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<document id="278B4C83C07AD458665F4CC3E73CCA3C" ID="10.11646/zootaxa.4221.3.6" ID-CLB-Dataset="37291" ID-DOI="10.5281/zenodo.250320" ID-GBIF-Dataset="d56da6d2-4950-4798-9057-d8875115930b" ID-ISSN="1175-5326" ID-Zenodo-Dep="250320" ID-ZooBank="ABD475CA-CD30-41D0-B9DF-3F6721186781" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484727834738" checkinUser="plazi" docAuthor="Trýzna, Miloš" docDate="2017" docId="03B08792DB2ECD00FF6E20818FEFFCFA" docLanguage="en" docName="zootaxa.4221.3.6.pdf" docOrigin="Zootaxa 4221 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tophoderes Dejean 1834" docType="treatment" docVersion="8" lastPageNumber="378" masterDocId="FF89FFEADB2FCD01FFF921348D05FFEA" masterDocTitle="Description of a new species of the genus Tophoderes Dejean (Coleoptera: Anthribidae) from east Madagascar, with images of all Madagascan species of the genus" masterLastPageNumber="385" masterPageNumber="377" pageNumber="378" updateTime="1698435854850" updateUser="ExternalLinkService">
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<mods:title id="3BC6DEC361A6FF8BA599F2893AA19ACA">Description of a new species of the genus Tophoderes Dejean (Coleoptera: Anthribidae) from east Madagascar, with images of all Madagascan species of the genus</mods:title>
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<mods:name id="E2130F3FB6E66B82CDFE5DBAC18E1840" type="personal">
<mods:role id="1CFDAD5EFB756266822094FA7CA02ADB">
<mods:roleTerm id="5717AB58D9BA77D8377E7826F16571DA">Author</mods:roleTerm>
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<mods:namePart id="CA3D3CF16A49BC92EE5302BAB4219AD3">Trýzna, Miloš</mods:namePart>
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<mods:typeOfResource id="1624A3546996D3243590B0DF6136031E">text</mods:typeOfResource>
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<mods:title id="EBAF12D355375CF32DB5FAE6B34F768E">Zootaxa</mods:title>
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<mods:date id="26D8172A75CE52FB7FF441153DC632E8">2017</mods:date>
<mods:detail id="7E0992A4A7BDF235D5B220EFB0103CA4" type="volume">
<mods:number id="022A115A54FE93CF0601FB54650BEBA7">4221</mods:number>
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<mods:number id="96809DB55975FD09315108B70C03C794">3</mods:number>
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<mods:classification id="5A1FE12D69C11D8BFB8A7E455C6C13B5">journal article</mods:classification>
<mods:identifier id="EF17AB474697FD77B0C65D3314EB74AF" type="CLB-Dataset">37291</mods:identifier>
<mods:identifier id="ECD141739130736F81241B33827D9FAC" type="DOI">10.5281/zenodo.250320</mods:identifier>
<mods:identifier id="C31BB8BC758107D9BD09D4E2CFBE2C80" type="GBIF-Dataset">d56da6d2-4950-4798-9057-d8875115930b</mods:identifier>
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<mods:identifier id="621EA4AC6D72FDFF7CD62EE484E0EEA2" type="Zenodo-Dep">250320</mods:identifier>
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<treatment id="03B08792DB2ECD00FF6E20818FEFFCFA" ID-DOI="http://doi.org/10.5281/zenodo.6051453" ID-GBIF-Taxon="127614447" ID-Zenodo-Dep="6051453" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA" httpUri="http://treatment.plazi.org/id/03B08792DB2ECD00FF6E20818FEFFCFA" lastPageNumber="378" pageId="1" pageNumber="378">
<subSubSection id="C303650FDB2ECD00FF6E20818FEFFCFA" pageId="1" pageNumber="378" type="nomenclature">
<paragraph id="8BA63684DB2ECD00FF6E20818CCEFE25" blockId="1.[151,459,437,463]" box="[151,459,437,463]" pageId="1" pageNumber="378">
<heading id="D0EE81E8DB2ECD00FF6E20818CCEFE25" bold="true" box="[151,459,437,463]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
<taxonomicName id="4C194D07DB2ECD00FF6E20818CCEFE25" ID-CoL="8HS7H" authority="Dejean, 1834" authorityName="Dejean" authorityYear="1834" box="[151,459,437,463]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="genus">
<emphasis id="B96DEA96DB2ECD00FF6E20818CCEFE25" bold="true" box="[151,459,437,463]" pageId="1" pageNumber="378">
<emphasis id="B96DEA96DB2ECD00FF6E20818C27FE25" bold="true" box="[151,290,437,463]" italics="true" pageId="1" pageNumber="378">Tophoderes</emphasis>
<bibRefCitation id="EF884B75DB2ECD00FED020818CCEFE25" author="Dejean" box="[297,459,437,463]" pageId="1" pageNumber="378" refString="Dejean, P. F. M. A. (1834) Catalogue des Coleopteres de la collection de M. le Comte Dejean. [2. ed.]. Mequignon-Marvis &amp; Sons, Paris (Fasc. 3), 177 - 256." type="book chapter" year="1834">Dejean, 1834</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA63684DB2ECD00FF6E20C88FEFFCFA" blockId="1.[151,1436,508,784]" pageId="1" pageNumber="378">
<materialsCitation id="3B713CD9DB2ECD00FF6E20C88FE3FCFA" ID-GBIF-Occurrence="1424788243" collectingDate="1922-04-01" collectingDateMax="1922-11-01" collectingDateMin="1922-04-01" country="Jordan" county="Figures" location="Eyes" municipality="For" pageId="1" pageNumber="378" specimenCount="1">
<emphasis id="B96DEA96DB2ECD00FF6E20C88C28FDFF" bold="true" box="[151,301,508,533]" pageId="1" pageNumber="378">Recognition.</emphasis>
Head comparatively long, rostrum longer, extended apically, with one central longitudinal carina, without depression in the middle.
<location id="8EC6605FDB2ECD00FDE123148F4BFDD2" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA:8EC6605FDB2ECD00FDE123148F4BFDD2" box="[536,590,544,568]" country="Jordan" county="Figures" municipality="For" name="Eyes" pageId="1" pageNumber="378">Eyes</location>
spherical, not emarginate.
<location id="8EC6605FDB2ECD00FC7823148EE8FDD2" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA:8EC6605FDB2ECD00FC7823148EE8FDD2" box="[897,1005,544,568]" country="Jordan" county="Figures" municipality="For" name="Antennae" pageId="1" pageNumber="378">Antennae</location>
of males reaching to posterior margin of pronotum or longer (two last antennomeres can extend beyond this margin), antennae of females shorter, not reaching posterior margin of pronotum.
<location id="8EC6605FDB2ECD00FD8F23538FCFFD6A" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA:8EC6605FDB2ECD00FD8F23538FCFFD6A" box="[630,714,615,640]" country="Jordan" county="Figures" municipality="For" name="Funicle" pageId="1" pageNumber="378">Funicle</location>
thick, robust, antennal club 3-segmented.
<location id="8EC6605FDB2ECD00FB32235C883CFD95" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA:8EC6605FDB2ECD00FB32235C883CFD95" box="[1227,1337,616,639]" country="Jordan" county="Figures" municipality="For" name="Pronotum" pageId="1" pageNumber="378">Pronotum</location>
with or without forward oriented spine on lateral side; slightly transverse, dorsal transverse carina distinct.
<location id="8EC6605FDB2ECD00FAE123B88899FD4F" LSID="urn:lsid:plazi:treatment:03B08792DB2ECD00FF6E20818FEFFCFA:8EC6605FDB2ECD00FAE123B88899FD4F" box="[1304,1436,652,677]" country="Jordan" county="Figures" municipality="For" name="Large-sized" pageId="1" pageNumber="378">Large-sized</location>
species, from
<quantity id="4CE19B61DB2ECD00FEC723848C94FD2D" box="[318,401,688,712]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="378" unit="mm" value="11.0">11 mm</quantity>
(small specimens of
<taxonomicName id="4C194D07DB2ECD00FD6C23858EECFD22" authority="Jordan, 1895" authorityName="Jordan" authorityYear="1895" box="[661,1001,687,712]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="sinuatocollis">
<emphasis id="B96DEA96DB2ECD00FD6C23858E40FD22" box="[661,837,687,712]" italics="true" pageId="1" pageNumber="378">T. sinuatocollis</emphasis>
<collectingCountry id="F30E7614DB2ECD00FCAB239B8EA5FD22" box="[850,928,687,712]" name="Jordan" pageId="1" pageNumber="378">Jordan</collectingCountry>
, 1895
</taxonomicName>
) to more than
<quantity id="4CE19B61DB2ECD00FB5A238489F2FD2D" box="[1187,1271,688,712]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="378" unit="mm" value="32.0">32 mm</quantity>
(
<taxonomicName id="4C194D07DB2ECD00FAF523858C6CFD07" authority="Waterhouse, 1875" authorityName="Waterhouse" authorityYear="1875" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="annulatus">
<emphasis id="B96DEA96DB2ECD00FAF52385889EFD22" box="[1292,1435,687,712]" italics="true" pageId="1" pageNumber="378">T. annulatus</emphasis>
Waterhouse, 1875
</taxonomicName>
;
<taxonomicName id="4C194D07DB2ECD00FE8223E18FAAFD06" authority="Alluaud, 1899" authorityName="Alluaud" authorityYear="1899" box="[379,687,724,749]" class="Insecta" family="Anthribidae" genus="Tophoderes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="murinus">
<emphasis id="B96DEA96DB2ECD00FE8223E18CFDFD06" box="[379,504,725,748]" italics="true" pageId="1" pageNumber="378">T. murinus</emphasis>
Alluaud, 1899
</taxonomicName>
).
<collectingMunicipality id="6BC2ACFEDB2ECD00FD3123E18FEAFD06" box="[712,751,725,748]" pageId="1" pageNumber="378">For</collectingMunicipality>
comparative purposes all known species of the genus in
<collectingCountry id="F30E7614DB2ECD00FF6E23C38C1BFCFA" box="[151,286,759,784]" name="Madagascar" pageId="1" pageNumber="378">Madagascar</collectingCountry>
are illustrated in
<figureCitation id="13222A01DB2ECD00FE1B23C38F6DFCFA" box="[482,616,759,784]" captionStart="FIGURES 1 – 4. 1 – 2" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[263,1327,220,1836]" captionTargetId="figure@2.[227,1360,193,1857]" captionTargetPageId="2" captionText="FIGURES 1 – 4. 1 – 2, Tophoderes banari Trýzna sp. nov.; 1, male holotype, dorsal habitus; 2, female allotype, dorsal habitus; 3 – 4, Tophoderes annulatus Waterhouse, 1875; 3, male, 29 mm, Ankarampotsy, Tantamala; 4, female, 29 mm, Ambositra." httpUri="https://zenodo.org/record/250321/files/figure.png" pageId="1" pageNumber="378">
<collectingCounty id="62C74E08DB2ECD00FE1B23C38F33FCFA" box="[482,566,759,784]" pageId="1" pageNumber="378">Figures</collectingCounty>
<date id="FFA71044DB2ECD00FDC423CC8F6DFCFA" box="[573,616,760,784]" pageId="1" pageNumber="378" value="1922-04-01" valueMax="1922-11-01" valueMin="1922-04-01">
<collectingDate id="EFE3E9ACDB2ECD00FDC423CC8F6DFCFA" box="[573,616,760,784]" pageId="1" pageNumber="378" value="1922-04-01" valueMax="1922-11-01" valueMin="1922-04-01">1–4</collectingDate>
</date>
</figureCitation>
and
<figureCitation id="13222A01DB2ECD00FD6723CC8FE3FCFA" box="[670,742,760,784]" captionStart-0="FIGURES 11 – 16. 11" captionStart-1="FIGURES 17 – 22. 17" captionStartId-0="5.[151,264,1914,1937]" captionStartId-1="6.[151,264,1842,1865]" captionTargetBox-0="[158,1403,231,1845]" captionTargetBox-1="[168,1417,213,1777]" captionTargetId-0="figure@5.[151,1436,193,1893]" captionTargetId-1="figure@6.[151,1436,193,1821]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 11 – 16. 11, Tophoderes frenatus (Klug, 1833), 29 mm, Andasibe-Mantadia Nat. Park, Analamazaotra forest; 12, T. lidmilae Trýzna &amp; Baňař, 2015, 27 mm, Montagne d´Ambre Nat. Park, Ambohitra env.; 13, T. funebris (Klug, 1833), 17 mm, Tamatave, forêts d´Alahakato; 14, T. acarinulus Wolfrum, 1959, 17 mm, Maroantsetra; 15, T. nubeculosus Fairmaire, 1888, 23 mm, Baie d´Antongil; 16, T. ferrugatus (Klug, 1833), 23 mm, Tananarivo env." captionText-1="FIGURES 17 – 22. 17, Tophoderes griseovarius Fairmaire, 1901, 18 mm, Madagascar Sud, vallée d´Ambolo; 18, T. griseipes Fairmaire, 1901, 24 mm, Anjiro; 19, T. murinus Alluaud, 1899, 28 mm, Chaines Anosyennes, Massif nord, Ranomandry; 20, T. sikorae Jordan, 1895, 14 mm, Moramanga; 21, T. fuscoareatus Wolfrum, 1959, 13 mm, Moramanga; 22, T. sinuatocollis Jordan, 1895, 16 mm, Moramanga." httpUri-0="https://zenodo.org/record/250324/files/figure.png" httpUri-1="https://zenodo.org/record/250325/files/figure.png" pageId="1" pageNumber="378">11–22</figureCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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