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<document id="C01BC107FA8CD86B4B9C5515F7ABC0F5" ID-CLB-Dataset="287450" ID-DOI="10.11646/zootaxa.5406.3.9" ID-GBIF-Dataset="561731c7-996f-492e-b694-38c6ca2cb014" ID-ISSN="1175-5326" ID-Zenodo-Dep="10628130" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1707284994355" checkinUser="plazi" docAuthor="Simone, Luiz Ricardo L. &amp; Monsecour, Kevin" docDate="2024" docId="038B2411FFECBE30FF3054B8FAA3FAF9" docLanguage="en" docName="zootaxa.5406.3.9.pdf" docOrigin="Zootaxa 5406 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5406.3.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmioconcha costorum Simone &amp; Monsecour 2024, new species" docType="treatment" docVersion="2" lastPageNumber="494" masterDocId="FFB25C69FFECBE32FFA7516EFFCAFF8E" masterDocTitle="Paratypes of Brazilian Cosmioconcha helenae (Gastropoda, Columbellidae) actually belong to a new species" masterLastPageNumber="494" masterPageNumber="492" pageNumber="492" updateTime="1707285816865" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CF9642477865FA2B9374CBA0360BAB48">Paratypes of Brazilian Cosmioconcha helenae (Gastropoda, Columbellidae) actually belong to a new species</mods:title>
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<mods:namePart id="5F5597574F7840D55AE25F17EA635952">Simone, Luiz Ricardo L.</mods:namePart>
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<mods:namePart id="6E5AD9BD80B1D4A6CF069BA2362A1357">Monsecour, Kevin</mods:namePart>
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<mods:date id="25BC1F6D45E79D7CA53C47EA9025D4FE">2024</mods:date>
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<mods:identifier id="1D5FB15BBD0F7A230A648465BD702BE7" type="DOI">10.11646/zootaxa.5406.3.9</mods:identifier>
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<treatment id="038B2411FFECBE30FF3054B8FAA3FAF9" ID-DOI="http://doi.org/10.5281/zenodo.10628128" ID-Zenodo-Dep="10628128" LSID="urn:lsid:plazi:treatment:038B2411FFECBE30FF3054B8FAA3FAF9" httpUri="http://treatment.plazi.org/id/038B2411FFECBE30FF3054B8FAA3FAF9" lastPageId="2" lastPageNumber="494" pageId="0" pageNumber="492">
<subSubSection id="C338C68CFFECBE32FF3054B8FBBEF99A" pageId="0" pageNumber="492" type="nomenclature">
<paragraph id="8B9D9507FFECBE32FF3054B8FD93FA7E" blockId="0.[151,1437,1458,2024]" box="[151,601,1494,1521]" pageId="0" pageNumber="492">
<heading id="D0D5226BFFECBE32FF3054B8FD93FA7E" bold="true" box="[151,601,1494,1521]" fontSize="11" level="1" pageId="0" pageNumber="492" reason="1">
<emphasis id="B9564915FFECBE32FF3054B8FD93FA7E" bold="true" box="[151,601,1494,1521]" pageId="0" pageNumber="492">
<taxonomicName id="4C22EE84FFECBE32FF3054B8FE08FA7F" authority="Simone &amp; Monsecour, 2024" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[151,450,1494,1521]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="0" pageNumber="492" phylum="Mollusca" rank="species" species="costorum" status="sp. nov.">
<emphasis id="B9564915FFECBE32FF3054B8FE08FA7F" bold="true" box="[151,450,1494,1521]" italics="true" pageId="0" pageNumber="492">Cosmioconcha costorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265F46EFFECBE32FE6D54B8FD93FA7E" box="[458,601,1494,1520]" pageId="0" pageNumber="492" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D9507FFECBE32FF305492FBBEF99A" blockId="0.[151,1437,1458,2024]" box="[151,1140,1532,1556]" pageId="0" pageNumber="492">
http://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="FF84AFD2FFECBE32FDD65492FBBEF99A" box="[625,1140,1532,1556]" pageId="0" pageNumber="492">93A5AEB0-BDC9-4C84-8EE9-801E1E060F2C</uuid>
</uri>
</paragraph>
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<subSubSection id="C338C68CFFECBE32FF30574EFB95F9D2" pageId="0" pageNumber="492" type="description">
<paragraph id="8B9D9507FFECBE32FF30574EFEBBF9B6" blockId="0.[151,1437,1458,2024]" box="[151,369,1568,1592]" pageId="0" pageNumber="492">
(
<figureCitation id="13198982FFECBE32FF38574EFEA3F9B6" box="[159,361,1568,1592]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="0" pageNumber="492">Figures. 1–6, 9–12</figureCitation>
)
</paragraph>
<paragraph id="8B9D9507FFECBE32FF30572AFB95F9D2" blockId="0.[151,1437,1458,2024]" box="[151,1119,1604,1628]" pageId="0" pageNumber="492">
<emphasis id="B9564915FFECBE32FF30572AFF2AF9D2" bold="true" box="[151,224,1604,1628]" inLineHeading="true" pageId="0" pageNumber="492" reason="1">Types.</emphasis>
<materialsCitation id="3B4A9F5AFFECBE32FF40572AFDE5F9D2" box="[231,559,1604,1628]" collectionCode="MZSP" pageId="0" pageNumber="492" specimenCode="MZSP 102539" specimenCount="1" typeStatus="holotype">
<typeStatus id="54992BA5FFECBE32FF40572AFE80F9D2" box="[231,330,1604,1628]" pageId="0" pageNumber="492" type="holotype">Holotype</typeStatus>
<specimenCode id="DB843D7CFFECBE32FEF6572AFE25F9D2" box="[337,495,1604,1628]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="492" type="Museum">MZSP 102539</specimenCode>
, shell
</materialsCitation>
.
<materialsCitation id="3B4A9F5AFFECBE32FD9E572AFB91F9D2" box="[569,1115,1604,1628]" collectionCode="MZSP" pageId="0" pageNumber="492" specimenCode="MZSP 102534, 1" specimenCount="1" typeStatus="paratype">
<typeStatus id="54992BA5FFECBE32FD9E572AFD51F9D2" box="[569,667,1604,1628]" pageId="0" pageNumber="492" type="paratype">Paratype</typeStatus>
:
<specimenCode id="DB843D7CFFECBE32FD03572AFC89F9D2" box="[676,835,1604,1628]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="492" type="Museum">MZSP 102534</specimenCode>
,
<specimenCode id="DB843D7CFFECBE32FCEA572AFC90F9D2" box="[845,858,1604,1628]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="492" type="Museum">1</specimenCode>
shell from type locality
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFECBE32FF605706FAB0F90E" box="[199,1402,1639,1664]" pageId="0" pageNumber="492" type="materials_examined">
<paragraph id="8B9D9507FFECBE32FF605706FAB0F90E" blockId="0.[151,1437,1458,2024]" box="[199,1402,1639,1664]" pageId="0" pageNumber="492">
<materialsCitation id="3B4A9F5AFFECBE32FF605706FABCF90E" box="[199,1398,1639,1664]" collectingDate="1968-05-01" collectionCode="RV" collectorName="Almirante Saldanha" country="Brazil" elevation="56" latitude="3.0" location="Amapa" longLatPrecision="1307" longitude="-49.466667" pageId="0" pageNumber="492" specimenCount="1" stateProvince="Amapa" typeStatus="holotype">
<emphasis id="B9564915FFECBE32FF605706FE94F90E" bold="true" box="[199,350,1640,1664]" inLineHeading="true" pageId="0" pageNumber="492" reason="1">
<typeStatus id="54992BA5FFECBE32FF605706FF35F90E" box="[199,255,1640,1664]" pageId="0" pageNumber="492">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F335D597FFECBE32FEC35706FE02F90E" box="[356,456,1640,1664]" name="Brazil" pageId="0" pageNumber="492">BRAZIL</collectingCountry>
.
<emphasis id="B9564915FFECBE32FE755709FDEFF9F1" bold="true" box="[466,549,1639,1663]" pageId="0" pageNumber="492">
<collectingRegion id="49E65BE5FFECBE32FE755709FDEFF9F1" box="[466,549,1639,1663]" country="Brazil" name="Amapa" pageId="0" pageNumber="492">Amapá</collectingRegion>
</emphasis>
; off
<location id="8EFDC3DCFFECBE32FDF05706FD6DF90E" LSID="urn:lsid:plazi:treatment:038B2411FFECBE30FF3054B8FAA3FAF9:8EFDC3DCFFECBE32FDF05706FD6DF90E" box="[599,679,1640,1664]" country="Brazil" latitude="3.0" longLatPrecision="1307" longitude="-49.466667" name="Amapa" pageId="0" pageNumber="492" stateProvince="Amapa">Amapá</location>
, ~
<geoCoordinate id="EE16F3C0FFECBE32FD675706FCC4F90E" box="[704,782,1640,1664]" degrees="3" direction="north" minutes="00" orientation="latitude" pageId="0" pageNumber="492" precision="925" value="3.0">3°00’N</geoCoordinate>
<geoCoordinate id="EE16F3C0FFECBE32FCB25706FCBDF90E" box="[789,887,1640,1664]" degrees="49" direction="west" minutes="28" orientation="longitude" pageId="0" pageNumber="492" precision="925" value="-49.466667">49°28’W</geoCoordinate>
,
<quantity id="4CDA38E2FFECBE32FC255706FC72F9F1" box="[898,952,1640,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="0" pageNumber="492" unit="m" value="56.0">
<elevation id="000F7234FFECBE32FC255706FC72F9F1" box="[898,952,1640,1664]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="0" pageNumber="492" unit="m" value="56.0">56 m</elevation>
</quantity>
[
<collectionCode id="ED330DC2FFECBE32FC625706FC21F9F1" box="[965,1003,1640,1663]" country="Italy" name="Collection of Leptospira Strains" pageId="0" pageNumber="492">RV</collectionCode>
<collectorName id="26D7F0D1FFECBE32FC485706FB0EF90E" box="[1007,1220,1640,1664]" pageId="0" pageNumber="492">Almirante Saldanha</collectorName>
col.,
<date id="FF9CB3C7FFECBE32FB585706FABAF90E" box="[1279,1392,1640,1664]" pageId="0" pageNumber="492" value="1968-05-01">
<collectingDate id="EFD84A2FFFECBE32FB585706FABAF90E" box="[1279,1392,1640,1664]" pageId="0" pageNumber="492" value="1968-05-01">1.05.1968</collectingDate>
</date>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFECBE32FF6057E2FB36F965" pageId="0" pageNumber="492" type="diagnosis">
<paragraph id="8B9D9507FFECBE32FF6057E2FB36F965" blockId="0.[151,1437,1458,2024]" pageId="0" pageNumber="492">
<emphasis id="B9564915FFECBE32FF6057E2FEF6F92D" box="[199,316,1676,1700]" inLineHeading="true" pageId="0" pageNumber="492" reason="1">
<emphasis id="B9564915FFECBE32FF6057E2FEFFF92A" bold="true" box="[199,309,1676,1700]" pageId="0" pageNumber="492">Diagnosis</emphasis>
.
</emphasis>
<taxonomicName id="4C22EE84FFECBE32FEE457E2FDFCF92A" box="[323,566,1676,1700]" pageId="0" pageNumber="492">
<emphasis id="B9564915FFECBE32FEE457E2FE28F92A" box="[323,482,1676,1700]" italics="true" pageId="0" pageNumber="492">Cosmioconcha</emphasis>
species
</taxonomicName>
of small size, fusiform, protoconch of 3 whorls, small subsutural shoulder, axial cords on all whorls, small nodes on the intervals of the subsutural band and the axials, spiral liration on lower half of last whorl, outer lip simply arched, inner lip shallowly concave with thin callus, anal notch small, canal open.
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFECBE30FF605796FE2DFF21" lastPageId="2" lastPageNumber="494" pageId="0" pageNumber="492" type="description">
<paragraph id="8B9D9507FFECBE33FF605796FCCCFEB1" blockId="0.[151,1437,1458,2024]" lastBlockId="1.[151,1436,151,319]" lastPageId="1" lastPageNumber="493" pageId="0" pageNumber="492">
<emphasis id="B9564915FFECBE32FF605796FE98F881" box="[199,338,1784,1808]" inLineHeading="true" pageId="0" pageNumber="492" reason="1">
<emphasis id="B9564915FFECBE32FF605796FE81F89E" bold="true" box="[199,331,1784,1808]" pageId="0" pageNumber="492">Description</emphasis>
.
</emphasis>
Shell of ~
<quantity id="4CDA38E2FFECBE32FE195796FE37F881" box="[446,509,1784,1807]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="0" pageNumber="492" unit="mm" value="7.0">7 mm</quantity>
, fusiform; walls thick. Color white, with light yellow band in lower half in spire whorls, and in middle third in last whorl (
<figureCitation id="13198982FFECBE32FDAE5672FD66F8BA" box="[521,684,1820,1844]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="0" pageNumber="492">Figs. 2, 3, 5, 6</figureCitation>
). Apex relatively blunt; spire ~57% of shell length, spire angle ~40°. Protoconch of 3 smooth whorls, successive larger, width of 0.6 mm; transition with teleoconch clear-cut, orthocline, sigmoid (
<figureCitation id="13198982FFECBE32FF5A560AFEB4F8F2" box="[253,382,1892,1916]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="0" pageNumber="492">Figs. 11, 12</figureCitation>
); occupying ~7% of shell length, ~19% of shell width. Teleoconch of ~5 whorls, each whorl with weakly convex, almost straight profile, with small subsutural shoulder, bearing small nodes in uniform intervals,
<quantity id="4CDA38E2FFECBE32FAC256E6FA56F82E" box="[1381,1436,1928,1952]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="0" pageNumber="492" unit="in" value="14.0">14 in</quantity>
penultimate whorl. Each subsutural node continuing inferiorly as axial, shallow rib, forming undulating sculpture; in spire, axial cords from suture to suture; in last whorl, axial ribs more nitid in superior half, in inferior half becoming lower, crossed by strong spiral liration, as ~15 spiral, narrow, smooth, uniform, crowded cords, covering entire inferior half, including canal (
<figureCitation id="13198982FFEDBE33FEEA51D5FE3AFF5D" box="[333,496,187,212]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="1" pageNumber="493">Figs. 2, 3, 5, 6</figureCitation>
). Aperture elliptical, oblique, not deflected, ~length twice exceeding width; occupying ~38% of shell length, ~40% of shell width; outer lip simply arched, weakly prosocline; ~10° with shell longitudinal axis (
<figureCitation id="13198982FFEDBE33FF38506DFEC9FE95" box="[159,259,259,284]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="1" pageNumber="493">Figs. 2, 5</figureCitation>
); inner lip shallowly concave, callus thin; anal notch small, producing subsutural shoulder. Canal ~12% of shell length; ~25% of shell width; rather opened. No umbilicus.
</paragraph>
<caption id="DF5DC58FFFEDBE33FF3F5664FBCCF823" ID-DOI="http://doi.org/10.5281/zenodo.10628132" ID-Zenodo-Dep="10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="1" pageNumber="493" startId="1.[152,270,1802,1826]" targetBox="[151,1436,344,1780]" targetPageId="1" targetType="figure">
<paragraph id="8B9D9507FFEDBE33FF3F5664FBCCF823" blockId="1.[151,1437,1802,1966]" pageId="1" pageNumber="493">
<emphasis id="B9564915FFEDBE33FF3F5664FD19F8AC" bold="true" box="[152,723,1802,1826]" pageId="1" pageNumber="493">
FIGURES 1–13:
<taxonomicName id="4C22EE84FFEDBE33FEFD5664FDE9F8AC" box="[346,547,1802,1826]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B9564915FFEDBE33FEFD5664FE3FF8AC" bold="true" box="[346,501,1802,1826]" italics="true" pageId="1" pageNumber="493">Cosmioconcha</emphasis>
spp
</taxonomicName>
type shells. 1–3
</emphasis>
.
<taxonomicName id="4C22EE84FFEDBE33FD465665FC95F8AC" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[737,863,1803,1826]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FD465665FC95F8AC" box="[737,863,1803,1826]" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm.
<emphasis id="B9564915FFEDBE33FE145643FE15F8CB" bold="true" box="[435,479,1837,1861]" pageId="1" pageNumber="493">4–6.</emphasis>
<taxonomicName id="4C22EE84FFEDBE33FE425643FD95F8CB" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[485,607,1837,1861]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FE425643FD95F8CB" box="[485,607,1837,1861]" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm.
<emphasis id="B9564915FFEDBE33FF30563EFF00F8E6" bold="true" box="[151,202,1872,1896]" pageId="1" pageNumber="493">7–10</emphasis>
. Types at same scale (= 3 mm).
<emphasis id="B9564915FFEDBE33FDBA563EFD89F8E6" bold="true" box="[541,579,1872,1896]" pageId="1" pageNumber="493">7–8</emphasis>
. Holotype of
<taxonomicName id="4C22EE84FFEDBE33FD76563EFC4CF8E6" authority="MZSP" authorityName="MZSP" box="[721,902,1872,1896]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEDBE33FD76563EFCF4F8E6" box="[721,830,1872,1896]" italics="true" pageId="1" pageNumber="493">C. helenae</emphasis>
MZSP
</taxonomicName>
102531, frontal and dorsal views.
<emphasis id="B9564915FFEDBE33FB4D563EFB3DF8E6" bold="true" box="[1258,1271,1872,1896]" pageId="1" pageNumber="493">9</emphasis>
. Holotype of
<taxonomicName id="4C22EE84FFEDBE33FA22563EFF3FF805" authorityName="Simone &amp; Monsecour" authorityYear="2024" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FA22563EFF3FF805" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
. 10. Paratype of
<taxonomicName id="4C22EE84FFEDBE33FE02561DFDEBF805" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[421,545,1907,1931]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FE02561DFDEBF805" box="[421,545,1907,1931]" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
.
<emphasis id="B9564915FFEDBE33FD89561DFDA7F805" bold="true" box="[558,621,1907,1931]" pageId="1" pageNumber="493">11–13</emphasis>
. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm.
<emphasis id="B9564915FFEDBE33FF3056F8FF65F820" bold="true" box="[151,175,1942,1966]" pageId="1" pageNumber="493">11</emphasis>
. Holotype of
<taxonomicName id="4C22EE84FFEDBE33FE9B56F8FE7DF820" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[316,439,1942,1966]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FE9B56F8FE7DF820" box="[316,439,1942,1966]" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
.
<emphasis id="B9564915FFEDBE33FE6356F8FE17F820" bold="true" box="[452,477,1942,1966]" pageId="1" pageNumber="493">12</emphasis>
. Paratype of
<taxonomicName id="4C22EE84FFEDBE33FDC256F8FD2AF820" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[613,736,1942,1966]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEDBE33FDC256F8FD2AF820" box="[613,736,1942,1966]" italics="true" pageId="1" pageNumber="493">C. costorum</emphasis>
</taxonomicName>
.
<emphasis id="B9564915FFEDBE33FD4A56F8FCCCF820" bold="true" box="[749,774,1942,1966]" pageId="1" pageNumber="493">13</emphasis>
. Holotype of
<taxonomicName id="4C22EE84FFEDBE33FC3456F8FBCAF820" box="[915,1024,1942,1966]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="1" pageNumber="493" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEDBE33FC3456F8FBCAF820" box="[915,1024,1942,1966]" italics="true" pageId="1" pageNumber="493">C. helenae</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B9D9507FFEEBE30FF6051F9FE2DFF21" blockId="2.[151,1437,151,1399]" box="[199,487,151,175]" pageId="2" pageNumber="494">
<emphasis id="B9564915FFEEBE30FF6051F9FEECFF21" box="[199,294,151,175]" inLineHeading="true" pageId="2" pageNumber="494" reason="1">
<emphasis id="B9564915FFEEBE30FF6051F9FED4FF21" bold="true" box="[199,286,151,175]" pageId="2" pageNumber="494">Habitat</emphasis>
.
</emphasis>
Dredged in
<specimenCount id="9D245E8EFFEEBE30FE0D51F9FE2DFF21" box="[426,487,151,175]" count="56" pageId="2" pageNumber="494" type="generic">
<quantity id="4CDA38E2FFEEBE30FE0D51F9FE29FF21" box="[426,483,151,175]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="494" unit="m" value="56.0">56 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFEEBE30FF6051D2FC75FEB1" pageId="2" pageNumber="494" type="etymology">
<paragraph id="8B9D9507FFEEBE30FF6051D2FC75FEB1" blockId="2.[151,1437,151,1399]" pageId="2" pageNumber="494">
<emphasis id="B9564915FFEEBE30FF6051D2FE83FF5D" box="[199,329,188,212]" inLineHeading="true" pageId="2" pageNumber="494" reason="1">
<emphasis id="B9564915FFEEBE30FF6051D2FE88FF5A" bold="true" box="[199,322,188,212]" pageId="2" pageNumber="494">Etymology</emphasis>
.
</emphasis>
the epithet is in honor of both researchers that dealt with the species, Fabio H. A. Costa, who described
<taxonomicName id="4C22EE84FFEEBE30FF30518EFEC0FF79" box="[151,266,223,247]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FF30518EFEC0FF79" box="[151,266,223,247]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
and designated the here studied
<typeStatus id="54992BA5FFEEBE30FDCF518EFD04FF76" box="[616,718,224,248]" pageId="2" pageNumber="494" type="paratype">paratypes</typeStatus>
(
<bibRefCitation id="EFB3E8F6FFEEBE30FD7B518EFCABFF76" author="Costa, F. H. A." box="[732,865,224,248]" pageId="2" pageNumber="494" pagination="95 - 99" refId="ref2013" refString="Costa, F. H. A. (1983) Anachis helenae, nova especie de Columbellidae do litoral Brasileiro (Mollusca: Gastropoda). Anais da Sociedade Nordestina de Zoologia, 1, 95 - 99." type="journal article" year="1983">Costa, 1983</bibRefCitation>
); and Paulo Marcio S. Costa, a Brazilian columbellid specialist who also have seen differences between southern and northern samples of previous concept of
<taxonomicName id="4C22EE84FFEEBE30FA86506AFA5CFE95" box="[1313,1430,259,283]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FA86506AFA5CFE95" box="[1313,1430,259,283]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
, additionally who designated the present generic combination (
<bibRefCitation id="EFB3E8F6FFEEBE30FC895046FC7EFEB1" author="Costa, P. M. S." box="[814,948,295,320]" pageId="2" pageNumber="494" refId="ref2053" refString="Costa, P. M. S. (2005) Estudo Taxonomico dos Representantes da Familia Columbellidae Swainson, 1840 (Mollusca, Caenogastropoda) da Costa Brasileira. PhD Dissertation. Universidade Federal do Rio de Janeiro, Rio de Janeiro, 293 pp." type="book" year="2005">Costa, 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFEEBE30FF605022FF2BFE09" pageId="2" pageNumber="494" type="description">
<paragraph id="8B9D9507FFEEBE30FF605022FF2BFE09" blockId="2.[151,1437,151,1399]" pageId="2" pageNumber="494">
<emphasis id="B9564915FFEEBE30FF605022FE1CFEEA" box="[199,470,331,356]" inLineHeading="true" pageId="2" pageNumber="494" reason="1">
<emphasis id="B9564915FFEEBE30FF605022FEA4FEEA" bold="true" box="[199,366,332,356]" pageId="2" pageNumber="494">Measurements</emphasis>
(in mm).
</emphasis>
<typeStatus id="54992BA5FFEEBE30FE7A5025FDF0FEED" box="[477,570,331,355]" pageId="2" pageNumber="494" type="holotype">holotype</typeStatus>
MZSP 102539 (
<figureCitation id="13198982FFEEBE30FD575025FC92FEEA" box="[752,856,331,356]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Figs. 1–3</figureCitation>
): 6.9 by 3.0;
<typeStatus id="54992BA5FFEEBE30FC4B5022FB82FEEA" box="[1004,1096,332,356]" pageId="2" pageNumber="494" type="paratype">paratype</typeStatus>
MZSP 102534 (
<figureCitation id="13198982FFEEBE30FB5A5025FAADFEED" box="[1277,1383,331,355]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Figs. 4–6</figureCitation>
): 6.2 by 2.5.
</paragraph>
</subSubSection>
<subSubSection id="C338C68CFFEEBE30FF6050FDFAA3FAF9" pageId="2" pageNumber="494" type="discussion">
<paragraph id="8B9D9507FFEEBE30FF6050FDFEEFFC49" blockId="2.[151,1437,151,1399]" pageId="2" pageNumber="494">
<emphasis id="B9564915FFEEBE30FF6050FDFE8CFE25" box="[199,326,403,427]" inLineHeading="true" pageId="2" pageNumber="494" reason="1">
<emphasis id="B9564915FFEEBE30FF6050FDFEF5FE25" bold="true" box="[199,319,403,427]" pageId="2" pageNumber="494">Discussion</emphasis>
.
</emphasis>
<taxonomicName id="4C22EE84FFEEBE30FEF550FDFDAAFE25" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[338,608,403,427]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FEF550FDFDAAFE25" box="[338,608,403,427]" italics="true" pageId="2" pageNumber="494">Cosmioconcha costorum</emphasis>
</taxonomicName>
is really very similar to
<taxonomicName id="4C22EE84FFEEBE30FC2050FAFC35FE25" box="[903,1023,403,427]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FC2050FAFC35FE25" box="[903,1023,403,427]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
, so much that both specimens were comprehensibly designated
<typeStatus id="54992BA5FFEEBE30FE1850D6FDEFFE5E" box="[447,549,440,464]" pageId="2" pageNumber="494" type="paratype">paratypes</typeStatus>
of that species by its author. However, a series of details indicate that both represent different entities. One of the main differences is that
<taxonomicName id="4C22EE84FFEEBE30FD6150B2FC82FE7D" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[710,840,476,499]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FD6150B2FC82FE7D" box="[710,840,476,499]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
has a multispiral protoconch (
<figureCitation id="13198982FFEEBE30FB2D50B5FAC0FE7A" box="[1162,1290,475,500]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Figs. 11, 12</figureCitation>
), of 3 whorls, while in
<taxonomicName id="4C22EE84FFEEBE30FF53536EFEACFD99" box="[244,358,511,535]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FF53536EFEACFD99" box="[244,358,511,535]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
the protoconch is paucispiral (
<figureCitation id="13198982FFEEBE30FD0B5091FD30FD96" box="[684,762,511,536]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Fig. 13</figureCitation>
), with 1.5 whorl. The size of both is also different,
<taxonomicName id="4C22EE84FFEEBE30FABD536EFA56FD99" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[1306,1436,512,535]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FABD536EFA56FD99" box="[1306,1436,512,535]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
is up to
<quantity id="4CDA38E2FFEEBE30FF54534AFEFFFDB5" box="[243,309,548,571]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="494" unit="mm" value="7.0">7 mm</quantity>
, while
<taxonomicName id="4C22EE84FFEEBE30FE22534AFE30FDB5" box="[389,506,547,571]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FE22534AFE30FDB5" box="[389,506,547,571]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
usually is over
<quantity id="4CDA38E2FFEEBE30FD0C534AFD3DFDB5" box="[683,759,548,572]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="494" unit="mm" value="10.0">10 mm</quantity>
with equivalent number of shell whorls (compare
<figureCitation id="13198982FFEEBE30FA84534DFA52FDB2" box="[1315,1432,547,572]" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Figs. 7–10</figureCitation>
, shells in the same scale). The shell profile of
<taxonomicName id="4C22EE84FFEEBE30FDD25326FD3CFDD1" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[629,758,584,607]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FDD25326FD3CFDD1" box="[629,758,584,607]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
is slightly more elongated and straight, while that of
<taxonomicName id="4C22EE84FFEEBE30FA8C5326FA56FDD1" box="[1323,1436,583,607]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FA8C5326FA56FDD1" box="[1323,1436,583,607]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
is slightly more globose, mainly in the last whorl, so much that the fist whorls have a slightly concave profile (
<figureCitation id="13198982FFEEBE30FAC05305FF09FD26" captionStart="FIGURES 1–13" captionStartId="1.[152,270,1802,1826]" captionTargetBox="[151,1436,344,1780]" captionTargetId="figure-107@1.[151,1436,341,1782]" captionTargetPageId="1" captionText="FIGURES 1–13: Cosmioconcha spp type shells. 1–3. C. costorum holotype MZSP 102539 (L 6.9 mm), frontal, right and dorsal views, scale = 1 mm. 4–6. C. costorum paratype MZSP 102534 (L 6.2 mm), frontal, right and dorsal views, scale = 1 mm. 7–10. Types at same scale (= 3 mm). 7–8. Holotype of C. helenae MZSP 102531, frontal and dorsal views. 9. Holotype of C. costorum. 10. Paratype of C. costorum. 11–13. Shell apex, arrows showing protoconch-teleoconch transition, scales= 0.5 mm. 11. Holotype of C. costorum. 12. Paratype of C. costorum. 13. Holotype of C. helenae." figureDoi="http://doi.org/10.5281/zenodo.10628132" httpUri="https://zenodo.org/record/10628132/files/figure.png" pageId="2" pageNumber="494">Figs. 7–8</figureCitation>
). The subsutural shoulder is much more pronounced in
<taxonomicName id="4C22EE84FFEEBE30FCBE53FEFC56FD29" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[793,924,656,679]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FCBE53FEFC56FD29" box="[793,924,656,679]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
, rendering the whorls profile flat. A shoulder is less developed in
<taxonomicName id="4C22EE84FFEEBE30FEF053DAFE00FD45" box="[343,458,691,715]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FEF053DAFE00FD45" box="[343,458,691,715]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
, which has whorls more convex. However, its
<typeStatus id="54992BA5FFEEBE30FC6553DDFBD5FD45" box="[962,1055,691,715]" pageId="2" pageNumber="494" type="holotype">holotype</typeStatus>
has a vestigial subsutural shoulder, but this has been shown a rare feature, as most specimens shows this much less-developed. Additionally, the lirations highly developed in the inferior half of last whorl, are not detected at all in superior half of
<taxonomicName id="4C22EE84FFEEBE30FBD75392FB39FC9D" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[1136,1267,764,787]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FBD75392FB39FC9D" box="[1136,1267,764,787]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
, but it is barely present in
<taxonomicName id="4C22EE84FFEEBE30FEA1524EFEBDFCB9" box="[262,375,799,823]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FEA1524EFEBDFCB9" box="[262,375,799,823]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
, overlying on axial sculpture. This discussion was enriched with examination of more specimens of
<taxonomicName id="4C22EE84FFEEBE30FF30522AFEC0FCD5" box="[151,266,835,859]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FF30522AFEC0FCD5" box="[151,266,835,859]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
that, as referred above, have been often collected, in the MZSP collection, or photos of which were accessed at some conchological web sites (e.g., Femorale, 2023). Geographically,
<taxonomicName id="4C22EE84FFEEBE30FC145206FBEDFCF1" box="[947,1063,871,895]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="helenae">
<emphasis id="B9564915FFEEBE30FC145206FBEDFCF1" box="[947,1063,871,895]" italics="true" pageId="2" pageNumber="494">C. helenae</emphasis>
</taxonomicName>
appears to be restricted to
<collectingRegion id="49E65BE5FFEEBE30FAF35209FEF8FC2D" country="Brazil" name="Rio de Janeiro" pageId="2" pageNumber="494">Rio de Janeiro region</collectingRegion>
, in shallow waters. Reported occurrences in other regions are apparently misidentifications, as the present reported one.
</paragraph>
<paragraph id="8B9D9507FFEEBE30FF6052BDFAA3FAF9" blockId="2.[151,1437,151,1399]" pageId="2" pageNumber="494">
<bibRefCitation id="EFB3E8F6FFEEBE30FF6052BDFEAEFC62" author="Pelorce, J." box="[199,356,979,1004]" pageId="2" pageNumber="494" pagination="4 - 21" refId="ref2155" refString="Pelorce, J. (2017) Les Columbellidae (Gastropoda: Neogastropoda) de la Guyane francaise. Xenophora Taxonomy, 14, 4 - 21." type="journal article" year="2017">Pelorce (2017)</bibRefCitation>
reviewed the
<taxonomicName id="4C22EE84FFEEBE30FE5C52BDFD52FC65" authorityName="Swainson" authorityYear="1840" box="[507,664,979,1003]" class="Gastropoda" family="Columbellidae" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="family">Columbellidae</taxonomicName>
from French
<collectingCountry id="F335D597FFEEBE30FC8D52BAFCB7FC62" box="[810,893,980,1004]" name="Guyana" pageId="2" pageNumber="494">Guyana</collectingCountry>
resulting in the description of several new species. He recognized six species of
<taxonomicName id="4C22EE84FFEEBE30FE775299FDA5FB81" authorityName="Dall" authorityYear="1913" box="[464,623,1015,1039]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="genus">
<emphasis id="B9564915FFEEBE30FE775299FDA5FB81" box="[464,623,1015,1039]" italics="true" pageId="2" pageNumber="494">Cosmioconcha</emphasis>
</taxonomicName>
, of which four were introduced as new species:
<taxonomicName id="4C22EE84FFEEBE30FBD35296FB3CFB81" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[1140,1270,1016,1039]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FBD35296FB3CFB81" box="[1140,1270,1016,1039]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C22EE84FFEEBE30FA235296FE0AFBBA" authority="Pelorce, 2017" authorityName="Pelorce" authorityYear="2017" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costattenuata">
<emphasis id="B9564915FFEEBE30FA235296FEEDFBBD" italics="true" pageId="2" pageNumber="494">C. costattenuata</emphasis>
<bibRefCitation id="EFB3E8F6FFEEBE30FE8A5575FE0AFBBA" author="Pelorce, J." box="[301,448,1051,1076]" pageId="2" pageNumber="494" pagination="4 - 21" refId="ref2155" refString="Pelorce, J. (2017) Les Columbellidae (Gastropoda: Neogastropoda) de la Guyane francaise. Xenophora Taxonomy, 14, 4 - 21." type="journal article" year="2017">
Pelorce,
<quantity id="4CDA38E2FFEEBE30FE2C5572FE0AFBBA" box="[395,448,1052,1076]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="2" pageNumber="494" unit="in" value="2017.0">2017</quantity>
</bibRefCitation>
</taxonomicName>
in being smaller, in having multispiral protoconch (that species has a 1.5 whorl protoconch), by fewer teleoconch whorl (that species has 6 whorls), by more globose outline (that species is more elongated) and in having much more pronounced axial ribs.
<taxonomicName id="4C22EE84FFEEBE30FDFD550AFD16FBF5" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[602,732,1124,1147]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FDFD550AFD16FBF5" box="[602,732,1124,1147]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C22EE84FFEEBE30FCCE550AFBB7FBF2" authority="Pelorce, 2017" authorityName="Pelorce" authorityYear="2017" box="[873,1149,1123,1148]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="flammea">
<emphasis id="B9564915FFEEBE30FCCE550AFC2EFBF5" box="[873,996,1123,1147]" italics="true" pageId="2" pageNumber="494">C. flammea</emphasis>
<bibRefCitation id="EFB3E8F6FFEEBE30FC4C550DFBB7FBF2" author="Pelorce, J." box="[1003,1149,1123,1148]" pageId="2" pageNumber="494" pagination="4 - 21" refId="ref2155" refString="Pelorce, J. (2017) Les Columbellidae (Gastropoda: Neogastropoda) de la Guyane francaise. Xenophora Taxonomy, 14, 4 - 21." type="journal article" year="2017">
Pelorce,
<quantity id="4CDA38E2FFEEBE30FBEF550AFBB7FBF2" box="[1096,1149,1124,1148]" metricMagnitude="1" metricUnit="m" metricValue="5.12318" pageId="2" pageNumber="494" unit="in" value="2017.0">2017</quantity>
</bibRefCitation>
</taxonomicName>
in being slightly larger, by more elongated outline, by shorter protoconch (that species has a 4-whorl protoconch) lacking a clear subsutural spiral incision; additionally,
<taxonomicName id="4C22EE84FFEEBE30FE2155C2FDC8FB4D" authorityName="Pelorce" authorityYear="2017" box="[390,514,1195,1219]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="flammea">
<emphasis id="B9564915FFEEBE30FE2155C2FDC8FB4D" box="[390,514,1195,1219]" italics="true" pageId="2" pageNumber="494">C. flammea</emphasis>
</taxonomicName>
is distinct in only showing large quantity of axial ribs, close located from each other, on the last whorls, while earlier whorls lack axial sculpture.
<taxonomicName id="4C22EE84FFEEBE30FCBE55BEFC51FB69" authorityName="Simone &amp; Monsecour" authorityYear="2024" box="[793,923,1232,1255]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FCBE55BEFC51FB69" box="[793,923,1232,1255]" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C22EE84FFEEBE30FB8F55BEFF05FA85" authority="Monsecour &amp; Monsecour, 2006" authorityName="Monsecour &amp; Monsecour" authorityYear="2006" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="rikae">
<emphasis id="B9564915FFEEBE30FB8F55BEFBB4FB69" box="[1064,1150,1231,1255]" italics="true" pageId="2" pageNumber="494">C. rikae</emphasis>
Monsecour &amp; Monsecour, 2006
</taxonomicName>
, from the Caribbean
<collectingCountry id="F335D597FFEEBE30FE0F559AFE36FA85" box="[424,508,1268,1291]" name="Panama" pageId="2" pageNumber="494">Panama</collectingCountry>
and French
<collectingCountry id="F335D597FFEEBE30FDDC559AFD1BFA82" box="[635,721,1268,1292]" name="Guyana" pageId="2" pageNumber="494">Guyana</collectingCountry>
, in having a shorter outline, and fewer axial ribs in entire teleoconch, including in last whorl, where that species lacks sculpture; both species share the same size and protoconch features.
<taxonomicName id="4C22EE84FFEEBE30FA235476FF30FADD" authorityName="Simone &amp; Monsecour" authorityYear="2024" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="costorum">
<emphasis id="B9564915FFEEBE30FA235476FF30FADD" italics="true" pageId="2" pageNumber="494">C. costorum</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C22EE84FFEEBE30FE2C5452FCF8FADD" authority="Monsecour &amp; Monsecour" authorityName="Monsecour &amp; Monsecour" box="[395,818,1339,1364]" class="Gastropoda" family="Columbellidae" genus="Cosmioconcha" kingdom="Animalia" order="Neogastropoda" pageId="2" pageNumber="494" phylum="Mollusca" rank="species" species="coltrorum">
<emphasis id="B9564915FFEEBE30FE2C5452FDDCFADD" box="[395,534,1339,1363]" italics="true" pageId="2" pageNumber="494">C. coltrorum</emphasis>
Monsecour &amp; Monsecour
</taxonomicName>
(in press) in being smaller, more globose (that species is slender), fewer teleoconch whorls, more spaced and more pronounced axial ribs, and less pronounced color pattern.
</paragraph>
</subSubSection>
</treatment>
</document>
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