slightly narrower fringe tips.
</paragraph>
<paragraph id="8BA2369E5B332A31FF0B3D3167E6F9B3" blockId="31.[188,1400,1892,2025]" lastBlockId="33.[189,1398,1450,1547]" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
SEGMENT 3. Present segment, and eight remaining ones, consist of one tergal and two sternal plates (
<figureCitation id="13262A1B5B332A0FFF733DD2641AF819" box="[197,341,1927,1953]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="31" pageNumber="32">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B332A0FFED63DD264FAF819" box="[352,437,1927,1953]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="31" pageNumber="32">12C, E</figureCitation>
). Sensory spots present in subdorsal and midlateral positions. Sensory spots on this and following segments slightly smaller than on preceding segments, but still with one long marginal hair. Glandular cell outlets
<typeStatus id="54A6883C5B332A0FFDB63D9A677CF851" box="[512,563,1999,2025]" pageId="31" pageNumber="32">type</typeStatus>
1 present in middorsal and in ventromedial positions. Perforation sites appear as band around segment, interrupted in middorsal and laterodorsal areas and in central part of sternal plate on this and following five segments (
<figureCitation id="13262A1B5B0D2A31FCB23F9866D9FA5F" box="[772,918,1485,1511]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC143F9866B7FA5F" box="[930,1016,1485,1511]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="33" pageNumber="34">12BE</figureCitation>
). Secondary fringe and posterior segment margin as on preceding segment.
</paragraph>
<caption id="DF6266165B0C2A30FF0B3DD26467F850" ID-DOI="http://doi.org/10.5281/zenodo.7222181" ID-Zenodo-Dep="7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="32" pageNumber="33" startId="32.[189,232,1927,1953]" targetBox="[217,1368,271,1892]" targetPageId="32">
<paragraph id="8BA2369E5B0C2A30FF0B3DD26467F850" blockId="32.[189,1398,1927,2024]" pageId="32" pageNumber="33">
<emphasis id="B969EA8C5B0C2A30FF0B3DD26458F819" bold="true" box="[189,279,1927,1953]" pageId="32" pageNumber="33">Fig. 11.</emphasis>
Line art illustrations of
<taxonomicName id="4C1D4D1D5B0C2A30FD863DD2665FF819" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[560,784,1927,1953]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="32" pageNumber="33" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B0C2A30FD863DD2665FF819" box="[560,784,1927,1953]" italics="true" pageId="32" pageNumber="33">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0C2A30FCA13DDD663EF81A" box="[791,881,1928,1954]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B0C2A30FCC13DD266C3F819" bold="true" box="[887,908,1927,1953]" pageId="32" pageNumber="33">A</emphasis>
. ♂, dorsal view.
<emphasis id="B969EA8C5B0C2A30FBE53DD26128F819" bold="true" box="[1107,1127,1927,1953]" pageId="32" pageNumber="33">B</emphasis>
. ♂, ventral view.
<emphasis id="B969EA8C5B0C2A30FA803DD26004F819" bold="true" box="[1334,1355,1927,1953]" pageId="32" pageNumber="33">C</emphasis>
. ♀, segments 1011, dorsal view.
<emphasis id="B969EA8C5B0C2A30FD9A3DFE670EF87D" bold="true" box="[556,577,1963,1989]" pageId="32" pageNumber="33">D</emphasis>
. ♀, segments 1011, ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B0D2A31FF0B3B5C6456FEFE" pageId="33" pageNumber="34" startId="33.[189,258,265,291]" targetBox="[383,1245,360,1403]" targetIsTable="true" targetPageId="33">
<paragraph id="8BA2369E5B0D2A31FF0B3B5C6456FEFE" blockId="33.[187,1398,265,326]" pageId="33" pageNumber="34">
<emphasis id="B969EA8C5B0D2A31FF0B3B5C6452FE9B" bold="true" box="[189,285,265,291]" pageId="33" pageNumber="34">Table 8.</emphasis>
Measurements from light microscopy of
<taxonomicName id="4C1D4D1D5B0D2A31FD403B5C669AFE9B" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[758,981,265,291]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="33" pageNumber="34" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B0D2A31FD403B5C669AFE9B" box="[758,981,265,291]" italics="true" pageId="33" pageNumber="34">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0D2A31FC6C3B5C6179FE9B" box="[986,1078,265,291]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
(in µm) from the Hikurangi Margin.
</paragraph>
</caption>
<paragraph id="8BA2369E5B0D2A31FE353B2C6135FAC3" pageId="33" pageNumber="34">
<table id="F91DC43E5B0DD5EFFEC93B3D6192FAC3" box="[383,1245,360,1403]" gridcols="3" gridrows="27" pageId="33" pageNumber="34">
<tr id="352D34DC5B0DD5EFFEC93B3D6192FE1D" box="[383,1245,360,421]" gridrow="0" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93B3D6747FE1D" box="[383,520,360,421]" gridcol="0" gridrow="0" pageId="33" pageNumber="34">
<emphasis id="B969EA8C5B0D2A31FE353B2C674BFE2B" bold="true" box="[387,516,377,403]" pageId="33" pageNumber="34">Character</emphasis>
</th>
<th id="76FC5DA05B0DD5EFFD103B3D66C7FE1D" box="[678,904,360,421]" gridcol="1" gridrow="0" pageId="33" pageNumber="34">
<emphasis id="B969EA8C5B0D2A31FD683B3D6626FE1D" bold="true" pageId="33" pageNumber="34">Holotype NIWA-159405 (</emphasis>
♂)
</th>
<th id="76FC5DA05B0DD5EFFC4C3B3D6192FE1D" box="[1018,1245,360,421]" gridcol="2" gridrow="0" pageId="33" pageNumber="34">
<emphasis id="B969EA8C5B0D2A31FB843B3D61F2FE1D" bold="true" pageId="33" pageNumber="34">Paratype NIWA-159406 (</emphasis>
♀)
</th>
</tr>
<tr id="352D34DC5B0DD5EFFEC93BE56192FE72" box="[383,1245,432,458]" gridrow="1" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93BE56747FE72" box="[383,520,432,458]" gridcol="0" gridrow="1" pageId="33" pageNumber="34">TL</th>
<td id="76FC5DA05B0DD5EFFD103BE566C7FE72" box="[678,904,432,458]" gridcol="1" gridrow="1" pageId="33" pageNumber="34">212</td>
<td id="76FC5DA05B0DD5EFFC4C3BE56192FE72" box="[1018,1245,432,458]" gridcol="2" gridrow="1" pageId="33" pageNumber="34">234</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93B806192FE57" box="[383,1245,469,495]" gridrow="2" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93B806747FE57" box="[383,520,469,495]" gridcol="0" gridrow="2" pageId="33" pageNumber="34">MSW-6</th>
<td id="76FC5DA05B0DD5EFFD103B8066C7FE57" box="[678,904,469,495]" gridcol="1" gridrow="2" pageId="33" pageNumber="34">60</td>
<td id="76FC5DA05B0DD5EFFC4C3B806192FE57" box="[1018,1245,469,495]" gridcol="2" gridrow="2" pageId="33" pageNumber="34">62</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93BAE6192FDAE" box="[383,1245,507,534]" gridrow="3" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93BAE6747FDAE" box="[383,520,507,534]" gridcol="0" gridrow="3" pageId="33" pageNumber="34">MSW-6/TL</th>
<td id="76FC5DA05B0DD5EFFD103BAE66C7FDAE" box="[678,904,507,534]" gridcol="1" gridrow="3" pageId="33" pageNumber="34">28%</td>
<td id="76FC5DA05B0DD5EFFC4C3BAE6192FDAE" box="[1018,1245,507,534]" gridcol="2" gridrow="3" pageId="33" pageNumber="34">27%</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC938746192FD84" box="[383,1245,545,572]" gridrow="4" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC938746747FD84" box="[383,520,545,572]" gridcol="0" gridrow="4" pageId="33" pageNumber="34">SW-10</th>
<td id="76FC5DA05B0DD5EFFD10387466C7FD84" box="[678,904,545,572]" gridcol="1" gridrow="4" pageId="33" pageNumber="34">45</td>
<td id="76FC5DA05B0DD5EFFC4C38746192FD84" box="[1018,1245,545,572]" gridcol="2" gridrow="4" pageId="33" pageNumber="34">47</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC938126192FDD9" box="[383,1245,583,609]" gridrow="5" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC938126747FDD9" box="[383,520,583,609]" gridcol="0" gridrow="5" pageId="33" pageNumber="34">SW-10/TL</th>
<td id="76FC5DA05B0DD5EFFD10381266C7FDD9" box="[678,904,583,609]" gridcol="1" gridrow="5" pageId="33" pageNumber="34">21%</td>
<td id="76FC5DA05B0DD5EFFC4C38126192FDD9" box="[1018,1245,583,609]" gridcol="2" gridrow="5" pageId="33" pageNumber="34">20%</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC938386192FD30" box="[383,1245,621,648]" gridrow="6" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC938386747FD30" box="[383,520,621,648]" gridcol="0" gridrow="6" pageId="33" pageNumber="34">S1</th>
<td id="76FC5DA05B0DD5EFFD10383866C7FD30" box="[678,904,621,648]" gridcol="1" gridrow="6" pageId="33" pageNumber="34">25</td>
<td id="76FC5DA05B0DD5EFFC4C38386192FD30" box="[1018,1245,621,648]" gridcol="2" gridrow="6" pageId="33" pageNumber="34">26</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC938C76192FD15" box="[383,1245,658,685]" gridrow="7" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC938C76747FD15" box="[383,520,658,685]" gridcol="0" gridrow="7" pageId="33" pageNumber="34">S2</th>
<td id="76FC5DA05B0DD5EFFD1038C766C7FD15" box="[678,904,658,685]" gridcol="1" gridrow="7" pageId="33" pageNumber="34">23</td>
<td id="76FC5DA05B0DD5EFFC4C38C76192FD15" box="[1018,1245,658,685]" gridcol="2" gridrow="7" pageId="33" pageNumber="34">25</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC938ED6192FD6A" box="[383,1245,696,722]" gridrow="8" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC938ED6747FD6A" box="[383,520,696,722]" gridcol="0" gridrow="8" pageId="33" pageNumber="34">S3</th>
<td id="76FC5DA05B0DD5EFFD1038ED66C7FD6A" box="[678,904,696,722]" gridcol="1" gridrow="8" pageId="33" pageNumber="34">28</td>
<td id="76FC5DA05B0DD5EFFC4C38ED6192FD6A" box="[1018,1245,696,722]" gridcol="2" gridrow="8" pageId="33" pageNumber="34">28</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC9388B6192FD40" box="[383,1245,734,760]" gridrow="9" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC9388B6747FD40" box="[383,520,734,760]" gridcol="0" gridrow="9" pageId="33" pageNumber="34">S4</th>
<td id="76FC5DA05B0DD5EFFD10388B66C7FD40" box="[678,904,734,760]" gridcol="1" gridrow="9" pageId="33" pageNumber="34">28</td>
<td id="76FC5DA05B0DD5EFFC4C388B6192FD40" box="[1018,1245,734,760]" gridcol="2" gridrow="9" pageId="33" pageNumber="34">29</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC939516192FCA7" box="[383,1245,772,799]" gridrow="10" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC939516747FCA7" box="[383,520,772,799]" gridcol="0" gridrow="10" pageId="33" pageNumber="34">S5</th>
<td id="76FC5DA05B0DD5EFFD10395166C7FCA7" box="[678,904,772,799]" gridcol="1" gridrow="10" pageId="33" pageNumber="34">28</td>
<td id="76FC5DA05B0DD5EFFC4C39516192FCA7" box="[1018,1245,772,799]" gridcol="2" gridrow="10" pageId="33" pageNumber="34">31</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC9397F6192FCFC" box="[383,1245,810,836]" gridrow="11" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC9397F6747FCFC" box="[383,520,810,836]" gridcol="0" gridrow="11" pageId="33" pageNumber="34">S6</th>
<td id="76FC5DA05B0DD5EFFD10397F66C7FCFC" box="[678,904,810,836]" gridcol="1" gridrow="11" pageId="33" pageNumber="34">32</td>
<td id="76FC5DA05B0DD5EFFC4C397F6192FCFC" box="[1018,1245,810,836]" gridcol="2" gridrow="11" pageId="33" pageNumber="34">33</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC9391A6192FCD2" box="[383,1245,847,874]" gridrow="12" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC9391A6747FCD2" box="[383,520,847,874]" gridcol="0" gridrow="12" pageId="33" pageNumber="34">S7</th>
<td id="76FC5DA05B0DD5EFFD10391A66C7FCD2" box="[678,904,847,874]" gridcol="1" gridrow="12" pageId="33" pageNumber="34">32</td>
<td id="76FC5DA05B0DD5EFFC4C391A6192FCD2" box="[1018,1245,847,874]" gridcol="2" gridrow="12" pageId="33" pageNumber="34">36</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC939206192FC28" box="[383,1245,885,912]" gridrow="13" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC939206747FC28" box="[383,520,885,912]" gridcol="0" gridrow="13" pageId="33" pageNumber="34">S8</th>
<td id="76FC5DA05B0DD5EFFD10392066C7FC28" box="[678,904,885,912]" gridcol="1" gridrow="13" pageId="33" pageNumber="34">35</td>
<td id="76FC5DA05B0DD5EFFC4C39206192FC28" box="[1018,1245,885,912]" gridcol="2" gridrow="13" pageId="33" pageNumber="34">37</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC939CE6192FC0D" box="[383,1245,923,949]" gridrow="14" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC939CE6747FC0D" box="[383,520,923,949]" gridcol="0" gridrow="14" pageId="33" pageNumber="34">S9</th>
<td id="76FC5DA05B0DD5EFFD1039CE66C7FC0D" box="[678,904,923,949]" gridcol="1" gridrow="14" pageId="33" pageNumber="34">40</td>
<td id="76FC5DA05B0DD5EFFC4C39CE6192FC0D" box="[1018,1245,923,949]" gridcol="2" gridrow="14" pageId="33" pageNumber="34">39</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC939946192FC63" box="[383,1245,961,987]" gridrow="15" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC939946747FC63" box="[383,520,961,987]" gridcol="0" gridrow="15" pageId="33" pageNumber="34">S10</th>
<td id="76FC5DA05B0DD5EFFD10399466C7FC63" box="[678,904,961,987]" gridcol="1" gridrow="15" pageId="33" pageNumber="34">40</td>
<td id="76FC5DA05B0DD5EFFC4C39946192FC63" box="[1018,1245,961,987]" gridcol="2" gridrow="15" pageId="33" pageNumber="34">38</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC939B26192FBB9" box="[383,1245,999,1025]" gridrow="16" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC939B26747FBB9" box="[383,520,999,1025]" gridcol="0" gridrow="16" pageId="33" pageNumber="34">S11</th>
<td id="76FC5DA05B0DD5EFFD1039B266C7FBB9" box="[678,904,999,1025]" gridcol="1" gridrow="16" pageId="33" pageNumber="34">23</td>
<td id="76FC5DA05B0DD5EFFC4C39B26192FBB9" box="[1018,1245,999,1025]" gridcol="2" gridrow="16" pageId="33" pageNumber="34">24</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93E596192FB9F" box="[383,1245,1036,1063]" gridrow="17" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93E596747FB9F" box="[383,520,1036,1063]" gridcol="0" gridrow="17" pageId="33" pageNumber="34">MD4 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103E5966C7FB9F" box="[678,904,1036,1063]" gridcol="1" gridrow="17" pageId="33" pageNumber="34">47</td>
<td id="76FC5DA05B0DD5EFFC4C3E596192FB9F" box="[1018,1245,1036,1063]" gridcol="2" gridrow="17" pageId="33" pageNumber="34">52</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93E676192FBF4" box="[383,1245,1074,1100]" gridrow="18" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93E676747FBF4" box="[383,520,1074,1100]" gridcol="0" gridrow="18" pageId="33" pageNumber="34">MD6 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103E6766C7FBF4" box="[678,904,1074,1100]" gridcol="1" gridrow="18" pageId="33" pageNumber="34">78</td>
<td id="76FC5DA05B0DD5EFFC4C3E676192FBF4" box="[1018,1245,1074,1100]" gridcol="2" gridrow="18" pageId="33" pageNumber="34">78</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93E0D6192FBCA" box="[383,1245,1112,1138]" gridrow="19" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93E0D6747FBCA" box="[383,520,1112,1138]" gridcol="0" gridrow="19" pageId="33" pageNumber="34">MD8 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103E0D66C7FBCA" box="[678,904,1112,1138]" gridcol="1" gridrow="19" pageId="33" pageNumber="34">82</td>
<td id="76FC5DA05B0DD5EFFC4C3E0D6192FBCA" box="[1018,1245,1112,1138]" gridcol="2" gridrow="19" pageId="33" pageNumber="34">86</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93E2B6192FB20" box="[383,1245,1150,1176]" gridrow="20" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93E2B6747FB20" box="[383,520,1150,1176]" gridcol="0" gridrow="20" pageId="33" pageNumber="34">LV6 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103E2B66C7FB20" box="[678,904,1150,1176]" gridcol="1" gridrow="20" pageId="33" pageNumber="34">27</td>
<td id="76FC5DA05B0DD5EFFC4C3E2B6192FB20" box="[1018,1245,1150,1176]" gridcol="2" gridrow="20" pageId="33" pageNumber="34">28</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93EF16192FB06" box="[383,1245,1188,1214]" gridrow="21" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93EF16747FB06" box="[383,520,1188,1214]" gridcol="0" gridrow="21" pageId="33" pageNumber="34">LV7 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103EF166C7FB06" box="[678,904,1188,1214]" gridcol="1" gridrow="21" pageId="33" pageNumber="34">30</td>
<td id="76FC5DA05B0DD5EFFC4C3EF16192FB06" box="[1018,1245,1188,1214]" gridcol="2" gridrow="21" pageId="33" pageNumber="34">29</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93E9C6192FB5C" box="[383,1245,1225,1252]" gridrow="22" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93E9C6747FB5C" box="[383,520,1225,1252]" gridcol="0" gridrow="22" pageId="33" pageNumber="34">LV8 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103E9C66C7FB5C" box="[678,904,1225,1252]" gridcol="1" gridrow="22" pageId="33" pageNumber="34">37</td>
<td id="76FC5DA05B0DD5EFFC4C3E9C6192FB5C" box="[1018,1245,1225,1252]" gridcol="2" gridrow="22" pageId="33" pageNumber="34">35</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93EBA6192FAB2" box="[383,1245,1263,1290]" gridrow="23" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93EBA6747FAB2" box="[383,520,1263,1290]" gridcol="0" gridrow="23" pageId="33" pageNumber="34">LV9 (ac)</th>
<td id="76FC5DA05B0DD5EFFD103EBA66C7FAB2" box="[678,904,1263,1290]" gridcol="1" gridrow="23" pageId="33" pageNumber="34">32</td>
<td id="76FC5DA05B0DD5EFFC4C3EBA6192FAB2" box="[1018,1245,1263,1290]" gridcol="2" gridrow="23" pageId="33" pageNumber="34">35</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93F406192FA97" box="[383,1245,1301,1327]" gridrow="24" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93F406747FA97" box="[383,520,1301,1327]" gridcol="0" gridrow="24" pageId="33" pageNumber="34">LTS</th>
<td id="76FC5DA05B0DD5EFFD103F4066C7FA97" box="[678,904,1301,1327]" gridcol="1" gridrow="24" pageId="33" pageNumber="34">172</td>
<td id="76FC5DA05B0DD5EFFC4C3F406192FA97" box="[1018,1245,1301,1327]" gridcol="2" gridrow="24" pageId="33" pageNumber="34">163</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93F6E6192FAED" box="[383,1245,1339,1365]" gridrow="25" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93F6E6747FAED" box="[383,520,1339,1365]" gridcol="0" gridrow="25" pageId="33" pageNumber="34">LTS/TL</th>
<td id="76FC5DA05B0DD5EFFD103F6E66C7FAED" box="[678,904,1339,1365]" gridcol="1" gridrow="25" pageId="33" pageNumber="34">81%</td>
<td id="76FC5DA05B0DD5EFFC4C3F6E6192FAED" box="[1018,1245,1339,1365]" gridcol="2" gridrow="25" pageId="33" pageNumber="34">70%</td>
</tr>
<tr id="352D34DC5B0DD5EFFEC93F346192FAC3" box="[383,1245,1377,1403]" gridrow="26" pageId="33" pageNumber="34">
<th id="76FC5DA05B0DD5EFFEC93F346747FAC3" box="[383,520,1377,1403]" gridcol="0" gridrow="26" pageId="33" pageNumber="34">LTAS</th>
<td id="76FC5DA05B0DD5EFFD103F3466C7FAC3" box="[678,904,1377,1403]" gridcol="1" gridrow="26" pageId="33" pageNumber="34">n/a</td>
<td id="76FC5DA05B0DD5EFFC4C3F346192FAC3" box="[1018,1245,1377,1403]" gridcol="2" gridrow="26" pageId="33" pageNumber="34">27</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA2369E5B0D2A31FF0B3C676795F90E" blockId="33.[188,1400,1586,1718]" pageId="33" pageNumber="34">
SEGMENT 4. With spine in middorsal position (
<figureCitation id="13262A1B5B0D2A31FD663C676671F9F4" box="[720,830,1586,1612]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Figs 11A</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FCFC3C676633F9F4" box="[842,892,1586,1612]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="33" pageNumber="34">12B</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC3E3C6766F1F9F4" box="[904,958,1586,1612]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="33" pageNumber="34">13D</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B0D2A31FB623C676048F9F4" box="[1236,1287,1586,1612]" pageId="33" pageNumber="34">type</typeStatus>
1 present in paradorsal and ventromedial positions. No other traits observed. Posterior segment margin straight, terminating in pectinate fringe with slightly slenderer fringe tips along ventral margin than on preceding segments, otherwise as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B0D2A31FF0B3C8867E7F8D9" blockId="33.[188,1399,1757,1889]" pageId="33" pageNumber="34">
SEGMENT 5. With tubes in lateroventral positions (
<figureCitation id="13262A1B5B0D2A31FC9B3C8866EEF94F" box="[813,929,1757,1783]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Figs 11B</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC023C8866A8F94F" box="[948,999,1757,1783]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="33" pageNumber="34">12C</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC4C3C886164F94F" box="[1018,1067,1757,1783]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="33" pageNumber="34">13F</figureCitation>
). Sensory spots present in subdorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B0D2A31FCBD3D5566D2F8A2" box="[779,925,1792,1818]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC1C3D55614FF8A2" box="[938,1024,1792,1818]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="33" pageNumber="34">12BC</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FBBB3D55612CF8A2" box="[1037,1123,1792,1818]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="33" pageNumber="34">13D, F</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B0D2A31FF083D7165BEF886" box="[190,241,1828,1854]" pageId="33" pageNumber="34">type</typeStatus>
1 present in middorsal and ventromedial positions. Perforation sites, secondary fringe and posterior segment margin as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B0D2A31FF0B3DDD61C2F851" blockId="33.[189,1399,1928,2025]" pageId="33" pageNumber="34">
SEGMENT 6. With spines in middorsal and lateroventral positions (
<figureCitation id="13262A1B5B0D2A31FC0E3DDD6108F81A" box="[952,1095,1928,1954]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Fig. 11AB</figureCitation>
). Sensory spots present in paradorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B0D2A31FCAE3DFE66E5F87D" box="[792,938,1963,1989]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="33" pageNumber="34">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B0D2A31FC0E3DFE6140F87D" box="[952,1039,1963,1989]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="33" pageNumber="34">13D, F</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B0D2A31FA9A3DF96010F87E" box="[1324,1375,1964,1990]" pageId="33" pageNumber="34">type</typeStatus>
1 present in paradorsal and ventromedial positions. Segment otherwise as segment 5.
</paragraph>
<paragraph id="8BA2369E5B0E2A32FF0B3B5C6177FE34" blockId="34.[189,1399,265,397]" pageId="34" pageNumber="35">
SEGMENT 7. With spines in lateroventral positions, and sensory spots in paradorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B0E2A32FE503B796731FEFE" box="[486,638,300,326]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="34" pageNumber="35">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B0E2A32FD273B79678DFEFE" box="[657,706,300,326]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="34" pageNumber="35">13F</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B0E2A32FC413B786165FEFF" box="[1015,1066,301,327]" pageId="34" pageNumber="35">type</typeStatus>
1 present in ventromedial positions only; no glands observed on dorsal side. Tips of pectinate fringe of posterior segment margin more slender than on preceding segments. Segment otherwise as segment 6.
</paragraph>
<paragraph id="8BA2369E5B0E2A32FF0B3BE16037FE49" blockId="34.[189,1400,436,498]" pageId="34" pageNumber="35">
SEGMENT 8. With spines in middorsal and lateroventral positions, and relatively long tubes (21 µm) in sublateral positions (
<figureCitation id="13262A1B5B0E2A32FE053B82670AFE49" box="[435,581,471,497]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="34" pageNumber="35">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B0E2A32FDE53B8267E7FE49" box="[595,680,471,497]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="34" pageNumber="35">12DE</figureCitation>
,
<figureCitation id="13262A1B5B0E2A32FD033B826642FE49" box="[693,781,471,497]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="34" pageNumber="35">13E, G</figureCitation>
). Sensory spots present in paradorsal positions only.
</paragraph>
<caption id="DF6266165B0E2A32FF083D1461F1F850" ID-DOI="http://doi.org/10.5281/zenodo.7222183" ID-Zenodo-Dep="7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="34" pageNumber="35" startId="34.[190,233,1857,1883]" targetBox="[196,1394,587,1819]" targetPageId="34">
<paragraph id="8BA2369E5B0E2A32FF083D1461F1F850" blockId="34.[189,1399,1857,2025]" pageId="34" pageNumber="35">
<emphasis id="B969EA8C5B0E2A32FF083D146450F8E3" bold="true" box="[190,287,1857,1883]" pageId="34" pageNumber="35">Fig. 12.</emphasis>
Light micrographs showing overview and details of
<taxonomicName id="4C1D4D1D5B0E2A32FC193D1761DAF8E4" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[943,1173,1858,1884]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="34" pageNumber="35" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B0E2A32FC193D1761DAF8E4" box="[943,1173,1858,1884]" italics="true" pageId="34" pageNumber="35">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0E2A32FB163D17604FF8E4" box="[1184,1280,1858,1884]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B0E2A32FABD3D146009F8E3" bold="true" box="[1291,1350,1857,1883]" pageId="34" pageNumber="35">AG</emphasis>
. ♂, holotype (NIWA-159405).
<emphasis id="B969EA8C5B0E2A32FDB43D306758F8C7" bold="true" box="[514,535,1893,1919]" pageId="34" pageNumber="35">H</emphasis>
. ♀, paratype (NIWA-159406).
<emphasis id="B969EA8C5B0E2A32FC393D3066EBF8C7" bold="true" box="[911,932,1893,1919]" pageId="34" pageNumber="35">A</emphasis>
. Dorsal overview.
<emphasis id="B969EA8C5B0E2A32FB3C3D3061D3F8C7" bold="true" box="[1162,1180,1893,1919]" pageId="34" pageNumber="35">B</emphasis>
. Segments 1 to 6, dorsal view.
<emphasis id="B969EA8C5B0E2A32FEE33DDD6426F81A" bold="true" box="[341,361,1928,1954]" pageId="34" pageNumber="35">C</emphasis>
. Segments 1 to 6, ventral view.
<emphasis id="B969EA8C5B0E2A32FD5F3DDD67B2F81A" bold="true" box="[745,765,1928,1954]" pageId="34" pageNumber="35">D</emphasis>
. Segments 6 to 11, dorsal view.
<emphasis id="B969EA8C5B0E2A32FB363DDD61DCF81A" bold="true" box="[1152,1171,1928,1954]" pageId="34" pageNumber="35">E</emphasis>
. Segments 6 to 11, ventral view.
<emphasis id="B969EA8C5B0E2A32FED73DFE643CF87D" bold="true" box="[353,371,1963,1989]" pageId="34" pageNumber="35">F</emphasis>
. Segments 9 to 11 of male, dorsal view.
<emphasis id="B969EA8C5B0E2A32FCD13DFE6631F87D" bold="true" box="[871,894,1963,1989]" pageId="34" pageNumber="35">G</emphasis>
. Segments 10 to 11 of male, dorsal view.
<emphasis id="B969EA8C5B0E2A32FF0B3D9A659BF851" bold="true" box="[189,212,1999,2025]" pageId="34" pageNumber="35">H</emphasis>
. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B0F2A33FF0B3B5C67B5FEFF" pageId="35" pageNumber="36" startId="35.[189,258,265,291]" targetBox="[206,1375,374,804]" targetIsTable="true" targetPageId="35">
<paragraph id="8BA2369E5B0F2A33FF0B3B5C67B5FEFF" blockId="35.[189,1398,265,327]" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FF0B3B5C646CFE9B" bold="true" box="[189,291,265,291]" pageId="35" pageNumber="36">Table 9.</emphasis>
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
<taxonomicName id="4C1D4D1D5B0F2A33FE0E3B7967D6FEFE" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[440,665,300,326]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="35" pageNumber="36" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B0F2A33FE0E3B7967D6FEFE" box="[440,665,300,326]" italics="true" pageId="35" pageNumber="36">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0F2A33FD163B7867B5FEFF" box="[672,762,301,327]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA2369E5B0F2A33FF783B2361DEFC9B" pageId="35" pageNumber="36">
<table id="F91DC43E5B0FD5EFFF783B236010FC9C" box="[206,1375,374,804]" gridcols="10" gridrows="12" pageId="35" pageNumber="36">
<tr id="352D34DC5B0FD5EFFF783B236010FE28" box="[206,1375,374,400]" gridrow="0" pageId="35" pageNumber="36">
<th id="76FC5DA05B0FD5EFFF783B2364EEFE28" box="[206,417,374,400]" gridcol="0" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FF783B2364EEFE28" bold="true" box="[206,417,374,400]" pageId="35" pageNumber="36">Position segment</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFE013B236759FE28" box="[439,534,374,400]" gridcol="1" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FE783B2364B0FE28" bold="true" box="[462,511,374,400]" pageId="35" pageNumber="36">MD</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFD863B2367C8FE28" box="[560,647,374,400]" gridcol="2" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FDFE3B236720FE28" bold="true" box="[584,623,374,400]" pageId="35" pageNumber="36">PD</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFD193B23679BFE28" box="[687,724,374,400]" gridcol="3" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FD193B23679BFE28" bold="true" box="[687,724,374,400]" pageId="35" pageNumber="36">SD</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFD443B23667DFE28" box="[754,818,374,400]" gridcol="4" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FCB43B236664FE28" bold="true" box="[770,811,374,400]" pageId="35" pageNumber="36">LD</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFCF13B2366F1FE28" box="[839,958,374,400]" gridcol="5" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FCDA3B2366D4FE28" bold="true" box="[876,923,374,400]" pageId="35" pageNumber="36">ML</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFC643B236115FE28" box="[978,1114,374,400]" gridcol="6" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FC643B236105FE28" bold="true" box="[978,1098,374,400]" pageId="35" pageNumber="36">SL LA</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFBC63B2361D9FE28" box="[1136,1174,374,400]" gridcol="7" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FBC63B2361D9FE28" bold="true" box="[1136,1174,374,400]" pageId="35" pageNumber="36">LV</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFB193B2361A7FE28" box="[1199,1256,374,400]" gridcol="8" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FB0E3B2361AEFE28" bold="true" box="[1208,1249,374,400]" pageId="35" pageNumber="36">VL</emphasis>
</th>
<th id="76FC5DA05B0FD5EFFABE3B236010FE28" box="[1288,1375,374,400]" gridcol="9" gridrow="0" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FAAD3B236003FE28" bold="true" box="[1307,1356,374,400]" pageId="35" pageNumber="36">VM</emphasis>
</th>
</tr>
<tr id="352D34DC5B0FD5EFFF783BF26010FE79" box="[206,1375,423,449]" gridrow="1" pageId="35" pageNumber="36" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76FC5DA05B0FD5EFFF783BF264EEFE79" box="[206,417,423,449]" gridcol="0" gridrow="1" pageId="35" pageNumber="36">1</th>
<td id="76FC5DA05B0FD5EFFE013BF26759FE79" box="[439,534,423,449]" gridcol="1" gridrow="1" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFD193BF2679BFE79" box="[687,724,423,449]" gridcol="3" gridrow="1" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFD443BF2667DFE79" box="[754,818,423,449]" gridcol="4" gridrow="1" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFB193BF261A7FE79" box="[1199,1256,423,449]" gridcol="8" gridrow="1" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFABE3BF26010FE79" box="[1288,1375,423,449]" gridcol="9" gridrow="1" pageId="35" pageNumber="36">ss</td>
</tr>
<tr id="352D34DC5B0FD5EFFF783B9F6010FE5D" box="[206,1375,458,485]" gridrow="2" pageId="35" pageNumber="36" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76FC5DA05B0FD5EFFF783B9F64EEFE5D" box="[206,417,458,485]" gridcol="0" gridrow="2" pageId="35" pageNumber="36">2</th>
<td id="76FC5DA05B0FD5EFFE013B9F6759FE5D" box="[439,534,458,485]" gridcol="1" gridrow="2" pageId="35" pageNumber="36">gco1,ss</td>
<td id="76FC5DA05B0FD5EFFD443B9F667DFE5D" box="[754,818,458,485]" gridcol="4" gridrow="2" pageId="35" pageNumber="36">tu,ss</td>
<td id="76FC5DA05B0FD5EFFB193B9F61A7FE5D" box="[1199,1256,458,485]" gridcol="8" gridrow="2" pageId="35" pageNumber="36">tu</td>
<td id="76FC5DA05B0FD5EFFABE3B9F6010FE5D" box="[1288,1375,458,485]" gridcol="9" gridrow="2" pageId="35" pageNumber="36">gco1,ss</td>
</tr>
<tr id="352D34DC5B0FD5EFFF783BBB6010FDB0" box="[206,1375,494,520]" gridrow="3" pageId="35" pageNumber="36" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF783BBB64EEFDB0" box="[206,417,494,520]" gridcol="0" gridrow="3" pageId="35" pageNumber="36">3</th>
<td id="76FC5DA05B0FD5EFFE013BBB6759FDB0" box="[439,534,494,520]" gridcol="1" gridrow="3" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFD193BBB679BFDB0" box="[687,724,494,520]" gridcol="3" gridrow="3" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFCF13BBB66F1FDB0" box="[839,958,494,520]" gridcol="5" gridrow="3" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFABE3BBB6010FDB0" box="[1288,1375,494,520]" gridcol="9" gridrow="3" pageId="35" pageNumber="36">gco1</td>
</tr>
<tr id="352D34DC5B0FD5EFFF7838446010FD93" box="[206,1375,529,555]" gridrow="4" pageId="35" pageNumber="36" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF78384464EEFD93" box="[206,417,529,555]" gridcol="0" gridrow="4" pageId="35" pageNumber="36">4</th>
<td id="76FC5DA05B0FD5EFFE0138446759FD93" box="[439,534,529,555]" gridcol="1" gridrow="4" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFD86384467C8FD93" box="[560,647,529,555]" gridcol="2" gridrow="4" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFABE38446010FD93" box="[1288,1375,529,555]" gridcol="9" gridrow="4" pageId="35" pageNumber="36">gco1</td>
</tr>
<tr id="352D34DC5B0FD5EFFF7838616010FDF7" box="[206,1375,564,591]" gridrow="5" pageId="35" pageNumber="36" rowspan-2="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF78386164EEFDF7" box="[206,417,564,591]" gridcol="0" gridrow="5" pageId="35" pageNumber="36">5</th>
<td id="76FC5DA05B0FD5EFFE0138616759FDF7" box="[439,534,564,591]" gridcol="1" gridrow="5" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFD193861679BFDF7" box="[687,724,564,591]" gridcol="3" gridrow="5" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFCF1386166F1FDF7" box="[839,958,564,591]" gridcol="5" gridrow="5" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFBC6386161D9FDF7" box="[1136,1174,564,591]" gridcol="7" gridrow="5" pageId="35" pageNumber="36">tu</td>
<td id="76FC5DA05B0FD5EFFABE38616010FDF7" box="[1288,1375,564,591]" gridcol="9" gridrow="5" pageId="35" pageNumber="36">gco1,ss</td>
</tr>
<tr id="352D34DC5B0FD5EFFF78380D6010FDCA" box="[206,1375,600,626]" gridrow="6" pageId="35" pageNumber="36" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF78380D64EEFDCA" box="[206,417,600,626]" gridcol="0" gridrow="6" pageId="35" pageNumber="36">6</th>
<td id="76FC5DA05B0FD5EFFE01380D6759FDCA" box="[439,534,600,626]" gridcol="1" gridrow="6" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFD86380D67C8FDCA" box="[560,647,600,626]" gridcol="2" gridrow="6" pageId="35" pageNumber="36">gco1,ss</td>
<td id="76FC5DA05B0FD5EFFCF1380D66F1FDCA" box="[839,958,600,626]" gridcol="5" gridrow="6" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFBC6380D61D9FDCA" box="[1136,1174,600,626]" gridcol="7" gridrow="6" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFABE380D6010FDCA" box="[1288,1375,600,626]" gridcol="9" gridrow="6" pageId="35" pageNumber="36">gco1,ss</td>
</tr>
<tr id="352D34DC5B0FD5EFFF78382E6010FD2E" box="[206,1375,635,662]" gridrow="7" pageId="35" pageNumber="36" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF78382E64EEFD2E" box="[206,417,635,662]" gridcol="0" gridrow="7" pageId="35" pageNumber="36">7</th>
<td id="76FC5DA05B0FD5EFFD86382E67C8FD2E" box="[560,647,635,662]" gridcol="2" gridrow="7" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFCF1382E66F1FD2E" box="[839,958,635,662]" gridcol="5" gridrow="7" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFBC6382E61D9FD2E" box="[1136,1174,635,662]" gridcol="7" gridrow="7" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFABE382E6010FD2E" box="[1288,1375,635,662]" gridcol="9" gridrow="7" pageId="35" pageNumber="36">gco1,ss</td>
</tr>
<tr id="352D34DC5B0FD5EFFF7838CA6010FD01" box="[206,1375,671,697]" gridrow="8" pageId="35" pageNumber="36" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-8="1">
<th id="76FC5DA05B0FD5EFFF7838CA64EEFD01" box="[206,417,671,697]" gridcol="0" gridrow="8" pageId="35" pageNumber="36">8</th>
<td id="76FC5DA05B0FD5EFFE0138CA6759FD01" box="[439,534,671,697]" gridcol="1" gridrow="8" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFD8638CA67C8FD01" box="[560,647,671,697]" gridcol="2" gridrow="8" pageId="35" pageNumber="36">gco1,ss</td>
<td id="76FC5DA05B0FD5EFFC6438CA6115FD01" box="[978,1114,671,697]" gridcol="6" gridrow="8" pageId="35" pageNumber="36">tu</td>
<td id="76FC5DA05B0FD5EFFBC638CA61D9FD01" box="[1136,1174,671,697]" gridcol="7" gridrow="8" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFABE38CA6010FD01" box="[1288,1375,671,697]" gridcol="9" gridrow="8" pageId="35" pageNumber="36">gco1</td>
</tr>
<tr id="352D34DC5B0FD5EFFF7838976010FD65" box="[206,1375,706,733]" gridrow="9" pageId="35" pageNumber="36" rowspan-1="1" rowspan-6="1">
<th id="76FC5DA05B0FD5EFFF78389764EEFD65" box="[206,417,706,733]" gridcol="0" gridrow="9" pageId="35" pageNumber="36">9</th>
<td id="76FC5DA05B0FD5EFFD86389767C8FD65" box="[560,647,706,733]" gridcol="2" gridrow="9" pageId="35" pageNumber="36">gco1,ss</td>
<td id="76FC5DA05B0FD5EFFD193897679BFD65" box="[687,724,706,733]" gridcol="3" gridrow="9" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFD443897667DFD65" box="[754,818,706,733]" gridcol="4" gridrow="9" pageId="35" pageNumber="36">tu</td>
<td id="76FC5DA05B0FD5EFFCF1389766F1FD65" box="[839,958,706,733]" gridcol="5" gridrow="9" pageId="35" pageNumber="36">si</td>
<td id="76FC5DA05B0FD5EFFBC6389761D9FD65" box="[1136,1174,706,733]" gridcol="7" gridrow="9" pageId="35" pageNumber="36">ac</td>
<td id="76FC5DA05B0FD5EFFB19389761A7FD65" box="[1199,1256,706,733]" gridcol="8" gridrow="9" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFABE38976010FD65" box="[1288,1375,706,733]" gridcol="9" gridrow="9" pageId="35" pageNumber="36">gco1</td>
</tr>
<tr id="352D34DC5B0FD5EFFF7838B36010FCB8" box="[206,1375,742,768]" gridrow="10" pageId="35" pageNumber="36" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76FC5DA05B0FD5EFFF7838B364EEFCB8" box="[206,417,742,768]" gridcol="0" gridrow="10" pageId="35" pageNumber="36">10</th>
<td id="76FC5DA05B0FD5EFFE0138B36759FCB8" box="[439,534,742,768]" gridcol="1" gridrow="10" pageId="35" pageNumber="36">gco1</td>
<td id="76FC5DA05B0FD5EFFD1938B3679BFCB8" box="[687,724,742,768]" gridcol="3" gridrow="10" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFD4438B3667DFCB8" box="[754,818,742,768]" gridcol="4" gridrow="10" pageId="35" pageNumber="36">tu(♂)</td>
<td id="76FC5DA05B0FD5EFFB1938B361A7FCB8" box="[1199,1256,742,768]" gridcol="8" gridrow="10" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFABE38B36010FCB8" box="[1288,1375,742,768]" gridcol="9" gridrow="10" pageId="35" pageNumber="36">gco1</td>
</tr>
<tr id="352D34DC5B0FD5EFFF78395C6010FC9C" box="[206,1375,777,804]" gridrow="11" pageId="35" pageNumber="36" rowspan-2="1" rowspan-4="1" rowspan-8="1" rowspan-9="1">
<th id="76FC5DA05B0FD5EFFF78395C64EEFC9C" box="[206,417,777,804]" gridcol="0" gridrow="11" pageId="35" pageNumber="36">11</th>
<td id="76FC5DA05B0FD5EFFE01395C6759FC9C" box="[439,534,777,804]" gridcol="1" gridrow="11" pageId="35" pageNumber="36">gco1×2</td>
<td id="76FC5DA05B0FD5EFFD19395C679BFC9C" box="[687,724,777,804]" gridcol="3" gridrow="11" pageId="35" pageNumber="36">ss</td>
<td id="76FC5DA05B0FD5EFFCF1395C66F1FC9C" box="[839,958,777,804]" gridcol="5" gridrow="11" pageId="35" pageNumber="36">pe ×3 (♂)</td>
<td id="76FC5DA05B0FD5EFFC64395C6115FC9C" box="[978,1114,777,804]" gridcol="6" gridrow="11" pageId="35" pageNumber="36">ltas(♀)</td>
<td id="76FC5DA05B0FD5EFFBC6395C61D9FC9C" box="[1136,1174,777,804]" gridcol="7" gridrow="11" pageId="35" pageNumber="36">lts</td>
</tr>
</table>
</paragraph>
<caption id="DF6266165B0F2A33FF0A3D6A648AF85E" ID-DOI="http://doi.org/10.5281/zenodo.7222185" ID-Zenodo-Dep="7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="35" pageNumber="36" startId="35.[188,231,1855,1881]" targetBox="[190,1396,857,1822]" targetPageId="35">
<paragraph id="8BA2369E5B0F2A33FF0A3D6A648AF85E" blockId="35.[187,1399,1855,2022]" pageId="35" pageNumber="36">
<emphasis id="B969EA8C5B0F2A33FF0A3D6A6456F8E1" bold="true" box="[188,281,1855,1881]" pageId="35" pageNumber="36">Fig. 13.</emphasis>
Scanning electron micrographs showing overviews and details of
<taxonomicName id="4C1D4D1D5B0F2A33FB843D6A605BF8E1" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1074,1300,1855,1881]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="35" pageNumber="36" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B0F2A33FB843D6A605BF8E1" box="[1074,1300,1855,1881]" italics="true" pageId="35" pageNumber="36">Echinoderes leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0F2A33FAAD3D156038F8E2" box="[1307,1399,1856,1882]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B0F2A33FF0B3D37659EF8C4" bold="true" box="[189,209,1890,1916]" pageId="35" pageNumber="36">A</emphasis>
. Lateroventral overview.
<emphasis id="B969EA8C5B0F2A33FDA53D376768F8C4" bold="true" box="[531,551,1890,1916]" pageId="35" pageNumber="36">B</emphasis>
. Segments 1 to 3, lateral view.
<emphasis id="B969EA8C5B0F2A33FC713D376693F8C4" bold="true" box="[967,988,1890,1916]" pageId="35" pageNumber="36">C</emphasis>
. Segments 1 to 3, ventral view.
<emphasis id="B969EA8C5B0F2A33FF0A3DD3659EF818" bold="true" box="[188,209,1926,1952]" pageId="35" pageNumber="36">D</emphasis>
. Segments 1 to 6, laterodorsal view.
<emphasis id="B969EA8C5B0F2A33FDCD3DD367C0F818" bold="true" box="[635,655,1926,1952]" pageId="35" pageNumber="36">E</emphasis>
. Segments 8 to 9, dorsal view.
<emphasis id="B969EA8C5B0F2A33FC4E3DD36145F818" bold="true" box="[1016,1034,1926,1952]" pageId="35" pageNumber="36">F</emphasis>
. Segments 3 to 7, ventral view.
<emphasis id="B969EA8C5B0F2A33FF0D3DFD659DF87A" bold="true" box="[187,210,1960,1986]" pageId="35" pageNumber="36">G</emphasis>
. Segments 7 to 9, laterodorsal view.
<emphasis id="B969EA8C5B0F2A33FD393DFC67EBF87B" bold="true" box="[655,676,1961,1987]" pageId="35" pageNumber="36">H</emphasis>
. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BA2369E5B082A34FF0B3B5C6470FEFF" blockId="36.[189,1398,265,327]" pageId="36" pageNumber="37">
Glandular cell outlets
<typeStatus id="54A6883C5B082A34FE7E3B5C64B4FE9B" box="[456,507,265,291]" pageId="36" pageNumber="37">type</typeStatus>
1 present in paradorsal and ventromedial positions. Segment otherwise as segment 7.
</paragraph>
<paragraph id="8BA2369E5B082A34FF0B3B3B67E6FD83" blockId="36.[188,1401,366,571]" pageId="36" pageNumber="37">
SEGMENT 9. With spines in lateroventral positions. Long tubes (20 µm) present in laterodorsal positions (
<figureCitation id="13262A1B5B082A34FF733BC76479FE14" box="[197,310,402,428]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11A</figureCitation>
,
<figureCitation id="13262A1B5B082A34FEF53BC764D8FE14" box="[323,407,402,428]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12D, F</figureCitation>
,
<figureCitation id="13262A1B5B082A34FE123BC764B3FE14" box="[420,508,402,428]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13E, G</figureCitation>
). Sensory spots located in paradorsal, subdorsal and ventrolateral positions (
<figureCitation id="13262A1B5B082A34FF703BE06415FE68" box="[198,346,437,464]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B082A34FEDF3BE364F8FE68" box="[361,439,438,464]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12EF</figureCitation>
,
<figureCitation id="13262A1B5B082A34FE733BE36751FE68" box="[453,542,438,464]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13E, G</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B082A34FCF63BE3663CFE68" box="[832,883,438,464]" pageId="36" pageNumber="37">type</typeStatus>
1 present paradorsally and ventromedially. Small sieve plates located in midlateral positions (
<figureCitation id="13262A1B5B082A34FCB43B8C6620FE4B" box="[770,879,473,499]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">Fig. 13G</figureCitation>
). Band of perforation site patches interrupted in subdorsal area instead of laterodorsally as on preceding segments. Secondary fringe and posterior segment margin as on preceding segment.
</paragraph>
<paragraph id="8BA2369E5B082A34FF0B38376076FCE9" blockId="36.[188,1399,610,850]" pageId="36" pageNumber="37">
SEGMENT 10. With well-developed laterodorsal tubes, present in males only, located near posterior segment margin (
<figureCitation id="13262A1B5B082A34FE3138D064BAFD18" box="[391,501,645,672]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11A</figureCitation>
,
<figureCitation id="13262A1B5B082A34FDB738D36760FD18" box="[513,559,646,672]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12F</figureCitation>
,
<figureCitation id="13262A1B5B082A34FD8C38D3673EFD18" box="[570,625,646,672]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13H</figureCitation>
). Females without tubes (
<figureCitation id="13262A1B5B082A34FC2C38D06148FD18" box="[922,1031,645,672]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11C</figureCitation>
,
<figureCitation id="13262A1B5B082A34FBA438D36106FD18" box="[1042,1097,646,672]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12H</figureCitation>
). Sensory spots present in subdorsal and ventrolateral positions (
<figureCitation id="13262A1B5B082A34FDCB38FC6640FD7B" box="[637,783,681,707]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B082A34FCAA38FC6624FD7B" box="[796,875,681,707]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12EF</figureCitation>
,
<figureCitation id="13262A1B5B082A34FCC138FC66E1FD7B" box="[887,942,681,707]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13H</figureCitation>
); subdorsal pair located rather close to paradorsal area. Glandular cell outlets
<typeStatus id="54A6883C5B082A34FD32389867F8FD5F" box="[644,695,717,743]" pageId="36" pageNumber="37">type</typeStatus>
1 present in middorsal and in ventromedial positions. Band of perforation site patches as on preceding segment but with additional patch present in paradorsal area. Posterior segment margin of tergal plate straight, without fringe tips (
<figureCitation id="13262A1B5B082A34FC6A39416107FC96" box="[988,1096,788,814]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">Fig. 13H</figureCitation>
); margins of sternal plates concave, reaching the posterior margin of the terminal segment, with short and narrow fringe tips.
</paragraph>
<paragraph id="8BA2369E5B082A34FF0B392C667AFB34" blockId="36.[188,1400,889,1165]" pageId="36" pageNumber="37">
SEGMENT 11. With lateral terminal spines (
<figureCitation id="13262A1B5B082A34FD7A392C662CFC2B" box="[716,867,889,915]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11AB</figureCitation>
,
<figureCitation id="13262A1B5B082A34FCC0392C66E4FC2B" box="[886,939,889,915]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12A</figureCitation>
,
<figureCitation id="13262A1B5B082A34FC0B392C66BCFC2B" box="[957,1011,889,915]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13A</figureCitation>
). Females with lateral terminal accessory spines (
<figureCitation id="13262A1B5B082A34FE3B39C86752FC0F" box="[397,541,925,951]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11CD</figureCitation>
,
<figureCitation id="13262A1B5B082A34FD9E39C86711FC0F" box="[552,606,925,951]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12H</figureCitation>
); males with three penile spines (
<figureCitation id="13262A1B5B082A34FC6B39C86103FC0F" box="[989,1100,925,951]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">Figs 12G</figureCitation>
,
<figureCitation id="13262A1B5B082A34FBE139C861C2FC0F" box="[1111,1165,925,951]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13H</figureCitation>
). Dorsal and ventral spines slender and tubular, with ventral ones longer than dorsal ones; median spines very stout, coneshaped (
<figureCitation id="13262A1B5B082A34FE9739B164DCFC46" box="[289,403,996,1022]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11A</figureCitation>
,
<figureCitation id="13262A1B5B082A34FE1439B16498FC46" box="[418,471,996,1022]" captionStart="Fig" captionStartId="34.[190,233,1857,1883]" captionTargetBox="[196,1394,587,1819]" captionTargetId="figure-92@34.[194,1404,586,1820]" captionTargetPageId="34" captionText="Fig. 12. Light micrographs showing overview and details of Echinoderes leduci sp. nov. AG. ♂, holotype (NIWA-159405). H. ♀, paratype (NIWA-159406). A. Dorsal overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 6, ventral view. D. Segments 6 to 11, dorsal view. E. Segments 6 to 11, ventral view. F. Segments 9 to 11 of male, dorsal view. G. Segments 10 to 11 of male, dorsal view. H. Segments 10 to 11 of female, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222183" httpUri="https://zenodo.org/record/7222183/files/figure.png" pageId="36" pageNumber="37">12G</figureCitation>
,
<figureCitation id="13262A1B5B082A34FE5039B16752FC46" box="[486,541,996,1022]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13H</figureCitation>
). Sensory spots present in subdorsal positions. Two unpaired glandular cell outlets
<typeStatus id="54A6883C5B082A34FEF13E5D6435FB9A" box="[327,378,1032,1058]" pageId="36" pageNumber="37">type</typeStatus>
1 present middorsally. Segment devoid of characteristic perforation sites patches, but with very short cuticular hair-like structures covering paradorsal area. Short fringes covering margins of tergal and sternal plates. Tergal extensions triangular (
<figureCitation id="13262A1B5B082A34FC893E1B669BFBD1" box="[831,980,1102,1129]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Figs 11A, C</figureCitation>
,
<figureCitation id="13262A1B5B082A34FC573E1A6157FBD1" box="[993,1048,1103,1129]" captionStart="Fig" captionStartId="35.[188,231,1855,1881]" captionTargetBox="[190,1396,857,1822]" captionTargetId="figure-0@35.[188,1398,856,1832]" captionTargetPageId="35" captionText="Fig. 13. Scanning electron micrographs showing overviews and details of Echinoderes leduci sp. nov. A. Lateroventral overview. B. Segments 1 to 3, lateral view. C. Segments 1 to 3, ventral view. D. Segments 1 to 6, laterodorsal view. E. Segments 8 to 9, dorsal view. F. Segments 3 to 7, ventral view. G. Segments 7 to 9, laterodorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222185" httpUri="https://zenodo.org/record/7222185/files/figure.png" pageId="36" pageNumber="37">13H</figureCitation>
). Sternal extensions rounded, not extending beyond tergal extensions (
<figureCitation id="13262A1B5B082A34FD2F3E276667FB34" box="[665,808,1138,1164]" captionStart="Fig" captionStartId="32.[189,232,1927,1953]" captionTargetBox="[217,1368,271,1892]" captionTargetId="figure-39@32.[216,1371,267,1892]" captionTargetPageId="32" captionText="Fig. 11. Line art illustrations of Echinoderes leduci sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♀, segments 1011, dorsal view. D. ♀, segments 1011, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222181" httpUri="https://zenodo.org/record/7222181/files/figure.png" pageId="36" pageNumber="37">Fig. 11B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30765155B082A34FF0B3EE1679AFA98" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BA2369E5B082A34FF0B3EE16419FB76" blockId="36.[189,342,1204,1230]" box="[189,342,1204,1230]" pageId="36" pageNumber="37">
<heading id="D0EA81F25B082A34FF0B3EE16419FB76" bold="true" box="[189,342,1204,1230]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B969EA8C5B082A34FF0B3EE16419FB76" bold="true" box="[189,342,1204,1230]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B082A34FF0B3EB6679AFA98" blockId="36.[189,1398,1251,1312]" pageId="36" pageNumber="37">
Canyons: Pahaua, Honeycomb,
<quantity id="4CE59B7B5B082A34FD833EB66786FB45" box="[565,713,1251,1277]" metricMagnitude="2" metricUnit="m" metricValue="8.415" metricValueMax="10.129999999999999" metricValueMin="6.7" pageId="36" pageNumber="37" unit="m" value="841.5" valueMax="1013.0" valueMin="670.0">6701013 m</quantity>
b.s.l. See
<figureCitation id="13262A1B5B082A34FCF43EB666C8FB45" box="[834,903,1251,1277]" captionStart="Fig" captionStartId="2.[189,232,1961,1987]" captionTargetBox="[189,1399,1136,1922]" captionTargetId="figure-333@2.[189,1399,1136,1923]" captionTargetPageId="2" captionText="Fig. 1. Map showing the sampling stations. Colour codes indicate slopes (red), canyons (green), and seamounts (yellow). Inset shows New Zealand with a frame marking the sampling area." figureDoi="http://doi.org/10.5281/zenodo.7222159" httpUri="https://zenodo.org/record/7222159/files/figure.png" pageId="36" pageNumber="37">Fig. 1</figureCitation>
for a geographic overview of stations and
<tableCitation id="C69F03255B082A34FF0B3F53645BFA98" box="[189,276,1286,1312]" captionStart="Table 1" captionStartId="3.[185,254,549,577]" captionTargetBox="[192,1910,658,1734]" captionTargetPageId="3" captionText="Table 1 (continued on next four pages). Coordinates and basic data on sampling stations, and identities, type status and catalogue numbers of species of Echinoderes Claparède, 1863 from each station." httpUri="http://table.plazi.org/id/DF6266165B2F2A13FF0F38706644FDDC" pageId="36" pageNumber="37" tableUuid="DF6266165B2F2A13FF0F38706644FDDC">Table 1</tableCitation>
for station and specimen information.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B082A35FF0B3F1D6674FB1F" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="discussion">
<paragraph id="8BA2369E5B082A34FF0B3F1D6111FADA" blockId="36.[189,1118,1352,1379]" box="[189,1118,1352,1379]" pageId="36" pageNumber="37">
<heading id="D0EA81F25B082A34FF0B3F1D6111FADA" bold="true" box="[189,1118,1352,1379]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B969EA8C5B082A34FF0B3F1D6111FADA" bold="true" box="[189,1118,1352,1379]" pageId="36" pageNumber="37">
Taxonomic remarks on
<taxonomicName id="4C1D4D1D5B082A34FE533F1D67A4FADB" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[485,747,1352,1379]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B082A34FE533F1D67A4FADB" bold="true" box="[485,747,1352,1379]" italics="true" pageId="36" pageNumber="37">Echinoderes dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B082A34FD443F1D6600FADA" box="[754,847,1352,1378]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1D4D1D5B082A34FC3B3F1D66B6FADB" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[909,1017,1352,1379]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B082A34FC3B3F1D66B6FADB" bold="true" box="[909,1017,1352,1379]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B082A34FBB63F1D6111FADA" box="[1024,1118,1352,1378]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B082A34FF0B3F226162F8E7" blockId="36.[188,1400,1399,1888]" pageId="36" pageNumber="37">
<taxonomicName id="4C1D4D1D5B082A34FF0B3F2264F3FA29" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,444,1399,1425]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B082A34FF0B3F2264F3FA29" box="[189,444,1399,1425]" italics="true" pageId="36" pageNumber="37">Echinoderes dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B082A34FE743F2D6754FA2A" box="[450,539,1400,1426]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1D4D1D5B082A34FDE63F2D67F6FA29" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[592,697,1399,1425]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B082A34FDE63F2D67F6FA29" box="[592,697,1399,1425]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B082A34FD093F2D6657FA2A" box="[703,792,1400,1426]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
can easily be recognized by the nature of their trunk appearance with cuticular hairs that are reduced to minute scales, and characteristic strong perforation sites present on segments 2 to 10. The complete lack of cuticular hairs is an uncommon feature among species of
<taxonomicName id="4C1D4D1D5B082A34FE833FB76488FA44" authorityName="Claparede" authorityYear="1863" box="[309,455,1506,1532]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B082A34FE833FB76488FA44" box="[309,455,1506,1532]" italics="true" pageId="36" pageNumber="37">Echinoderes</emphasis>
</taxonomicName>
. Within
<taxonomicName id="4C1D4D1D5B082A34FD9A3FB76796FA44" authorityName="Carus" authorityYear="1885" box="[556,729,1506,1532]" class="Cyclorhagida" family="Echinoderidae" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="family">Echinoderidae</taxonomicName>
, the species peculiar lack of cuticular hairs is shared only with
<taxonomicName id="4C1D4D1D5B082A34FF4F3C5067A6F9A7" authority="Sorensen et al., 2013" authorityName="Sorensen" authorityYear="2013" box="[249,745,1541,1568]" class="Cyclorhagida" family="Echinoderidae" genus="Meristoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="glaber">
<emphasis id="B969EA8C5B082A34FF4F3C5064A7F9A7" box="[249,488,1541,1567]" italics="true" pageId="36" pageNumber="37">Meristoderes glaber</emphasis>
<bibRefCitation id="EF8C4B6F5B082A34FE463C5067A6F9A7" author="Sorensen M. V. &amp; Rho H. S. &amp; Min W. &amp; Kim D. &amp; Chang CY" box="[496,745,1541,1568]" pageId="36" pageNumber="37" pagination="291 - 319" refId="ref53041" refString="Sorensen M. V., Rho H. S., Min W., Kim D. &amp; Chang CY. 2013. Occurrence of the newly described kinorhynch genus Meristoderes (Cyclorhagida: Echinoderidae) in Korea, with the description of four new species. Helgoland Marine Research 67: 291 - 319. https: // doi. org / 10.1007 / s 10152 - 012 - 0323 - 2" type="journal article" year="2013">
Sørensen
<emphasis id="B969EA8C5B082A34FDD23C5367D0F9A7" box="[612,671,1541,1567]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
, 2013
</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EF8C4B6F5B082A34FD4F3C5066BDF9A7" author="Sorensen M. V. &amp; Rho H. S. &amp; Min W. &amp; Kim D. &amp; Chang CY" box="[761,1010,1541,1567]" pageId="36" pageNumber="37" pagination="291 - 319" refId="ref53041" refString="Sorensen M. V., Rho H. S., Min W., Kim D. &amp; Chang CY. 2013. Occurrence of the newly described kinorhynch genus Meristoderes (Cyclorhagida: Echinoderidae) in Korea, with the description of four new species. Helgoland Marine Research 67: 291 - 319. https: // doi. org / 10.1007 / s 10152 - 012 - 0323 - 2" type="journal article" year="2013">
Sørensen
<emphasis id="B969EA8C5B082A34FCD93C5366E5F9A7" box="[879,938,1541,1567]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
2013
</bibRefCitation>
). Resemblance with
<taxonomicName id="4C1D4D1D5B082A34FB513C536013F9A7" box="[1255,1372,1541,1567]" class="Cyclorhagida" family="Echinoderidae" genus="Meristoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="glaber">
<emphasis id="B969EA8C5B082A34FB513C536013F9A7" box="[1255,1372,1541,1567]" italics="true" pageId="36" pageNumber="37">M. glaber</emphasis>
</taxonomicName>
is only superficial though, and the lack of lateral spines on segment6, together with the presence of glandular cell outlets
<typeStatus id="54A6883C5B082A34FEF23C186438F9DF" box="[324,375,1613,1639]" pageId="36" pageNumber="37">type</typeStatus>
2 instead of tubes on segment
<quantity id="4CE59B7B5B082A34FD543C186640F9DE" box="[738,783,1612,1639]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="36" pageNumber="37" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C1D4D1D5B082A34FCA03C1866C5F9DE" box="[790,906,1612,1638]" class="Cyclorhagida" family="Echinoderidae" genus="Meristoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="glaber">
<emphasis id="B969EA8C5B082A34FCA03C1866C5F9DE" box="[790,906,1612,1638]" italics="true" pageId="36" pageNumber="37">M. glaber</emphasis>
</taxonomicName>
easily distinguish it from
<taxonomicName id="4C1D4D1D5B082A34FB0B3C18600AF9DE" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1213,1349,1612,1638]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B082A34FB0B3C18600AF9DE" box="[1213,1349,1612,1638]" italics="true" pageId="36" pageNumber="37">E. dalzottoi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B082A34FF0B3C246467F932" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,296,1648,1674]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci">
<emphasis id="B969EA8C5B082A34FF0B3C246467F932" box="[189,296,1648,1674]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
. The new species can also be distinguished from each other without any difficulty. Despite the number of similarities suggesting a close relationship, such as presence of lateroventral spines on segments 69, lateroventral tubes on segment 5, sublateral tubes on segment 8 and laterodorsal tubes on segment 9, it is possible to distinguish the two species based on several characters. The main difference is the number of middorsal spines, present on segments 4, 6 and
<quantity id="4CE59B7B5B082A34FC633CAB614BF8A0" box="[981,1028,1790,1816]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="36" pageNumber="37" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C1D4D1D5B082A34FBB93CAA6135F8A0" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1039,1146,1790,1816]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci">
<emphasis id="B969EA8C5B082A34FBB93CAA6135F8A0" box="[1039,1146,1790,1816]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
in contrast to spines on segments 4 and 6 only in
<taxonomicName id="4C1D4D1D5B082A34FDA93D7667E8F884" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[543,679,1826,1852]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B082A34FDA93D7667E8F884" box="[543,679,1826,1852]" italics="true" pageId="36" pageNumber="37">E. dalzottoi</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="4C1D4D1D5B082A34FCEB3D766688F884" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[861,967,1826,1852]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci">
<emphasis id="B969EA8C5B082A34FCEB3D766688F884" box="[861,967,1826,1852]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
lacks subdorsal tubes on segment 2 (present in
<taxonomicName id="4C1D4D1D5B082A34FEF73D136486F8E7" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[321,457,1861,1887]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B082A34FEF73D136486F8E7" box="[321,457,1861,1887]" italics="true" pageId="36" pageNumber="37">E. dalzottoi</emphasis>
</taxonomicName>
), but has laterodorsal tubes on this segment instead.
</paragraph>
<paragraph id="8BA2369E5B082A35FF0B3DD2605FFE0D" blockId="36.[189,1399,1927,2025]" lastBlockId="37.[189,1399,265,437]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Having tubes on segment
<quantity id="4CE59B7B5B082A34FE413DD26769F819" box="[503,550,1927,1953]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="36" pageNumber="37" unit="in" value="2.0">2 in</quantity>
only laterodorsal and ventrolateral positions as in
<taxonomicName id="4C1D4D1D5B082A34FB3D3DDD61BBF819" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1163,1268,1927,1953]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B082A34FB3D3DDD61BBF819" box="[1163,1268,1927,1953]" italics="true" pageId="36" pageNumber="37">E. leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B082A34FB4B3DDD6015F81A" box="[1277,1370,1928,1954]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
is quite a rare character, and only shared with two other species of
<taxonomicName id="4C1D4D1D5B082A34FC733DFE6118F87D" authorityName="Claparede" authorityYear="1863" box="[965,1111,1963,1989]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B082A34FC733DFE6118F87D" box="[965,1111,1963,1989]" italics="true" pageId="36" pageNumber="37">Echinoderes</emphasis>
</taxonomicName>
, i.e.,
<taxonomicName id="4C1D4D1D5B082A34FB2B3DF9600AF87D" authorityName="Grzelak &amp; Sorensen" authorityYear="2017" box="[1181,1349,1963,1989]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="daenerysae">
<emphasis id="B969EA8C5B082A34FB2B3DF9600AF87D" box="[1181,1349,1963,1989]" italics="true" pageId="36" pageNumber="37">E. daenerysae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B082A34FF093D9A673EF850" authority="Huys &amp; Coomans, 1989" authorityName="Huys &amp; Coomans" authorityYear="1989" box="[191,625,1998,2025]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="36" pageNumber="37" phylum="Kinorhyncha" rank="species" species="higginsi">
<emphasis id="B969EA8C5B082A34FF093D9A640FF850" box="[191,320,1998,2024]" italics="true" pageId="36" pageNumber="37">E. higginsi</emphasis>
<bibRefCitation id="EF8C4B6F5B082A34FEFC3D9A673EF850" author="Huys R. &amp; Coomans A." box="[330,625,1998,2025]" pageId="36" pageNumber="37" pagination="211 - 221" refId="ref50825" refString="Huys R. &amp; Coomans A. 1989. Echinoderes higginsi sp. n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparede. Zoologica Scripta 18: 211 - 221. https: // doi. org / 10.1111 / j. 1463 - 6409.1989. tb 00446. x" type="journal article" year="1989">Huys &amp; Coomans, 1989</bibRefCitation>
</taxonomicName>
(although a laterodorsal pair was not mentioned in the original description of the latter species) (
<bibRefCitation id="EF8C4B6F5B092A35FDF13B5C662CFE9B" author="Huys R. &amp; Coomans A." box="[583,867,265,291]" pageId="37" pageNumber="38" pagination="211 - 221" refId="ref50825" refString="Huys R. &amp; Coomans A. 1989. Echinoderes higginsi sp. n. (Kinorhyncha, Cyclorhagida) from the southern North Sea with a key to the genus Echinoderes Claparede. Zoologica Scripta 18: 211 - 221. https: // doi. org / 10.1111 / j. 1463 - 6409.1989. tb 00446. x" type="journal article" year="1989">Huys &amp; Coomans 1989</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B092A35FCC63B5C61E8FE9B" author="Grzelak K. &amp; Sorensen M. V." box="[880,1191,265,291]" pageId="37" pageNumber="38" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen 2018</bibRefCitation>
). Neither of them can be confused with
<taxonomicName id="4C1D4D1D5B092A35FE083B7B6769FEFF" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[446,550,301,327]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="leduci">
<emphasis id="B969EA8C5B092A35FE083B7B6769FEFF" box="[446,550,301,327]" italics="true" pageId="37" pageNumber="38">E. leduci</emphasis>
</taxonomicName>
, however. Except for the dense cuticular hair covering observed in both species,
<taxonomicName id="4C1D4D1D5B092A35FE953B066483FED4" authorityName="Grzelak &amp; Sorensen" authorityYear="2017" box="[291,460,338,364]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="daenerysae">
<emphasis id="B969EA8C5B092A35FE953B066483FED4" box="[291,460,338,364]" italics="true" pageId="37" pageNumber="38">E. daenerysae</emphasis>
</taxonomicName>
possesses only two middorsal spines on segments 6 and 8, while
<taxonomicName id="4C1D4D1D5B092A35FB5B3B06603FFED4" authorityName="Huys &amp; Coomans" authorityYear="1989" box="[1261,1392,338,364]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="higginsi">
<emphasis id="B969EA8C5B092A35FB5B3B06603FFED4" box="[1261,1392,338,364]" italics="true" pageId="37" pageNumber="38">E. higginsi</emphasis>
</taxonomicName>
, although having the same number and arrangement of middorsal spines as
<taxonomicName id="4C1D4D1D5B092A35FB933B2261C2FE28" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1061,1165,374,400]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="leduci">
<emphasis id="B969EA8C5B092A35FB933B2261C2FE28" box="[1061,1165,374,400]" italics="true" pageId="37" pageNumber="38">E. leduci</emphasis>
</taxonomicName>
, is characterized by long, conspicuous spinous tergal extensions of segment 11 and the lack of tubes on segment 9.
</paragraph>
<paragraph id="8BA2369E5B092A35FF0B3B88661FFD4F" blockId="37.[188,1399,477,759]" pageId="37" pageNumber="38">
<taxonomicName id="4C1D4D1D5B092A35FF0B3B8864F1FE4F" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,446,477,503]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B092A35FF0B3B8864F1FE4F" box="[189,446,477,503]" italics="true" pageId="37" pageNumber="38">Echinoderes dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B092A35FE703B8B676DFE40" box="[454,546,478,504]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
is also easily distinguished from all other congeners by its combination of spines and tubes. Middorsal spines being restricted to segments 4 and 6 only is rare and shared only with four other species, i.e.,
<taxonomicName id="4C1D4D1D5B092A35FDAE387267D7FDF8" authorityName="Sorensen" authorityYear="2014" box="[536,664,550,576]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="astridae">
<emphasis id="B969EA8C5B092A35FDAE387267D7FDF8" box="[536,664,550,576]" italics="true" pageId="37" pageNumber="38">E. astridae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B092A35FD1F38726607FDF8" authorityName="Higgins" authorityYear="1982" box="[681,840,550,576]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="bispinosus">
<emphasis id="B969EA8C5B092A35FD1F38726607FDF8" box="[681,840,550,576]" italics="true" pageId="37" pageNumber="38">E. bispinosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B092A35FCEF38726696FDF8" box="[857,985,551,576]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="uozumii">
<emphasis id="B969EA8C5B092A35FCEF38726696FDF8" box="[857,985,551,576]" italics="true" pageId="37" pageNumber="38">E. uozumii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C1D4D1D5B092A35FBA9387261EDFDF8" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1055,1186,550,576]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="landersi" status="sp. nov.">
<emphasis id="B969EA8C5B092A35FBA9387261EDFDF8" box="[1055,1186,550,576]" italics="true" pageId="37" pageNumber="38">E. landersi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B092A35FB1C38726048FDF9" box="[1194,1287,551,577]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
(
<bibRefCitation id="EF8C4B6F5B092A35FAAE387365B2FDDD" author="Higgins R. P." pageId="37" pageNumber="38" pagination="305 - 316" refId="ref50573" refString="Higgins R. P. 1982. Three new species of Kinorhyncha from Bermuda. Transactions of the American Microscopical Society 101: 305 - 316. https: // doi. org / 10.2307 / 3225748" type="journal article" year="1982">Higgins 1982</bibRefCitation>
; Sørensen 2014; Yamasaki
<emphasis id="B969EA8C5B092A35FDF438196732FDDD" box="[578,637,587,613]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
2020). However,
<taxonomicName id="4C1D4D1D5B092A35FCE338196699FDDD" authorityName="Sorensen" authorityYear="2014" box="[853,982,587,613]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="astridae">
<emphasis id="B969EA8C5B092A35FCE338196699FDDD" box="[853,982,587,613]" italics="true" pageId="37" pageNumber="38">E. astridae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B092A35FC50381961CCFDDD" authorityName="Higgins" authorityYear="1982" box="[998,1155,587,613]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="bispinosus">
<emphasis id="B969EA8C5B092A35FC50381961CCFDDD" box="[998,1155,587,613]" italics="true" pageId="37" pageNumber="38">E. bispinosus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B092A35FB7638196071FDDD" box="[1216,1342,588,613]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="uozumii">
<emphasis id="B969EA8C5B092A35FB7638196071FDDD" box="[1216,1342,588,613]" italics="true" pageId="37" pageNumber="38">E. uozumii</emphasis>
</taxonomicName>
lack dorsal and lateral tubes on segment 2, and have instead two pairs of glandular cell outlet
<typeStatus id="54A6883C5B092A35FB76382561BCFD32" box="[1216,1267,624,650]" pageId="37" pageNumber="38">type</typeStatus>
2 a gland
<typeStatus id="54A6883C5B092A35FF0A38C165A0FD16" box="[188,239,660,686]" pageId="37" pageNumber="38">type</typeStatus>
that is absent in
<taxonomicName id="4C1D4D1D5B092A35FE0C38C0670DFD16" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[442,578,660,686]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B092A35FE0C38C0670DFD16" box="[442,578,660,686]" italics="true" pageId="37" pageNumber="38">E. dalzottoi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1D4D1D5B092A35FDE738C16604FD16" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[593,843,660,686]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B092A35FDE738C16604FD16" box="[593,843,660,686]" italics="true" pageId="37" pageNumber="38">Echinoderes landersi</emphasis>
</taxonomicName>
, in turn, is characterized by four pairs of tubes on segment 2 and three pairs of tubes on segment 8 a so far unique tube combination (see Remarks in the description of
<taxonomicName id="4C1D4D1D5B092A35FE06388B677FFD4F" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[432,560,733,759]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="landersi">
<emphasis id="B969EA8C5B092A35FE06388B677FFD4F" box="[432,560,733,759]" italics="true" pageId="37" pageNumber="38">E. landersi</emphasis>
</taxonomicName>
above for more details).
</paragraph>
<paragraph id="8BA2369E5B092A35FF0B394A6674FB1F" blockId="37.[188,1399,799,1191]" pageId="37" pageNumber="38">
The combination of tubes in subdorsal, sublateral and ventrolateral positions on segment 2 is also quite rare. Numerous species present combinations of two or four pairs of tubes on this segment, but only four species have three pairs, among which only
<taxonomicName id="4C1D4D1D5B092A35FD0F393C617AFC3B" authority="Pardos et al., 1998" authorityName="Pardos" authorityYear="1998" box="[697,1077,872,899]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="hispanicus">
<emphasis id="B969EA8C5B092A35FD0F393C6618FC3A" box="[697,855,872,898]" italics="true" pageId="37" pageNumber="38">E. hispanicus</emphasis>
<bibRefCitation id="EF8C4B6F5B092A35FCEB393D617AFC3B" author="Pardos F. &amp; Higgins R. P. &amp; Benito J." box="[861,1077,872,899]" pageId="37" pageNumber="38" pagination="195 - 208" refId="ref52069" refString="Pardos F., Higgins R. P. &amp; Benito J. 1998. Two new Echinoderes (Kinorhyncha, Cyclorhagida) including a reevaluation of kinorhynch taxonomic characters. Zoologischer Anzeiger 237: 195 - 208." type="journal article" year="1998">
Pardos
<emphasis id="B969EA8C5B092A35FC04393C66A9FC3A" box="[946,998,872,898]" italics="true" pageId="37" pageNumber="38">et al</emphasis>
., 1998
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B092A35FBDC393C64CDFC1F" authority="Higgins &amp; Kristensen, 1988" authorityName="Higgins &amp; Kristensen" authorityYear="1988" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="peterseni">
<emphasis id="B969EA8C5B092A35FBDC393C61B8FC3A" box="[1130,1271,873,898]" italics="true" pageId="37" pageNumber="38">E. peterseni</emphasis>
<bibRefCitation id="EF8C4B6F5B092A35FB4B393D64CDFC1F" author="Higgins R. P. &amp; Kristensen R. M." pageId="37" pageNumber="38" pagination="1 - 56" refId="ref50656" refString="Higgins R. P. &amp; Kristensen R. M. 1988. Kinorhyncha from Disko Island, West Greenland. Smithsonian Contributions to Zoology 458: 1 - 56. https: // doi. org / 10.5479 / si. 00810282.458" type="journal article" year="1988">Higgins &amp; Kristensen, 1988</bibRefCitation>
</taxonomicName>
have tube arrangements as in
<taxonomicName id="4C1D4D1D5B092A35FD5739DB6627FC1F" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[737,872,909,935]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B092A35FD5739DB6627FC1F" box="[737,872,909,935]" italics="true" pageId="37" pageNumber="38">E. dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B092A35FCD939D86687FC1F" box="[879,968,909,935]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
(
<bibRefCitation id="EF8C4B6F5B092A35FC6139D86054FC1F" author="Higgins R. P. &amp; Kristensen R. M." box="[983,1307,909,935]" pageId="37" pageNumber="38" pagination="1 - 56" refId="ref50656" refString="Higgins R. P. &amp; Kristensen R. M. 1988. Kinorhyncha from Disko Island, West Greenland. Smithsonian Contributions to Zoology 458: 1 - 56. https: // doi. org / 10.5479 / si. 00810282.458" type="journal article" year="1988">Higgins &amp; Kristensen 1988</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B092A35FA9139D8640DFC74" author="Pardos F. &amp; Higgins R. P. &amp; Benito J." pageId="37" pageNumber="38" pagination="195 - 208" refId="ref52069" refString="Pardos F., Higgins R. P. &amp; Benito J. 1998. Two new Echinoderes (Kinorhyncha, Cyclorhagida) including a reevaluation of kinorhynch taxonomic characters. Zoologischer Anzeiger 237: 195 - 208." type="journal article" year="1998">
Pardos
<emphasis id="B969EA8C5B092A35FF0A39E765B5FC73" box="[188,250,945,971]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
1998
</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B092A35FEE739E467C2FC74" author="Grzelak K. &amp; Sorensen M. V." box="[337,653,945,972]" pageId="37" pageNumber="38" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen 2018</bibRefCitation>
). However, both species also have three middorsal spines and lack laterodorsal tubes on segment 9, which easily distinguish them from
<taxonomicName id="4C1D4D1D5B092A35FBA3398261D3FC48" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1045,1180,982,1008]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B092A35FBA3398261D3FC48" box="[1045,1180,982,1008]" italics="true" pageId="37" pageNumber="38">E. dalzottoi</emphasis>
</taxonomicName>
. Even considering that sublateral tubes might be absent in some specimens of
<taxonomicName id="4C1D4D1D5B092A35FC3E39AE6140FBAC" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[904,1039,1018,1044]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B092A35FC3E39AE6140FBAC" box="[904,1039,1018,1044]" italics="true" pageId="37" pageNumber="38">E. dalzottoi</emphasis>
</taxonomicName>
(a variation in the occurrence of tubes has recently been reported for a number of echinoderid species; see, e.g.,
<bibRefCitation id="EF8C4B6F5B092A35FB3F3E4A6444FBE5" author="Grzelak K. &amp; Sorensen M. V." pageId="37" pageNumber="38" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen (2018)</bibRefCitation>
and
<bibRefCitation id="EF8C4B6F5B092A35FEF53E1667EEFBE5" author="Yamasaki H. &amp; Dal Zotto M." box="[323,673,1091,1118]" pageId="37" pageNumber="38" pagination="189 - 205" refId="ref53781" refString="Yamasaki H. &amp; Dal Zotto M. 2019. Investigation of echinoderid kinorhynchs described 90 years ago: redescription of Echinoderes capitatus (Zelinka, 1928) and Echinoderes ferrugineus Zelinka, 1928. Zoologischer Anzeiger 282: 189 - 205. https: // doi. org / 10.1016 / j. jcz. 2019.05.013" type="journal article" year="2019">Yamasaki &amp; Dal Zotto (2019)</bibRefCitation>
for further details), none of the species having only subdorsal and ventrolateral tubes on segment 2 can be confused with
<taxonomicName id="4C1D4D1D5B092A35FCD33E3C66A3FB3A" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[869,1004,1128,1154]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="dalzottoi">
<emphasis id="B969EA8C5B092A35FCD33E3C66A3FB3A" box="[869,1004,1128,1154]" italics="true" pageId="37" pageNumber="38">E. dalzottoi</emphasis>
</taxonomicName>
due to their significantly different middorsal spine patterns and trunk cuticle appearance.
</paragraph>
</subSubSection>
</treatment>
</document>