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379 lines
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<document id="9C9C258C99B95BF919C3B88399BA0106" ID-DOI="10.5281/zenodo.189740" ID-GBIF-Dataset="8dc3930d-f5da-496f-a86f-f0da1543811a" ID-ISSN="1175-5326" ID-Zenodo-Dep="189740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461104606930" checkinUser="plazi" docAuthor="Walter, David Evans" docDate="2009" docId="039F612DFFBFFA7640AAD7EFFED8FC28" docLanguage="en" docName="zt02206p044.pdf" docOrigin="Zootaxa 2206" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Gymnodamaeus Kulczynski 1902" docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FFA61955FFBCFA72403DD778FFA2FFB8" masterDocTitle="Genera of Gymnodamaeidae (Acari: Oribatida: Plateremaeoidea) of Canada, with notes on some nomenclatorial problems" masterLastPageNumber="44" masterPageNumber="23" pageNumber="26" updateTime="1698592501701" updateUser="plazi">
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<mods:title id="B68EC4CA5C729CFFF0EA202EBF9DDA3A">Genera of Gymnodamaeidae (Acari: Oribatida: Plateremaeoidea) of Canada, with notes on some nomenclatorial problems</mods:title>
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<mods:namePart id="C327F55FC391EF3BE2CD89E28D29D949">Walter, David Evans</mods:namePart>
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<mods:title id="5DC0C772AB4CBE79894DE9E3D4A3B000">Zootaxa</mods:title>
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<mods:part id="3A6A17D0BEBE5B618A8540F64336C31E">
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<mods:date id="A0078A06FC7A07C69420BDECF8D68F20">2009</mods:date>
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<mods:detail id="16CE4ACCBEE35179F045576CE7B1847A" type="volume">
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<mods:number id="1B260AFEA44F084C971509A0211F5882">2206</mods:number>
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<mods:identifier id="E24B09B1FA74173538B1639FF49C5833" type="DOI">10.5281/zenodo.189740</mods:identifier>
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<mods:identifier id="01F5BC6455C0AD4D0FC8230CA736DCEA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="674A0FFBFDF7F01F45FBA3E257D11A98" type="Zenodo-Dep">189740</mods:identifier>
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<treatment id="039F612DFFBFFA7640AAD7EFFED8FC28" ID-DOI="http://doi.org/10.5281/zenodo.6222996" ID-GBIF-Taxon="119622858" ID-Zenodo-Dep="6222996" LSID="urn:lsid:plazi:treatment:039F612DFFBFFA7640AAD7EFFED8FC28" httpUri="http://treatment.plazi.org/id/039F612DFFBFFA7640AAD7EFFED8FC28" lastPageId="4" lastPageNumber="27" pageId="3" pageNumber="26">
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<subSubSection id="C32C83B0FFBFFA7140AAD7EFFD94FF09" box="[151,566,151,177]" pageId="3" pageNumber="26" type="nomenclature">
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<paragraph id="8B89D03BFFBFFA7140AAD7EFFD94FF09" blockId="3.[151,566,151,177]" box="[151,566,151,177]" pageId="3" pageNumber="26">
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<heading id="D0C16757FFBFFA7140AAD7EFFD94FF09" bold="true" box="[151,566,151,177]" fontSize="11" level="1" pageId="3" pageNumber="26" reason="1">
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<taxonomicName id="4C36ABB8FFBFFA7140AAD7EFFD94FF09" ID-CoL="4RTB" authority="Kulczynski, 1902" authorityName="Kulczynski" authorityYear="1902" box="[151,566,151,177]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9420C29FFBFFA7140AAD7EFFD94FF09" bold="true" box="[151,566,151,177]" pageId="3" pageNumber="26">
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<emphasis id="B9420C29FFBFFA7140AAD7EFFEF8FF09" bold="true" box="[151,346,151,177]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus</emphasis>
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<bibRefCitation id="EFA7ADCAFFBFFA71415CD7EFFD94FF09" author="Kulczynski" box="[353,566,151,177]" pageId="3" pageNumber="43" refString="Kulczynski, V. (1902) Species Oribatinarum (Oudms.) (Damaeinarum Michael) in Galicia collectae. Bulletin of the International Academy of Science, Cracow, 2, 89 - 96." type="journal article" year="1902">Kulczynski, 1902</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C83B0FFBFFA7140AAD79DFE6FFF44" box="[151,461,229,252]" pageId="3" pageNumber="26" type="reference_group">
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<paragraph id="8B89D03BFFBFFA7140AAD79DFE6FFF44" blockId="3.[151,702,229,284]" box="[151,461,229,252]" pageId="3" pageNumber="26">
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<taxonomicName id="4C36ABB8FFBFFA7140AAD79DFE6FFF44" ID-CoL="8T5PR" authority="Ewing, 1917" authorityName="Ewing" authorityYear="1917" box="[151,461,229,252]" class="Arachnida" family="Gymnodamaeidae" genus="Heterodamaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
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<emphasis id="B9420C29FFBFFA7140AAD79DFE9DFF44" box="[151,319,229,252]" italics="true" pageId="3" pageNumber="26">Heterodamaeus</emphasis>
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<bibRefCitation id="EFA7ADCAFFBFFA714178D79DFE6FFF44" author="Ewing" box="[325,461,229,252]" pageId="3" pageNumber="43" refString="Ewing, H. E. (1917) A synopsis of the genera of beetle mites with special reference to the North American fauna. Annals of the Entomological Society of America, 10, 117 - 132." type="journal article" year="1917">Ewing, 1917</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C83B0FFBFFA7140AAD67DFD1CFEA4" box="[151,702,261,284]" pageId="3" pageNumber="26" type="materials_examined">
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<paragraph id="8B89D03BFFBFFA7140AAD67DFD1CFEA4" blockId="3.[151,702,229,284]" box="[151,702,261,284]" pageId="3" pageNumber="26">
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<typeStatus id="548D6E99FFBFFA7140AAD67DFF6EFEA4" box="[151,204,261,284]" pageId="3" pageNumber="26">Type</typeStatus>
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species:
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<taxonomicName id="4C36ABB8FFBFFA714110D67DFD1CFEA4" authority="C.L. Koch, 1835" authorityName="C.L. Koch" authorityYear="1835" box="[301,702,261,284]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
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<emphasis id="B9420C29FFBFFA714110D67DFDA7FEA4" box="[301,517,261,284]" italics="true" pageId="3" pageNumber="26">Damaeus bicostatus</emphasis>
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<bibRefCitation id="EFA7ADCAFFBFFA714231D67DFD1CFEA4" author="Koch" box="[524,702,261,284]" pageId="3" pageNumber="43" refString="Koch, C. L. (1835 - 1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Vol. I - XL, Regensburg." type="book" year="1835">C.L. Koch, 1835</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32C83B0FFBFFA7140AAD633FD18FCF9" pageId="3" pageNumber="26" type="diagnosis">
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<paragraph id="8B89D03BFFBFFA7140AAD633FD18FCF9" blockId="3.[151,1437,331,2032]" pageId="3" pageNumber="26">
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<emphasis id="B9420C29FFBFFA7140AAD633FEBBFEDD" bold="true" box="[151,281,331,357]" pageId="3" pageNumber="26">Diagnosis:</emphasis>
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<taxonomicName id="4C36ABB8FFBFFA71411FD634FE5AFEDE" box="[290,504,332,358]" class="Arachnida" family="Gymnodamaeidae" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="family">Gymnodamaeidae</taxonomicName>
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with a ventral plate typically lacking an anogenital bridge (
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<figureCitation id="130DCCBEFFBFFA714486D634FA8CFEDE" box="[1211,1326,332,358]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Figs. 4, 7</figureCitation>
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, arrows), but with lateral pits; genital plates with smooth median margin and bearing 7 pairs of setae, seta
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<emphasis id="B9420C29FFBFFA714522D60AFA8CFE33" box="[1311,1326,370,395]" italics="true" pageId="3" pageNumber="26">g</emphasis>
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2 inserted posteriad
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<emphasis id="B9420C29FFBFFA714134D6E4FEBAFE0D" box="[265,280,412,437]" italics="true" pageId="3" pageNumber="26">g</emphasis>
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1; 2 pairs of adanal setae. Legs without retrotecta on distal segments (
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<figureCitation id="130DCCBEFFBFFA714406D6E4FB23FE0E" box="[1083,1153,412,438]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Fig. 8</figureCitation>
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), femur IV with 2 setae. Prodorsum without X-shaped raised cerotegumental ridges (
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<figureCitation id="130DCCBEFFBFFA714362D6BDFBAFFE67" box="[863,1037,453,479]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Figs. 1–2, 5–6</figureCitation>
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), with spherical pustules (
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<figureCitation id="130DCCBEFFBFFA71457ED6BDFA29FE67" box="[1347,1419,453,479]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Fig. 3</figureCitation>
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); sensillus (
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<emphasis id="B9420C29FFBFFA714130D694FE88FDBD" box="[269,298,492,517]" italics="true" pageId="3" pageNumber="26">bo</emphasis>
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) with long stalk and elongate, barbed head (
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<figureCitation id="130DCCBEFFBFFA714305D694FC23FDBE" box="[824,897,492,518]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Fig. 2</figureCitation>
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); bothridial rim simple, bothridial ridges well developed, arising anteriad bothridia, and ending in strong apophyses bearing the spine-like interlamellar seta (
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<emphasis id="B9420C29FFBFFA71409CD542FF1BFDEB" box="[161,185,570,595]" italics="true" pageId="3" pageNumber="26">in</emphasis>
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) and forming enantiophysis with median prodorsal tubercle (
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<figureCitation id="130DCCBEFFBFFA714396D542FBA3FDEC" box="[939,1025,570,596]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Figs. 1</figureCitation>
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arrow, 2); exobothridial seta (
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<emphasis id="B9420C29FFBFFA71454AD542FA30FDEB" box="[1399,1426,570,595]" italics="true" pageId="3" pageNumber="26">ex</emphasis>
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) inserted posteriad
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<emphasis id="B9420C29FFBFFA714141D519FE36FDC2" box="[380,404,609,634]" italics="true" pageId="3" pageNumber="26">in</emphasis>
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; rostral seta (
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<emphasis id="B9420C29FFBFFA71427CD519FDF8FDC2" box="[577,602,609,634]" italics="true" pageId="3" pageNumber="26">ro</emphasis>
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) inserted posteriad lamellar seta; lamellar seta (
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<emphasis id="B9420C29FFBFFA714488D519FB69FDC2" box="[1205,1227,609,634]" italics="true" pageId="3" pageNumber="26">le</emphasis>
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) curves ventrad, inserted at anterior base of strong transverse prelamellar ridge (
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<figureCitation id="130DCCBEFFBFFA7143B9D5FFFBB6FD19" box="[900,1044,647,673]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Figs. 1–2, 6</figureCitation>
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); pedotectum II rounded distally. Notogaster flattened, with 5 pairs of more or less sessile setae (
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<emphasis id="B9420C29FFBFFA7143B0D5D6FC3EFD7F" box="[909,924,686,711]" italics="true" pageId="3" pageNumber="26">h</emphasis>
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1-2,
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<emphasis id="B9420C29FFBFFA7143FCD5D6FC72FD7F" box="[961,976,686,711]" italics="true" pageId="3" pageNumber="26">p</emphasis>
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1-3), raised rim, crenulate outer margin, and posterior median nipple-like process between setae
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<emphasis id="B9420C29FFBFFA71435CD5AFFCD2FD48" box="[865,880,727,752]" italics="true" pageId="3" pageNumber="26">h</emphasis>
|
|||
|
|
1; without anterior median tubercular patch; cerotegument with spherical pustules and usually with a pattern of raised ridges or cells and parallel marginal folds (
|
|||
|
|
<figureCitation id="130DCCBEFFBFFA7140DCD45FFE8AFCF9" box="[225,296,807,833]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Fig. 5</figureCitation>
|
|||
|
|
); sejugal tubercle well developed.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32C83B0FFBFFA7140FBD436FA38F857" pageId="3" pageNumber="26" type="discussion">
|
|||
|
|
<paragraph id="8B89D03BFFBFFA7140FBD436FE30FB7C" blockId="3.[151,1437,331,2032]" pageId="3" pageNumber="26">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7140FBD436FEEFFCD0" bold="true" box="[198,333,846,872]" pageId="3" pageNumber="26">Comments</emphasis>
|
|||
|
|
: As discussed in
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714205D436FD7BFCD0" author="Woas" box="[568,729,846,872]" pageId="3" pageNumber="44" refString="Woas, S. (1992) Beitrag zur Revision der Gymnodamaeidae Grandjean, 1954 (Acari, Oribatei). Andrias, Karlsruhe, 9, 121 - 161." type="journal article" year="1992">Woas (1992)</bibRefCitation>
|
|||
|
|
, the actual identity of the
|
|||
|
|
<typeStatus id="548D6E99FFBFFA71440BD436FBCBFCD0" box="[1078,1129,846,872]" pageId="3" pageNumber="26">type</typeStatus>
|
|||
|
|
species
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7144E6D436FAD8FCDF" box="[1243,1402,846,871]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7144E6D436FAD8FCDF" box="[1243,1402,846,871]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is contentious: the location of the
|
|||
|
|
<typeStatus id="548D6E99FFBFFA714234D40DFDD2FC37" box="[521,624,885,911]" pageId="3" pageNumber="26" type="holotype">holotype</typeStatus>
|
|||
|
|
is unknown (
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714333D40DFC5FFC37" author="Marshall" box="[782,1021,885,911]" pageId="3" pageNumber="43" refString="Marshall, V. G., Reeves, R. M. & Norton, R. A. (1987) Catalogue of the Oribatida (Acari) of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139. 418 pp." type="book" year="1987">
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA714333D40DFCDAFC37" box="[782,888,885,911]" name="Marshall Islands" pageId="3" pageNumber="26">Marshall</collectingCountry>
|
|||
|
|
<emphasis id="B9420C29FFBFFA714342D40DFC18FC36" box="[895,954,885,910]" italics="true" pageId="3" pageNumber="26">et al.</emphasis>
|
|||
|
|
1987
|
|||
|
|
</bibRefCitation>
|
|||
|
|
) and two similar forms that vary in the presence (e.g.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714157D4E3FD81FC0D" author="Willmann" box="[362,547,923,949]" pageId="3" pageNumber="44" refString="Willmann, C. (1931) Moosmilben oder Oribatiden (Cryptostigmata). In: Dahl, F. (Ed), Die Tierwelt Deutschlands, Bd. 22, vol. 5, Gustav Fischer, Jena, 79 - 200." type="journal article" year="1931">Willmann 1931</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714212D4E3FD11FC0D" author="Woas" box="[559,691,923,949]" pageId="3" pageNumber="44" refString="Woas, S. (1992) Beitrag zur Revision der Gymnodamaeidae Grandjean, 1954 (Acari, Oribatei). Andrias, Karlsruhe, 9, 121 - 161." type="journal article" year="1992">Woas 1992</bibRefCitation>
|
|||
|
|
) or absence (e.g.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7143BCD4E3FBEEFC0D" author="Grandjean" box="[897,1100,923,949]" pageId="3" pageNumber="43" refString="Grandjean, F. (1954 a) Observations sur les Oribates (28 e serie). Bulletin du Museum National d'Histoire Naturelle (2), 26, 204 - 211." type="journal article" year="1954" yearSuffix="a">Grandjean 1954a</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714465D4E3FAB3FC0D" author="Paschoal" box="[1112,1297,923,949]" pageId="3" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">Paschoal 1982a</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714523D4E3FF7BFC64" author="Weigmann" pageId="3" pageNumber="44" refString="Weigmann, G. (2006) Hornmilben (Oribatida). Die Tierwelt Deutschlands 76 Teil Goeke & Evers, Keltern, Germany." type="book chapter" year="2006">Weigmann 2006</bibRefCitation>
|
|||
|
|
) of a narrow anogenital bridge are known.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714331D4BAFC12FC64" author="Woas" box="[780,944,962,988]" pageId="3" pageNumber="44" refString="Woas, S. (1992) Beitrag zur Revision der Gymnodamaeidae Grandjean, 1954 (Acari, Oribatei). Andrias, Karlsruhe, 9, 121 - 161." type="journal article" year="1992">Woas (1992)</bibRefCitation>
|
|||
|
|
argued that although it may represent intraspecific variation, the form with a narrow anogenital bridge is likely to be the true
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7144B6D491FA80FBBA" box="[1163,1314,1001,1026]" class="Arachnida" family="Damaeidae" genus="Damaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7144B6D491FA80FBBA" box="[1163,1314,1001,1026]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
because it is present in
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA71410FD377FE24FB91" box="[306,390,1039,1065]" name="Poland" pageId="3" pageNumber="26">Poland</collectingCountry>
|
|||
|
|
near the site of its presumed
|
|||
|
|
<typeStatus id="548D6E99FFBFFA7142D0D377FC83FB91" box="[749,801,1039,1065]" pageId="3" pageNumber="26">type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7143ACD368FB76FB91" box="[913,1236,1040,1065]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7143ACD368FB76FB91" box="[913,1236,1040,1065]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has been shown with a smooth notogaster (e.g.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA71422CD34EFD71FBE8" author="Paschoal" box="[529,723,1078,1104]" pageId="3" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">Paschoal 1982a</bibRefCitation>
|
|||
|
|
), but it actually has a U-shaped cerotegumental ridge with outer branches formed from certotegument (e.g.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7142F0D325FCF3FBCF" author="Woas" box="[717,849,1117,1143]" pageId="3" pageNumber="44" refString="Woas, S. (1992) Beitrag zur Revision der Gymnodamaeidae Grandjean, 1954 (Acari, Oribatei). Andrias, Karlsruhe, 9, 121 - 161." type="journal article" year="1992">Woas 1992</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714360D325FB82FBCF" author="Weigmann" box="[861,1056,1117,1143]" pageId="3" pageNumber="44" refString="Weigmann, G. (2006) Hornmilben (Oribatida). Die Tierwelt Deutschlands 76 Teil Goeke & Evers, Keltern, Germany." type="book chapter" year="2006">Weigmann 2006</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714416D325FAD3FBCF" author="Weigmann" box="[1067,1393,1117,1143]" pageId="3" pageNumber="44" refString="Weigmann, G. & Mourek, J. (2008) Contribution to the Central European Gymnodamaeus species G. barbarossa and G. bicostatus (Acari, Oribatida, Gymnodamaeidae). Zoosystematics and Evolution, 84, 255 - 264." type="journal article" year="2008">Weigmann & Mourek 2008</bibRefCitation>
|
|||
|
|
). In slide-flattened specimens these ridges can be difficult to observe (and are absent or obscure in recently moulted individuals).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89D03BFFBFFA7140F8D3A9FA36FA14" blockId="3.[151,1437,331,2032]" pageId="3" pageNumber="26">
|
|||
|
|
About a dozen species of
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA71423FD3A9FD66FB52" box="[514,708,1233,1258]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA71423FD3A9FD66FB52" box="[514,708,1233,1258]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are known from North
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA7143D5D3A9FBF6FB53" box="[1000,1108,1233,1259]" name="United States of America" pageId="3" pageNumber="26">America</collectingCountry>
|
|||
|
|
, but according to
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA71450DD3A9FF50FAA9" author="Paschoal" pageId="3" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">Paschoal (1982a)</bibRefCitation>
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7140C7D380FE30FAA9" box="[250,402,1272,1297]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7140C7D380FE30FAA9" box="[250,402,1272,1297]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
does not occur in North
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA714287D38FFCBDFAA9" box="[698,799,1271,1297]" name="United States of America" pageId="3" pageNumber="26">America</collectingCountry>
|
|||
|
|
and, except as quarantine interceptions (e.g.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714508D38FFF7AFA80" author="Hammer" pageId="3" pageNumber="43" refString="Hammer, M. (1969) Investigations on Oribatids found at plant quarantine stations in U. S. A. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 132, 63 - 78." type="journal article" year="1969">Hammer 1969</bibRefCitation>
|
|||
|
|
), previous records refer to other species. For example, the record of
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA714418D266FB61FA8F" box="[1061,1219,1310,1335]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA714418D266FB61FA8F" box="[1061,1219,1310,1335]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Alberta on the
|
|||
|
|
<emphasis id="B9420C29FFBFFA7140AAD23DFD86FAE6" box="[151,548,1349,1374]" italics="true" pageId="3" pageNumber="26">
|
|||
|
|
Diversity of
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA714117D23DFE3DFAE6" box="[298,415,1349,1374]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="order">Oribatida</taxonomicName>
|
|||
|
|
in
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA7141FBD23DFD86FAE6" box="[454,548,1349,1374]" name="Canada" pageId="3" pageNumber="26">Canada</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
website (
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7142ABD23DFBABFAE7" author="Behan-Pelletier" box="[662,1033,1349,1375]" pageId="3" pageNumber="43" refString="Behan-Pelletier, V. M. & Eamer, B. (2004) Diversity of Oribatida in Canada, Canadian Biodiversity Information Facility, Ottawa, Ontario, Canada. Available from: http: // www. cbif. gc. ca / spp _ pages / mites / phps / index _ e. php (Accessed 13 May 2009)." type="journal article" year="2004">Behan-Pelletier & Eamer 2004</bibRefCitation>
|
|||
|
|
) is based on the
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7144EFD23DFF52FA3D" author="Powell" pageId="3" pageNumber="44" refString="Powell, J. M. & Skaley, L. S. (1975) Arthropods from forest litter under lodgepole pine infected with comandra blister rust. Northern Forest Research Centre (Edmonton), Information Report NOR-X- 130, 33 pp." type="book" year="1975">Powell & Skaley (1975)</bibRefCitation>
|
|||
|
|
study listed in
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7141F9D213FD52FA3D" box="[452,752,1387,1413]" pageId="3" pageNumber="43" refString="Marshall, V. G., Reeves, R. M. & Norton, R. A. (1987) Catalogue of the Oribatida (Acari) of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139. 418 pp." type="book">
|
|||
|
|
<collectingCountry id="F32190ABFFBFFA7141F9D213FD98FA3D" box="[452,570,1387,1413]" name="Marshall Islands" pageId="3" pageNumber="26">Marshall</collectingCountry>
|
|||
|
|
<emphasis id="B9420C29FFBFFA714279D214FD29FA3D" box="[580,651,1388,1413]" italics="true" pageId="3" pageNumber="26">et al.</emphasis>
|
|||
|
|
(1987)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, but
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714309D213FBBFFA3D" author="Paschoal" box="[820,1053,1387,1413]" pageId="3" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">Paschoal (1982a)</bibRefCitation>
|
|||
|
|
must have considered this a misidentification, since the specimen was designated as the
|
|||
|
|
<typeStatus id="548D6E99FFBFFA714368D2EAFC19FA14" box="[853,955,1426,1452]" pageId="3" pageNumber="26" type="holotype">holotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7143DED2EAFA36FA14" authority="Paschoal." authorityName="Paschoal." box="[995,1428,1426,1452]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="taedaceus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7143DED2EAFABEFA13" box="[995,1308,1426,1451]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus taedaceus</emphasis>
|
|||
|
|
Paschoal.
|
|||
|
|
</taxonomicName>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89D03BFFBFFA7140F8D2C1FEF0F81B" blockId="3.[151,1437,331,2032]" pageId="3" pageNumber="26">
|
|||
|
|
With the exception of three Paschoal species (see below), and one record for
|
|||
|
|
<emphasis id="B9420C29FFBFFA714471D2C1FBC1FA6A" box="[1100,1123,1465,1490]" italics="true" pageId="3" pageNumber="26">G.</emphasis>
|
|||
|
|
sp. nr.
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA714486D2C1FEE8FA41" authority="Oswald & Minty 1971" authorityName="Oswald & Minty" authorityYear="1971" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA714486D2C1FA93FA6A" box="[1211,1329,1465,1490]" italics="true" pageId="3" pageNumber="26">bicostatus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA71457CD2C1FEE3FA41" author="Oswald" pageId="3" pageNumber="43" refString="Oswald, E. T. & Minty. L. W. (1971) Soil acarine fauna of southeastern Manitoba II. Riparian communities. Manitoba." type="book" year="1971">Oswald & Minty 1971</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, other Canadian records for
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7142AED298FCF2FA41" box="[659,848,1504,1529]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7142AED298FCF2FA41" box="[659,848,1504,1529]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
have been attributed to
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA71445AD298FA3AFA41" authority="Hammer, 1952" authorityName="Hammer" authorityYear="1952" box="[1127,1432,1503,1529]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA71445AD298FB42FA41" box="[1127,1248,1504,1529]" italics="true" pageId="3" pageNumber="26">G. ornatus</emphasis>
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7144DAD2A7FA3AFA41" author="Hammer" box="[1255,1432,1503,1529]" pageId="3" pageNumber="43" refString="Hammer, M. (1952) Investigations on the microfauna of Northern Canada. Part I: Oribatidae. Acta Arctica, 4, 1 - 108." type="journal article" year="1952">Hammer, 1952</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA7140AAD17EFEF1F998" author="Hammer" box="[151,339,1542,1568]" pageId="3" pageNumber="43" refString="Hammer, M. (1952) Investigations on the microfauna of Northern Canada. Part I: Oribatidae. Acta Arctica, 4, 1 - 108." type="journal article" year="1952">Hammer (1952)</bibRefCitation>
|
|||
|
|
described this species from Reindeer Station based on
|
|||
|
|
<specimenCount id="9D301BB2FFBFFA7143E6D17EFBC9F998" box="[987,1131,1542,1568]" pageId="3" pageNumber="26" type="generic">7 specimens</specimenCount>
|
|||
|
|
of which I have observed two
|
|||
|
|
<typeStatus id="548D6E99FFBFFA7140F1D155FE97F9FF" box="[204,309,1581,1607]" pageId="3" pageNumber="26" type="syntype">syntypes</typeStatus>
|
|||
|
|
on a slide in the CNC. She distinguished
|
|||
|
|
<emphasis id="B9420C29FFBFFA71430ED155FC0EF9FE" box="[819,940,1581,1606]" italics="true" pageId="3" pageNumber="26">G. o r n a t u s</emphasis>
|
|||
|
|
from
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7143C6D155FB34F9FE" box="[1019,1174,1581,1606]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7143C6D155FB34F9FE" box="[1019,1174,1581,1606]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the smooth ridges forming “oval and crescent-shaped areas” (p. 28) on the notogaster and by the longer lamellar setae that project beyond the tip of the rostrum. Length of the lamellar seta, however, is difficult to judge from a dorsal slide mount (compare
|
|||
|
|
<figureCitation id="130DCCBEFFBFFA7141A3D1D9FE41F903" box="[414,483,1697,1723]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,271,1797,1821]" captionTargetBox="[167,1422,185,1773]" captionTargetId="figure@5.[167,1424,185,1773]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Gymnodamaeus cf. ornatus Hammer, Alberta, Canada; scale bars = 0.1 mm, except Fig. 3 = 1 μm. 1, Prodorsum showing interlamellar seta (in) on apophysis at median edge of bothridial ridge and opposed to median prodorsal tubercle (larger arrow); 2, Prodorsum and anterior notogaster; 3, Spherical crystalline notogastral pustules; 4, Ano-genital region, arrow points to soft cuticle between genital and anal openings; 5, Dorsal habitus view showing Ushaped raised notogastral ridge pattern; 6, Rostrum from anterior view; 7, Light micrograph of ano-genital region, arrow points to preanal organ; 8, Genu IV (DIC micrograph) showing normal joint, i. e. lacking retrotecta, between femur (top) and tibia." httpUri="https://zenodo.org/record/189741/files/figure.png" pageId="3" pageNumber="26">Fig. 2</figureCitation>
|
|||
|
|
to 6). Also, her illustration (fig. 35, p. 98) shows a more elaborate cell pattern on the notogaster than I usually find on Canadian specimens. The
|
|||
|
|
<typeStatus id="548D6E99FFBFFA7143A5D1BFFBA0F959" box="[920,1026,1735,1761]" pageId="3" pageNumber="26" type="syntype">syntypes</typeStatus>
|
|||
|
|
are flattened, but as far as can be discerned show the more typical form having one central U-shaped cell with incomplete internal ridges and two pairs of arms forming partial (rarely complete) lateral cells. The oval and crescentic areas within the Ushaped ridges appear to be artefacts caused by smooth areas without pustules and not by raised ridges; complete lateral cells are not visible on the
|
|||
|
|
<typeStatus id="548D6E99FFBFFA714285D01AFC85F8C4" box="[696,807,1890,1916]" pageId="3" pageNumber="26" type="syntype">syntypes</typeStatus>
|
|||
|
|
, but I have seen a few specimens with one present asymmetrically.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89D03BFFBFFA7140F8D0C8FA38F857" blockId="3.[151,1437,331,2032]" pageId="3" pageNumber="26">
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA7140F8D0C8FE47F871" box="[197,485,1968,1993]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA7140F8D0C8FE47F871" box="[197,485,1968,1993]" italics="true" pageId="3" pageNumber="26">Gymnodamaeus ornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C36ABB8FFBFFA714222D0C8FD1AF871" box="[543,696,1968,1993]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFBFFA714222D0C8FD1AF871" box="[543,696,1968,1993]" italics="true" pageId="3" pageNumber="26">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are very similar (see
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFBFFA714387D0D7FAA1F871" author="Weigmann" box="[954,1283,1967,1993]" pageId="3" pageNumber="44" refString="Weigmann, G. & Mourek, J. (2008) Contribution to the Central European Gymnodamaeus species G. barbarossa and G. bicostatus (Acari, Oribatida, Gymnodamaeidae). Zoosystematics and Evolution, 84, 255 - 264." type="journal article" year="2008">Weigmann & Mourek 2008</bibRefCitation>
|
|||
|
|
) and may be conspecific. Since this problem cannot be resolved at this time, I accept Woas’ (1992) diagnosis of
|
|||
|
|
<emphasis id="B9420C29FFBFFA7145BFD0AEFA38F857" box="[1410,1434,2006,2031]" italics="true" pageId="3" pageNumber="26">G.</emphasis>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32C83B0FFB8FA7640AAD7E0FE24FCA4" pageId="4" pageNumber="27" type="reference_group">
|
|||
|
|
<paragraph id="8B89D03BFFB8FA7640AAD7E0FD63FE9E" blockId="4.[151,1437,152,912]" pageId="4" pageNumber="27">
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA7640AAD7E0FEACFF09" ID-CoL="3HRJV" box="[151,270,152,177]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA7640AAD7E0FEACFF09" box="[151,270,152,177]" italics="true" pageId="4" pageNumber="27">bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as having a narrow anogenital bridge and refer the very similar mite that lacks the anogenital bridge to
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA76408AD7C7FD87FF60" ID-CoL="3HRK7" authority="Hammer, 1952" authorityName="Hammer" authorityYear="1952" box="[183,549,190,216]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA76408AD7C7FF6CFF60" box="[183,206,191,216]" italics="true" pageId="4" pageNumber="27">G.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="B9420C29FFB8FA76413ED7C7FEC3FF60" box="[259,353,191,216]" italics="true" pageId="4" pageNumber="27">ornatus</emphasis>
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA764157D7C6FD87FF60" author="Hammer" box="[362,549,190,216]" pageId="4" pageNumber="43" refString="Hammer, M. (1952) Investigations on the microfauna of Northern Canada. Part I: Oribatidae. Acta Arctica, 4, 1 - 108." type="journal article" year="1952">Hammer, 1952</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. I have
|
|||
|
|
<emphasis id="B9420C29FFB8FA7642BAD7C7FD63FF60" box="[647,705,191,216]" italics="true" pageId="4" pageNumber="27">cox1</emphasis>
|
|||
|
|
mitochondrial DNA sequences of the barcode region for
|
|||
|
|
<specimenCount id="9D301BB2FFB8FA7645B0D7C6FEB0FF47" pageId="4" pageNumber="27" type="generic">4 specimens</specimenCount>
|
|||
|
|
from an Alberta population of
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA7642B7D79DFC8FFF46" ID-CoL="3HRK7" box="[650,813,229,255]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA7642B7D79DFD03FF46" box="[650,673,229,254]" italics="true" pageId="4" pageNumber="27">G.</emphasis>
|
|||
|
|
cf.
|
|||
|
|
<emphasis id="B9420C29FFB8FA7642EFD79DFC8FFF46" box="[722,813,229,254]" italics="true" pageId="4" pageNumber="27">ornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and they may be informative if comparisons can be made to European populations of
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA76421ED674FD18FE9D" ID-CoL="3HRJV" box="[547,698,268,293]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA76421ED674FD18FE9D" box="[547,698,268,293]" italics="true" pageId="4" pageNumber="27">G. bicostatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B89D03BFFB8FA7640F8D64AFE24FCA4" blockId="4.[151,1437,152,912]" pageId="4" pageNumber="27">
|
|||
|
|
The
|
|||
|
|
<typeStatus id="548D6E99FFB8FA7640C4D64AFEFDFEF4" box="[249,351,306,332]" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
|
|||
|
|
slides and only known specimens of
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA764329D64AFBB4FEF4" ID-CoL="3HRKC" authority="Paschoal" authorityName="Paschoal" box="[788,1046,306,332]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="saltuensis">
|
|||
|
|
<emphasis id="B9420C29FFB8FA764329D64AFC05FEF3" box="[788,935,306,331]" italics="true" pageId="4" pageNumber="27">G. saltuensis</emphasis>
|
|||
|
|
Paschoal
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA764473D64AFB41FEF3" ID-CoL="6L6V7" box="[1102,1251,306,331]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="taedaceus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA764473D64AFB41FEF3" box="[1102,1251,306,331]" italics="true" pageId="4" pageNumber="27">G. taedaceus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were located in the CNC: both were labelled “
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA76423FD621FC3DFECB" ID-CoL="3HRJV" authority="Koch" authorityName="Koch" box="[514,927,345,371]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA76423FD621FCE2FECA" box="[514,832,345,370]" italics="true" pageId="4" pageNumber="27">Gymnodamaeus bicostatus</emphasis>
|
|||
|
|
(Koch)
|
|||
|
|
</taxonomicName>
|
|||
|
|
” but were identified by the collection data recorded in
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA76411ED6F8FE6FFE22" author="Paschoal" box="[291,461,384,410]" pageId="4" pageNumber="44" refString="Paschoal, A. D. (1975) A revision of the Gymnodamaeidae (Acari: Oribatei): A numerical taxonomic study. Dissertation, Ohio State Univiversity, Columbus, Ohio, 374 pp." type="book" year="1975">
|
|||
|
|
Paschoal
|
|||
|
|
<date id="FF88F6FBFFB8FA7641AFD6F8FE6FFE22" box="[402,461,384,410]" pageId="4" pageNumber="27" value="1975" valueMax="1982">1975</date>
|
|||
|
|
</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA76423AD6F8FDF3FE22" author="Paschoal" box="[519,593,384,410]" pageId="4" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">1982a</bibRefCitation>
|
|||
|
|
. Both mites were badly crushed, perhaps the reason that no illustrations were provided for the species (
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA764279D6DEFCAAFE78" author="Paschoal" box="[580,776,422,448]" pageId="4" pageNumber="44" refString="Paschoal, A. D. (1975) A revision of the Gymnodamaeidae (Acari: Oribatei): A numerical taxonomic study. Dissertation, Ohio State Univiversity, Columbus, Ohio, 374 pp." type="book" year="1975">Paschoal 1975</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA764323D6DEFCCCFE78" author="Paschoal" box="[798,878,422,448]" pageId="4" pageNumber="44" refString="Paschoal, A. D. (1982 a) A revision of the genus Gymnodamaeus (Acari, Oribatei, Gymnodamaeidae), with descriptions of nine new species. Revista Brasileira de Entomologia, 26, 113 - 132." type="journal article" year="1982" yearSuffix="a">1982a</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA7643B2D6DEFB7BFE78" author="Powell" box="[911,1241,422,448]" pageId="4" pageNumber="44" refString="Powell, J. M. & Skaley, L. S. (1975) Arthropods from forest litter under lodgepole pine infected with comandra blister rust. Northern Forest Research Centre (Edmonton), Information Report NOR-X- 130, 33 pp." type="book" year="1975">Powell & Skaley (1975)</bibRefCitation>
|
|||
|
|
published the “
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA764099D6B5FDEAFE5F" ID-CoL="3HRJV" authority="Koch" authorityName="Koch" box="[164,584,461,487]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="bicostatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA764099D6B5FE45FE5E" box="[164,487,461,486]" italics="true" pageId="4" pageNumber="27">Gymnodamaeus bicostatus</emphasis>
|
|||
|
|
(Koch)
|
|||
|
|
</taxonomicName>
|
|||
|
|
” record in a pine blister rust study in Alberta which was subsequently used by
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA7640CAD68CFE57FDB6" box="[247,501,500,526]" pageId="4" pageNumber="43" refString="Marshall, V. G., Reeves, R. M. & Norton, R. A. (1987) Catalogue of the Oribatida (Acari) of continental United States and Canada. Memoirs of the Entomological Society of Canada, 139. 418 pp." type="book">
|
|||
|
|
<collectingCountry id="F32190ABFFB8FA7640CAD68CFEFDFDB6" box="[247,351,500,526]" name="Marshall Islands" pageId="4" pageNumber="27">Marshall</collectingCountry>
|
|||
|
|
<emphasis id="B9420C29FFB8FA76415BD68CFE03FDB5" box="[358,417,500,525]" italics="true" pageId="4" pageNumber="27">et al.</emphasis>
|
|||
|
|
(1987)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
and
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA764213D68CFC08FDB6" author="Behan-Pelletier" box="[558,938,500,526]" pageId="4" pageNumber="43" refString="Behan-Pelletier, V. M. & Eamer, B. (2004) Diversity of Oribatida in Canada, Canadian Biodiversity Information Facility, Ottawa, Ontario, Canada. Available from: http: // www. cbif. gc. ca / spp _ pages / mites / phps / index _ e. php (Accessed 13 May 2009)." type="journal article" year="2004">Behan-Pelletier & Eamer (2004)</bibRefCitation>
|
|||
|
|
to report this European species in
|
|||
|
|
<collectingCountry id="F32190ABFFB8FA764500D68CFA3BFDB6" box="[1341,1433,500,526]" name="Canada" pageId="4" pageNumber="27">Canada</collectingCountry>
|
|||
|
|
. I consider these species to be
|
|||
|
|
<emphasis id="B9420C29FFB8FA76422DD562FD58FD8B" box="[528,762,538,563]" italics="true" pageId="4" pageNumber="27">species inquirenda</emphasis>
|
|||
|
|
for the following reasons: the
|
|||
|
|
<typeStatus id="548D6E99FFB8FA7644BDD562FB67FD8C" box="[1152,1221,538,564]" pageId="4" pageNumber="27">types</typeStatus>
|
|||
|
|
, and only known specimens, of these two species are both badly crushed; one is a male and one a female; most of the major characters in the descriptions were misinterpretations of thick areas of cuticle (mostly ridges or tubercles on the surface of the mite) as apodemata; many other characters call for qualitative judgements; and there is no way to distinguish the descriptions of these mites from each other or from
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA76443DD5CDFBD8FD76" ID-CoL="3HRK7" box="[1024,1146,693,718]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="ornatus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA76443DD5CDFBD8FD76" box="[1024,1146,693,718]" italics="true" pageId="4" pageNumber="27">G. ornatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<bibRefCitation id="EFA7ADCAFFB8FA7644B4D5CDFF47FD4E" author="Bayartogtokh" pageId="4" pageNumber="43" refString="Bayartogtokh, B. & Schatz, H. (2009) Two new species of the genus Gymnodamaeus (Acari: Oribatida: Gymnodamaeidae) from Tyrol (Austria), with remarks on diversity and distribution of the known species. Revue Suisse de Zoologie, 116, 31 - 51." type="journal article" year="2009">Bayartogtokh & Schatz (2009)</bibRefCitation>
|
|||
|
|
similarly considered these species to be unrecognizable, and suggested that they should be treated as a
|
|||
|
|
<emphasis id="B9420C29FFB8FA7640AAD47AFEDDFCA3" box="[151,383,770,795]" italics="true" pageId="4" pageNumber="27">species inquirendae</emphasis>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C32C83B0FFB8FA7640F8D451FED8FC28" pageId="4" pageNumber="27" type="description">
|
|||
|
|
<paragraph id="8B89D03BFFB8FA7640F8D451FED8FC28" blockId="4.[151,1437,152,912]" pageId="4" pageNumber="27">
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA7640F8D451FDC1FCFB" authority="Paschoal" authorityName="Paschoal" box="[197,611,809,835]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="victoriae">
|
|||
|
|
<emphasis id="B9420C29FFB8FA7640F8D451FE50FCFA" box="[197,498,809,834]" italics="true" pageId="4" pageNumber="27">Gymnodamaeus victoriae</emphasis>
|
|||
|
|
Paschoal
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be identified, but is transferred to
|
|||
|
|
<emphasis id="B9420C29FFB8FA76440DD451FB7AFCFA" box="[1072,1240,809,834]" italics="true" pageId="4" pageNumber="27">Roynortonella</emphasis>
|
|||
|
|
below.
|
|||
|
|
<typeStatus id="548D6E99FFB8FA764508D451FAD9FCFB" box="[1333,1403,809,835]" pageId="4" pageNumber="27">Types</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA7640AAD428FDE8FCD2" authority="Paschoal" authorityName="Paschoal" box="[151,586,848,874]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="gregarious">
|
|||
|
|
<emphasis id="B9420C29FFB8FA7640AAD428FE79FCD1" box="[151,475,848,873]" italics="true" pageId="4" pageNumber="27">Gymnodamaeus gregarious</emphasis>
|
|||
|
|
Paschoal
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA7642B9D428FC34FCD2" authority="Paschoal" authorityName="Paschoal" box="[644,918,848,874]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="orbicularis">
|
|||
|
|
<emphasis id="B9420C29FFB8FA7642B9D428FC84FCD1" box="[644,806,848,873]" italics="true" pageId="4" pageNumber="27">G. orbicularis</emphasis>
|
|||
|
|
Paschoal
|
|||
|
|
</taxonomicName>
|
|||
|
|
are in good condition, and clearly members of
|
|||
|
|
<taxonomicName id="4C36ABB8FFB8FA76408BD40FFED1FC28" box="[182,371,887,912]" class="Arachnida" family="Gymnodamaeidae" genus="Gymnodamaeus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B9420C29FFB8FA76408BD40FFED1FC28" box="[182,371,887,912]" italics="true" pageId="4" pageNumber="27">Gymnodamaeus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|