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<document id="1527B9C40C5DBE00204E907117F18A80" ID-CLB-Dataset="20775" ID-DOI="10.11646/zootaxa.3656.1.1" ID-GBIF-Dataset="d2c3e582-04bb-4d44-8261-b62f1b0db73a" ID-ISSN="1175­5334" ID-Zenodo-Dep="6039126" ID-ZooBank="73E59AED-B16D-430C-B611-EB5CFEAF7FDD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629857720847" checkinUser="felipe" docAuthor="FÖLDVÁRI, MIHÁLY" docDate="2013" docId="03A487FDFFB9FFD71D13FF56FE7E4EB7" docLanguage="en" docName="zt03656p121.pdf" docOrigin="Zootaxa 3656 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3656.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Eudorylas bisetosus" docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FF9DFF85FF96FFF81F5FFFC1FFB94A2B" masterDocTitle="Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae)" masterLastPageNumber="121" masterPageNumber="1" pageNumber="48" updateTime="1699143883713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A6D49DE3EBA8656FE125FFC39A110C52">Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae)</mods:title>
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<treatment id="03A487FDFFB9FFD71D13FF56FE7E4EB7" ID-DOI="http://doi.org/10.5281/zenodo.6302798" ID-GBIF-Taxon="193278014" ID-Zenodo-Dep="6302798" LSID="urn:lsid:plazi:treatment:03A487FDFFB9FFD71D13FF56FE7E4EB7" httpUri="http://treatment.plazi.org/id/03A487FDFFB9FFD71D13FF56FE7E4EB7" lastPageNumber="48" pageId="47" pageNumber="48">
<subSubSection id="C3176560FFB9FFD71D13FF56FC5E4A99" box="[588,999,151,178]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="8BB236EBFFB9FFD71D13FF56FC5E4A99" blockId="47.[588,999,151,210]" box="[588,999,151,178]" pageId="47" pageNumber="48">
<taxonomicName id="4C0D4D68FFB9FFD71D13FF56FC5E4A99" ID-CoL="3C2R9" authority="(Hardy, 1962)" baseAuthorityName="Hardy" baseAuthorityYear="1962" box="[588,999,151,178]" class="Insecta" family="Pipunculidae" genus="Eudorylas" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bisetosus">
<emphasis id="B979EAF9FFB9FFD71D13FF56FC854A9A" bold="true" box="[588,828,151,177]" italics="true" pageId="47" pageNumber="48">Eudorylas bisetosus</emphasis>
(
<bibRefCitation id="EF9C4B1AFFB9FFD71C14FF59FC664A99" author="Hardy" box="[843,991,152,178]" firstAuthor="Hardy" pageId="47" pageNumber="48" pagination="241 - 269" refId="ref88057" refString="Hardy, D. E. (1962) Preliminary studies on the Pipunculidae of Madagascar (Diptera). Verhandlungen der Naturforschenden Gesellschaft in Basel, 73, 241 - 269." type="journal article" year="1962">Hardy, 1962</bibRefCitation>
)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71D92FF7BFCDF4AF9" box="[717,870,186,210]" pageId="47" pageNumber="48" type="description">
<paragraph id="8BB236EBFFB9FFD71D92FF7BFCDF4AF9" blockId="47.[588,999,151,210]" box="[717,870,186,210]" pageId="47" pageNumber="48">
(
<figureCitation id="13362A6EFFB9FFD71D8AFF7BFCE74AF9" box="[725,862,186,210]" captionStart="FIGURE 15" captionStartId="46.[151,250,1843,1866]" captionTargetBox="[297,1295,184,1820]" captionTargetId="figure-117@46.[276,1315,179,1829]" captionTargetPageId="46" captionText="FIGURE 15. Male genitalia of Eudorylas barracloughi sp. nov. (A–F) and E. bisetosus (G–L).—A: outer surstylus with phallic guide, attachment of phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: surstyli, dorsal view, E: sperm pump with ejaculatory apodeme, F: ventral view with phallic guide and phallus (without ST8), G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme, L: ventral view with phallic guide and subepandrial sclerite (without ST8). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6039188" httpUri="https://zenodo.org/record/6039188/files/figure.png" pageId="47" pageNumber="48">Figs 15G–L</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71FC8FEC3FD1D4B33" box="[151,676,258,280]" pageId="47" pageNumber="48" type="reference_group">
<paragraph id="8BB236EBFFB9FFD71FC8FEC3FD1D4B33" blockId="47.[151,676,258,280]" box="[151,676,258,280]" pageId="47" pageNumber="48">
<treatmentCitationGroup id="AB1D11C5FFB9FFD71FC8FEC3FD1D4B33" box="[151,676,258,280]" pageId="47" pageNumber="48">
<treatmentCitation id="0AAC10FAFFB9FFD71FC8FEC3FD194B33" author="Hardy, D. E." box="[151,672,258,280]" page="255" pageId="47" pageNumber="48" year="1962">
<taxonomicName id="4C0D4D68FFB9FFD71FC8FEC3FD194B33" ID-CoL="8DD5L" authority="Hardy 1962: 255" authorityName="Hardy" authorityPageNumber="255" authorityYear="1962" box="[151,672,258,280]" class="Insecta" family="Pipunculidae" genus="Pipunculus" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="bisetosus" subGenus="Eudorylas">
<emphasis id="B979EAF9FFB9FFD71FC8FEC3FE524B33" box="[151,491,258,280]" italics="true" pageId="47" pageNumber="48">Pipunculus (Eudorylas) bisetosus</emphasis>
<bibRefCitation id="EF9C4B1AFFB9FFD71EAEFEC3FD194B33" author="Hardy" box="[497,672,258,280]" firstAuthor="Hardy" page="255" pageId="47" pageNumber="48" pagination="241 - 269" refId="ref88057" refString="Hardy, D. E. (1962) Preliminary studies on the Pipunculidae of Madagascar (Diptera). Verhandlungen der Naturforschenden Gesellschaft in Basel, 73, 241 - 269." type="journal article" year="1962">Hardy 1962: 255</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71FC8FE87FB2E4B8D" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BB236EBFFB9FFD71FC8FE87FB2E4B8D" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">Diagnosis: Third antennal segment very long acuminate. Two strong bristles on 3rd tibia at the apex anteriorly. T5 4 times as long as ST8 (hidden). Epandrium wide at base, pointed structure around cerci. SES distinct, with minute hairs; gonopods hairy, small, symmetric; phallic guide with numerous hairs in ventral view.</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71FC8FE72FA374BE1" box="[151,1422,434,459]" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="8BB236EBFFB9FFD71FC8FE72FA374BE1" blockId="47.[151,1436,325,1180]" box="[151,1422,434,459]" pageId="47" pageNumber="48">
<typeStatus id="54B68849FFB9FFD71FC8FE72FF684BE0" box="[151,209,435,459]" pageId="47" pageNumber="48">Type</typeStatus>
material:
<materialsCitation id="3B653CB6FFB9FFD71E1DFE72FA334BE1" ID-GBIF-Occurrence="3462316377" box="[322,1418,434,459]" collectingDate="18. I. 58" collectionCode="BMNH" collectorName="F. Keiser" country="Madagascar" location="Ambalamanakana" municipality="Fia" pageId="47" pageNumber="48" specimenCount="1" specimenCount-male="1" stateProvince="[Central] Madagascar" typeStatus="holotype">
<emphasis id="B979EAF9FFB9FFD71E1DFE72FE664BE0" bold="true" box="[322,479,435,459]" pageId="47" pageNumber="48">
<collectingCountry id="F31A767BFFB9FFD71E1DFE72FE634BE0" box="[322,474,435,459]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
:
</emphasis>
<specimenCount id="9D0BFD62FFB9FFD71EBAFE73FDB04BE1" box="[485,521,434,458]" pageId="47" pageNumber="48" type="male">1♂</specimenCount>
, HT,
<collectingRegion id="49C9F809FFB9FFD71D15FE73FCF94BE1" box="[586,832,434,458]" pageId="47" pageNumber="48">[Central] Madagascar</collectingRegion>
,
<collectingMunicipality id="6BD6AC91FFB9FFD71C14FE73FCC84BE1" box="[843,881,434,458]" pageId="47" pageNumber="48">Fia</collectingMunicipality>
,
<location id="8ED26030FFB9FFD71C22FE73FBF34BE1" LSID="urn:lsid:plazi:treatment:03A487FDFFB9FFD71D13FF56FE7E4EB7:8ED26030FFB9FFD71C22FE73FBF34BE1" box="[893,1098,434,458]" country="Madagascar" municipality="Fia" name="Ambalamanakana" pageId="47" pageNumber="48" stateProvince="[Central] Madagascar">Ambalamanakana</location>
,
<collectingDate id="EFF7E9C3FFB9FFD71B0AFE73FB1F4BE1" box="[1109,1190,434,458]" pageId="47" pageNumber="48">18.I.58</collectingDate>
,
<collectorName id="26F8533DFFB9FFD71BEDFE72FAA04BE1" box="[1202,1305,434,458]" pageId="47" pageNumber="48">F. Keiser</collectorName>
(
<collectionCode id="ED1CAE2EFFB9FFD71A77FE73FA3A4BE1" box="[1320,1411,434,458]" pageId="47" pageNumber="48">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71FC8FE17FEC24E53" pageId="47" pageNumber="48" type="description">
<paragraph id="8BB236EBFFB9FFD71FC8FE17FF694BC5" blockId="47.[151,1436,325,1180]" box="[151,208,470,494]" pageId="47" pageNumber="48">Male</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FE3AFD384871" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">
<emphasis id="B979EAF9FFB9FFD71FC8FE3AFF6D4838" box="[151,212,507,531]" italics="true" pageId="47" pageNumber="48">Head</emphasis>
. Third antennal segment very long acuminate; yellow. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 2.5 times ocellar triangle. Occiput, lower half silvery pollinose, upper half shining brown.
</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FDA7FE7548ED" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">
<emphasis id="B979EAF9FFB9FFD71FC8FDA7FF514855" box="[151,232,614,638]" italics="true" pageId="47" pageNumber="48">Thorax</emphasis>
. Humeri yellow. Mesonotum (viewed obliquely from front) subshining brown, yellow from the side. Scutellum yellow-brown, with 6–7 pairs of hairs (as long as f3 at widest). Dorsocentral hairs weakly developed. Halter yellow, knob brown.
</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FD12FAE4491F" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">Legs. Completely yellow. Ventroapical row of 5–7 small, black spines on f2; no hairs or spines on f1,3. Subapical (distal) spines on first four tibiae present. No anteromedial hairs on 3rd tibia, but two strong bristles at the apex anteriorly. Hind trochanter with minute whitish hairs on ventral side. Pulvilli shorter than last tarsal segment.</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FC81FB754973" blockId="47.[151,1436,325,1180]" box="[151,1228,832,856]" pageId="47" pageNumber="48">Wing. Cross-vein R-M at 2/5 of discal cell. Pterostigma fully coloured. Hairs on tegula missing.</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FCA5FC4749C3" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">
<emphasis id="B979EAF9FFB9FFD71FC8FCA5FEB84957" box="[151,257,868,892]" italics="true" pageId="47" pageNumber="48">Abdomen</emphasis>
. Viewed obliquely from front T5 subshining brown. T1–4 brownish yellow, hind margins with yellow markings laterally, sides yellow. Hairs dispersed, darkening towards tip of T5, as long as width of t3 at base. 2–3 yellow, lateral hairs on first tergite. Postabdomen in dorsal view: T5 4 times as long as ST8. Genitalia without dissection: ST8 hidden, dark brown, SS and EP yellow, m.a. small and round.
</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FC35FAF24E7F" blockId="47.[151,1436,325,1180]" pageId="47" pageNumber="48">
Genitalia. Surstyli subsymmetrical; epandrium wide at base, pointed structure around cerci (
<figureCitation id="13362A6EFFB9FFD71BBBFC35FAE84E27" box="[1252,1361,1012,1036]" captionStart="FIGURE 15" captionStartId="46.[151,250,1843,1866]" captionTargetBox="[297,1295,184,1820]" captionTargetId="figure-117@46.[276,1315,179,1829]" captionTargetPageId="46" captionText="FIGURE 15. Male genitalia of Eudorylas barracloughi sp. nov. (A–F) and E. bisetosus (G–L).—A: outer surstylus with phallic guide, attachment of phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: surstyli, dorsal view, E: sperm pump with ejaculatory apodeme, F: ventral view with phallic guide and phallus (without ST8), G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme, L: ventral view with phallic guide and subepandrial sclerite (without ST8). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6039188" httpUri="https://zenodo.org/record/6039188/files/figure.png" pageId="47" pageNumber="48">Fig. 15H</figureCitation>
). SES distinct, with minute hairs; gonopods small, symmetric, with hairs; phallic guide with numerous hairs in ventral view (
<figureCitation id="13362A6EFFB9FFD71F83FBFDFEF84E7F" box="[220,321,1084,1108]" captionStart="FIGURE 15" captionStartId="46.[151,250,1843,1866]" captionTargetBox="[297,1295,184,1820]" captionTargetId="figure-117@46.[276,1315,179,1829]" captionTargetPageId="46" captionText="FIGURE 15. Male genitalia of Eudorylas barracloughi sp. nov. (A–F) and E. bisetosus (G–L).—A: outer surstylus with phallic guide, attachment of phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: surstyli, dorsal view, E: sperm pump with ejaculatory apodeme, F: ventral view with phallic guide and phallus (without ST8), G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme, L: ventral view with phallic guide and subepandrial sclerite (without ST8). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6039188" httpUri="https://zenodo.org/record/6039188/files/figure.png" pageId="47" pageNumber="48">Fig. 15L</figureCitation>
); phallus trifid; ejaculatory apodeme broadening towards tip; sperm pump linear (
<figureCitation id="13362A6EFFB9FFD71B88FBFDFA864E7F" box="[1239,1343,1084,1108]" captionStart="FIGURE 15" captionStartId="46.[151,250,1843,1866]" captionTargetBox="[297,1295,184,1820]" captionTargetId="figure-117@46.[276,1315,179,1829]" captionTargetPageId="46" captionText="FIGURE 15. Male genitalia of Eudorylas barracloughi sp. nov. (A–F) and E. bisetosus (G–L).—A: outer surstylus with phallic guide, attachment of phallus and outer gonopod, lateral view, B: dorsal view, C: inner surstylus, lateral view, D: surstyli, dorsal view, E: sperm pump with ejaculatory apodeme, F: ventral view with phallic guide and phallus (without ST8), G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: dorsal view, I: inner surstylus, lateral view, J: surstyli, dorsal view, K: sperm pump with ejaculatory apodeme, L: ventral view with phallic guide and subepandrial sclerite (without ST8). Scale bars: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.6039188" httpUri="https://zenodo.org/record/6039188/files/figure.png" pageId="47" pageNumber="48">Fig. 15K</figureCitation>
).
</paragraph>
<paragraph id="8BB236EBFFB9FFD71FC8FBA1FEC24E53" blockId="47.[151,1436,325,1180]" box="[151,379,1120,1144]" pageId="47" pageNumber="48">
Female
<emphasis id="B979EAF9FFB9FFD71FB6FBA1FEBC4E53" bold="true" box="[233,261,1120,1144]" pageId="47" pageNumber="48">—</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3176560FFB9FFD71FC8FB45FE7E4EB7" box="[151,455,1156,1180]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BB236EBFFB9FFD71FC8FB45FE7E4EB7" blockId="47.[151,1436,325,1180]" box="[151,455,1156,1180]" pageId="47" pageNumber="48">
<emphasis id="B979EAF9FFB9FFD71FC8FB45FE814EB7" box="[151,312,1156,1180]" italics="true" pageId="47" pageNumber="48">Distribution—</emphasis>
<collectingCountry id="F31A767BFFB9FFD71E66FB45FE7D4EB7" box="[313,452,1156,1180]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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