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<document id="D9F23A52D3910F2E9A175635633C196F" ID-CLB-Dataset="7024" ID-DOI="10.1080/00222933.2013.791950" ID-GBIF-Dataset="8b15385e-73ff-4e17-aa18-4ab5bd25713b" ID-ISSN="1464-5262" ID-Zenodo-Dep="4632183" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616525030300" checkinUser="carolina" docAuthor="Mašán, Peter &amp; Halliday, Bruce" docDate="2013" docId="038687DAD512002FFDB17C1CFB107E06" docLanguage="en" docName="JNATHIST.48.7-8.403-412.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 48 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2013.791950" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Chaetodellus Mašán &amp; Halliday 2013, gen. nov." docType="treatment" docUuid="A8AF4FCD-F56A-4B4D-B5C5-B1ABD18CD965" docUuidSource="ZooBank" docVersion="8" lastPageNumber="409" masterDocId="FFBFFFA2D5160027FF867D3AFF837F59" masterDocTitle="A new genus and species of soil mite (Acari: Pachylaelapidae: Pachylaelapinae) from the Canary Islands" masterLastPageNumber="412" masterPageNumber="403" pageNumber="405" updateTime="1698934716162" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D2D15BDFABE259D0D6CED9503CDE26D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1D57532FD991C5A9D23560E86FB0BC2C">
<mods:title id="7AAE5A1A0B18846CE11121900E6EA69E">A new genus and species of soil mite (Acari: Pachylaelapidae: Pachylaelapinae) from the Canary Islands</mods:title>
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<mods:namePart id="63B1844B2B208BA99729E60E67F5CCDC">Mašán, Peter</mods:namePart>
<mods:affiliation id="AB389115D924EAD99D4DC108D9986AB7">Institute of Zoology, Slovak Academy of Sciences, Bratislava, Slovakia;</mods:affiliation>
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<mods:namePart id="E0EAE4505D3854C9D49FDF088D2BD1F3">Halliday, Bruce</mods:namePart>
<mods:affiliation id="ED17D0627C2CA6BA5B85C02314E9400E">CSIRO Ecosystem Sciences, Canberra, Australia</mods:affiliation>
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<mods:title id="A70235C7256D39CD26F3C04D315F7745">Journal of Natural History</mods:title>
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<mods:part id="F656E8FDD053FC7F7D635B80C7AEDE5F">
<mods:date id="F7D76FE88D537FABDDC3275408AC600E">2013</mods:date>
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<mods:title id="712F2EAD74922E1EF2748F3E02D3A52A">J. Nat. Hist.</mods:title>
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<mods:number id="AAC4838C970DEAA79183452E246A79C8">2013-12-18</mods:number>
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<mods:number id="D662166D63CED54C5663FBC4CBFB4757">48</mods:number>
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<mods:number id="F620F9CF439808CD0EA5C18E5B196AF9">7 - 8</mods:number>
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<mods:classification id="91200BE7A035899A78692F835B7FC680">journal article</mods:classification>
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<treatment id="038687DAD512002FFDB17C1CFB107E06" ID-DOI="http://doi.org/10.5281/zenodo.4748154" ID-GBIF-Taxon="181972648" ID-Zenodo-Dep="4748154" LSID="urn:lsid:zoobank.org:act:A8AF4FCD-F56A-4B4D-B5C5-B1ABD18CD965" httpUri="http://treatment.plazi.org/id/038687DAD512002FFDB17C1CFB107E06" lastPageId="8" lastPageNumber="409" pageId="4" pageNumber="405">
<subSubSection id="C3356547D5120023FDB17C1CFCB37E66" box="[567,816,294,320]" pageId="4" pageNumber="405" type="nomenclature">
<paragraph id="8B9036CCD5120023FDB17C1CFCB37E66" blockId="4.[567,816,294,351]" box="[567,816,294,320]" pageId="4" pageNumber="405">
<heading id="D0D881A0D5120023FDB17C1CFCB37E66" box="[567,816,294,320]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="405" reason="2">
<taxonomicName id="4C2F4D4FD5120023FDB17C1CFD467E66" ID-CoL="8XKCQ" authority="Mašán &amp; Halliday, 2013" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[567,709,294,319]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="405" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95BEADED5120023FDB17C1CFD467E66" bold="true" box="[567,709,294,319]" italics="true" pageId="4" pageNumber="405">Chaetodellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857A5D5120023FD4A7C12FCB37E66" box="[716,816,296,320]" pageId="4" pageNumber="405" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3356547D5120023FDFE7C7DFD6B7E06" box="[632,744,327,351]" pageId="4" pageNumber="405" type="description">
<paragraph id="8B9036CCD5120023FDFE7C7DFD6B7E06" blockId="4.[567,816,294,351]" box="[632,744,327,351]" pageId="4" pageNumber="405">
(
<figureCitation id="13142A49D5120023FD077C7DFD5C7E06" box="[641,735,327,351]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3356547D5120023FF557CBDFD337EC7" box="[211,688,391,415]" pageId="4" pageNumber="405" type="type_taxon">
<paragraph id="8B9036CCD5120023FF557CBDFD337EC7" blockId="4.[211,688,391,415]" box="[211,688,391,415]" pageId="4" pageNumber="405">
<heading id="D0D881A0D5120023FF557CBDFD337EC7" box="[211,688,391,415]" fontSize="10" level="3" pageId="4" pageNumber="405" reason="5">
<typeStatus id="5494886ED5120023FF557CBDFE897EC6" box="[211,266,391,415]" pageId="4" pageNumber="405">Type</typeStatus>
species
<taxonomicName id="4C2F4D4FD5120023FEE37CBDFDD67EC6" ID-CoL="8XX6R" authority="Mašán &amp; Halliday, 2013" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[357,597,391,415]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="comatus" status="sp. nov.">
<emphasis id="B95BEADED5120023FEE37CBDFDD67EC6" box="[357,597,391,415]" italics="true" pageId="4" pageNumber="405">Chaetodellus comatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857A5D5120023FDDA7CBDFD337EC7" box="[604,688,391,414]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3356547D5120023FF557CDEFD6E7A19" pageId="4" pageNumber="405" type="diagnosis">
<paragraph id="8B9036CCD5120023FF557CDEFE2A7EA2" blockId="4.[211,1172,483,1344]" box="[211,425,483,507]" pageId="4" pageNumber="405">
<heading id="D0D881A0D5120023FF557CDEFE2A7EA2" box="[211,425,483,507]" fontSize="10" level="3" pageId="4" pageNumber="405" reason="5">
<emphasis id="B95BEADED5120023FF557CDEFE2A7EA2" box="[211,425,483,507]" italics="true" pageId="4" pageNumber="405">Diagnosis (female)</emphasis>
</heading>
</paragraph>
<paragraph id="8B9036CCD5120023FF557F31FEAF7D9B" blockId="4.[211,1172,483,1344]" pageId="4" pageNumber="405">
<emphasis id="B95BEADED5120023FF557F31FE0E7D7A" box="[211,397,523,547]" italics="true" pageId="4" pageNumber="405">Dorsal idiosoma</emphasis>
(
<figureCitation id="13142A49D5120023FE227F31FDCF7D7A" box="[420,588,523,547]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1A, G</figureCitation>
). Dorsal shield oval-shaped, completely covering idiosoma, margins encroaching slightly onto ventral idiosoma; with 29 pairs of smooth pointed setae; setae strongly differentiated in length, j1 and J5 minute; J1 widely separated, markedly anterolateral to J2; z6 markedly anterolateral to j6. Shield with approximately 12 pairs of pore-like structures, some of them enlarged, suboval to slit-like.
</paragraph>
<paragraph id="8B9036CCD5120023FF787FF0FE547CB8" blockId="4.[211,1172,483,1344]" pageId="4" pageNumber="405">
<emphasis id="B95BEADED5120023FF787FF0FE457DBB" box="[254,454,714,738]" italics="true" pageId="4" pageNumber="405">Ventral idiosoma</emphasis>
(
<figureCitation id="13142A49D5120023FE607FF0FDE67DBB" box="[486,613,714,738]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1B</figureCitation>
). Presternal shields absent. Peritremes and peritrematal shields well-developed, peritrematal shields fused to exopodal shields, produced posteriorly well behind coxae IV. Genitoventral shield relatively large, trapezoidal, with straight posterior margin and three pairs of setae (st5, Jv1 and Zv2). Metapodal shields absent or fused with peritrematal–exopodal–parapodal complex (but this complex with no outlines of the fusion). Anal shield free, wide, subtriangular, bearing three short setae (one pair of paranals and one postanal). All ventral idiosomal setae smooth and pointed. Unsclerotized integument with five pairs of setae (r6, Jv2−Jv4 and Zv3).
</paragraph>
<paragraph id="8B9036CCD5120023FF787ED3FD307B18" blockId="4.[211,1172,483,1344]" pageId="4" pageNumber="405">
<emphasis id="B95BEADED5120023FF787ED3FD887B58" box="[254,523,1001,1025]" italics="true" pageId="4" pageNumber="405">Spermathecal apparatus</emphasis>
(
<figureCitation id="13142A49D5120023FD987ED3FD127B58" box="[542,657,1001,1025]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1F</figureCitation>
). Sperm access system with tubular components opening onto coxae III; tubular components long, fine for whole length, slightly and regularly broadened towards the coxa.
</paragraph>
<paragraph id="8B9036CCD5120023FF787970FE6A7B99" blockId="4.[211,1172,483,1344]" pageId="4" pageNumber="405">
<emphasis id="B95BEADED5120023FF787970FEB97B38" box="[254,314,1098,1121]" italics="true" pageId="4" pageNumber="405">Legs.</emphasis>
Legs normal in length, shorter than idiosoma. Tarsus II with all distal setae fine and pointed, not modified into spurs (
<figureCitation id="13142A49D5120023FD457953FCB57BD8" box="[707,822,1129,1153]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1E</figureCitation>
). Chaetotaxy I-II-III-IV: coxa 2-2-2-1, trochanter 6-5-5-5, femur 13-11-6-6, genu 12 (2-5
<emphasis id="B95BEADED5120023FCD579B3FCDC7BF8" bold="true" box="[851,863,1161,1185]" italics="true" pageId="4" pageNumber="405">/</emphasis>
3-2)-11-7-7, tibia 12-10-7-7; tarsi II–IV with 18 setae.
</paragraph>
<paragraph id="8B9036CCD5120023FF7879F3FD6E7A19" blockId="4.[211,1172,483,1344]" pageId="4" pageNumber="405">
<emphasis id="B95BEADED5120023FF7879F3FE137BB8" box="[254,400,1225,1249]" italics="true" pageId="4" pageNumber="405">
<taxonomicName id="4C2F4D4FD5120023FF7879F3FE0F7BB8" authorityName="Krefft" authorityYear="1871" box="[254,396,1225,1249]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="405" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Corniculi short, horn-like. Palp segments without projections. Epistome with wide base and medial projection on distal margin; lateral margins of base rounded and denticulate, central projection with short narrow neck and furcate apex with large and small denticles (
<figureCitation id="13142A49D5120023FDEF7812FD5E7A19" box="[617,733,1320,1344]" captionStart="Figure 1" captionStartId="5.[143,210,1594,1616]" captionTargetBox="[144,1104,161,1561]" captionTargetId="figure-6@5.[144,1104,161,1561]" captionTargetPageId="5" captionText="Figure 1. Chaetodellus spp., females. (A–F). Chaetodellus comatus sp. nov. (A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G) Chaetodellus meganalis, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E)." figureDoi="http://doi.org/10.5281/zenodo.4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="4" pageNumber="405">Figure 1D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3356547D5120021FF5578BFFC107B00" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405" type="etymology">
<paragraph id="8B9036CCD5120023FF5578BFFEC87AC4" blockId="4.[211,1172,1413,1540]" box="[211,331,1413,1437]" pageId="4" pageNumber="405">
<heading id="D0D881A0D5120023FF5578BFFEC87AC4" box="[211,331,1413,1437]" fontSize="10" level="3" pageId="4" pageNumber="405" reason="5">
<emphasis id="B95BEADED5120023FF5578BFFEC87AC4" box="[211,331,1413,1437]" italics="true" pageId="4" pageNumber="405">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9036CCD5120023FF557896FCC3795D" blockId="4.[211,1172,1413,1540]" pageId="4" pageNumber="405">
The name of the new genus is derived from the Greek
<emphasis id="B95BEADED5120023FCCC7897FC127A9C" box="[842,913,1453,1477]" italics="true" pageId="4" pageNumber="405">chaeto</emphasis>
-, referring to the long dorsal shield setae, in combination with the diminutive suffix -
<emphasis id="B95BEADED5120023FC2B78F7FC5C7ABC" box="[941,991,1485,1509]" italics="true" pageId="4" pageNumber="405">ellus</emphasis>
, as used in the names of other pachylaelapine taxa. Gender masculine.
</paragraph>
<paragraph id="8B9036CCD5120023FF557B70FE227938" blockId="4.[211,1171,1609,1704]" box="[211,417,1609,1633]" pageId="4" pageNumber="405">
<heading id="D0D881A0D5120023FF557B70FE227938" box="[211,417,1609,1633]" fontSize="10" level="3" pageId="4" pageNumber="405" reason="5">
<emphasis id="B95BEADED5120023FF557B70FE227938" box="[211,417,1609,1633]" italics="true" pageId="4" pageNumber="405">Notes on the genus</emphasis>
</heading>
</paragraph>
<paragraph id="8B9036CCD5120021FF557B4AFD297DA4" blockId="4.[211,1171,1609,1704]" lastBlockId="6.[211,1172,167,1020]" lastPageId="6" lastPageNumber="407" pageId="4" pageNumber="405">
The new genus is diagnosed by the following combination of characters: (1) dorsal shield expanded to encroach slightly onto the ventral surface of the idiosoma; (2) dorsal shield with 29 pairs of setae, which vary widely in length; (3) vertical setae j1 minute; (4) clunal setae J5 minute; (5) dorsocentral setae J
<quantity id="4CD79B29D5100021FC4C7DFDFC707F86" box="[970,1011,199,223]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="407" unit="in" value="1.0">1 in</quantity>
unusual position markedly anterolateral to J2; (6) dorsal setae z
<quantity id="4CD79B29D5100021FCAC7DDDFCD47FA6" box="[810,855,231,255]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="6" pageNumber="407" unit="in" value="6.0">6 in</quantity>
unusual position markedly anterolateral to j6; (7) strongly reduced number of setae on unsclerotized ventral integument, R setae absent; (8) genitoventral shield with three pairs of setae including Zv2; (9) tarsus II with uniform fine, pointed setae, without modified spur-like setae; (10) separate metapodal shields absent; and (11) sperm access system with long spermathecal tubes opened onto coxae III. Some of these character states occur in other genera of
<taxonomicName id="4C2F4D4FD5100021FE0E7C9CFDBD7EE7" authorityName="Berlese" authorityYear="1913" box="[392,574,422,446]" pageId="6" pageNumber="404" rank="subFamily" subFamily="Pachylaelapinae">Pachylaelapinae</taxonomicName>
, but infrequently, and not in combination (especially the combination of characters in points 4 and 8 is unique among the all known pachylaelapid genera). For example
<emphasis id="B95BEADED5100021FD0F7CDDFCC17EA6" box="[649,834,487,511]" italics="true" pageId="6" pageNumber="407">Neopachylaelaps</emphasis>
and
<taxonomicName id="4C2F4D4FD5100021FC0F7CDDFBD17EA6" authorityName="Masan" authorityYear="2007" box="[905,1106,487,511]" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FC0F7CDDFBD17EA6" box="[905,1106,487,511]" italics="true" pageId="6" pageNumber="407">Pachyglobolaelaps</emphasis>
</taxonomicName>
have 29 pairs of dorsal shield setae; seta J1 is anterolateral to J
<quantity id="4CD79B29D5100021FCE27F3CFC137D47" box="[868,912,518,542]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="407" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C2F4D4FD5100021FC1E7F3CFBE27D47" authorityName="Masan" authorityYear="2007" box="[920,1121,518,542]" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FC1E7F3CFBE27D47" box="[920,1121,518,542]" italics="true" pageId="6" pageNumber="407">Pachyglobolaelaps</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F4D4FD5100021FF557F1CFEFA7D67" authorityName="Berlese" authorityYear="1903" box="[211,377,550,574]" class="Arachnida" family="Pachylaelapidae" genus="Sphaerolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FF557F1CFEFA7D67" box="[211,377,550,574]" italics="true" pageId="6" pageNumber="407">Sphaerolaelaps</emphasis>
</taxonomicName>
; spur-like distal setae are also absent from tarsi II in
<taxonomicName id="4C2F4D4FD5100021FC6E7F1CFE367D07" authority=", Pachysphaerolaelaps" authorityName="Pachysphaerolaelaps" class="Arachnida" family="Pachylaelapidae" genus="Elaphrolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FC6E7F1CFB0F7D67" box="[1000,1164,550,574]" italics="true" pageId="6" pageNumber="407">Elaphrolaelaps</emphasis>
,
<emphasis id="B95BEADED5100021FF557F7CFE367D07" box="[211,437,582,606]" italics="true" pageId="6" pageNumber="407">Pachysphaerolaelaps</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D4FD5100021FE457F7CFDC07D07" authorityName="Tragardh" authorityYear="1908" box="[451,579,582,606]" class="Arachnida" family="Pachylaelapidae" genus="Paralaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FE457F7CFDC07D07" box="[451,579,582,606]" italics="true" pageId="6" pageNumber="407">Paralaelaps</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2F4D4FD5100021FDFF7F7CFC9C7D07" authorityName="Berlese" authorityYear="1903" box="[633,799,582,606]" class="Arachnida" family="Pachylaelapidae" genus="Sphaerolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FDFF7F7CFC9C7D07" box="[633,799,582,606]" italics="true" pageId="6" pageNumber="407">Sphaerolaelaps</emphasis>
</taxonomicName>
; dorsal shield setae J5 are minute in
<emphasis id="B95BEADED5100021FF777F5CFE297D27" box="[241,426,614,638]" italics="true" pageId="6" pageNumber="407">Neopachylaelaps</emphasis>
,
<taxonomicName id="4C2F4D4FD5100021FE3E7F5CFDA97D27" authorityName="Berlese" authorityYear="1910" box="[440,554,614,638]" class="Arachnida" family="Pachylaelapidae" genus="Olopachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FE3E7F5CFDA97D27" box="[440,554,614,638]" italics="true" pageId="6" pageNumber="407">Olopachys</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D4FD5100021FDBC7F5CFD397D27" authorityName="Masan" authorityYear="2007" box="[570,698,614,638]" class="Arachnida" family="Pachylaelapidae" genus="Pachydellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FDBC7F5CFD397D27" box="[570,698,614,638]" italics="true" pageId="6" pageNumber="407">Pachydellus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2F4D4FD5100021FD4F7F5CFC117D27" authorityName="Masan" authorityYear="2007" box="[713,914,614,638]" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FD4F7F5CFC117D27" box="[713,914,614,638]" italics="true" pageId="6" pageNumber="407">Pachyglobolaelaps</emphasis>
</taxonomicName>
and
<emphasis id="B95BEADED5100021FC4D7F5CFBE37D27" box="[971,1120,614,638]" italics="true" pageId="6" pageNumber="407">
<taxonomicName id="4C2F4D4FD5100021FC4D7F5CFBDF7D27" authorityName="Berlese" authorityYear="1888" box="[971,1116,614,638]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">Pachylaelaps</taxonomicName>
.
</emphasis>
The combination of a tubular sperm access system and minute setae J5 is found in the genus
<taxonomicName id="4C2F4D4FD5100021FE9A7F9CFE667DE7" authorityName="Masan" authorityYear="2007" box="[284,485,678,702]" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FE9A7F9CFE667DE7" box="[284,485,678,702]" italics="true" pageId="6" pageNumber="407">Pachyglobolaelaps</emphasis>
</taxonomicName>
, which also has dorsal setae of widely varying lengths. The new genus may be easily distinguished from the other related genera by the characters introduced in the identification key below.
</paragraph>
<caption id="DF506644D5130022FF097B00FC1379F1" ID-DOI="http://doi.org/10.5281/zenodo.4632185" ID-Zenodo-Dep="4632185" httpUri="https://zenodo.org/record/4632185/files/figure.png" pageId="5" pageNumber="406" startId="5.[143,210,1594,1616]" targetBox="[144,1104,161,1561]" targetPageId="5">
<paragraph id="8B9036CCD5130022FF097B00FC1379F1" blockId="5.[143,1103,1594,1704]" pageId="5" pageNumber="406">
Figure 1.
<taxonomicName id="4C2F4D4FD5130022FF7F7B01FEFA7909" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[249,377,1595,1616]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="406" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5130022FF7F7B01FEFA7909" box="[249,377,1595,1616]" italics="true" pageId="5" pageNumber="406">Chaetodellus</emphasis>
</taxonomicName>
spp., females. (A–F).
<taxonomicName id="4C2F4D4FD5130022FDD87B01FCBB7909" authority="Mašán &amp; Halliday, 2013" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[606,824,1595,1616]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="comatus" status="sp. nov.">
<emphasis id="B95BEADED5130022FDD87B01FCBB7909" box="[606,824,1595,1616]" italics="true" pageId="5" pageNumber="406">Chaetodellus comatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857A5D5130022FCC67B01FC0D7909" box="[832,910,1595,1616]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
(A) Dorsal shield; (B) ventral idiosoma; (C) sternal shield, anterior portion; (D) epistome; (E) tarsus II, distal section; (F) tubes of sperm access system opened onto coxae III. (G)
<taxonomicName id="4C2F4D4FD5130022FCD97B4FFBCA79D3" authorityName="Mašán &amp; Halliday" authorityYear="2013" baseAuthorityName="Halliday" baseAuthorityYear="2005" box="[863,1097,1653,1674]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="meganalis">
<emphasis id="B95BEADED5130022FCD97B4FFBCA79D3" box="[863,1097,1653,1674]" italics="true" pageId="5" pageNumber="406">Chaetodellus meganalis</emphasis>
</taxonomicName>
, dorsal shield. Scale bars: 100 µm (A, B, F, G), 50 µm (C, D) and 25 µm (E).
</paragraph>
</caption>
<paragraph id="8B9036CCD5100021FF787E3CFD407CA5" blockId="6.[211,1172,167,1020]" pageId="6" pageNumber="407">
<taxonomicName id="4C2F4D4FD5100021FF787E3CFCDD7C44" authority="Samsinak and Daniel, 1978" authorityName="Samsinak and Daniel" authorityYear="1978" box="[254,862,773,798]" class="Arachnida" family="Pachylaelapidae" genus="Elaphrolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="B95BEADED5100021FF787E3CFD9A7C47" box="[254,537,774,798]" italics="true" pageId="6" pageNumber="407">Elaphrolaelaps nepalensis</emphasis>
<bibRefCitation id="EFBE4B3DD5100021FDA57E3FFCDD7C44" author="Samsinak K &amp; Daniel M" box="[547,862,773,798]" pageId="6" pageNumber="407" pagination="91 - 100" refId="ref6130" refString="Samsinak K, Daniel M. 1978. Five new species of mesostigmatic mites from Geotrupes beetles collected in the Himalayan Region of Nepal (Acari: Mesostigmata). Int J Acarol. 4: 91 - 100." type="journal article" year="1978">Samšiňák and Daniel, 1978</bibRefCitation>
</taxonomicName>
has some features in common with the new genus – 29 pairs of dorsal shield setae, dorsal shield setae strongly differentiated in length, setae j1 and J5 minute, dorsal shield expanded onto the ventral idiosoma, and tarsus II without spur-like setae. However, the ventral idiosoma of
<taxonomicName id="4C2F4D4FD5100021FF7E7EBCFE0E7CC4" authorityName="Samsinak and Daniel" authorityYear="1978" box="[248,397,901,925]" class="Arachnida" family="Pachylaelapidae" genus="Elaphrolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="B95BEADED5100021FF7E7EBCFE0E7CC4" box="[248,397,901,925]" italics="true" pageId="6" pageNumber="407">E. nepalensis</emphasis>
</taxonomicName>
differs markedly from that of
<taxonomicName id="4C2F4D4FD5100021FC807EBFFC177CC4" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[774,916,901,925]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FC807EBFFC177CC4" box="[774,916,901,925]" italics="true" pageId="6" pageNumber="407">Chaetodellus</emphasis>
</taxonomicName>
– Zv2 is not on the genitoventral shield, the unsclerotized ventral integument has eight or nine pairs of setae, and the peritrematal shields are strongly expanded posteriorly to reach the posterior margin of the genitoventral shield.
</paragraph>
<paragraph id="8B9036CCD5100021FF55797BFC107B00" blockId="6.[211,1171,1089,1217]" box="[211,915,1089,1113]" pageId="6" pageNumber="407">
<emphasis id="B95BEADED5100021FF55797BFC107B00" bold="true" box="[211,915,1089,1113]" pageId="6" pageNumber="407">
Key to world genera and subgenera of
<taxonomicName id="4C2F4D4FD5100021FDF2797BFCA47B00" authorityName="Berlese" authorityYear="1913" box="[628,807,1089,1113]" pageId="6" pageNumber="404" rank="subFamily" subFamily="Pachylaelapinae">Pachylaelapinae</taxonomicName>
(females)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3356547D510002FFE817953FB107E06" lastPageId="8" lastPageNumber="409" pageId="6" pageNumber="407" type="key">
<key id="FFBE61FDD510002FFE817953FB107E06" lastPageId="8" lastPageNumber="409" pageId="6" pageNumber="407">
<paragraph id="8B9036CCD5100021FE817953FB107B99" blockId="6.[211,1171,1089,1217]" pageId="6" pageNumber="407">
<keyLead id="30DE95F9D5100021FE817953FB107B99" pageId="6" pageNumber="407">
<keyStep id="30DB2E69D5100021FE817953FB107B99" pageId="6" pageNumber="407">1. Tarsus II without thick spur-like setae.................................... 2 Tarsus II with at least distal seta pl1 modified, thick and spur-like, pl2 may also be so modified...................................................... 7</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5100021FE8179E2FB107A69" blockId="6.[263,1171,1240,1328]" pageId="6" pageNumber="407">
<keyLead id="30DE95F9D5100021FE8179E2FB107A69" pageId="6" pageNumber="407">
<keyStep id="30DB2E69D5100021FE8179E2FB107A69" pageId="6" pageNumber="407">
2. Dorsal shield with about 80 pairs of setae...........................................................................
<emphasis id="B95BEADED5100021FC9C79C3FC7F7A48" box="[794,1020,1273,1297]" italics="true" pageId="6" pageNumber="407">Pachysphaerolaelaps</emphasis>
<bibRefCitation id="EFBE4B3DD5100021FB8579C2FB107A49" author="Masan P" box="[1027,1171,1272,1296]" pageId="6" pageNumber="407" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Dorsal shield with at most 37 pairs of setae............................... 3
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5100021FE817872FB107AF9" blockId="6.[263,1171,1352,1440]" pageId="6" pageNumber="407">
<keyLead id="30DE95F9D5100021FE817872FB107AF9" pageId="6" pageNumber="407">
<keyStep id="30DB2E69D5100021FE817872FB107AF9" pageId="6" pageNumber="407">
3. Some dorsal shield setae modified, elongated and thickened, with club-like tips..........................................
<taxonomicName id="4C2F4D4FD5100021FCD37852FED77AF9" ID-CoL="8XPYB" authority="Berlese, 1903 All" authorityName="Berlese" authorityYear="1903" class="Arachnida" family="Pachylaelapidae" genus="Sphaerolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FCD37852FC787AD9" box="[853,1019,1384,1408]" italics="true" pageId="6" pageNumber="407">Sphaerolaelaps</emphasis>
<bibRefCitation id="EFBE4B3DD5100021FB847852FB107AD9" author="Berlese A" box="[1026,1171,1384,1408]" pageId="6" pageNumber="407" pagination="235 - 252" refId="ref5739" refString="Berlese A. 1903. Acari nuovi. Manipulus I. Redia. 1: 235 - 252." type="journal article" year="1903">Berlese, 1903</bibRefCitation>
All
</taxonomicName>
dorsal shield setae uniform, smooth and pointed...................... 4
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5100021FE817882FB107956" blockId="6.[263,1171,1464,1552]" pageId="6" pageNumber="407">
<keyLead id="30DE95F9D5100021FE817882FB107956" pageId="6" pageNumber="407">
<keyStep id="30DB2E69D5100021FE817882FB107956" pageId="6" pageNumber="407">
4. Unsclerotized idiosomal integument strongly hypertrichous................................................................
<taxonomicName id="4C2F4D4FD5100021FCE778E2FE4E7956" ID-CoL="8XNLM" authority="Tragardh, 1908 Unsclerotized" authorityName="Tragardh" authorityYear="1908" class="Arachnida" family="Pachylaelapidae" genus="Paralaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5100021FCE778E2FC627AA9" box="[865,993,1496,1520]" italics="true" pageId="6" pageNumber="407">Paralaelaps</emphasis>
<bibRefCitation id="EFBE4B3DD5100021FC6E78EDFB107AB6" author="Tragardh I" box="[1000,1171,1495,1519]" pageId="6" pageNumber="407" pagination="31 - 57" refId="ref6170" refString="Tragardh I. 1908. Acari. In: Sjostedt Y, editor. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostfrikas 1905 - 1906, 3. Band, 20 (3): 31 - 57 + Plates 2 - 3." type="journal article" year="1908">Trägårdh, 1908</bibRefCitation>
Unsclerotized
</taxonomicName>
integument never hypertrichous, with 5−21 pairs of setae.... 5
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5100021FE817B1DFB107926" blockId="6.[263,1172,1575,1663]" pageId="6" pageNumber="407">
<keyLead id="30DE95F9D5100021FE817B1DFB107926" pageId="6" pageNumber="407">
<keyStep id="30DB2E69D5100021FE817B1DFB107926" pageId="6" pageNumber="407">
5. Genitoventral shield with three pairs of setae (st5, Jv1, Zv2)....................................................................
<taxonomicName id="4C2F4D4FD5100021FC1C7B7DFBAB7906" ID-CoL="8XKCQ" authority="Mašán &amp; Halliday, 2013" authorityName="Mašán &amp; Halliday" authorityYear="2013" box="[922,1064,1607,1631]" class="Arachnida" family="Pachylaelapidae" genus="Chaetodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="407" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B95BEADED5100021FC1C7B7DFBAB7906" box="[922,1064,1607,1631]" italics="true" pageId="6" pageNumber="407">Chaetodellus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26857A5D5100021FBA97B72FB177906" box="[1071,1172,1608,1632]" pageId="6" pageNumber="407" rank="genus">gen. nov.</taxonomicNameLabel>
Genitoventral shield with two pairs of setae (Zv2 off the shield)............ 6
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF457D9DFBCC7E46" blockId="7.[195,1103,167,287]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF457D9DFBCC7E46" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF457D9DFBCC7E46" pageId="7" pageNumber="408">
6. Sternal shield with deep incision in posterior margin, at least to the level between setae st1 and st2........
<taxonomicName id="4C2F4D4FD5110020FDF47DF2FEBD7FA6" ID-CoL="8XRM6" authority="Elsen, 1974 Sternal" authorityName="Elsen" authorityYear="1974" class="Arachnida" family="Pachylaelapidae" genus="Elaphrolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subGenus" subGenus="Incisosternum">
<emphasis id="B95BEADED5110020FDF47DF2FC957FB9" box="[626,790,200,224]" italics="true" pageId="7" pageNumber="408">Elaphrolaelaps</emphasis>
(
<emphasis id="B95BEADED5110020FCA37DF2FC3C7F86" box="[805,959,200,223]" italics="true" pageId="7" pageNumber="408">Incisosternum</emphasis>
)
<bibRefCitation id="EFBE4B3DD5110020FC497DFDFBCC7F86" author="Elsen P" box="[975,1103,199,223]" pageId="7" pageNumber="408" pagination="470 - 476" refId="ref5825" refString="Elsen P. 1974. Incisosternum nov. subgen., un nouveau sous-genre pour Elaphrolaelaps sternalis Ryke, 1959 et description d'une espece nouvelle de ce sous-genre (Acarina - Pachylaelapidae). Rev Zool Afr. 88: 470 - 476." type="journal article" year="1974">Elsen, 1974</bibRefCitation>
Sternal
</taxonomicName>
shield with no posterior incision.................................................................
<taxonomicName id="4C2F4D4FD5110020FDD17C3DFBCC7E46" ID-CoL="8XL5W" authority="(Elaphrolaelaps) Berlese, 1910" authorityName="Berlese" authorityYear="1910" box="[599,1103,263,287]" class="Arachnida" family="Pachylaelapidae" genus="Elaphrolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subGenus" subGenus="Elaphrolaelaps">
<emphasis id="B95BEADED5110020FDD17C3DFD787E46" box="[599,763,263,287]" italics="true" pageId="7" pageNumber="408">Elaphrolaelaps</emphasis>
(
<emphasis id="B95BEADED5110020FC8C7C3DFC2D7E46" box="[778,942,263,287]" italics="true" pageId="7" pageNumber="408">Elaphrolaelaps</emphasis>
)
<bibRefCitation id="EFBE4B3DD5110020FC387C3DFBCC7E46" author="Berlese A" box="[958,1103,263,287]" pageId="7" pageNumber="408" pagination="242 - 271" refId="ref5780" refString="Berlese A. 1910. Lista di nuove specie e nuovi generi di Acari. Redia. 6: 242 - 271." type="journal article" year="1910">Berlese, 1910</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF457C02FBCC7ED6" blockId="7.[195,1103,311,399]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF457C02FBCC7ED6" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF457C02FBCC7ED6" pageId="7" pageNumber="408">7. Anal and genitoventral shields fused to form a genitoventrianal shield bearing three to five pairs of pre-anal setae and three circumanal setae............. 8 Anal shield free, subtriangular, bearing only three circumanal setae........ 9</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF457C9CFBCC7D07" blockId="7.[195,1103,422,606]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF457C9CFBCC7D07" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF457C9CFBCC7D07" pageId="7" pageNumber="408">
8. Setae J5 minute (shorter than
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), subconical; sperm access system with tubular components opened onto coxae IV......................................................................
<taxonomicName id="4C2F4D4FD5110020FD3C7CDDFEB87D47" ID-CoL="8X7MJ" authority="(Olopachys) Berlese, 1910 Setae J" authorityName="Berlese" authorityYear="1910" class="Arachnida" family="Pachylaelapidae" genus="Olopachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subGenus" subGenus="Olopachys">
<emphasis id="B95BEADED5110020FD3C7CDDFCAF7EA6" box="[698,812,487,511]" italics="true" pageId="7" pageNumber="408">Olopachys</emphasis>
(
<emphasis id="B95BEADED5110020FCBA7CDDFC2D7EA6" box="[828,942,487,511]" italics="true" pageId="7" pageNumber="408">Olopachys</emphasis>
)
<bibRefCitation id="EFBE4B3DD5110020FC387CDCFBCC7EA7" author="Berlese A" box="[958,1103,486,510]" pageId="7" pageNumber="408" pagination="242 - 271" refId="ref5780" refString="Berlese A. 1910. Lista di nuove specie e nuovi generi di Acari. Redia. 6: 242 - 271." type="journal article" year="1910">Berlese, 1910</bibRefCitation>
Setae J
</taxonomicName>
5 not reduced in length, needle-like; sperm access system with saccular components opened onto coxae IV..........................................................................
<taxonomicName id="4C2F4D4FD5110020FD0F7F7CFBCC7D07" ID-CoL="8XS2D" authority="Masan, 2007" authorityName="Masan" authorityYear="2007" box="[649,1103,582,606]" class="Arachnida" family="Pachylaelapidae" genus="Olopachys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subGenus" subGenus="Olopachylaella">
<emphasis id="B95BEADED5110020FD0F7F7CFD787D07" box="[649,763,582,606]" italics="true" pageId="7" pageNumber="408">Olopachys</emphasis>
(
<emphasis id="B95BEADED5110020FC8D7F7CFC2C7D07" box="[779,943,582,606]" italics="true" pageId="7" pageNumber="408">Olopachylaella</emphasis>
)
<bibRefCitation id="EFBE4B3DD5110020FC397F7CFBCC7D07" author="Masan P" box="[959,1103,582,606]" pageId="7" pageNumber="408" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF457F4CFBD37DF7" blockId="7.[195,1104,630,686]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF457F4CFBD37DF7" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF457F4CFBD37DF7" pageId="7" pageNumber="408">9. Tarsus II with one distal seta (pl1) modified, thick and spur-like.......... 10 Tarsus II with two distal setae (pl1, pl2) modified, thick and spur-like..... 13</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF307FFFFBD37CE4" blockId="7.[182,1104,709,957]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF307FFFFBD37CE4" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF307FFFFBD37CE4" pageId="7" pageNumber="408">
10. Idiosoma subglobular; dorsal shield strongly vaulted, expanded ventrally and covering margins of ventral surface; setae J5 inserted with some other marginal dorsal shield setae on ventral expansion of the shield; unsclerotized idiosomal integument with seven pairs of setae........................................................................
<taxonomicName id="4C2F4D4FD5110020FD697E7FFEDB7C24" ID-CoL="8XNH4" authority="Masan, 2007 Idiosoma" authorityName="Masan" authorityYear="2007" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5110020FD697E7FFC3B7C04" box="[751,952,837,861]" italics="true" pageId="7" pageNumber="408">Pachyglobolaelaps</emphasis>
<bibRefCitation id="EFBE4B3DD5110020FC397E7FFBCC7C04" author="Masan P" box="[959,1103,837,861]" pageId="7" pageNumber="408" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Idiosoma
</taxonomicName>
dorsoventrally flattened; dorsal shield flat or slightly convex, covering only dorsal surface; setae J5 inserted dorsally; unsclerotized integument with more than seven pairs of setae (usually 10–15 pairs)................. 11
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF307EEEFBD37BF5" blockId="7.[182,1104,980,1196]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF307EEEFBD37BF5" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF307EEEFBD37BF5" pageId="7" pageNumber="408">
11. Setae J5 minute (shorter than
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), subconical; tubular components of sperm access system concealed in coxae III (in
<taxonomicName id="4C2F4D4FD5110020FC827ECFFCF97B54" box="[772,890,1013,1037]" class="Arachnida" family="Pachylaelapidae" genus="Pachyglobolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="sculptus">
<emphasis id="B95BEADED5110020FC827ECFFCF97B54" box="[772,890,1013,1037]" italics="true" pageId="7" pageNumber="408">P. sculptus</emphasis>
</taxonomicName>
group, extra-coxal section very short and with opening slightly protruding above inner surface of the coxa)........................................
<taxonomicName id="4C2F4D4FD5110020FCBE790EFEC37B35" ID-CoL="8XNGY" authority="Masan, 2007 Setae J" authorityName="Masan" authorityYear="2007" class="Arachnida" family="Pachylaelapidae" genus="Pachydellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5110020FCBE790EFC3B7B15" box="[824,952,1076,1100]" italics="true" pageId="7" pageNumber="408">Pachydellus</emphasis>
<bibRefCitation id="EFBE4B3DD5110020FC38790EFBCC7B15" author="Masan P" box="[958,1103,1076,1100]" pageId="7" pageNumber="408" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Setae J
</taxonomicName>
5 not reduced in length, needle-like; tubular components of sperm access system otherwise formed, with long and fine tubes simply opened onto coxae III............................................................... 12
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF3079FEFBCC7A02" blockId="7.[182,1104,1220,1372]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF3079FEFBCC7A02" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF3079FEFBCC7A02" pageId="7" pageNumber="408">
12. Long-legged forms, legs I and IV markedly longer than dorsal shield; genu IV with eight setae (2-4
<emphasis id="B95BEADED5110020FE4879DEFE597BA5" bold="true" box="[462,474,1252,1276]" italics="true" pageId="7" pageNumber="408">/</emphasis>
1-1) and tibia IV with nine setae (2-3
<emphasis id="B95BEADED5110020FCF079DEFC017BA5" bold="true" box="[886,898,1252,1276]" italics="true" pageId="7" pageNumber="408">/</emphasis>
2-2)........................................................
<emphasis id="B95BEADED5110020FD74783EFC347A45" box="[754,951,1284,1308]" italics="true" pageId="7" pageNumber="408">Pachylaelapsoides</emphasis>
<bibRefCitation id="EFBE4B3DD5110020FC387839FBCC7A42" author="Masan P" box="[958,1103,1283,1307]" pageId="7" pageNumber="408" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Short-legged forms, legs I and IV shorter than dorsal shield; genu IV (1-4
<emphasis id="B95BEADED5110020FB90781EFBA17A65" bold="true" box="[1046,1058,1316,1340]" italics="true" pageId="7" pageNumber="408">/</emphasis>
1-1) and tibia IV (1-3
<emphasis id="B95BEADED5110020FE2E787EFE377A05" bold="true" box="[424,436,1348,1372]" italics="true" pageId="7" pageNumber="408">/</emphasis>
2-1) with seven setae............
<taxonomicName id="4C2F4D4FD5110020FCB7787EFBCC7A02" ID-CoL="8W4VB" authority="Berlese, 1904" authorityName="Berlese" authorityYear="1904" box="[817,1103,1347,1372]" class="Arachnida" family="Pachylaelapidae" genus="Onchodellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B95BEADED5110020FCB7787EFC347A05" box="[817,951,1348,1372]" italics="true" pageId="7" pageNumber="408">Onchodellus</emphasis>
<bibRefCitation id="EFBE4B3DD5110020FC3B7879FBCC7A02" author="Berlese A" box="[957,1103,1347,1371]" pageId="7" pageNumber="408" pagination="299 - 474" refId="ref5758" refString="Berlese A. 1904. Illustrazione iconografica degli Acari mirmecofili. Redia. 1: 299 - 474 + Plates VII-XX." type="journal article" year="1904">Berlese, 1904</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8B9036CCD5110020FF307849FBD37933" blockId="7.[182,1104,1395,1642]" pageId="7" pageNumber="408">
<keyLead id="30DE95F9D5110020FF307849FBD37933" pageId="7" pageNumber="408">
<keyStep id="30DB2E69D5110020FF307849FBD37933" pageId="7" pageNumber="408">
13. Dorsal shield with 29 pairs of setae (S1 absent); chaetotaxy of some leg segments deficient: femur I with 12 setae (2-5
<emphasis id="B95BEADED5110020FD7D78A9FC847AF2" bold="true" box="[763,775,1427,1451]" italics="true" pageId="7" pageNumber="408">/</emphasis>
3-2), genu I and tibia I with 11 setae (2-5
<emphasis id="B95BEADED5110020FEFD7889FE047A92" bold="true" box="[379,391,1459,1483]" italics="true" pageId="7" pageNumber="408">/</emphasis>
2-2), genu III with six setae (1-4
<emphasis id="B95BEADED5110020FD747889FD7D7A92" bold="true" box="[754,766,1459,1483]" italics="true" pageId="7" pageNumber="408">/</emphasis>
0-1); R setae on unsclerotized integument absent...........................
<emphasis id="B95BEADED5110020FD7978E9FC3B7AB2" box="[767,952,1491,1515]" italics="true" pageId="7" pageNumber="408">Neopachylaelaps</emphasis>
<bibRefCitation id="EFBE4B3DD5110020FC3878E9FBCC7AB2" author="Masan P" box="[958,1103,1491,1515]" pageId="7" pageNumber="408" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Dorsal shield with 30 pairs of setae (S1 present); leg chaetotaxy normal: femur I (2-5
<emphasis id="B95BEADED5110020FEAD7B29FEB47972" bold="true" box="[299,311,1555,1579]" italics="true" pageId="7" pageNumber="408">/</emphasis>
4-2) and genu I (2-6
<emphasis id="B95BEADED5110020FD9F7B29FDA67972" bold="true" box="[537,549,1555,1579]" italics="true" pageId="7" pageNumber="408">/</emphasis>
3-2) with 13 setae, tibia I with 12 setae (2-5
<emphasis id="B95BEADED5110020FB897B29FB987972" bold="true" box="[1039,1051,1555,1579]" italics="true" pageId="7" pageNumber="408">/</emphasis>
3-2), genu III with seven setae (1-4
<emphasis id="B95BEADED5110020FDD57B09FDDC7912" bold="true" box="[595,607,1587,1611]" italics="true" pageId="7" pageNumber="408">/</emphasis>
1-1); at least one R seta on unsclerotized integument present..................................................... 14
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</paragraph>
<paragraph id="8B9036CCD51E002FFF7C7D9DFB107E06" blockId="8.[250,1172,167,351]" pageId="8" pageNumber="409">
<keyLead id="30DE95F9D51E002FFF7C7D9DFB107E06" pageId="8" pageNumber="409">
<keyStep id="30DB2E69D51E002FFF7C7D9DFB107E06" pageId="8" pageNumber="409">
14. Setae J5 not reduced in length, needle-like; posterolateral margins of dorsal shield with one pair of slit-like hypertrophied gland pores (gdS4)........................................
<taxonomicName id="4C2F4D4FD51E002FFD0C7DDDFE067E46" ID-CoL="8XRTC" authority="Masan, 2007 Setae J" authorityName="Masan" authorityYear="2007" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="subGenus" subGenus="Longipachylaelaps">
<emphasis id="B95BEADED51E002FFD0C7DDDFC9B7FA6" box="[650,792,231,255]" italics="true" pageId="8" pageNumber="409">Pachylaelaps</emphasis>
(
<emphasis id="B95BEADED51E002FFCA17DDDFC707FA6" box="[807,1011,231,255]" italics="true" pageId="8" pageNumber="409">Longipachylaelaps</emphasis>
)
<bibRefCitation id="EFBE4B3DD51E002FFB857DDDFB107FA6" author="Masan P" box="[1027,1171,231,255]" pageId="8" pageNumber="409" refId="ref6092" refString="Masan P. 2007. A review of the family Pachylaelapidae in Slovakia, with systematics and ecology of European Species (Acari: Mesostigmata: Eviphidoide) a. Bratislava: NOI Press; 247 pp." type="book" year="2007">Mašán, 2007</bibRefCitation>
Setae J
</taxonomicName>
5 minute (shorter than
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), subconical; posterolateral margins of dorsal shield with two pairs of slit-like hypertrophied gland pores (gdZ1, gdS4)............................
<taxonomicName id="4C2F4D4FD51E002FFD417C7DFB107E06" ID-CoL="8W4W5" authority="(Pachylaelaps) Berlese, 1888" authorityName="Berlese" authorityYear="1888" box="[711,1171,327,351]" class="Arachnida" family="Pachylaelapidae" genus="Pachylaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="409" phylum="Arthropoda" rank="subGenus" subGenus="Pachylaelaps">
<emphasis id="B95BEADED51E002FFD417C7DFCD67E06" box="[711,853,327,351]" italics="true" pageId="8" pageNumber="409">Pachylaelaps</emphasis>
(
<emphasis id="B95BEADED51E002FFCE27C7DFC717E06" box="[868,1010,327,351]" italics="true" pageId="8" pageNumber="409">Pachylaelaps</emphasis>
)
<bibRefCitation id="EFBE4B3DD51E002FFB847C7DFB107E06" author="Berlese A" box="[1026,1171,327,351]" pageId="8" pageNumber="409" refId="ref5703" refString="Berlese A. 1888. Acari, Myriopoda et Scorpiones hucusque in Italia reperta 51. 17 text pages + Plates 1 - 10. (Reprint by Junk, The Hague, 1979)." type="book" year="1888">Berlese, 1888</bibRefCitation>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</key>
</subSubSection>
</treatment>
</document>
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