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<document id="B85E0350D0DDBAC98253CDE9E2F7F594" ID-DOI="10.5281/zenodo.5324736" ID-ISSN="0253-116X" ID-Zenodo-Dep="5324736" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630150166855" checkinUser="carolina" docAuthor="Guéorguiev, B." docDate="2011" docId="038887CD6E637F2AFF26FA92FC5C33FE" docLanguage="en" docName="LinzerbiolBeitr.2011.43.1.501-547.pdf" docOrigin="Linzer biologische Beiträge 43 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Feronidius) incommodus SCHAUM 1858" docType="treatment" docVersion="3" lastPageNumber="506" masterDocId="FFB1FFB56E677F2FFF94FFB8FF953631" masterDocTitle="New and interesting records of Carabid Beetles from South-East Europe, South-West and Central Asia, with taxonomic notes on Pterostichini and Zabrini (Coleoptera, Carabidae)" masterLastPageNumber="547" masterPageNumber="501" pageNumber="505" updateTime="1699164681049" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="05F3D15628AF8D8BA543183225681C45" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8C29B0943E799CCFAB52A4F9D4F4A2A9">
<mods:title id="9C3F11426E0C945F6B601AD5F25E328A">New and interesting records of Carabid Beetles from South-East Europe, South-West and Central Asia, with taxonomic notes on Pterostichini and Zabrini (Coleoptera, Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="3910308D5726C0C0C98D4C910A1E9FCD" type="personal">
<mods:role id="133D730639303DD4D3CBCDAD3D1AC3D5">
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<mods:namePart id="C31C948E49AE503D6358D2AC6BD5B288">Guéorguiev, B.</mods:namePart>
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<mods:typeOfResource id="61D1FE1A84B1B930E572163995E4AFC9">text</mods:typeOfResource>
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<mods:title id="A444746DFB82854F5E1B2FDF0A3FB75B">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="AE8D063718B592B2EBF8059B39E1F2DB">
<mods:date id="37FC38779E4560E6A8B871C1CFE4EECD">2011</mods:date>
<mods:detail id="6F5D13521EF863F27978D8B5F7137B8A" type="pubDate">
<mods:number id="56CEC97518116F92BCBBDFFB08F2E6F6">2011-07-25</mods:number>
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<mods:detail id="E6EF3905C325A2AD7D4CD471F6A31DE6" type="volume">
<mods:number id="0D0BE675413F4EC8991848C9D9A302C3">43</mods:number>
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<mods:number id="784FAC5AE1CFEC0FDDFB902320E20329">1</mods:number>
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<mods:start id="8E337D49374CCD6A78E7349659DCF995">501</mods:start>
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<mods:classification id="3797E31EF361765ADAAAFB5F4D8BFD44">journal article</mods:classification>
<mods:identifier id="0AD7303506F4975FDCB704C13D79EB72" type="DOI">10.5281/zenodo.5324736</mods:identifier>
<mods:identifier id="0D7935DF481C5A8AD42B1CD079362480" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="F511E840E2276ED6B1D14428E1115EFF" type="Zenodo-Dep">5324736</mods:identifier>
</mods:mods>
<treatment id="038887CD6E637F2AFF26FA92FC5C33FE" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE" httpUri="http://treatment.plazi.org/id/038887CD6E637F2AFF26FA92FC5C33FE" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<subSubSection id="C33B65506E637F2BFF26FA92FD703370" box="[178,741,1322,1345]" pageId="4" pageNumber="505" type="nomenclature">
<paragraph id="8B9E36DB6E637F2BFF26FA92FD703370" blockId="4.[178,741,1322,1345]" box="[178,741,1322,1345]" pageId="4" pageNumber="505">
<heading id="D0D681B76E637F2BFF26FA92FD703370" box="[178,741,1322,1345]" fontSize="8" level="2" pageId="4" pageNumber="505" reason="2">
<taxonomicName id="4C214D586E637F2BFF26FA92FD703370" ID-CoL="4Q3Q3" authority="SCHAUM 1858" authorityName="SCHAUM" authorityYear="1858" box="[178,741,1322,1345]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="incommodus" subGenus="Feronidius">
<emphasis id="B955EAC96E637F2BFF26FA92FD703370" bold="true" box="[178,741,1322,1345]" pageId="4" pageNumber="505">
<emphasis id="B955EAC96E637F2BFF26FA92FEA63371" bold="true" box="[178,307,1322,1344]" italics="true" pageId="4" pageNumber="505">Pterostichus</emphasis>
(
<emphasis id="B955EAC96E637F2BFED6FA92FE203371" bold="true" box="[322,437,1322,1344]" italics="true" pageId="4" pageNumber="505">Feronidius</emphasis>
)
<emphasis id="B955EAC96E637F2BFE50FA92FDDF3371" bold="true" box="[452,586,1322,1344]" italics="true" pageId="4" pageNumber="505">incommodus</emphasis>
SCHAUM 1858
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B65506E637F2BFF26FAEAFCCA33AB" pageId="4" pageNumber="505" type="reference_group">
<paragraph id="8B9E36DB6E637F2BFF26FAEAFD2C3357" blockId="4.[178,863,1362,1460]" box="[178,697,1362,1382]" pageId="4" pageNumber="505">
<taxonomicName id="4C214D586E637F2BFF26FAEAFD2C3357" ID-CoL="4Q3Q3" authority="SCHAUM 1858: 470" authorityName="SCHAUM" authorityPageNumber="470" authorityYear="1858" box="[178,697,1362,1382]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="incommodus" subGenus="Steropus">
<emphasis id="B955EAC96E637F2BFF26FAEAFEB63357" box="[178,291,1362,1382]" italics="true" pageId="4" pageNumber="505">Pterostichus</emphasis>
(
<emphasis id="B955EAC96E637F2BFEA5FAEAFE153357" box="[305,384,1362,1382]" italics="true" pageId="4" pageNumber="505">Steropus</emphasis>
)
<emphasis id="B955EAC96E637F2BFE19FAEAFD953357" box="[397,512,1362,1382]" italics="true" pageId="4" pageNumber="505">incommodus</emphasis>
SCHAUM 1858: 470
</taxonomicName>
</paragraph>
<paragraph id="8B9E36DB6E637F2BFF26FAD4FD3E33B1" blockId="4.[178,863,1362,1460]" box="[178,683,1388,1408]" pageId="4" pageNumber="505">
<taxonomicName id="4C214D586E637F2BFF26FAD4FDDE33B1" ID-CoL="4Q629" authority="APFELBECK 1899: 290" authorityName="APFELBECK" authorityPageNumber="290" authorityYear="1899" box="[178,587,1388,1408]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E637F2BFF26FAD4FEE033B1" box="[178,373,1388,1408]" italics="true" pageId="4" pageNumber="505">Pterostichus serbicus</emphasis>
APFELBECK 1899: 290
</taxonomicName>
,
<emphasis id="B955EAC96E637F2BFDC1FAD4FD3E33B1" bold="true" box="[597,683,1388,1408]" pageId="4" pageNumber="505">
<taxonomicNameLabel id="A26657B26E637F2BFDC1FAD4FD3E33B1" box="[597,683,1388,1408]" pageId="4" pageNumber="505">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9E36DB6E637F2BFF26FA3EFCCA33AB" blockId="4.[178,863,1362,1460]" box="[178,863,1414,1434]" pageId="4" pageNumber="505">
<taxonomicName id="4C214D586E637F2BFF26FA3EFCCA33AB" ID-CoL="4Q69P" authority="APFLEBECK 1904: 265" authorityName="APFLEBECK" authorityYear="1904" box="[178,863,1414,1434]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="variety" species="serbicus" subGenus="Pterostichus" variety="unistriatus">
<emphasis id="B955EAC96E637F2BFF26FA3EFEB633AB" box="[178,291,1414,1434]" italics="true" pageId="4" pageNumber="505">Pterostichus</emphasis>
(
<emphasis id="B955EAC96E637F2BFEA5FA3EFE3733AB" box="[305,418,1414,1434]" italics="true" pageId="4" pageNumber="505">Pterostichus</emphasis>
)
<emphasis id="B955EAC96E637F2BFE3BFA3EFE6E33AB" box="[431,507,1414,1434]" italics="true" pageId="4" pageNumber="505">serbicus</emphasis>
var.
<emphasis id="B955EAC96E637F2BFDBDFA3EFD1F33AB" box="[553,650,1414,1434]" italics="true" pageId="4" pageNumber="505">unistriatus</emphasis>
APFLEBECK 1904: 265
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C33B65506E637F2AFF26FA18FC943786" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="description">
<paragraph id="8B9E36DB6E637F2BFF26FA18FCD23385" blockId="4.[178,863,1362,1460]" box="[178,839,1440,1460]" pageId="4" pageNumber="505">
<emphasis id="B955EAC96E637F2BFF26FA18FE923385" box="[178,263,1440,1460]" italics="true" pageId="4" pageNumber="505">Platysma</emphasis>
(
<taxonomicName id="4C214D586E637F2BFE80FA18FE103385" authorityName="BONELLI" authorityYear="1810" box="[276,389,1440,1460]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="genus">
<emphasis id="B955EAC96E637F2BFE80FA18FE103385" box="[276,389,1440,1460]" italics="true" pageId="4" pageNumber="505">Pterostichus</emphasis>
</taxonomicName>
)
<emphasis id="B955EAC96E637F2BFE06FA18FE703385" box="[402,485,1440,1460]" italics="true" pageId="4" pageNumber="505">serbicum</emphasis>
<taxonomicName id="4C214D586E637F2BFE7FFA18FCD23385" authority="LUTSCHNIK 1915: 29" authorityName="LUTSCHNIK" authorityPageNumber="29" authorityYear="1915" box="[491,839,1440,1460]" pageId="4" pageNumber="505" rank="variety" variety="karageorgii">
var.
<emphasis id="B955EAC96E637F2BFD80FA18FDEA3385" box="[532,639,1440,1460]" italics="true" pageId="4" pageNumber="505">karageorgii</emphasis>
LUTSCHNIK 1915: 29
</taxonomicName>
</paragraph>
<paragraph id="8B9E36DB6E637F2BFF26FA7AFD8E33E7" blockId="4.[178,1085,1474,1608]" box="[178,539,1474,1494]" pageId="4" pageNumber="505">
<taxonomicName id="4C214D586E637F2BFF26FA7AFD8E33E7" authority="APFELBECK 1899" authorityName="APFELBECK" authorityYear="1899" box="[178,539,1474,1494]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="505" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E637F2BFF26FA7AFEE033E7" box="[178,373,1474,1494]" italics="true" pageId="4" pageNumber="505">Pterostichus serbicus</emphasis>
APFELBECK 1899
</taxonomicName>
</paragraph>
<paragraph id="8B9E36DB6E637F2BFF26FA61FC4F302A" blockId="4.[178,1085,1474,1608]" pageId="4" pageNumber="505">
T y p e m a t e r i a l 1,
<specimenCount id="9D27FD526E637F2BFE48FA61FDC333DC" box="[476,598,1497,1517]" pageId="4" pageNumber="505" type="generic" typeStatus="syntypes">1 syntypes</specimenCount>
, labelled: &quot;AK-Palanka Suha plan.&quot; [typeset white label] / &quot;
<emphasis id="B955EAC96E637F2BFE91FA48FEC43035" box="[261,337,1520,1540]" italics="true" pageId="4" pageNumber="505">serbicus</emphasis>
det. Apflb.&quot; [white label, &quot;
<emphasis id="B955EAC96E637F2BFDC7FA48FD0A3035" box="[595,671,1520,1540]" italics="true" pageId="4" pageNumber="505">serbicus</emphasis>
&quot; handwritten and red underlined, remaining typeset inscription] / &quot;Collectie C. &amp; O. Vogt Acq. 1960&quot; [typeset white label] (ZMAN).
</paragraph>
<paragraph id="8B9E36DB6E637F2BFF26F9A5FD293000" blockId="4.[178,1085,1474,1608]" box="[178,700,1565,1585]" pageId="4" pageNumber="505">T y p e l o c a l i t y.&quot;AkPalankainSerbien(Hilf)&quot;.</paragraph>
<paragraph id="8B9E36DB6E637F2AFF26F98CFC943786" blockId="4.[178,1085,1474,1608]" lastBlockId="5.[178,1086,215,439]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<materialsCitation id="3B493C866E637F2AFF26F98CFE6536DA" collectingDate="1906-04-04" collectionCode="NMNHS" country="Bulgaria" lastPageId="5" lastPageNumber="506" location="Dragalevtsi Village" municipality="Vitosha Mt." pageId="4" pageNumber="505" specimenCount="1">
O t h e r m a t e r i a l e x m a i n e d.
<collectingCountry id="F336764B6E637F2BFDE6F98CFD7E3079" box="[626,747,1588,1608]" name="Bulgaria" pageId="4" pageNumber="505">BULGARIA</collectingCountry>
.
<collectingMunicipality id="6BFAACA16E637F2BFD68F98CFCE03079" box="[764,885,1588,1608]" pageId="4" pageNumber="505">Vitosha Mt.</collectingMunicipality>
, around
<location id="8EFE60006E637F2AFC40F98CFF6236DA" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E637F2AFC40F98CFF6236DA" country="Bulgaria" lastPageId="5" lastPageNumber="506" municipality="Vitosha Mt." name="Dragalevtsi Village" pageId="4" pageNumber="505">Dragalevtsi Village</location>
,
<date id="FF9F101B6E627F2AFE95FF6FFECB36DA" box="[257,350,215,235]" pageId="5" pageNumber="506" value="1906-04-04">
<collectingDate id="EFDBE9F36E627F2AFE95FF6FFECB36DA" box="[257,350,215,235]" pageId="5" pageNumber="506" value="1906-04-04">4.IV.1906</collectingDate>
</date>
, 1 (
<collectionCode id="ED30AE1E6E627F2AFE07FF6FFE7F36DA" box="[403,490,215,235]" pageId="5" pageNumber="506">NMNHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFE6EFF6FFE833733" collectingDate="1911-07-20" collectionCode="NMNHS" country="Bulgaria" location="Germanski Monastery" municipality="Lozenska Planina Mt." pageId="5" pageNumber="506" specimenCount="1">
<collectingMunicipality id="6BFAACA16E627F2AFE6EFF6FFD5036DA" box="[506,709,215,235]" pageId="5" pageNumber="506">Lozenska Planina Mt.</collectingMunicipality>
,
<location id="8EFE60006E627F2AFD59FF6FFC0F36DA" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFD59FF6FFC0F36DA" box="[717,922,215,235]" country="Bulgaria" municipality="Lozenska Planina Mt." name="Germanski Monastery" pageId="5" pageNumber="506">Germanski Monastery</location>
,
<date id="FF9F101B6E627F2AFC30FF6FFB8036DA" box="[932,1045,215,235]" pageId="5" pageNumber="506" value="1911-07-20">
<collectingDate id="EFDBE9F36E627F2AFC30FF6FFB8036DA" box="[932,1045,215,235]" pageId="5" pageNumber="506" value="1911-07-20">20.VII.1911</collectingDate>
</date>
, 1 (
<collectionCode id="ED30AE1E6E627F2AFF2CFF56FE9A3733" box="[184,271,238,258]" pageId="5" pageNumber="506">NMNHS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFEB2FF56FC683733" box="[294,1021,238,258]" collectingDate="1963-10-12" collectionCode="R, MIZ" country="Bulgaria" county="Bulgaria" location="Jskrec" municipality="Stara Planina" pageId="5" pageNumber="506" specimenCount="1">
&quot;
<collectingCounty id="62FF4E576E627F2AFEB9FF56FEEA3733" box="[301,383,238,258]" pageId="5" pageNumber="506">Bulgaria</collectingCounty>
,
<collectingMunicipality id="6BFAACA16E627F2AFE1BFF56FD853733" box="[399,528,238,258]" pageId="5" pageNumber="506">Stara Planina</collectingMunicipality>
,
<location id="8EFE60006E627F2AFDB4FF56FDCC3733" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFDB4FF56FDCC3733" box="[544,601,238,258]" country="Bulgaria" county="Bulgaria" municipality="Stara Planina" name="Jskrec" pageId="5" pageNumber="506">Jskrec</location>
<date id="FF9F101B6E627F2AFDF1FF56FD533733" box="[613,710,238,258]" pageId="5" pageNumber="506" value="1963-10-12">
<collectingDate id="EFDBE9F36E627F2AFDF1FF56FD533733" box="[613,710,238,258]" pageId="5" pageNumber="506" value="1963-10-12">12.X.1963</collectingDate>
</date>
leg.
<collectionCode id="ED30AE1E6E627F2AFD6BFF56FC843733" box="[767,785,238,258]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="506">R</collectionCode>
. Bielawski&quot;, 1 (
<collectionCode id="ED30AE1E6E627F2AFC5CFF56FC633733" box="[968,1014,238,258]" pageId="5" pageNumber="506">MIZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFB99FF56FE13371E" collectingDate="2004-04-03" collectingDateMax="2004-07-22" collectingDateMin="2004-04-03" collectionCode="NMNHS" collectorName="E. Migliaccio" country="Bulgaria" elevation="630" location="Marsh" municipality="Dragomansko" pageId="5" pageNumber="506" specimenCount="1" stateProvince="Sofiya-Grad">
<collectingRegion id="49E5F8396E627F2AFB99FF56FF6D3729" country="Bulgaria" name="Sofiya-Grad" pageId="5" pageNumber="506">Sofia District</collectingRegion>
,
<collectingMunicipality id="6BFAACA16E627F2AFE9CFEBCFE123729" box="[264,391,260,280]" pageId="5" pageNumber="506">Dragomansko</collectingMunicipality>
blato
<location id="8EFE60006E627F2AFE59FEBCFD9C3729" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFE59FEBCFD9C3729" box="[461,521,260,280]" country="Bulgaria" municipality="Dragomansko" name="Marsh" pageId="5" pageNumber="506" stateProvince="Sofiya-Grad">Marsh</location>
,
<quantity id="4CD99B3E6E627F2AFD8DFEBCFDCE3729" box="[537,603,260,280]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="506" unit="m" value="630.0">
<elevation id="000CD1E86E627F2AFD8DFEBCFDCE3729" box="[537,603,260,280]" metricMagnitude="2" metricUnit="m" metricValue="6.3" pageId="5" pageNumber="506" unit="m" value="630.0">630 m</elevation>
</quantity>
,
<date id="FF9F101B6E627F2AFDFFFEBCFC983729" box="[619,781,260,280]" pageId="5" pageNumber="506" value="2004-07-22" valueMax="2004-04-03" valueMin="2004-07-22">
<collectingDate id="EFDBE9F36E627F2AFDFFFEBCFC983729" box="[619,781,260,280]" pageId="5" pageNumber="506" value="2004-07-22" valueMax="2004-04-03" valueMin="2004-07-22">22.VII-3.IV.2004</collectingDate>
</date>
,
<collectingMethod id="52604ECC6E627F2AFC89FEBCFCED3729" box="[797,888,260,280]" pageId="5" pageNumber="506">soil traps</collectingMethod>
, 1, 2, leg.
<collectorName id="26D4530D6E627F2AFBBDFEBCFE81371E" pageId="5" pageNumber="506">E. Migliaccio</collectorName>
(
<collectionCode id="ED30AE1E6E627F2AFEBDFEA3FE15371E" box="[297,384,283,303]" pageId="5" pageNumber="506">NMNHS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFE03FEA3FC81371E" box="[407,788,283,303]" collectionCode="ZMAN" country="Croatia" location="Dalmatia" pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFE03FEA3FE6A371E" box="[407,511,283,303]" name="Croatia" pageId="5" pageNumber="506">CROATIA</collectingCountry>
. &quot;
<location id="8EFE60006E627F2AFD83FEA3FDE5371E" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFD83FEA3FDE5371E" box="[535,624,283,303]" country="Croatia" name="Dalmatia" pageId="5" pageNumber="506">Dalmatia</location>
&quot;, 2 s. (
<collectionCode id="ED30AE1E6E627F2AFD51FEA3FC98371E" box="[709,781,283,303]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" lsid="urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="5" pageNumber="506">ZMAN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFCB1FEA3FE043777" collectionCode="NMNHS" country="Hungary" location="Umgb." pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFCB1FEA3FC0F371E" box="[805,922,283,303]" name="Hungary" pageId="5" pageNumber="506">HUNGARY</collectingCountry>
. &quot;
<location id="8EFE60006E627F2AFC26FEA3FC66371E" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFC26FEA3FC66371E" box="[946,1011,283,303]" country="Hungary" name="Umgb." pageId="5" pageNumber="506">Umgb.</location>
B-Pest Diener&quot;, 1 (
<collectionCode id="ED30AE1E6E627F2AFEA0FE8AFE1E3777" box="[308,395,306,326]" pageId="5" pageNumber="506">NMNHS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFE08FE8AFE9C376C" collectionCode="HNHM" country="Kosovo" location="Banjska" municipality="Novipazar" pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFE08FE8AFE6B3777" box="[412,510,306,326]" name="Kosovo" pageId="5" pageNumber="506">KOSOVO</collectingCountry>
. &quot;
<collectingMunicipality id="6BFAACA16E627F2AFD85FE8AFDE03777" box="[529,629,306,326]" pageId="5" pageNumber="506">Novipazar</collectingMunicipality>
:
<location id="8EFE60006E627F2AFD14FE8AFD5E3777" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFD14FE8AFD5E3777" box="[640,715,306,326]" country="Kosovo" municipality="Novipazar" name="Banjska" pageId="5" pageNumber="506">Banjska</location>
&quot; / &quot;Akad. Balk. Exp. Csiki 1916&quot;, 1 s. (
<collectionCode id="ED30AE1E6E627F2AFF2CFEF1FE96376C" box="[184,259,329,349]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="506">HNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFE87FEF1FCC5376C" box="[275,848,329,349]" collectionCode="HNHM" country="Kosovo" location="Albania Ipek" pageId="5" pageNumber="506" specimenCount="1">
&quot;
<location id="8EFE60006E627F2AFE8EFEF1FE01376C" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFE8EFEF1FE01376C" box="[282,404,329,349]" country="Kosovo" name="Albania Ipek" pageId="5" pageNumber="506">Albania Ipek</location>
&quot; / &quot;Akad. Balk. Exp. Csiki 1917&quot;, 1 s. (
<collectionCode id="ED30AE1E6E627F2AFD6BFEF1FCDF376C" box="[767,842,329,349]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="506">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFCCDFEF1FE183742" collectionCode="MIZ" country="Romania" location="ROMANIA" pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFCCDFEF1FC5C376C" box="[857,969,329,349]" name="Romania" pageId="5" pageNumber="506">ROMANIA</collectingCountry>
. &quot;Biharfüred Hungaria&quot;, 1 (
<collectionCode id="ED30AE1E6E627F2AFECCFEE7FE133742" box="[344,390,351,371]" pageId="5" pageNumber="506">MIZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFE0BFEE7FCD63742" box="[415,835,351,371]" collectionCode="MIZ" country="Romania" location="Mts. Bihar Hungaria" pageId="5" pageNumber="506" specimenCount="1">
&quot;
<location id="8EFE60006E627F2AFE33FEE7FDEE3742" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFE33FEE7FDEE3742" box="[423,635,351,371]" country="Romania" name="Mts. Bihar Hungaria" pageId="5" pageNumber="506">Mts. Bihar Hungaria</location>
&quot;, 2, 1 (
<collectionCode id="ED30AE1E6E627F2AFC99FEE7FCAE3742" box="[781,827,351,371]" pageId="5" pageNumber="506">MIZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFCC0FEE7FEEC37BB" collectionCode="MIZ" country="Serbia" location="SERBIA" pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFCC0FEE7FC3D3742" box="[852,936,351,371]" name="Serbia" pageId="5" pageNumber="506">SERBIA</collectingCountry>
. &quot;Fruška Gora Syrmien.&quot;, 1 (
<collectionCode id="ED30AE1E6E627F2AFED0FECEFEE737BB" box="[324,370,374,394]" pageId="5" pageNumber="506">MIZ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFE16FECEFCF937BB" box="[386,876,374,394]" collectionCode="MNHUB" collectorName="Dr. Maertens" country="Serbia" location="Serbien" pageId="5" pageNumber="506" specimenCount="1" stateProvince="Paracin">
&quot;
<location id="8EFE60006E627F2AFE1EFECEFE4437BB" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFE1EFECEFE4437BB" box="[394,465,374,394]" country="Serbia" name="Serbien" pageId="5" pageNumber="506" stateProvince="Paracin">Serbien</location>
<collectingRegion id="49E5F8396E627F2AFE43FECEFD8E37BB" box="[471,539,374,394]" country="Serbia" name="Paracin" pageId="5" pageNumber="506">Paracin</collectingRegion>
1918
<collectorName id="26D4530D6E627F2AFDC0FECEFD5A37BB" box="[596,719,374,394]" pageId="5" pageNumber="506">Dr. Maertens</collectorName>
&quot;, 2 s. (
<collectionCode id="ED30AE1E6E627F2AFC98FECEFCF337BB" box="[780,870,374,394]" pageId="5" pageNumber="506">MNHUB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFCE1FECEFDA03790" collectingDate="1957-03" collectionCode="HNHM" country="Serbia" location="Jugoslavia" pageId="5" pageNumber="506" specimenCount="1" stateProvince="Nis">
&quot;
<location id="8EFE60006E627F2AFCE9FECEFC4A37BB" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFCE9FECEFC4A37BB" box="[893,991,374,394]" country="Serbia" name="Jugoslavia" pageId="5" pageNumber="506" stateProvince="Nis">Jugoslavia</location>
<collectingRegion id="49E5F8396E627F2AFC70FECEFB9337BB" box="[996,1030,374,394]" country="Serbia" name="Nis" pageId="5" pageNumber="506">Nis</collectingRegion>
,
<date id="FF9F101B6E627F2AFB84FECEFF453790" pageId="5" pageNumber="506" value="1957-03">
<collectingDate id="EFDBE9F36E627F2AFB84FECEFF453790" pageId="5" pageNumber="506" value="1957-03">1957 III.</collectingDate>
</date>
&quot; / &quot;legit dr. Lenczy&quot;, 18 s. (
<collectionCode id="ED30AE1E6E627F2AFE70FE35FDBA3790" box="[484,559,397,417]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="506">HNHM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B493C866E627F2AFDD6FE35FE9C3786" collectingDate="1957-04" collectionCode="HNHM" country="Serbia" location="Jugoslavia" pageId="5" pageNumber="506" specimenCount="1" stateProvince="Nis">
&quot;
<location id="8EFE60006E627F2AFDDDFE35FD3E3790" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFDDDFE35FD3E3790" box="[585,683,397,417]" country="Serbia" name="Jugoslavia" pageId="5" pageNumber="506" stateProvince="Nis">Jugoslavia</location>
<collectingRegion id="49E5F8396E627F2AFD20FE35FD433790" box="[692,726,397,417]" country="Serbia" name="Nis" pageId="5" pageNumber="506">Nis</collectingRegion>
,
<date id="FF9F101B6E627F2AFD76FE35FCAD3790" box="[738,824,397,417]" pageId="5" pageNumber="506" value="1957-04">
<collectingDate id="EFDBE9F36E627F2AFD76FE35FCAD3790" box="[738,824,397,417]" pageId="5" pageNumber="506" value="1957-04">1957 IV.</collectingDate>
</date>
&quot; / &quot;legit dr. Lenczy&quot;, 17 s. (
<collectionCode id="ED30AE1E6E627F2AFF2CFE1BFE963786" box="[184,259,419,439]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="506">HNHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B493C866E627F2AFE86FE1BFD683786" box="[274,765,419,439]" collectionCode="V, ZMAN" country="Slovakia" location="Presb. Com." pageId="5" pageNumber="506" specimenCount="1">
<collectingCountry id="F336764B6E627F2AFE86FE1BFE1C3786" box="[274,393,419,439]" name="Slovakia" pageId="5" pageNumber="506">SLOVAKIA</collectingCountry>
. &quot;
<location id="8EFE60006E627F2AFE0EFE1BFD983786" LSID="urn:lsid:plazi:treatment:038887CD6E637F2AFF26FA92FC5C33FE:8EFE60006E627F2AFE0EFE1BFD983786" box="[410,525,419,439]" country="Slovakia" name="Presb. Com." pageId="5" pageNumber="506">Presb. Com.</location>
<collectionCode id="ED30AE1E6E627F2AFD85FE1BFDB13786" box="[529,548,419,439]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="506" type="Museum">V</collectionCode>
. Zoufal.&quot;, 1 (
<collectionCode id="ED30AE1E6E627F2AFD3AFE1BFD633786" box="[686,758,419,439]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34975" lsid="urn:lsid:biocol.org:col:34975" name="Instituut voor Taxonomische Zoologie, Zoologisch Museum" pageId="5" pageNumber="506">ZMAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33B65506E627F2AFF26FE7FFD54335E" pageId="5" pageNumber="506" type="reference_group">
<paragraph id="8B9E36DB6E627F2AFF26FE7FFE4434D2" blockId="5.[178,1085,454,739]" pageId="5" pageNumber="506">
<taxonomicName id="4C214D586E627F2AFF26FE7FFEB737ED" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[178,290,454,476]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFF26FE7FFF5437ED" box="[178,193,455,476]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFF5AFE7EFEB737ED" box="[206,290,454,476]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
was defined by APFELBECK (1899, 1904) on the ground that it is distinct from both
<taxonomicName id="4C214D586E627F2AFF72FE5CFE6D37C8" ID-CoL="4Q3Q4" authority="(CREUTZER 1799)" baseAuthorityName="CREUTZER" baseAuthorityYear="1799" box="[230,504,483,505]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="melas">
<emphasis id="B955EAC96E627F2AFF72FE5CFF6037C8" box="[230,245,484,505]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFE96FE5BFEA837C8" box="[258,317,483,505]" italics="true" pageId="5" pageNumber="506">melas</emphasis>
(CREUTZER 1799)
</taxonomicName>
and
<taxonomicName id="4C214D586E627F2AFDBEFE5CFCCE37C8" ID-CoL="4Q3Q2" authority="(DEJEAN 1828)" baseAuthorityName="DEJEAN" baseAuthorityYear="1828" box="[554,859,483,505]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="hungaricus">
<emphasis id="B955EAC96E627F2AFDBEFE5CFDAC37C8" box="[554,569,484,505]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFDD2FE5BFD2D37C8" box="[582,696,483,505]" italics="true" pageId="5" pageNumber="506">hungaricus</emphasis>
(DEJEAN 1828)
</taxonomicName>
. At the same time, the author did not compare the new form with
<taxonomicName id="4C214D586E627F2AFDF9FDB9FC9C3427" ID-CoL="4Q3Q3" authorityName="SCHAUM" authorityYear="1858" box="[621,777,512,534]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFDF9FDB9FDEE3427" box="[621,635,513,534]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFD1DFDB8FC9C3427" box="[649,777,512,534]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
. Characters for the separation of
<taxonomicName id="4C214D586E627F2AFF45FDA6FED13405" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[209,324,542,564]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFF45FDA6FF753402" box="[209,224,542,563]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFF64FDA6FED13405" box="[240,324,542,564]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
from the other two species were size, shape and proportion of body, as well as pronotum with characteristic shape and presence of shallow and short basal impressions. So far, records for this species were noted only for
<collectingCountry id="F336764B6E627F2AFCA9FDE0FCEB345F" box="[829,894,600,622]" name="Serbia" pageId="5" pageNumber="506">Serbia</collectingCountry>
(data summarized by
<bibRefCitation id="EFB04B2A6E627F2AFF41FDCDFE4634BA" author="CURCIC S &amp; BRAJKOVIC M" box="[213,467,629,651]" pageId="5" pageNumber="506" refId="ref25258" refString="CURCIC S. B., BRAJKOVIC M. M. &amp; B. P. M CURCIC (2007): The Carabids of Serbia. - Institute of Zoology, Monographs, Vol. XI, Institute of Zoology - Faculty of Biology - University of Belgrade, Committee for Karst and Speleology - Serbian Academy of Sciences and Arts in Belgrade, Department of Conservation Biology, Vegetation- and Landscape Ecology, Faculty of Life Sciences, University of Vienna, and UNESCO MAB Committee Serbia in Belgrade, Belgrade-Vienna, 1083 pp., 681 maps." type="book" year="2007">ĆURČIĆ et al. 2007: 249</bibRefCitation>
, 755) and
<collectingCountry id="F336764B6E627F2AFDD2FDCDFD0834BA" box="[582,669,629,651]" name="Bulgaria" pageId="5" pageNumber="506">Bulgaria</collectingCountry>
(GUÉORGUIEV &amp; MUILWYJK 2000). On the other hand, faunistic data for
<taxonomicName id="4C214D586E627F2AFD97FD2CFD0A3498" ID-CoL="4Q3Q3" authorityName="SCHAUM" authorityYear="1858" box="[515,671,659,681]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFD97FD2CFD873498" box="[515,530,660,681]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFD8BFD2BFD0A3498" box="[543,671,659,681]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
from the Balkans were cited for
<collectingCountry id="F336764B6E627F2AFC7AFD2BFBA83499" box="[1006,1085,659,680]" name="Kosovo" pageId="5" pageNumber="506">Kosovo</collectingCountry>
(
<bibRefCitation id="EFB04B2A6E627F2AFF2DFD08FEF034F7" author="CSIKI E" box="[185,357,688,710]" pageId="5" pageNumber="506" pagination="208 - 288" refId="ref25219" refString="CSIKI E. (1940): XVI. Coleopteren, pp. 208 - 288. - In: Explorationes zoologicae ab E. Csiki in Albania peractae. A Magyar Tudomanyos Akademia Balkan-kutatasaiak Tudomanyos Eredmenyei 1940." type="journal article" year="1940">CSIKI 1940: 216</bibRefCitation>
;
<bibRefCitation id="EFB04B2A6E627F2AFEE7FD08FDF934F7" author="CURCIC S &amp; BRAJKOVIC M" box="[371,620,688,710]" pageId="5" pageNumber="506" refId="ref25258" refString="CURCIC S. B., BRAJKOVIC M. M. &amp; B. P. M CURCIC (2007): The Carabids of Serbia. - Institute of Zoology, Monographs, Vol. XI, Institute of Zoology - Faculty of Biology - University of Belgrade, Committee for Karst and Speleology - Serbian Academy of Sciences and Arts in Belgrade, Department of Conservation Biology, Vegetation- and Landscape Ecology, Faculty of Life Sciences, University of Vienna, and UNESCO MAB Committee Serbia in Belgrade, Belgrade-Vienna, 1083 pp., 681 maps." type="book" year="2007">ĆURČIĆ et al. 2007: 247</bibRefCitation>
, 751),
<collectingCountry id="F336764B6E627F2AFD22FD08FC9834F7" box="[694,781,688,710]" name="Bulgaria" pageId="5" pageNumber="506">Bulgaria</collectingCountry>
(HIEKE &amp; WRASE 1988) and
<collectingCountry id="F336764B6E627F2AFF26FD75FF6634D2" box="[178,243,717,739]" name="Serbia" pageId="5" pageNumber="506">Serbia</collectingCountry>
(
<bibRefCitation id="EFB04B2A6E627F2AFE94FD75FE5034D2" author="GUEORGUIEV B" box="[256,453,717,739]" pageId="5" pageNumber="506" pagination="73 - 92" refId="ref25667" refString="GUEORGUIEV B. (2008): New data on the ground beetles (Coleoptera: Carabidae) of Serbia. - Historia naturalis bulgarica 19: 73 - 92." type="journal article" year="2008">GUÉORGUIEV 2008</bibRefCitation>
).
</paragraph>
<paragraph id="8B9E36DB6E627F2AFF26FD4BFD54335E" blockId="5.[178,1086,755,1391]" pageId="5" pageNumber="506">
However, the study of material originally identified as
<taxonomicName id="4C214D586E627F2AFD79FD4BFC1C3538" ID-CoL="4Q3Q3" authorityName="SCHAUM" authorityYear="1858" box="[749,905,755,777]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFD79FD4BFD6E3539" box="[749,763,755,776]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFC9DFD4BFC1C3538" box="[777,905,755,777]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C214D586E627F2AFC3BFD4BFBB53538" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[943,1056,755,777]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFC3BFD4BFC283539" box="[943,957,755,776]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFC58FD4BFBB53538" box="[972,1056,755,777]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
as well as undetermined material established suspicion about the identity of the latter one. Study of the genitalia of male
<typeStatus id="549A88796E627F2AFE62FC96FDD03572" box="[502,581,814,835]" pageId="5" pageNumber="506" type="syntype">syntype</typeStatus>
of
<taxonomicName id="4C214D586E627F2AFDF8FC96FD753572" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[620,736,813,835]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFDF8FC96FDEE3572" box="[620,635,814,835]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFD18FC95FD753572" box="[652,736,813,835]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
and genitalia of specimens of
<taxonomicName id="4C214D586E627F2AFBB3FC96FEA73551" ID-CoL="4Q3Q3" authorityName="SCHAUM" authorityYear="1858" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFBB3FC96FBA33572" box="[1063,1078,814,835]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFF26FCF2FEA73551" box="[178,306,842,864]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
from
<collectingCountry id="F336764B6E627F2AFEE7FCF2FE5B3551" box="[371,462,842,864]" name="Slovakia" pageId="5" pageNumber="506">Slovakia</collectingCountry>
,
<collectingCountry id="F336764B6E627F2AFE4DFCF3FDA13551" box="[473,564,843,864]" name="Hungary" pageId="5" pageNumber="506">Hungary</collectingCountry>
,
<collectingCountry id="F336764B6E627F2AFDD4FCF2FD1F3551" box="[576,650,842,864]" name="Croatia" pageId="5" pageNumber="506">Croatia</collectingCountry>
and
<collectingCountry id="F336764B6E627F2AFD2BFCF2FC833551" box="[703,790,842,864]" name="Bulgaria" pageId="5" pageNumber="506">Bulgaria</collectingCountry>
demonstrated that the structure of the aedeagus and shape of the left parameres in all samples are identical and that both taxa in question represent single species. Main character that was used to distinguish these forms i.e. number of basal impressions of the pronotum (
<bibRefCitation id="EFB04B2A6E627F2AFCFAFC1AFBA33589" author="SCHATZMAYR A" box="[878,1078,930,952]" pageId="5" pageNumber="506" pagination="1 - 80" refId="ref27302" refString="SCHATZMAYR A. (1942): Bestimmungs-Tabellen europaischer Kafer. II. Fam. Carabidae. Subfam. Pterostichinae. 65. Gattungen: Pterostichus BON. u. Tapinopterus SCHAUM. - Koleopterologische Rundschau 27: 1 - 80." type="journal article" year="1942">SCHATZMAYR 1942</bibRefCitation>
) is irrelevant. For example, two of the
<specimenCount id="9D27FD526E627F2AFDD6FC07FD7235E4" box="[578,743,959,981]" pageId="5" pageNumber="506" type="generic">three specimens</specimenCount>
from Dragomansko blato Marsh have clear, moderately impressed outer impression. However, the third specimen has no outer impression. Both
<typeStatus id="549A88796E627F2AFE3DFC43FD973221" box="[425,514,1019,1040]" pageId="5" pageNumber="506" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C214D586E627F2AFDBEFC43FD083221" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[554,669,1018,1040]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFDBEFC43FDAC3221" box="[554,569,1019,1040]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFDDDFC42FD083221" box="[585,669,1018,1040]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
have distinct inner basal impression as well as shallow but distinct outer basal one at each side of pronotum. Variation
<emphasis id="B955EAC96E627F2AFF26FB8DFEA1327A" box="[178,308,1077,1099]" italics="true" pageId="5" pageNumber="506">karageorgeii</emphasis>
(LUTSCHNIK), replacement name for variation
<emphasis id="B955EAC96E627F2AFC88FB8DFC1D327B" box="[796,904,1077,1098]" italics="true" pageId="5" pageNumber="506">unistriatus</emphasis>
APFELBECK, was defined for a local form of
<taxonomicName id="4C214D586E627F2AFE76FBEBFDC23259" ID-CoL="4Q629" authorityName="APFELBECK" authorityYear="1899" box="[482,599,1106,1128]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="serbicus">
<emphasis id="B955EAC96E627F2AFE76FBEBFE643259" box="[482,497,1107,1128]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFD97FBEAFDC23259" box="[515,599,1106,1128]" italics="true" pageId="5" pageNumber="506">serbicus</emphasis>
</taxonomicName>
without outer basal impressions (
<bibRefCitation id="EFB04B2A6E627F2AFC52FBEAFF7C32B4" author="APFELBECK V" pageId="5" pageNumber="506" refId="ref24610" refString="APFELBECK V. (1904): Die Kaferfauna der Balkanhalbinsel, mit Berucksichtigung Klein- Asiens und der Insel Kreta. Erster Band: Familienreihe Caraboidea. - Berlin: R. Friedlander und Sohn, ix + 422 pp." type="book" year="1904">APFELBECK 1904</bibRefCitation>
). This form was described from Novi Pazar, a place situated very close to the first known citation of
<taxonomicName id="4C214D586E627F2AFEE5FB35FD9B3293" ID-CoL="4Q3Q3" authorityName="SCHAUM" authorityYear="1858" box="[369,526,1164,1186]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFEE5FB35FEEA3293" box="[369,383,1165,1186]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFE1AFB34FD9B3293" box="[398,526,1164,1186]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
from the Balkan Peninsula (
<bibRefCitation id="EFB04B2A6E627F2AFCA3FB34FC713293" author="CSIKI E" box="[823,996,1164,1186]" pageId="5" pageNumber="506" pagination="208 - 288" refId="ref25219" refString="CSIKI E. (1940): XVI. Coleopteren, pp. 208 - 288. - In: Explorationes zoologicae ab E. Csiki in Albania peractae. A Magyar Tudomanyos Akademia Balkan-kutatasaiak Tudomanyos Eredmenyei 1940." type="journal article" year="1940">CSIKI 1940: 216</bibRefCitation>
). On the other hand,
<bibRefCitation id="EFB04B2A6E627F2AFEBAFB12FE6F32F1" author="HURKA K" box="[302,506,1194,1216]" pageId="5" pageNumber="506" refId="ref26042" refString="HURKA K. (1996): Carabidae of the Czech and Slovak Republics. - Kabourek, Zlin, 565 pp." type="book" year="1996">HŮRKA (1996: 284)</bibRefCitation>
has noted that sometimes specimens from Moravia and
<collectingCountry id="F336764B6E627F2AFF26FB7FFE9F32EC" box="[178,266,1223,1245]" name="Slovakia" pageId="5" pageNumber="506">Slovakia</collectingCountry>
have &quot;vague and very shallow&quot; second basal impression at each side of the pronotum. Besides that the number of the basal impression and the expression of outer ones vary, variability exist too in the punctuation of the inner impressions. Such penotypic variances have intra-specific character that not allows formation of geographical races or vicariant species. This conclusion is supported from lack of any appreciable differences in the morphology of the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C33B65506E627F2AFF26FA38FC5C33FE" pageId="5" pageNumber="506" type="description">
<paragraph id="8B9E36DB6E627F2AFF26FA38FC5C33FE" blockId="5.[178,1085,1407,1487]" pageId="5" pageNumber="506">
<taxonomicName id="4C214D586E627F2AFF26FA38FEC133A4" authorityName="SCHAUM" authorityYear="1858" box="[178,340,1407,1429]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFF26FA38FF5433A4" box="[178,193,1408,1429]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFF40FAC7FEC133A4" box="[212,340,1407,1429]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
is a rare species with Balkan - Pannonian distribution which in the Balkan Peninsula occurs mainly at lowlands in the basins of the big rivers of Danube, Sava and Morava. First records of
<taxonomicName id="4C214D586E627F2AFD85FA02FD3833FE" authorityName="SCHAUM" authorityYear="1858" box="[529,685,1465,1487]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="incommodus">
<emphasis id="B955EAC96E627F2AFD85FA02FDB533FE" box="[529,544,1466,1487]" italics="true" pageId="5" pageNumber="506">P</emphasis>
.
<emphasis id="B955EAC96E627F2AFDB9FA01FD3833FE" box="[557,685,1465,1487]" italics="true" pageId="5" pageNumber="506">incommodus</emphasis>
</taxonomicName>
from
<collectingCountry id="F336764B6E627F2AFD7FFA01FCA033FE" box="[747,821,1465,1487]" name="Croatia" pageId="5" pageNumber="506">Croatia</collectingCountry>
and
<collectingCountry id="F336764B6E627F2AFCF2FA01FC5133FE" box="[870,964,1465,1487]" name="Romania" pageId="5" pageNumber="506">Romania</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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