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<document id="E46899D3415B1F2A60C5C5FE9E3AE382" ID-CLB-Dataset="230909" ID-DOI="10.5281/zenodo.7560404" ID-GBIF-Dataset="95dd82fb-b63c-41cf-b40c-39d13e9363bb" ID-ISSN="0035-418X" ID-Zenodo-Dep="7560404" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1641409684862" checkinUser="carolina" docAuthor="Yin, Zi-Wei &amp; Li, Lizhen" docDate="2013" docId="03A3879F6C56FFACFF7CF8CDFBBEFA9F" docLanguage="en" docName="RevSuisseZool.120.4.563-573.pdf.imf" docOrigin="Revue suisse de Zoologie 120 (4)" docStyle="DocumentStyle:0846B485C0C87F6EB7648DEEBF34B77F.2:RevSuisseZool.2011-2014.journal_article" docStyleId="0846B485C0C87F6EB7648DEEBF34B77F" docStyleName="RevSuisseZool.2011-2014.journal_article" docStyleVersion="2" docTitle="Nomuraius Hlavac 2003" docType="treatment" docVersion="5" lastPageNumber="565" masterDocId="FF9AFFE76C57FFAEFFEBFA6DFF9CFF9B" masterDocTitle="Taxonomic notes on the genus Nomuraius Hlaváč (Staphylinidae: Pselaphinae)" masterLastPageNumber="573" masterPageNumber="563" pageNumber="564" updateTime="1699295947412" updateUser="ExternalLinkService">
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<mods:title id="CEFAEEF70DF425061A0825C7984679D0">Taxonomic notes on the genus Nomuraius Hlaváč (Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="F9843B734ED0BF7C6557C3D46837055B">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="7082425DF153F677D7072C044AA13269">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road., Xuhui District, Shanghai, 200234, P. R. China.</mods:affiliation>
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<mods:namePart id="AA8CEA6D43F1B18EBB15792A5BAFA446">Li, Lizhen</mods:namePart>
<mods:affiliation id="806F18D7976648567D9495AD68F35F1D">Department of Biology, College of Life and Environmental Sciences, Shanghai Normal University, 100 Guilin Road., Xuhui District, Shanghai, 200234, P. R. China.</mods:affiliation>
<mods:nameIdentifier id="B88C1E2120A5FD6F6E8983FA7929AF9C" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:title id="0778BB2E58EB34999E733176C0E0525C">Revue suisse de Zoologie</mods:title>
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<mods:date id="9EC7E834B736FD5F5CC0FF0D25D4B27F">2013</mods:date>
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<mods:number id="05A34D181974FEB6A0B9A6CD1470063B">2013-12-31</mods:number>
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<mods:number id="C1AC1D42B41277A7E0092675FB34ACFD">120</mods:number>
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<mods:number id="1BBF6D6AB9EAB66E3769F3552F32D84E">4</mods:number>
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<mods:identifier id="27B4814C13CB5BA2FD59A70DD3F2835D" type="CLB-Dataset">230909</mods:identifier>
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<treatment id="03A3879F6C56FFACFF7CF8CDFBBEFA9F" ID-DOI="http://doi.org/10.5281/zenodo.7560406" ID-GBIF-Taxon="209100137" ID-Zenodo-Dep="7560406" LSID="urn:lsid:plazi:treatment:03A3879F6C56FFACFF7CF8CDFBBEFA9F" httpUri="http://treatment.plazi.org/id/03A3879F6C56FFACFF7CF8CDFBBEFA9F" lastPageId="2" lastPageNumber="565" pageId="1" pageNumber="564">
<subSubSection id="C31065026C56FFAFFF7CF8CDFE36FD22" box="[151,426,672,697]" pageId="1" pageNumber="564" type="nomenclature">
<paragraph id="8BB536896C56FFAFFF7CF8CDFE36FD22" blockId="1.[151,1073,672,727]" box="[151,426,672,697]" pageId="1" pageNumber="564">
<heading id="D0FD81E56C56FFAFFF7CF8CDFE36FD22" box="[151,426,672,697]" fontSize="10" level="2" pageId="1" pageNumber="564" reason="2">
<taxonomicName id="4C0A4D0A6C56FFAFFF7CF8CDFE36FD22" ID-CoL="8XN84" authority="Hlavac, 2003" authorityName="Hlavac" authorityYear="2003" box="[151,426,672,697]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C56FFAFFF7CF8CDFE8FFD23" bold="true" box="[151,275,672,696]" italics="true" pageId="1" pageNumber="564">Nomuraius</emphasis>
<bibRefCitation id="EF9B4B786C56FFAFFEF0F8CDFE36FD22" author="HLAVAC, P." box="[283,426,672,697]" firstAuthor="Hlavac" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="1" pageNumber="564" pagination="283 - 297" part="38" refId="ref4436" refString="HLAVAC, P. 2003. A taxonomic revision of the Tyrini of the Oriental Region. II. - Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." title="A taxonomic revision of the Tyrini of the Oriental Region. II. - Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2003">Hlaváč, 2003</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31065026C56FFAFFF7CF8AFFE21FD4D" box="[151,445,705,727]" pageId="1" pageNumber="564" type="reference_group">
<paragraph id="8BB536896C56FFAFFF7CF8AFFE21FD4D" blockId="1.[151,1073,672,727]" box="[151,445,705,727]" pageId="1" pageNumber="564">
<treatmentCitationGroup id="AB1A11A76C56FFAFFF7CF8AFFE21FD4D" box="[151,445,705,727]" pageId="1" pageNumber="564">
<treatmentCitation id="0AAB10986C56FFAFFF7CF8AFFE2AFD4C" author="HLAVAC, P." box="[151,438,705,727]" page="290" pageId="1" pageNumber="564" year="2003">
<taxonomicName id="4C0A4D0A6C56FFAFFF7CF8AFFE2AFD4C" ID-CoL="8XN84" authority="Hlavac, 2003: 290" authorityName="Hlavac" authorityPageNumber="290" authorityYear="2003" box="[151,438,705,727]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C56FFAFFF7CF8AFFE9DFD4C" box="[151,257,706,727]" italics="true" pageId="1" pageNumber="564">Nomuraius</emphasis>
<bibRefCitation id="EF9B4B786C56FFAFFEECF8ACFE2AFD4C" author="HLAVAC, P." box="[263,438,705,727]" firstAuthor="Hlavac" journalOrPublisher="Annales de la Societe Entomologique de France" page="290" pageId="1" pageNumber="564" pagination="283 - 297" part="38" refId="ref4436" refString="HLAVAC, P. 2003. A taxonomic revision of the Tyrini of the Oriental Region. II. - Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae). Annales de la Societe Entomologique de France 38: 283 - 297." title="A taxonomic revision of the Tyrini of the Oriental Region. II. - Systematic study on the genus Pselaphodes and its allied genera (Coleoptera: Staphylinidae: Pselaphinae)" type="journal article" year="2003">Hlaváč, 2003: 290</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31065026C56FFAFFE29F8ACFBADFD4D" box="[450,1073,705,727]" pageId="1" pageNumber="564" type="type_taxon">
<paragraph id="8BB536896C56FFAFFE29F8ACFBADFD4D" blockId="1.[151,1073,672,727]" box="[450,1073,705,727]" pageId="1" pageNumber="564">
<typeStatus id="54B1882B6C56FFAFFE29F8ACFE77FD4D" box="[450,491,705,726]" pageId="1" pageNumber="564">type</typeStatus>
species:
<taxonomicName id="4C0A4D0A6C56FFAFFDA9F8AFFC35FD4C" ID-CoL="47P94" authority="Hlavac, 2002" authorityName="Hlavac" authorityYear="2002" box="[578,937,705,727]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B97EEA9B6C56FFAFFDA9F8AFFCB8FD4C" box="[578,804,706,727]" italics="true" pageId="1" pageNumber="564">Nomuraius vietnamicus</emphasis>
Hlaváč, 2002
</taxonomicName>
by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C31065026C56FFAFFF08F885FF77FC24" pageId="1" pageNumber="564" type="diagnosis">
<paragraph id="8BB536896C56FFAFFF08F885FF77FC24" blockId="1.[151,1074,743,959]" pageId="1" pageNumber="564">DIAGNOSIS: Head with distinct frontal rostrum; with small, nude vertexal foveae; lacking frontal fovea; lacking ocular-mandibular carinae; maxillary palpi asymmetric, palpomeres I short, II distinctly pedunculate at bases, roundly expanded to shortly projecting laterally, palpomeres III-IV shortly pedunculate at bases, distinctly projecting laterally. Pronotum with nude median- and lateral antebasal foveae weakly to distinctly marked, lacking antebasal sulcus. Abdomen with tergite IV longest.</paragraph>
</subSubSection>
<subSubSection id="C31065026C56FFACFF08F9A2FD4EFEA0" lastPageId="2" lastPageNumber="565" pageId="1" pageNumber="564" type="description">
<paragraph id="8BB536896C56FFAFFF08F9A2FBAEFB18" blockId="1.[151,1074,974,1659]" pageId="1" pageNumber="564">
REDESCRIPTION: Length
<quantity id="4CF29B6C6C56FFAFFDEAF9A3FD3DFC7D" box="[513,673,974,998]" metricMagnitude="-3" metricUnit="m" metricValue="2.905" metricValueMax="3.11" metricValueMin="2.7" pageId="1" pageNumber="564" unit="mm" value="2.905" valueMax="3.11" valueMin="2.7">2.70-3.11 mm</quantity>
. Head with short, narrow frontal rostrum, rostrum slightly prominent, antennal tubercles weak; nude vertexal foveae small and nude, lacking frontal fovea, lateral postantennal pits small; with 11 antennomeres, clubs formed by apical three antennomeres; maxillary palpi asymmetric, palpomere II widest near apices, roundly expanded to slightly projecting laterally, III-IV widest near middle, distinctly projecting laterally. Gular flat, shallow foveae separated.
</paragraph>
<paragraph id="8BB536896C56FFAFFF08FEE1FEC9FAE4" blockId="1.[151,1074,974,1659]" pageId="1" pageNumber="564">Pronotum with small punctiform to large round median- and lateral antebasal foveae, median longitudinal sulcus present, lacking antebasal sulcus; lateral procoxal foveae deep. Each elytron with two basal foveae; with one subbasal fovea; discal striae extending from outer basal foveae posteriorly beyond elytral midpoint. Thorax with two median mesoventral foveae widely separated; large lateral mesoventral foveae forked, anterior branch larger than posterior branch; lateral mesocoxal foveae present; lacking median and lateral metaventral foveae; posterior margin of metaventrite deeply notched medially.</paragraph>
<paragraph id="8BB536896C56FFAFFF08FFE5FEB7FA25" blockId="1.[151,1074,974,1659]" pageId="1" pageNumber="564">Legs with tasomeres I short, II greatly lobed, extending beyond midpoint of tasomeres III.</paragraph>
<paragraph id="8BB536896C56FFAFFF08FFABFBAEF9E0" blockId="1.[151,1074,974,1659]" pageId="1" pageNumber="564">Abdomen with tergite IV longest, more than twice length of next tergite, VI shortest, V and VII subequal in length; tergite IV with mediobasal foveae moved laterally to form large round pockets at end of basal sulcus, with two small basolateral foveae; tergites V-VII each with one pair of small basolateral foveae. Sternite IV longest, as long as V-VII combined along midlength, lacking mediobasal foveae, with large pockets formed by basolateral foveae; V-VII with basolateral foveae weakly indicated.</paragraph>
<paragraph id="8BB536896C55FFACFF3FFA8EFD4EFEA0" blockId="2.[136,1059,227,315]" pageId="2" pageNumber="565">Males with posterior half of head, apical portions of pronotum, protibiae and metatibiae variously modified. Aedeagus with asymmetric median lobe; elongate parameres symmetric; diaphragm opening nearly oval.</paragraph>
</subSubSection>
<subSubSection id="C31065026C55FFACFF3FFB23FDF3FE1E" pageId="2" pageNumber="565" type="distribution">
<paragraph id="8BB536896C55FFACFF3FFB23FDF3FE1E" blockId="2.[136,1059,333,390]" pageId="2" pageNumber="565">
DISTRIBUTION: Three species are known from South
<collectingCountry id="F31D76196C55FFACFCC9FB20FCFEFEFE" box="[802,866,333,357]" name="China" pageId="2" pageNumber="565">China</collectingCountry>
(
<taxonomicName id="4C0A4D0A6C55FFACFC9EFB22FC7EFEFE" box="[885,994,334,358]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="sinicus">
<emphasis id="B97EEA9B6C55FFACFC9EFB22FC7EFEFE" box="[885,994,334,358]" italics="true" pageId="2" pageNumber="565">N. sinicus</emphasis>
</taxonomicName>
), and north
<collectingCountry id="F31D76196C55FFACFF2CFB00FEBDFE1E" box="[199,289,365,389]" name="Vietnam" pageId="2" pageNumber="565">Vietnam</collectingCountry>
(
<taxonomicName id="4C0A4D0A6C55FFACFED9FB02FE2DFE1E" box="[306,433,366,390]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="piaoacus">
<emphasis id="B97EEA9B6C55FFACFED9FB02FE2DFE1E" box="[306,433,366,390]" italics="true" pageId="2" pageNumber="565">N. piaoacus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A4D0A6C55FFACFE2BFB02FDC3FE1E" box="[448,607,366,390]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B97EEA9B6C55FFACFE2BFB02FDC3FE1E" box="[448,607,366,390]" italics="true" pageId="2" pageNumber="565">N. vietnamicus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31065026C55FFACFF3FFBF5FE21FD3D" pageId="2" pageNumber="565" type="discussion">
<paragraph id="8BB536896C55FFACFF3FFBF5FC27FDEB" blockId="2.[136,1059,408,624]" pageId="2" pageNumber="565">
COMPARATIVE NOTES: Among the members of the
<taxonomicName id="4C0A4D0A6C55FFACFCF9FBF5FC05FE2B" authorityName="Westwood" authorityYear="1870" box="[786,921,408,432]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFCF9FBF5FC05FE2B" box="[786,921,408,432]" italics="true" pageId="2" pageNumber="565">Pselaphodes</emphasis>
</taxonomicName>
complex of genera,
<taxonomicName id="4C0A4D0A6C55FFACFF34FBD4FECAFE4B" box="[223,342,441,464]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFF34FBD4FECAFE4B" box="[223,342,441,464]" italics="true" pageId="2" pageNumber="565">Nomuraius</emphasis>
</taxonomicName>
shares only with
<taxonomicName id="4C0A4D0A6C55FFACFDF5FBD5FC85FE4A" authority="Hlavac" authorityName="Hlavac" authorityYear="2002" box="[542,793,440,465]" class="Insecta" family="Staphylinidae" genus="Taiwanophodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFDF5FBD5FD58FE4B" box="[542,708,440,464]" italics="true" pageId="2" pageNumber="565">Taiwanophodes</emphasis>
Hlaváč
</taxonomicName>
the greatly lobed tarsomeres II. The two genera can be readily separated by the vertexal foveae being nude, the absence of a frontal fovea, and the pronotum lacking an antebasal sulcus connecting the nude median- and lateral antebasal foveae in
<taxonomicName id="4C0A4D0A6C55FFACFCB1F874FC4DFDAB" box="[858,977,537,560]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFCB1F874FC4DFDAB" box="[858,977,537,560]" italics="true" pageId="2" pageNumber="565">Nomuraius</emphasis>
</taxonomicName>
, while
<taxonomicName id="4C0A4D0A6C55FFACFF63F855FEB2FDCB" authorityName="Hlavac" authorityYear="2002" box="[136,302,568,592]" class="Insecta" family="Staphylinidae" genus="Taiwanophodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFF63F855FEB2FDCB" box="[136,302,568,592]" italics="true" pageId="2" pageNumber="565">Taiwanophodes</emphasis>
</taxonomicName>
has setose vertexal and frontal foveae, and the pronotal median- and lateral antebasal foveae are setose and are connected by an antebasal sulcus.
</paragraph>
<paragraph id="8BB536896C55FFACFF63F8E3FE21FD3D" blockId="2.[136,445,654,678]" box="[136,445,654,678]" pageId="2" pageNumber="565">
<heading id="D0FD81E56C55FFACFF63F8E3FE21FD3D" box="[136,445,654,678]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="565" reason="2">
<emphasis id="B97EEA9B6C55FFACFF63F8E3FE21FD3D" bold="true" box="[136,445,654,678]" pageId="2" pageNumber="565">
Key to males of
<taxonomicName id="4C0A4D0A6C55FFACFEAAF8E3FE21FD3D" box="[321,445,654,678]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA9B6C55FFACFEAAF8E3FE21FD3D" bold="true" box="[321,445,654,678]" italics="true" pageId="2" pageNumber="565">Nomuraius</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31065026C55FFACFF63F8D8FBBEFA9F" pageId="2" pageNumber="565" type="key">
<keyStep id="30FE2E2C6C55FFACFF63F8D8FBBEFC52" pageId="2" pageNumber="565">
<paragraph id="8BB536896C55FFACFF63F8D8FBBFFCD0" blockId="2.[136,1059,693,1284]" pageId="2" pageNumber="565">
<keyLead id="30FB95BC6C55FFACFF63F8D8FBBFFCD0" pageId="2" pageNumber="565">
1a Head with broad, deep cavity at posterior half (
<figureCitation id="13312A0C6C55FFACFD3CF8D8FCB6FD56" box="[727,810,693,717]" captionStart="FIG" captionStartId="3.[583,596,1118,1139]" captionTargetBox="[166,1060,237,1086]" captionTargetId="figure-849@3.[151,1073,229,1094]" captionTargetPageId="3" captionText="FIG. 1 Male habitus of Nomuraius Hlaváč. (A) N. piaoacus sp. n. (B) N. sinicus sp. n. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7560408" httpUri="https://zenodo.org/record/7560408/files/figure.png" pageId="2" pageNumber="565">Fig. 1A</figureCitation>
); antennomeres X much shorter than XI (
<figureCitation id="13312A0C6C55FFACFDEFF8B9FDC0FD77" box="[516,604,724,748]" captionStart="FIG" captionStartId="4.[568,581,1159,1180]" captionTargetBox="[145,1048,227,1125]" captionTargetId="figure-412@4.[136,1058,227,1134]" captionTargetPageId="4" captionText="FIG. 2 Male diagnostic features of Nomuraius piaoacus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560412" httpUri="https://zenodo.org/record/7560412/files/figure.png" pageId="2" pageNumber="565">Fig. 2A</figureCitation>
); pronotum strongly modified at anterior portion (
<figureCitation id="13312A0C6C55FFACFE7FF899FE75FC97" box="[404,489,756,780]" captionStart="FIG" captionStartId="4.[568,581,1159,1180]" captionTargetBox="[145,1048,227,1125]" captionTargetId="figure-412@4.[136,1058,227,1134]" captionTargetPageId="4" captionText="FIG. 2 Male diagnostic features of Nomuraius piaoacus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560412" httpUri="https://zenodo.org/record/7560412/files/figure.png" pageId="2" pageNumber="565">Fig. 2B</figureCitation>
); metatibiae simple, not expanded medially (
<figureCitation id="13312A0C6C55FFACFF36F97EFEB2FCB0" box="[221,302,787,811]" captionStart="FIG" captionStartId="4.[568,581,1159,1180]" captionTargetBox="[145,1048,227,1125]" captionTargetId="figure-412@4.[136,1058,227,1134]" captionTargetPageId="4" captionText="FIG. 2 Male diagnostic features of Nomuraius piaoacus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560412" httpUri="https://zenodo.org/record/7560412/files/figure.png" pageId="2" pageNumber="565">Fig. 2H</figureCitation>
). (north
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).......................................................
<taxonomicName id="4C0A4D0A6C55FFACFD8EF959FC37FCD0" ID-CoL="47P92" authority="Yin &amp; Li" authorityName="Yin &amp; Li" box="[613,939,819,843]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="piaoacus" status="sp. nov.">
<emphasis id="B97EEA9B6C55FFACFD8EF959FCDFFCD0" box="[613,835,820,843]" italics="true" pageId="2" pageNumber="565">Nomuraius piaoacus</emphasis>
Yin &amp; Li
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<paragraph id="8BB536896C55FFACFF63F93FFBBEFC52" blockId="2.[136,1059,693,1284]" pageId="2" pageNumber="565">
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1b Head concaved only at posterior margin (
<figureCitation id="13312A0C6C55FFACFD64F93FFD7FFCF1" box="[655,739,850,874]" captionStart="FIG" captionStartId="3.[583,596,1118,1139]" captionTargetBox="[166,1060,237,1086]" captionTargetId="figure-849@3.[151,1073,229,1094]" captionTargetPageId="3" captionText="FIG. 1 Male habitus of Nomuraius Hlaváč. (A) N. piaoacus sp. n. (B) N. sinicus sp. n. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7560408" httpUri="https://zenodo.org/record/7560408/files/figure.png" pageId="2" pageNumber="565">Figs 1B</figureCitation>
,
<figureCitation id="13312A0C6C55FFACFD1AF93FFC8DFCF1" box="[753,785,850,874]" captionStart="FIG" captionStartId="7.[583,596,1229,1250]" captionTargetBox="[167,1058,238,1195]" captionTargetId="figure-559@7.[151,1073,227,1201]" captionTargetPageId="7" captionText="FIG. 4 Nomuraius vietnamicus Hlaváč, 2003. (A) Male habitus (non-type). (B) Holotype habitus. (C) Type label. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7560418" httpUri="https://zenodo.org/record/7560418/files/figure.png" pageId="2" pageNumber="565">4A</figureCitation>
); antennomeres X slightly shorter than XI (
<figureCitation id="13312A0C6C55FFACFE35F91FFDAFFC11" box="[478,563,882,906]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Figs 3A</figureCitation>
,
<figureCitation id="13312A0C6C55FFACFDAAF91FFDFEFC11" box="[577,610,882,906]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">5A</figureCitation>
); pronotum only slightly modified at anterior portion (
<figureCitation id="13312A0C6C55FFACFE5AF9FCFD95FC32" box="[433,521,913,937]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Figs 3B</figureCitation>
,
<figureCitation id="13312A0C6C55FFACFDF0F9FCFDA6FC32" box="[539,570,913,937]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">5B</figureCitation>
); metatibiae expanded medially near apices (
<figureCitation id="13312A0C6C55FFACFECDF9DCFEE0FC53" box="[294,380,945,969]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Figs 3H</figureCitation>
,
<figureCitation id="13312A0C6C55FFACFE61F9DCFE37FC53" box="[394,427,945,969]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">5H</figureCitation>
)........................................... 2
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<keyStep id="30FE2E2C6C55FFACFF63F9BDFBBEFA9F" pageId="2" pageNumber="565">
<paragraph id="8BB536896C55FFACFF63F9BDFBBFFBFD" blockId="2.[136,1059,693,1284]" pageId="2" pageNumber="565">
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2a Pronotum and elytra relatively more elongate (
<figureCitation id="13312A0C6C55FFACFD25F9BDFCBCFC73" box="[718,800,976,1000]" captionStart="FIG" captionStartId="3.[583,596,1118,1139]" captionTargetBox="[166,1060,237,1086]" captionTargetId="figure-849@3.[151,1073,229,1094]" captionTargetPageId="3" captionText="FIG. 1 Male habitus of Nomuraius Hlaváč. (A) N. piaoacus sp. n. (B) N. sinicus sp. n. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7560408" httpUri="https://zenodo.org/record/7560408/files/figure.png" pageId="2" pageNumber="565">Fig. 1B</figureCitation>
); metatibiae less expanded medially near apices (
<figureCitation id="13312A0C6C55FFACFDDFF99DFD14FB9C" box="[564,648,1008,1032]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Fig. 3H</figureCitation>
); tergite VIII about as long as wide (
<figureCitation id="13312A0C6C55FFACFEF7FE62FEF4FBBC" box="[284,360,1039,1063]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Fig. 3I</figureCitation>
); apical portion of aedeagal median lobe bent leftward dorsoventrally (
<figureCitation id="13312A0C6C55FFACFE95FE43FE72FBDD" box="[382,494,1070,1094]" captionStart="FIG" captionStartId="5.[583,596,1136,1157]" captionTargetBox="[166,1064,228,1094]" captionTargetId="figure-628@5.[152,1074,226,1098]" captionTargetPageId="5" captionText="FIG. 3 Male diagnostic features of Nomuraius sinicus sp. n. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560416" httpUri="https://zenodo.org/record/7560416/files/figure.png" pageId="2" pageNumber="565">Fig. 3L, N</figureCitation>
). (South
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)..............................................
<taxonomicName id="4C0A4D0A6C55FFACFD97FE22FC37FBFD" ID-CoL="47P93" authority="Yin &amp; Li" authorityName="Yin &amp; Li" box="[636,939,1102,1126]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="sinicus" status="sp. nov.">
<emphasis id="B97EEA9B6C55FFACFD97FE22FCD8FBFD" box="[636,836,1103,1126]" italics="true" pageId="2" pageNumber="565">Nomuraius sinicus</emphasis>
Yin &amp; Li
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<paragraph id="8BB536896C55FFACFF63FE00FBBEFA9F" blockId="2.[136,1059,693,1284]" pageId="2" pageNumber="565">
<keyLead id="30FB95BC6C55FFACFF63FE00FBBEFA9F" pageId="2" pageNumber="565">
2b Pronotum and elytra relatively shorter (
<figureCitation id="13312A0C6C55FFACFD4DFE00FC9DFB1E" box="[678,769,1133,1157]" captionStart="FIG" captionStartId="7.[583,596,1229,1250]" captionTargetBox="[167,1058,238,1195]" captionTargetId="figure-559@7.[151,1073,227,1201]" captionTargetPageId="7" captionText="FIG. 4 Nomuraius vietnamicus Hlaváč, 2003. (A) Male habitus (non-type). (B) Holotype habitus. (C) Type label. Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7560418" httpUri="https://zenodo.org/record/7560418/files/figure.png" pageId="2" pageNumber="565">Fig. 4A</figureCitation>
); metatibiae more expanded medially near apices (
<figureCitation id="13312A0C6C55FFACFDD9FEE0FD19FB3E" box="[562,645,1165,1189]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">Fig. 5H</figureCitation>
); tergite VIII much wider than long (
<figureCitation id="13312A0C6C55FFACFEFDFEC1FEFDFB5F" box="[278,353,1196,1220]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">Fig. 5I</figureCitation>
); apical portion of aedeagal median lobe bent rightward dorsoventrally (
<figureCitation id="13312A0C6C55FFACFE95FEA1FE72FB7F" box="[382,494,1228,1252]" captionStart="FIG" captionStartId="8.[568,581,1175,1196]" captionTargetBox="[140,1059,231,1132]" captionTargetId="figure-529@8.[139,1060,227,1136]" captionTargetPageId="8" captionText="FIG. 5 Male diagnostic features of Nomuraius vietnamicus Hlaváč, 2003. (A) Antenna. (B) Pronotum. (C) Metaventral process, in lateral view. (D) Protrochanter and profemur. (E) Apical portion of protibia. (F) Mesotrochanter and mesofemur. (G) metatrochanter and metafemur. (H) Apical portion of metatibia. (I) Tergite VIII. (J) Sternite VIII. (K) Sternite IX. (L) Aedeagus, in dorsal view. (M) Same, in lateral view. (N) Same, in ventral view. Scales [mm]: A, B, D, F, G = 0.3; C, I, J, L, M, N = 0.2; H, K = 0.1, E = 0.05." figureDoi="http://doi.org/10.5281/zenodo.7560422" httpUri="https://zenodo.org/record/7560422/files/figure.png" pageId="2" pageNumber="565">Fig. 5L, N</figureCitation>
). (north
<collectingCountry id="F31D76196C55FFACFDA6FEA1FD3BFB7F" box="[589,679,1228,1252]" name="Vietnam" pageId="2" pageNumber="565">Vietnam</collectingCountry>
: Vinh Phu)..................................................
<taxonomicName id="4C0A4D0A6C55FFACFD38FE81FBBEFA9F" ID-CoL="47P94" authority="Hlavac" authorityName="Hlavac" box="[723,1058,1259,1284]" class="Insecta" family="Staphylinidae" genus="Nomuraius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="565" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B97EEA9B6C55FFACFD38FE81FC53FA98" box="[723,975,1260,1283]" italics="true" pageId="2" pageNumber="565">Nomuraius vietnamicus</emphasis>
Hlaváč
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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