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<mods:title id="4DE19DB035B5FB0D8CD5182AD52038A5">Systematics and biogeography of Hemidactylus homoeolepis Blanford, 1881 (Squamata: Gekkonidae), with the description of a new species from Arabia</mods:title>
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<mods:namePart id="2BD4BC05F69E109633C1056B5A739142">Carranza, Salvador</mods:namePart>
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<taxonomicName id="4C1B4D03FFA67D66B2E5FC7DFE1AFC4F" box="[151,423,978,1004]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFA67D66B2E5FC7DFE1AFC4F" bold="true" box="[151,423,978,1004]" italics="true" pageId="7" pageNumber="508">Hemidactylus minutus</emphasis>
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(
<figureCitation id="13202A05FFA67D66B2EDFC5AFEB7FBAD" box="[159,266,1013,1038]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="9.[151,250,1921,1943]" captionStartId-1="10.[151,250,1921,1943]" captionStartId-2="11.[151,250,1290,1312]" captionStartId-3="12.[151,250,1801,1823]" captionTargetBox-2="[196,1375,263,1264]" captionTargetBox-3="[220,1356,204,1774]" captionTargetId-2="figure@11.[176,1411,175,1285]" captionTargetId-3="figure@12.[219,1371,193,1775]" captionTargetPageId-2="11" captionTargetPageId-3="12" captionText-0="FIGURE 2. Maximum likelihood phylogenetic tree rooted with H. masirahensis and H. inexpectatus (outgroups not shown). Black dots on the nodes indicate ML bootstrap values ≥ 70 and BI posterior probabilities ≥ 0.95. Mean ages and 95 % highest posterior density intervals of the nodes estimated with BEAST dating analysis are respectively indicated above and below the nodes. Clades are colored according to their geographic location marked on the map depicted in Fig. 1." captionText-1="FIGURE 3. Parsimony networks corresponding to RAG 1, MC 1 R and c-mos nDNA sequence variation. Lines represent a mutational step, circles haplotypes and dots unsampled haplotypes. The size of circles is proportional to the number of individuals. Samples from the same species are similarly colored but with different tones for different geographic locations. For correspondences of sample and location codes see Appendix I." captionText-2="FIGURE 4. Size and shape differences between Hemidactylus minutus sp. nov. (blue lines and squares) from mainland Arabia and H. homoeolepis from Socotra (red lines and rectangles). The total contribution of the first three components of the PCA analysis to explain the total morphological variation is also given. See material and methods for details." captionText-3="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri-2="https://zenodo.org/record/228116/files/figure.png" httpUri-3="https://zenodo.org/record/228117/files/figure.png" pageId="7" pageNumber="508">Figs. 2–5</figureCitation>
, Table 1,2)
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<emphasis id="B96FEA92FFA67D66B2E5FB92FE12FBF0" box="[151,431,1085,1107]" italics="true" pageId="7" pageNumber="508">Hemidactylus homoeolepis</emphasis>
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: 103
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(part.); 
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: 279
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(part.); 
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: 419
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(part.); 
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: 50
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(part.); 
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: 515
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(part.); 
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: 115
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(part.); van der 
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: 111
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(part.); 
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<bibRefCitation id="EF8A4B71FFA67D66B3B5FBD5FD2FFB33" author="Razzetti" box="[455,658,1146,1168]" pageId="7" pageNumber="508" refString="Razzetti, E., Sindaco, R., Grieco, C., Pella, F., Ziliani, U., Pupin, F., Riservato, E., Pellitteri-Rosa, D., Suleiman, A. S., Al- Aseily, B. A., Carugati, C., Boncompagni, E. &amp; Fasola, M. (2011) Annotated checklist and distribution of the Socotran Archipelago Herpetofauna (Reptilia). Zootaxa, 2826, 1 - 44." type="journal article" year="2011">
Razzetti 
<emphasis id="B96FEA92FFA67D66B050FBD4FDEBFB33" box="[546,598,1146,1168]" italics="true" pageId="7" pageNumber="508">et al.</emphasis>
2011
</bibRefCitation>
: 8
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(part.); 
<treatmentCitation id="0ABA1091FFA67D66B08EFBD5FB94FB33" author="Carranza" box="[764,1065,1146,1168]" httpUri="http://treatment.plazi.org/id/03E36252C511FFD0F39BFF3AFF72FB2C" page="47" pageId="7" pageNumber="508" year="2012">
<bibRefCitation id="EF8A4B71FFA67D66B08EFBD5FBBBFB33" author="Carranza" box="[764,1030,1146,1168]" pageId="7" pageNumber="508" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold, 2012</bibRefCitation>
: 47
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(part.); 
<treatmentCitation id="0ABA1091FFA67D66B60CFBD5FAF2FB33" author="Smid" box="[1150,1359,1146,1168]" page="7" pageId="7" pageNumber="508" year="2013">
<bibRefCitation id="EF8A4B71FFA67D66B60CFBD5FA85FB33" author="Smid" box="[1150,1336,1146,1168]" pageId="7" pageNumber="508" refString="Smid, J., Carranza, S., Kratochvil, L., Gvozdik, V., Nasher, A. K. &amp; Moravec, J. (2013 a) Out of Arabia: A complex biogeographic history of multiple vicariance and dispersal events in the gecko genus Hemidactylus (Reptilia: Gekkonidae). PloS one, 8 (5), e 64018. http: // dx. doi. org / 10.1371 / journal. pone. 0064018" type="journal article" year="2013" yearSuffix="a">
Šmíd 
<emphasis id="B96FEA92FFA67D66B6C9FBD4FB54FB33" box="[1211,1257,1146,1168]" italics="true" pageId="7" pageNumber="508">et al</emphasis>
. 2013a
</bibRefCitation>
: 7
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(part.); 
<treatmentCitation id="0ABA1091FFA67D66B2B5FB36FE36FB0C" author="Gardner" box="[199,395,1177,1199]" page="134" pageId="7" pageNumber="508" year="2013">
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: 134
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(part.).
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<emphasis id="B96FEA92FFA67D66B2E5FB73FEB6FB56" bold="true" box="[151,267,1244,1269]" pageId="7" pageNumber="508">
<typeStatus id="54A08822FFA67D66B2E5FB73FEBBFB56" box="[151,262,1244,1269]" pageId="7" pageNumber="508" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="ED0AAE45FFA67D66B361FB73FEC3FB57" LSID="urn:lsid:biocol.org:col:34665" box="[275,382,1244,1268]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="508">NHMUK</collectionCode>
2013.901 (sample code S7676), male, from Asylah, 
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, 
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, 
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WGS84, collected on the 
<date id="FFA51040FFA67D66B327FB50FDA1FABB" box="[341,540,1279,1304]" pageId="7" pageNumber="508" value="2010-10-09">9th October 2010</date>
by 
<collectingCountry id="F30C7610FFA67D66B038FB50FD13FABB" box="[586,686,1279,1304]" name="El Salvador" pageId="7" pageNumber="508">Salvador</collectingCountry>
Carranza and Fèlix Amat (
<figureCitation id="13202A05FFA67D66B195FB50FB96FABB" box="[999,1067,1279,1304]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="7" pageNumber="508">Fig. 5</figureCitation>
B, D-E, Morphobank M999494- M9999).
</paragraph>
<paragraph id="8BA43680FFA67D66B2B5FAE8FC31FA6E" blockId="7.[151,1436,1244,2025]" pageId="7" pageNumber="508">
<emphasis id="B96FEA92FFA67D66B2B5FAE8FEFBFAC3" bold="true" box="[199,326,1351,1376]" pageId="7" pageNumber="508">
<typeStatus id="54A08822FFA67D66B2B5FAE8FEFCFAC3" box="[199,321,1351,1376]" pageId="7" pageNumber="508" type="paratype">Paratypes</typeStatus>
.
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<collectionCode id="ED0AAE45FFA67D66B33DFAE7FE04FAC3" LSID="urn:lsid:biocol.org:col:34665" box="[335,441,1352,1376]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="508">NHMUK</collectionCode>
2013.902 (sample code S7657), 
<collectionCode id="ED0AAE45FFA67D66B15CFAE7FC25FAC3" LSID="urn:lsid:biocol.org:col:34665" box="[814,920,1352,1376]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="508">NHMUK</collectionCode>
2013.903 (sample code S7673), 
<collectionCode id="ED0AAE45FFA67D66B77DFAE8FAD8FAC3" box="[1295,1381,1351,1376]" pageId="7" pageNumber="508">ONHM</collectionCode>
3716 (sample code S7668) and 
<collectionCode id="ED0AAE45FFA67D66B3BAFAC3FDBBFA26" box="[456,518,1388,1413]" httpUri="http://grbio.org/cool/fx6k-enrh" name="Natal Museum" pageId="7" pageNumber="508">NMP</collectionCode>
<date id="FFA51040FFA67D66B074FAC3FD9AFA26" box="[518,551,1388,1413]" pageId="7" pageNumber="508">
6 
<accessionNumber id="9448AB63FFA67D66B061FAC3FD9AFA26" box="[531,551,1388,1413]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="7" pageNumber="508" type="EnaNcbi">V</accessionNumber>
</date>
74885 (sample code S7664), All 
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are males, except the latter one (female), and have the same collection data as the 
<typeStatus id="54A08822FFA67D66B166FA20FCCBFA0B" box="[788,886,1423,1448]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
(Morphobank M100008-M100011, M100012- M100017, M100000-M100007, M100039-M100043, respectively).
</paragraph>
<paragraph id="8BA43680FFA67D66B2B5FA78FE84F99B" blockId="7.[151,1436,1244,2025]" pageId="7" pageNumber="508">
<emphasis id="B96FEA92FFA67D66B2B5FA78FE1AFA53" bold="true" box="[199,423,1495,1520]" pageId="7" pageNumber="508">Referred material.</emphasis>
Twenty specimens for genetic analyses (five of them not included in the morphological analyses; Appendix I) and 40 voucher specimens for morphological analyses (Appendix II), including the 
<typeStatus id="54A08822FFA67D66B748FA53FA21F9B6" box="[1338,1436,1532,1557]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
and 
<typeStatus id="54A08822FFA67D66B2B4F98FFE88F99B" box="[198,309,1568,1592]" pageId="7" pageNumber="508" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C301650BFFA67D66B2B5F9EBFDC7F923" pageId="7" pageNumber="508" type="etymology">
<paragraph id="8BA43680FFA67D66B2B5F9EBFDC7F923" blockId="7.[151,1436,1244,2025]" pageId="7" pageNumber="508">
<emphasis id="B96FEA92FFA67D66B2B5F9EBFEF3F9FE" bold="true" box="[199,334,1604,1629]" pageId="7" pageNumber="508">Etymology.</emphasis>
The species epithet “ 
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<emphasis id="B96FEA92FFA67D66B031F9EAFD21F9FF" box="[579,668,1605,1628]" italics="true" pageId="7" pageNumber="508">minutus</emphasis>
</taxonomicName>
” is a Latin adjective that refers to the small size of this species, the smallest 
<taxonomicName id="4C1B4D03FFA67D66B288F9C8FE29F923" box="[250,404,1639,1664]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96FEA92FFA67D66B288F9C8FE29F923" box="[250,404,1639,1664]" italics="true" pageId="7" pageNumber="508">Hemidactylus</emphasis>
</taxonomicName>
in mainland Arabia.
</paragraph>
</subSubSection>
<subSubSection id="C301650BFFA67D6BB2B5F923FDB5F8EF" lastPageId="10" lastPageNumber="511" pageId="7" pageNumber="508" type="diagnosis">
<paragraph id="8BA43680FFA67D66B2B5F923FC56F8DE" blockId="7.[151,1436,1244,2025]" pageId="7" pageNumber="508">
<emphasis id="B96FEA92FFA67D66B2B5F923FEFCF906" bold="true" box="[199,321,1676,1701]" pageId="7" pageNumber="508">Diagnosis.</emphasis>
A small 
<taxonomicName id="4C1B4D03FFA67D66B3D7F923FD82F906" box="[421,575,1676,1701]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96FEA92FFA67D66B3D7F923FD82F906" box="[421,575,1676,1701]" italics="true" pageId="7" pageNumber="508">Hemidactylus</emphasis>
</taxonomicName>
characterized by the following combination of morphological characters: (1) maximum recorded snout-vent length, SVL 
<quantity id="4CE39B65FFA67D66B0E7F900FD41F964" box="[661,764,1711,1736]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="7" pageNumber="508" unit="mm" value="34.6">34.6 mm</quantity>
(mean 29.2 ± 
<quantity id="4CE39B65FFA67D66B1D8F91FFBB5F96B" box="[938,1032,1712,1736]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="508" unit="mm" value="3.4">3.4 mm</quantity>
); (2) absence of enlarged tubercles anywhere on the body; (3) expanded subcaudal scales beginning some way from tail base; (4) head narrow and low (
<quantity id="4CE39B65FFA67D66B2ECF958FE97F8AC" box="[158,298,1783,1808]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" metricValueMax="6.7" metricValueMin="4.2" pageId="7" pageNumber="508" unit="mm" value="5.45" valueMax="6.7" valueMin="4.2">4.2–6.7 mm</quantity>
in width and 
<quantity id="4CE39B65FFA67D66B3A0F957FDE1F8AC" box="[466,604,1784,1808]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.7" metricValueMin="2.4" pageId="7" pageNumber="508" unit="mm" value="3.05" valueMax="3.7" valueMin="2.4">2.4–3.7 mm</quantity>
in height); (5) relatively short snout (2.0– 
<quantity id="4CE39B65FFA67D66B639F957FB18F8AC" box="[1099,1189,1784,1808]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="508" unit="mm" value="2.9">2.9 mm</quantity>
nostril–eye); (4) 4–6 preanal pores, PAP (mean 5.8 ± 0.5); (6) four or five lamellae under the first posterior toe, LP1st (4.5 ± 0.5); (7) seven to nine, but most usually eight lamellae under the fourth posterior toe, LP4th (mean LP4th 8.0 ± 0.2); (8) weakly contrasted black and white banded pattern on the ventral part of tail.
</paragraph>
<paragraph id="8BA43680FFA67D69B2B5F828FF63FEC7" blockId="7.[151,1436,1244,2025]" lastBlockId="8.[151,1437,151,357]" lastPageId="8" lastPageNumber="509" pageId="7" pageNumber="508">
<taxonomicName id="4C1B4D03FFA67D66B2B5F828FE76F803" box="[199,459,1927,1952]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFA67D66B2B5F828FE76F803" box="[199,459,1927,1952]" italics="true" pageId="7" pageNumber="508">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFA67D66B3A9F828FD83F803" box="[475,574,1927,1952]" pageId="7" pageNumber="508" rank="species">
<emphasis id="B96FEA92FFA67D66B3A9F828FD83F803" bold="true" box="[475,574,1927,1952]" pageId="7" pageNumber="508">sp. nov.</emphasis>
</taxonomicNameLabel>
is morphologically similar to its sister taxon 
<taxonomicName id="4C1B4D03FFA67D66B6F8F826FAFEF803" box="[1162,1347,1927,1952]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA67D66B6F8F826FAFEF803" box="[1162,1347,1927,1952]" italics="true" pageId="7" pageNumber="508">H. homoeolepis</emphasis>
</taxonomicName>
but is differentiated from it by its smaller adult size (mean SVL 
<quantity id="4CE39B65FFA67D66B16DF803FC38F867" box="[799,901,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="7" pageNumber="508" unit="mm" value="29.2">29.2 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA67D66B67BF803FBD2F867" box="[1033,1135,1964,1989]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="7" pageNumber="508" unit="mm" value="34.6">34.6 mm</quantity>
, compared with mean 
<quantity id="4CE39B65FFA67D66B719F803FF78F84B" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="7" pageNumber="508" unit="mm" value="37.1">37.1 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA67D66B33CF87FFE0AF84B" box="[334,439,2000,2025]" metricMagnitude="-2" metricUnit="m" metricValue="4.68" pageId="7" pageNumber="508" unit="mm" value="46.8">46.8 mm</quantity>
; 
<tableCitation id="C699033BFFA67D66B3B7F87FFDA1F84B" box="[453,540,2000,2025]" captionStart="TABLE 2" captionStartId="8.[151,242,404,427]" captionTargetBox="[159,1381,534,1238]" captionTargetPageId="8" captionText="TABLE 2. Mean, standard deviation (SD) and range of the morphological variables examined for H. minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra." httpUri="http://table.plazi.org/id/DF646608FFA97D69B2E5FE3BFD44FE6C" pageId="7" pageNumber="508" tableUuid="DF646608FFA97D69B2E5FE3BFD44FE6C">Table 2</tableCitation>
), higher number of PAP (80.8% with six PAP and none with three, compared with 66.7% with four PAP and 3.8% with three; 
<figureCitation id="13202A05FFA97D69B0C1FF38FD4EFF13" box="[691,755,151,176]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">Fig. 5</figureCitation>
F versus 
<figureCitation id="13202A05FFA97D69B12BFF38FC24FF13" box="[857,921,151,176]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">Fig. 5</figureCitation>
M, 
<tableCitation id="C699033BFFA97D69B1B3FF38FBA8FF13" box="[961,1045,151,176]" captionStart="TABLE 2" captionStartId="8.[151,242,404,427]" captionTargetBox="[159,1381,534,1238]" captionTargetPageId="8" captionText="TABLE 2. Mean, standard deviation (SD) and range of the morphological variables examined for H. minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra." httpUri="http://table.plazi.org/id/DF646608FFA97D69B2E5FE3BFD44FE6C" pageId="8" pageNumber="509" tableUuid="DF646608FFA97D69B2E5FE3BFD44FE6C">Table 2</tableCitation>
), fewer lamellae under the first toe (55.1% with four lamellae, 44.9% with five lamellae, compared with 0.0% with four lamellae, 97.5% with five lamellae and 2.5% with six lamellae), and fewer lamellae under the fourth toe (97.3% with either seven or eight lamellae, compared with 100% with nine to eleven lamellae; 
<figureCitation id="13202A05FFA97D69B103FEABFC0BFEBE" box="[881,950,260,285]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">Fig. 5</figureCitation>
G, versus 
<figureCitation id="13202A05FFA97D69B65CFEABFBC9FEBE" box="[1070,1140,260,285]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">Fig. 5</figureCitation>
N; 
<tableCitation id="C699033BFFA97D69B6EEFEABFB4BFEBF" box="[1180,1270,260,285]" captionStart="TABLE 2" captionStartId="8.[151,242,404,427]" captionTargetBox="[159,1381,534,1238]" captionTargetPageId="8" captionText="TABLE 2. Mean, standard deviation (SD) and range of the morphological variables examined for H. minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra." httpUri="http://table.plazi.org/id/DF646608FFA97D69B2E5FE3BFD44FE6C" pageId="8" pageNumber="509" tableUuid="DF646608FFA97D69B2E5FE3BFD44FE6C">Table 2</tableCitation>
), and the less contrasted black and white banded pattern on the tail, especially on the ventral part (
<figureCitation id="13202A05FFA97D69B635FE88FB34FEE3" box="[1095,1161,295,320]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">Fig. 5</figureCitation>
B, E compared with 
<figureCitation id="13202A05FFA97D69B702FE88FF1BFEC6" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="8" pageNumber="509">
Fig. 
<date id="FFA51040FFA97D69B2E5FEE3FF1BFEC6" box="[151,166,332,357]" pageId="8" pageNumber="509">5</date>
</figureCitation>
I, L).
</paragraph>
<caption id="DF646608FFA97D69B2E5FE3BFD44FE6C" ID-Table-UUID="DF646608FFA97D69B2E5FE3BFD44FE6C" httpUri="http://table.plazi.org/id/DF646608FFA97D69B2E5FE3BFD44FE6C" pageId="8" pageNumber="509" targetBox="[159,1381,534,1238]" targetIsTable="true" targetPageId="8">
<paragraph id="8BA43680FFA97D69B2E5FE3BFD44FE6C" blockId="8.[151,1436,404,464]" pageId="8" pageNumber="509">
<emphasis id="B96FEA92FFA97D69B2E5FE3BFEBBFE08" bold="true" box="[151,262,404,427]" pageId="8" pageNumber="509">TABLE 2</emphasis>
. Mean, standard deviation (SD) and range of the morphological variables examined for 
<taxonomicName id="4C1B4D03FFA97D69B6B7FE3AFA86FE0F" box="[1221,1339,405,428]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFA97D69B6B7FE3AFA86FE0F" box="[1221,1339,405,428]" italics="true" pageId="8" pageNumber="509">H. minutus</emphasis>
</taxonomicName>
<emphasis id="B96FEA92FFA97D69B736FE3BFA21FE08" bold="true" box="[1348,1436,404,427]" pageId="8" pageNumber="509">
<taxonomicNameLabel id="A25C57E9FFA97D69B736FE3BFA21FE08" box="[1348,1436,404,427]" pageId="8" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from mainland Arabia and 
<taxonomicName id="4C1B4D03FFA97D69B3C5FE16FDE2FE6C" box="[439,607,439,464]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA97D69B3C5FE16FDE2FE6C" box="[439,607,439,464]" italics="true" pageId="8" pageNumber="509">H. homoeolepis</emphasis>
</taxonomicName>
from Socotra.
</paragraph>
</caption>
<paragraph id="8BA43680FFA97D69B0B5FE42FBB0F81D" blockId="8.[711,1254,492,514]" lastBlockId="8.[151,1437,1309,1982]" pageId="8" pageNumber="509">
<taxonomicName id="4C1B4D03FFA97D69B0B5FE42FC88FDA1" box="[711,821,493,514]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFA97D69B0B5FE42FC88FDA1" box="[711,821,493,514]" italics="true" pageId="8" pageNumber="509">H. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFA97D69B14EFE43FC30FDA2" box="[828,909,492,513]" pageId="8" pageNumber="509" rank="species">
<emphasis id="B96FEA92FFA97D69B14EFE43FC30FDA2" bold="true" box="[828,909,492,513]" pageId="8" pageNumber="509">sp. nov.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="4C1B4D03FFA97D69B634FE42FB5BFDA1" box="[1094,1254,492,514]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="homoeolepis">H. homoeolepis</taxonomicName>
<taxonomicName id="4C1B4D03FFA97D69B2B5FAB2FE7CFA96" box="[199,449,1309,1334]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFA97D69B2B5FAB2FE7CFA96" box="[199,449,1309,1334]" italics="true" pageId="8" pageNumber="509">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFA97D69B3BAFAB2FD9FFA95" box="[456,546,1309,1334]" pageId="8" pageNumber="509" rank="species">
<emphasis id="B96FEA92FFA97D69B3BAFAB2FD9FFA95" bold="true" box="[456,546,1309,1334]" pageId="8" pageNumber="509">sp. nov.</emphasis>
</taxonomicNameLabel>
differs morphologically from neighboring populations of 
<taxonomicName id="4C1B4D03FFA97D69B6C2FAB1FA21FA95" box="[1200,1436,1309,1334]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="paucituberculatus">
<emphasis id="B96FEA92FFA97D69B6C2FAB1FA21FA95" box="[1200,1436,1309,1334]" italics="true" pageId="8" pageNumber="509">H. paucituberculatus</emphasis>
</taxonomicName>
(formerly 
<taxonomicName id="4C1B4D03FFA97D69B378FAEDFE07FAF9" box="[266,442,1345,1370]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA97D69B378FAEDFE07FAF9" box="[266,442,1345,1370]" italics="true" pageId="8" pageNumber="509">H. homoeolepis</emphasis>
</taxonomicName>
) from South 
<collectingCountry id="F30C7610FFA97D69B021FAEDFD25FAF9" box="[595,664,1346,1370]" name="Oman" pageId="8" pageNumber="509">Oman</collectingCountry>
in its rather smaller adult size (SVL mean 
<quantity id="4CE39B65FFA97D69B60DFAEDFB5BFAFA" box="[1151,1254,1346,1370]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="8" pageNumber="509" unit="mm" value="29.2">29.2 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA97D69B719FAEEFF78FADE" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="8" pageNumber="509" unit="mm" value="34.6">34.6 mm</quantity>
, compared with mean 
<quantity id="4CE39B65FFA97D69B3B5FAC9FD93FADE" box="[455,558,1382,1406]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="8" pageNumber="509" unit="mm" value="32.2">32.2 mm</quantity>
, maximum 38.4), absence of enlarged tubercles anywhere on the body (some enlarged tubercles present on the lateral sides of the second half of body), and expanded subcaudal scales beginning some way from tail base (expanded subcaudal scales usually extending almost to tail base). It differs from 
<taxonomicName id="4C1B4D03FFA97D69B2A4FA7DFE33FA49" box="[214,398,1489,1514]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="masirahensis">
<emphasis id="B96FEA92FFA97D69B2A4FA7DFE33FA49" box="[214,398,1489,1514]" italics="true" pageId="8" pageNumber="509">H. masirahensis</emphasis>
</taxonomicName>
(formerly 
<taxonomicName id="4C1B4D03FFA97D69B078FA7DFD06FA49" box="[522,699,1489,1514]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA97D69B078FA7DFD06FA49" box="[522,699,1489,1514]" italics="true" pageId="8" pageNumber="509">H. homoeolepis</emphasis>
</taxonomicName>
) endemic from Masirah Island, 
<collectingCountry id="F30C7610FFA97D69B65EFA7DFBCEFA49" box="[1068,1139,1490,1514]" name="Oman" pageId="8" pageNumber="509">Oman</collectingCountry>
, by its smaller adult body size (SVL mean 
<quantity id="4CE39B65FFA97D69B326FA59FE07F9AE" box="[340,442,1526,1550]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="8" pageNumber="509" unit="mm" value="29.2">29.2 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA97D69B04DFA5AFD1BF9AE" box="[575,678,1525,1550]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="8" pageNumber="509" unit="mm" value="34.6">34.6 mm</quantity>
, compared with mean 
<quantity id="4CE39B65FFA97D69B1D7FA59FBB1F9AE" box="[933,1036,1526,1550]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="8" pageNumber="509" unit="mm" value="32.2">32.2 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA97D69B6E3FA5AFB59F9AD" box="[1169,1252,1525,1550]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="509" unit="mm" value="45.0">45 mm</quantity>
), higher number of PAP in males (96.1% with five or six pores, compared with 100% with four pores), fewer lamellae under the first toe of pes (four or five lamellae, compared with six lamellae), and under the fourth toe of pes (from seven to nine lamellae, compared with 10-11 lamellae), absence of enlarged tubercles anywhere in the body (enlarged tubercles present on lateral sides of body), expanded subcaudal scales beginning some way from tail base (expanded subcaudal scales usually extending almost to tail base), and less contrasted black and white banded pattern on the tail, especially on the ventral part (dark bands of the tail very conspicuous and marked, especially on the underside of tail). It differs from 
<taxonomicName id="4C1B4D03FFA97D69B3E4F95DFDF8F8AA" box="[406,581,1778,1801]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="B96FEA92FFA97D69B3E4F95DFDF8F8AA" box="[406,581,1778,1801]" italics="true" pageId="8" pageNumber="509">H. inexpectatus</emphasis>
</taxonomicName>
(formerly 
<taxonomicName id="4C1B4D03FFA97D69B0CCF95DFCD0F8A9" box="[702,877,1777,1802]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="509" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA97D69B0CCF95DFCD0F8A9" box="[702,877,1777,1802]" italics="true" pageId="8" pageNumber="509">H. homoeolepis</emphasis>
</taxonomicName>
) endemic to 
<collectingCountry id="F30C7610FFA97D69B673F95DFBF5F8A9" box="[1025,1096,1778,1802]" name="Oman" pageId="8" pageNumber="509">Oman</collectingCountry>
, by its smaller adult body size (SVL mean 
<quantity id="4CE39B65FFA97D69B353F8B9FE3AF88E" box="[289,391,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="2.92" pageId="8" pageNumber="509" unit="mm" value="29.2">29.2 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA97D69B07EF8BAFDCEF88E" box="[524,627,1813,1838]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="8" pageNumber="509" unit="mm" value="34.6">34.6 mm</quantity>
, compared with mean 
<quantity id="4CE39B65FFA97D69B100F8BAFC65F88E" box="[882,984,1813,1838]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="8" pageNumber="509" unit="mm" value="37.5">37.5 mm</quantity>
, maximum 
<quantity id="4CE39B65FFA97D69B62FF8B9FB7BF88D" box="[1117,1222,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="4.41" pageId="8" pageNumber="509" unit="mm" value="44.1">44.1 mm</quantity>
), higher number of PAP in males (96.1% with five or six pores, compared with 100% with four PAP), fewer lamellae under the first toe of pes (four or five lamellae, compared with six lamellae), and under the fourth toe of pes (from seven to nine lamellae, compared with 10–11 lamellae), absence of enlarged tubercles anywhere on the body (presence of conical or weakly keeled and extensive tubercles on the body, nape, hind legs and tail).
</paragraph>
<paragraph id="8BA43680FFA97D69B2EDFDB9FAEAFB14" pageId="8" pageNumber="509">
<table id="F91BC420FFA9829EB2EDFDB9FAD8FB75" box="[159,1381,534,1238]" gridcols="5" gridrows="16" pageId="8" pageNumber="509">
<tr id="352B34C2FFA9829EB2EDFDB9FAD8FD8F" box="[159,1381,534,556]" gridrow="0" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFDB9FDCEFD8F" box="[159,627,534,556]" gridcol="0" gridrow="0" pageId="8" pageNumber="509">Variable code</th>
<th id="76FA5DBEFFA9829EB0B5FDB9FC87FD8F" box="[711,826,534,556]" gridcol="1" gridrow="0" pageId="8" pageNumber="509">mean ± SD</th>
<th id="76FA5DBEFFA9829EB1D2FDB9FBB8FD8F" box="[928,1029,534,556]" gridcol="2" gridrow="0" pageId="8" pageNumber="509">range</th>
<th id="76FA5DBEFFA9829EB634FDB9FB0AFD8F" box="[1094,1207,534,556]" gridcol="3" gridrow="0" pageId="8" pageNumber="509">mean ± SD</th>
<th id="76FA5DBEFFA9829EB68DFDB9FAD8FD8F" box="[1279,1381,534,556]" gridcol="4" gridrow="0" pageId="8" pageNumber="509">range</th>
</tr>
<tr id="352B34C2FFA9829EB2EDFDEFFAD8FDF5" box="[159,1381,576,598]" gridrow="1" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFDEFFDCEFDF5" box="[159,627,576,598]" gridcol="0" gridrow="1" pageId="8" pageNumber="509">Snot-vent length SVL</th>
<td id="76FA5DBEFFA9829EB0B5FDEFFC87FDF5" box="[711,826,576,598]" gridcol="1" gridrow="1" pageId="8" pageNumber="509">29.2 ± 3.4</td>
<td id="76FA5DBEFFA9829EB1D2FDEFFBB8FDF5" box="[928,1029,576,598]" gridcol="2" gridrow="1" pageId="8" pageNumber="509">20.5–34.6</td>
<td id="76FA5DBEFFA9829EB634FDEFFB0AFDF5" box="[1094,1207,576,598]" gridcol="3" gridrow="1" pageId="8" pageNumber="509">37.1 ± 4.0</td>
<td id="76FA5DBEFFA9829EB68DFDEFFAD8FDF5" box="[1279,1381,576,598]" gridcol="4" gridrow="1" pageId="8" pageNumber="509">29.2–46.8</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFDC6FAD8FDDC" box="[159,1381,617,639]" gridrow="2" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFDC6FDCEFDDC" box="[159,627,617,639]" gridcol="0" gridrow="2" pageId="8" pageNumber="509">Trunk length TRL</th>
<td id="76FA5DBEFFA9829EB0B5FDC6FC87FDDC" box="[711,826,617,639]" gridcol="1" gridrow="2" pageId="8" pageNumber="509">12.48 ± 1.7</td>
<td id="76FA5DBEFFA9829EB1D2FDC6FBB8FDDC" box="[928,1029,617,639]" gridcol="2" gridrow="2" pageId="8" pageNumber="509">8.8–14.9</td>
<td id="76FA5DBEFFA9829EB634FDC6FB0AFDDC" box="[1094,1207,617,639]" gridcol="3" gridrow="2" pageId="8" pageNumber="509">15.9 ± 2.0</td>
<td id="76FA5DBEFFA9829EB68DFDC6FAD8FDDC" box="[1279,1381,617,639]" gridcol="4" gridrow="2" pageId="8" pageNumber="509">12.1–19.8</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFD3DFAD8FD0B" box="[159,1381,658,680]" gridrow="3" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFD3DFDCEFD0B" box="[159,627,658,680]" gridcol="0" gridrow="3" pageId="8" pageNumber="509">Tail length TL</th>
<td id="76FA5DBEFFA9829EB0B5FD3DFC87FD0B" box="[711,826,658,680]" gridcol="1" gridrow="3" pageId="8" pageNumber="509">28.5 ± 4.4</td>
<td id="76FA5DBEFFA9829EB1D2FD3DFBB8FD0B" box="[928,1029,658,680]" gridcol="2" gridrow="3" pageId="8" pageNumber="509">23.5–36.0</td>
<td id="76FA5DBEFFA9829EB634FD3DFB0AFD0B" box="[1094,1207,658,680]" gridcol="3" gridrow="3" pageId="8" pageNumber="509">32.1 ± 8.9</td>
<td id="76FA5DBEFFA9829EB68DFD3DFAD8FD0B" box="[1279,1381,658,680]" gridcol="4" gridrow="3" pageId="8" pageNumber="509">17.0–44.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFD13FAD8FD71" box="[159,1381,700,722]" gridrow="4" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFD13FDCEFD71" box="[159,627,700,722]" gridcol="0" gridrow="4" pageId="8" pageNumber="509">Head length HL</th>
<td id="76FA5DBEFFA9829EB0B5FD13FC87FD71" box="[711,826,700,722]" gridcol="1" gridrow="4" pageId="8" pageNumber="509">7.6 ± 0.8</td>
<td id="76FA5DBEFFA9829EB1D2FD13FBB8FD71" box="[928,1029,700,722]" gridcol="2" gridrow="4" pageId="8" pageNumber="509">6.2–9.6</td>
<td id="76FA5DBEFFA9829EB634FD13FB0AFD71" box="[1094,1207,700,722]" gridcol="3" gridrow="4" pageId="8" pageNumber="509">9.4 ± 1.1</td>
<td id="76FA5DBEFFA9829EB68DFD13FAD8FD71" box="[1279,1381,700,722]" gridcol="4" gridrow="4" pageId="8" pageNumber="509">7.4–12.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFD4AFAD8FD58" box="[159,1381,741,763]" gridrow="5" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFD4AFDCEFD58" box="[159,627,741,763]" gridcol="0" gridrow="5" pageId="8" pageNumber="509">Head width HW</th>
<td id="76FA5DBEFFA9829EB0B5FD4AFC87FD58" box="[711,826,741,763]" gridcol="1" gridrow="5" pageId="8" pageNumber="509">5.2 ± 0.6</td>
<td id="76FA5DBEFFA9829EB1D2FD4AFBB8FD58" box="[928,1029,741,763]" gridcol="2" gridrow="5" pageId="8" pageNumber="509">4.2–6.7</td>
<td id="76FA5DBEFFA9829EB634FD4AFB0AFD58" box="[1094,1207,741,763]" gridcol="3" gridrow="5" pageId="8" pageNumber="509">6.9 ± 0.9</td>
<td id="76FA5DBEFFA9829EB68DFD4AFAD8FD58" box="[1279,1381,741,763]" gridcol="4" gridrow="5" pageId="8" pageNumber="509">5.3–9.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFCA1FAD8FC87" box="[159,1381,782,804]" gridrow="6" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFCA1FDCEFC87" box="[159,627,782,804]" gridcol="0" gridrow="6" pageId="8" pageNumber="509">Head height HH</th>
<td id="76FA5DBEFFA9829EB0B5FCA1FC87FC87" box="[711,826,782,804]" gridcol="1" gridrow="6" pageId="8" pageNumber="509">3.0 ± 0.4</td>
<td id="76FA5DBEFFA9829EB1D2FCA1FBB8FC87" box="[928,1029,782,804]" gridcol="2" gridrow="6" pageId="8" pageNumber="509">2.4–3.7</td>
<td id="76FA5DBEFFA9829EB634FCA1FB0AFC87" box="[1094,1207,782,804]" gridcol="3" gridrow="6" pageId="8" pageNumber="509">4.2 ± 0.5</td>
<td id="76FA5DBEFFA9829EB68DFCA1FAD8FC87" box="[1279,1381,782,804]" gridcol="4" gridrow="6" pageId="8" pageNumber="509">3.4–5.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFC97FAD8FCED" box="[159,1381,824,846]" gridrow="7" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFC97FDCEFCED" box="[159,627,824,846]" gridcol="0" gridrow="7" pageId="8" pageNumber="509">Orbital diameter OD</th>
<td id="76FA5DBEFFA9829EB0B5FC97FC87FCED" box="[711,826,824,846]" gridcol="1" gridrow="7" pageId="8" pageNumber="509">1.9 ± 0.2</td>
<td id="76FA5DBEFFA9829EB1D2FC97FBB8FCED" box="[928,1029,824,846]" gridcol="2" gridrow="7" pageId="8" pageNumber="509">1.4–2.3</td>
<td id="76FA5DBEFFA9829EB634FC97FB0AFCED" box="[1094,1207,824,846]" gridcol="3" gridrow="7" pageId="8" pageNumber="509">2.5 ± 0.3</td>
<td id="76FA5DBEFFA9829EB68DFC97FAD8FCED" box="[1279,1381,824,846]" gridcol="4" gridrow="7" pageId="8" pageNumber="509">1.7–3.1</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFCCEFAD8FCD4" box="[159,1381,865,887]" gridrow="8" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFCCEFDCEFCD4" box="[159,627,865,887]" gridcol="0" gridrow="8" pageId="8" pageNumber="509">Nostril–eye distance NE</th>
<td id="76FA5DBEFFA9829EB0B5FCCEFC87FCD4" box="[711,826,865,887]" gridcol="1" gridrow="8" pageId="8" pageNumber="509">2.4 ± 0.2</td>
<td id="76FA5DBEFFA9829EB1D2FCCEFBB8FCD4" box="[928,1029,865,887]" gridcol="2" gridrow="8" pageId="8" pageNumber="509">2.0–2.9</td>
<td id="76FA5DBEFFA9829EB634FCCEFB0AFCD4" box="[1094,1207,865,887]" gridcol="3" gridrow="8" pageId="8" pageNumber="509">3.2 ± 0.4</td>
<td id="76FA5DBEFFA9829EB68DFCCEFAD8FCD4" box="[1279,1381,865,887]" gridcol="4" gridrow="8" pageId="8" pageNumber="509">2.7–3.9</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFC25FAD8FC03" box="[159,1381,906,928]" gridrow="9" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFC25FDCEFC03" box="[159,627,906,928]" gridcol="0" gridrow="9" pageId="8" pageNumber="509">Anterior interorbital distance IO1</th>
<td id="76FA5DBEFFA9829EB0B5FC25FC87FC03" box="[711,826,906,928]" gridcol="1" gridrow="9" pageId="8" pageNumber="509">2.6 ± 0.4</td>
<td id="76FA5DBEFFA9829EB1D2FC25FBB8FC03" box="[928,1029,906,928]" gridcol="2" gridrow="9" pageId="8" pageNumber="509">2.0–3.5</td>
<td id="76FA5DBEFFA9829EB634FC25FB0AFC03" box="[1094,1207,906,928]" gridcol="3" gridrow="9" pageId="8" pageNumber="509">3.6 ± 0.4</td>
<td id="76FA5DBEFFA9829EB68DFC25FAD8FC03" box="[1279,1381,906,928]" gridcol="4" gridrow="9" pageId="8" pageNumber="509">2.9–4.5</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFC1BFAD8FC69" box="[159,1381,948,970]" gridrow="10" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFC1BFDCEFC69" box="[159,627,948,970]" gridcol="0" gridrow="10" pageId="8" pageNumber="509">Posterior interorbital distance IO2</th>
<td id="76FA5DBEFFA9829EB0B5FC1BFC87FC69" box="[711,826,948,970]" gridcol="1" gridrow="10" pageId="8" pageNumber="509">3.6 ± 0.4</td>
<td id="76FA5DBEFFA9829EB1D2FC1BFBB8FC69" box="[928,1029,948,970]" gridcol="2" gridrow="10" pageId="8" pageNumber="509">2.8–4.5</td>
<td id="76FA5DBEFFA9829EB634FC1BFB0AFC69" box="[1094,1207,948,970]" gridcol="3" gridrow="10" pageId="8" pageNumber="509">4.8 ± 0.7</td>
<td id="76FA5DBEFFA9829EB68DFC1BFAD8FC69" box="[1279,1381,948,970]" gridcol="4" gridrow="10" pageId="8" pageNumber="509">3.3–6.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFC72FAD8FC50" box="[159,1381,989,1011]" gridrow="11" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFC72FDCEFC50" box="[159,627,989,1011]" gridcol="0" gridrow="11" pageId="8" pageNumber="509">Preanal pores PAP</th>
<td id="76FA5DBEFFA9829EB0B5FC72FC87FC50" box="[711,826,989,1011]" gridcol="1" gridrow="11" pageId="8" pageNumber="509">5.8 ± 0.5</td>
<td id="76FA5DBEFFA9829EB1D2FC72FBB8FC50" box="[928,1029,989,1011]" gridcol="2" gridrow="11" pageId="8" pageNumber="509">4.0–6.0</td>
<td id="76FA5DBEFFA9829EB634FC72FB0AFC50" box="[1094,1207,989,1011]" gridcol="3" gridrow="11" pageId="8" pageNumber="509">4.3 ± 0.9</td>
<td id="76FA5DBEFFA9829EB68DFC72FAD8FC50" box="[1279,1381,989,1011]" gridcol="4" gridrow="11" pageId="8" pageNumber="509">3.0–6.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFBA9FAD8FBBF" box="[159,1381,1030,1052]" gridrow="12" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFBA9FDCEFBBF" box="[159,627,1030,1052]" gridcol="0" gridrow="12" pageId="8" pageNumber="509">Mean number of supra-labial scales SL</th>
<td id="76FA5DBEFFA9829EB0B5FBA9FC87FBBF" box="[711,826,1030,1052]" gridcol="1" gridrow="12" pageId="8" pageNumber="509">9.2 ± 0.5</td>
<td id="76FA5DBEFFA9829EB1D2FBA9FBB8FBBF" box="[928,1029,1030,1052]" gridcol="2" gridrow="12" pageId="8" pageNumber="509">8.0–10.0</td>
<td id="76FA5DBEFFA9829EB634FBA9FB0AFBBF" box="[1094,1207,1030,1052]" gridcol="3" gridrow="12" pageId="8" pageNumber="509">8.7 ± 0.7</td>
<td id="76FA5DBEFFA9829EB68DFBA9FAD8FBBF" box="[1279,1381,1030,1052]" gridcol="4" gridrow="12" pageId="8" pageNumber="509">8.0–10.5</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFB9FFAD8FBE5" box="[159,1381,1072,1094]" gridrow="13" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFB9FFDCEFBE5" box="[159,627,1072,1094]" gridcol="0" gridrow="13" pageId="8" pageNumber="509">Mean number of infra-labial scales IL</th>
<td id="76FA5DBEFFA9829EB0B5FB9FFC87FBE5" box="[711,826,1072,1094]" gridcol="1" gridrow="13" pageId="8" pageNumber="509">7.6 ± 0.5</td>
<td id="76FA5DBEFFA9829EB1D2FB9FFBB8FBE5" box="[928,1029,1072,1094]" gridcol="2" gridrow="13" pageId="8" pageNumber="509">6.5–8.5</td>
<td id="76FA5DBEFFA9829EB634FB9FFB0AFBE5" box="[1094,1207,1072,1094]" gridcol="3" gridrow="13" pageId="8" pageNumber="509">7.9 ± 0.5</td>
<td id="76FA5DBEFFA9829EB68DFB9FFAD8FBE5" box="[1279,1381,1072,1094]" gridcol="4" gridrow="13" pageId="8" pageNumber="509">6.0–9.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFBF6FAD8FB2D" box="[159,1381,1113,1166]" gridrow="14" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFBF6FDCEFB2D" box="[159,627,1113,1166]" gridcol="0" gridrow="14" pageId="8" pageNumber="509">Mean number of lamellae under the LP1st first toe</th>
<td id="76FA5DBEFFA9829EB0B5FBF6FC87FB2D" box="[711,826,1113,1166]" gridcol="1" gridrow="14" pageId="8" pageNumber="509">4.5 ± 0.5</td>
<td id="76FA5DBEFFA9829EB1D2FBF6FBB8FB2D" box="[928,1029,1113,1166]" gridcol="2" gridrow="14" pageId="8" pageNumber="509">4.0–5.0</td>
<td id="76FA5DBEFFA9829EB634FBF6FB0AFB2D" box="[1094,1207,1113,1166]" gridcol="3" gridrow="14" pageId="8" pageNumber="509">5.0 ± 0.2</td>
<td id="76FA5DBEFFA9829EB68DFBF6FAD8FB2D" box="[1279,1381,1113,1166]" gridcol="4" gridrow="14" pageId="8" pageNumber="509">5.0–6.0</td>
</tr>
<tr id="352B34C2FFA9829EB2EDFB0EFAD8FB75" box="[159,1381,1185,1238]" gridrow="15" pageId="8" pageNumber="509">
<th id="76FA5DBEFFA9829EB2EDFB0EFDCEFB75" box="[159,627,1185,1238]" gridcol="0" gridrow="15" pageId="8" pageNumber="509">Mean number of lamellae under the LP4th fourth toe</th>
<td id="76FA5DBEFFA9829EB0B5FB0EFC87FB75" box="[711,826,1185,1238]" gridcol="1" gridrow="15" pageId="8" pageNumber="509">8.0 ± 0.2</td>
<td id="76FA5DBEFFA9829EB1D2FB0EFBB8FB75" box="[928,1029,1185,1238]" gridcol="2" gridrow="15" pageId="8" pageNumber="509">7.5–9.0</td>
<td id="76FA5DBEFFA9829EB634FB0EFB0AFB75" box="[1094,1207,1185,1238]" gridcol="3" gridrow="15" pageId="8" pageNumber="509">9.9 ± 0.5</td>
<td id="76FA5DBEFFA9829EB68DFB0EFAD8FB75" box="[1279,1381,1185,1238]" gridcol="4" gridrow="15" pageId="8" pageNumber="509">9.0–11.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA43680FFA87D68B2B8FF7DFA89FEA0" blockId="9.[202,1332,210,488]" box="[202,1332,210,259]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B2B8FF7DFB96FF5B" bold="true" box="[202,1067,210,248]" pageId="9" pageNumber="510">
<figureCitation id="13202A05FFA87D68B2B8FF7DFEF5FF52" box="[202,328,210,241]" captionStart="FIGURE 2" captionStartId="9.[151,250,1921,1943]" captionText="FIGURE 2. Maximum likelihood phylogenetic tree rooted with H. masirahensis and H. inexpectatus (outgroups not shown). Black dots on the nodes indicate ML bootstrap values ≥ 70 and BI posterior probabilities ≥ 0.95. Mean ages and 95 % highest posterior density intervals of the nodes estimated with BEAST dating analysis are respectively indicated above and below the nodes. Clades are colored according to their geographic location marked on the map depicted in Fig. 1." pageId="9" pageNumber="510">Figure 2</figureCitation>
S2166
</emphasis>
<taxonomicName id="4C1B4D03FFA87D68B6E5FF5FFA89FEA0" box="[1175,1332,239,259]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="masirahensis">
<emphasis id="B96FEA92FFA87D68B6E5FF5FFA89FEA0" bold="true" box="[1175,1332,239,259]" italics="true" pageId="9" pageNumber="510">H. masirahensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BA43680FFA87D68B677FF50FB8DFEAE" blockId="9.[202,1332,210,488]" box="[1029,1072,255,269]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B677FF50FB8DFEAE" bold="true" box="[1029,1072,255,269]" pageId="9" pageNumber="510">S7735</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1ADFEBAFC57FE9A" blockId="9.[202,1332,210,488]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1ADFEBAFBB7FE80" bold="true" box="[991,1034,277,291]" pageId="9" pageNumber="510">S7710</emphasis>
<taxonomicName id="4C1B4D03FFA87D68B6E4FEB8FA94FE89" box="[1174,1321,278,298]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="B96FEA92FFA87D68B6E4FEB8FA94FE89" bold="true" box="[1174,1321,278,298]" italics="true" pageId="9" pageNumber="510">H. inexpectatus</emphasis>
</taxonomicName>
<emphasis id="B96FEA92FFA87D68B1CDFE84FC57FE9A" bold="true" box="[959,1002,299,313]" pageId="9" pageNumber="510">S7707</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1A7FEEEFC58FED8" blockId="9.[202,1332,210,488]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1A7FEEEFC52FEEC" bold="true" box="[981,1007,321,335]" pageId="9" pageNumber="510">JS8</emphasis>
<emphasis id="B96FEA92FFA87D68B6E5FE95FC58FED8" bold="true" pageId="9" pageNumber="510">
<collectingCountry id="F30C7610FFA87D68B6E5FE95FB77FEE9" box="[1175,1226,314,330]" name="Yemen" pageId="9" pageNumber="510">Yemen</collectingCountry>
JS6 (mainland) JS5
</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1A7FE2DFC42FE33" blockId="9.[202,1332,210,488]" box="[981,1023,386,400]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1A7FE2DFC42FE33" bold="true" box="[981,1023,386,400]" pageId="9" pageNumber="510">S7657</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B0D4FE26FBBDFE05" blockId="9.[202,1332,210,488]" box="[678,1024,393,422]" pageId="9" pageNumber="510">
<geoCoordinate id="EE2F5047FFA87D68B0D4FE26FC60FE05" box="[678,989,393,422]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-1.71">
<emphasis id="B96FEA92FFA87D68B0D4FE26FC60FE05" bold="true" box="[678,989,393,422]" pageId="9" pageNumber="510">1.71 S</emphasis>
</geoCoordinate>
7676
</paragraph>
<paragraph id="8BA43680FFA87D68B0FEFE03FBB4FE1F" blockId="9.[202,1332,210,488]" box="[652,1033,428,444]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B0FEFE03FBB4FE1F" bold="true" box="[652,1033,428,444]" pageId="9" pageNumber="510">
0.85 - 
<geoCoordinate id="EE2F5047FFA87D68B0C8FE03FD64FE1F" box="[698,729,428,444]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-2.73">2.73</geoCoordinate>
S7673
</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1ACFE6BFBB4FE71" blockId="9.[202,1332,210,488]" box="[990,1033,452,466]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1ACFE6BFBB4FE71" bold="true" box="[990,1033,452,466]" pageId="9" pageNumber="510">S7668</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1ACFE75FBB4FE4B" blockId="9.[202,1332,210,488]" box="[990,1033,474,488]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1ACFE75FBB4FE4B" bold="true" box="[990,1033,474,488]" pageId="9" pageNumber="510">S7664</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B312FD12FEC2FD6E" blockId="9.[327,1317,701,1883]" box="[352,383,701,717]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B312FD12FEC2FD6E" bold="true" box="[352,383,701,717]" pageId="9" pageNumber="510">4.37</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B335FD4FFE2EFD53" blockId="9.[327,1317,701,1883]" box="[327,403,736,752]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B335FD4FFE2EFD53" bold="true" box="[327,403,736,752]" pageId="9" pageNumber="510">2.43 - 6.64</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B11FFD5AFC2AFCA0" blockId="9.[327,1317,701,1883]" box="[877,919,757,771]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B11FFD5AFC2AFCA0" bold="true" box="[877,919,757,771]" pageId="9" pageNumber="510">S3562</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B177FD53FC20FCBA" blockId="9.[327,1317,701,1883]" box="[773,925,764,793]" pageId="9" pageNumber="510">
<geoCoordinate id="EE2F5047FFA87D68B177FD53FCC6FCBA" box="[773,891,764,793]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.23">
<emphasis id="B96FEA92FFA87D68B177FD53FCC6FCBA" bold="true" box="[773,891,764,793]" pageId="9" pageNumber="510">0.23 S</emphasis>
</geoCoordinate>
2831
</paragraph>
<paragraph id="8BA43680FFA87D68B099FCB0FC25FC8C" blockId="9.[327,1317,701,1883]" box="[747,920,799,815]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B099FCB0FC25FC8C" bold="true" box="[747,920,799,815]" pageId="9" pageNumber="510">
0.10 - 
<geoCoordinate id="EE2F5047FFA87D68B16BFCB0FC85FC8C" box="[793,824,799,815]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.39">0.39</geoCoordinate>
S5305
</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1F9FC98FC0BFCE6" blockId="9.[327,1317,701,1883]" box="[907,950,823,837]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1F9FC98FC0BFCE6" bold="true" box="[907,950,823,837]" pageId="9" pageNumber="510">S5189</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1F4FCE3FC0CFCF9" blockId="9.[327,1317,701,1883]" box="[902,945,844,858]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1F4FCE3FC0CFCF9" bold="true" box="[902,945,844,858]" pageId="9" pageNumber="510">S5306</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1E7FCCDFC02FCD3" blockId="9.[327,1317,701,1883]" box="[917,959,866,880]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1E7FCCDFC02FCD3" bold="true" box="[917,959,866,880]" pageId="9" pageNumber="510">S5353</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1F9FCD7FC08FC25" blockId="9.[327,1317,701,1883]" box="[907,949,888,902]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1F9FCD7FC08FC25" bold="true" box="[907,949,888,902]" pageId="9" pageNumber="510">S5154</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1E8FC21FC06FC11" blockId="9.[327,1317,701,1883]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1E8FC21FC06FC11" bold="true" pageId="9" pageNumber="510">S5106 S5373</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1E1FC16FC00FAF3" blockId="9.[327,1317,701,1883]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1E1FC16FC6FFC50" bold="true" box="[915,978,953,1011]" pageId="9" pageNumber="510">
<accessionNumber id="9448AB63FFA87D68B1E1FC16FC7AFC64" box="[915,967,953,967]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10575" pageId="9" pageNumber="510" type="EnaNcbi">S10575</accessionNumber>
S3422 S3308
</emphasis>
<geoCoordinate id="EE2F5047FFA87D68B0F4FC5EFC13FBAA" box="[646,942,1009,1033]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-1.35">
<emphasis id="B96FEA92FFA87D68B0F4FC5EFC13FBAA" bold="true" box="[646,942,1009,1033]" pageId="9" pageNumber="510">1.35 S</emphasis>
</geoCoordinate>
3264 
<emphasis id="B96FEA92FFA87D68B01EFBBBFC6BFBBC" bold="true" box="[620,982,1041,1060]" pageId="9" pageNumber="510">
0.75 - 
<geoCoordinate id="EE2F5047FFA87D68B0E8FBBBFD04FB87" box="[666,697,1044,1060]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-2.08">2.08</geoCoordinate>
S5142
</emphasis>
<geoCoordinate id="EE2F5047FFA87D68B142FB8AFC04FB97" box="[816,953,1061,1077]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.5">
<emphasis id="B96FEA92FFA87D68B142FB8AFC04FB97" bold="true" box="[816,953,1061,1077]" pageId="9" pageNumber="510">0.50 S</emphasis>
</geoCoordinate>
3347 
<accessionNumber id="9448AB63FFA87D68B1BAFB93FC41FBE9" box="[968,1020,1084,1098]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10574" pageId="9" pageNumber="510" type="EnaNcbi">S10574</accessionNumber>
<emphasis id="B96FEA92FFA87D68B15CFBE7FC5CFBC3" bold="true" box="[814,993,1096,1120]" pageId="9" pageNumber="510">0.22 - S3444</emphasis>
<geoCoordinate id="EE2F5047FFA87D68B15CFBF2FC13FBD5" box="[814,942,1117,1142]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.83">
<emphasis id="B96FEA92FFA87D68B15CFBF2FC13FBD5" bold="true" box="[814,942,1117,1142]" pageId="9" pageNumber="510">0.83 S</emphasis>
</geoCoordinate>
5234 
<accessionNumber id="9448AB63FFA87D68B1B7FBD1FC45FB2F" box="[965,1016,1150,1164]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10629" pageId="9" pageNumber="510" type="EnaNcbi">S10629</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1C6FB3BFC55FB01" box="[948,1000,1172,1186]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10572" pageId="9" pageNumber="510" type="EnaNcbi">S10572</accessionNumber>
S3281 
<accessionNumber id="9448AB63FFA87D68B1C1FB10FC5AFB6E" box="[947,999,1215,1229]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10573" pageId="9" pageNumber="510" type="EnaNcbi">S10573</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1ABFB7AFBB1FB40" box="[985,1036,1237,1251]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10576" pageId="9" pageNumber="510" type="EnaNcbi">S10576</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1A9FB44FBB3FB5A" box="[987,1038,1259,1273]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10803" pageId="9" pageNumber="510" type="EnaNcbi">S10803</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1BCFAAEFBBCFAAC" box="[974,1025,1281,1295]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10579" pageId="9" pageNumber="510" type="EnaNcbi">S10579</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1C5FAB9FC56FA87" box="[951,1003,1302,1316]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10577" pageId="9" pageNumber="510" type="EnaNcbi">S10577</accessionNumber>
<accessionNumber id="9448AB63FFA87D68B1B1FA83FC4AFA99" box="[963,1015,1324,1338]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10578" pageId="9" pageNumber="510" type="EnaNcbi">S10578</accessionNumber>
<taxonomicName id="4C1B4D03FFA87D68B6E5FA8DFA98FA96" box="[1175,1317,1313,1333]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="510" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFA87D68B6E5FA8DFA98FA96" bold="true" box="[1175,1317,1313,1333]" italics="true" pageId="9" pageNumber="510">H. homoeolepis</emphasis>
</taxonomicName>
<emphasis id="B96FEA92FFA87D68B1E1FAEDFC00FAF3" bold="true" box="[915,957,1346,1360]" pageId="9" pageNumber="510">S5624</emphasis>
</paragraph>
<paragraph id="8BA43680FFA87D68B1ECFAF7FC6DF8F8" blockId="9.[327,1317,701,1883]" pageId="9" pageNumber="510">
<emphasis id="B96FEA92FFA87D68B1ECFAF7FC63FADF" bold="true" pageId="9" pageNumber="510">S5060 24Hhomo</emphasis>
<emphasis id="B96FEA92FFA87D68B65DFAC2FC75FA32" bold="true" pageId="9" pageNumber="510">
<collectingCountry id="F30C7610FFA87D68B65DFAC2FBD8FADE" box="[1071,1125,1389,1405]" name="Yemen" pageId="9" pageNumber="510">Socotra</collectingCountry>
S3289
</emphasis>
<emphasis id="B96FEA92FFA87D68B65DFA2DFC69FA04" bold="true" pageId="9" pageNumber="510">
<collectingCountry id="F30C7610FFA87D68B65DFA2DFBE3FA31" box="[1071,1118,1410,1426]" name="Iceland" pageId="9" pageNumber="510">Island</collectingCountry>
S5648
</emphasis>
<geoCoordinate id="EE2F5047FFA87D68B154FA09FC02FA1E" box="[806,959,1446,1469]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.52">
<emphasis id="B96FEA92FFA87D68B154FA09FC02FA1E" bold="true" box="[806,959,1446,1469]" pageId="9" pageNumber="510">0.52 S</emphasis>
</geoCoordinate>
3398 
<emphasis id="B96FEA92FFA87D68B156FA66FC64FA70" bold="true" box="[804,985,1477,1497]" pageId="9" pageNumber="510">0.29 - S5430</emphasis>
<geoCoordinate id="EE2F5047FFA87D68B156FA71FC0AFA4A" box="[804,951,1499,1518]" direction="south" orientation="latitude" pageId="9" pageNumber="510" precision="555" value="-0.82">
<emphasis id="B96FEA92FFA87D68B156FA71FC0AFA4A" bold="true" box="[804,951,1499,1518]" pageId="9" pageNumber="510">0.82 S</emphasis>
</geoCoordinate>
5097 S3380 
<accessionNumber id="9448AB63FFA87D68B1D9F9A9FC62F9B7" box="[939,991,1542,1556]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10630" pageId="9" pageNumber="510" type="EnaNcbi">S10630</accessionNumber>
SPM002057 
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S3807 SPM001511 S3257 
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<emphasis id="B96FEA92FFA87D68B2E5F82EFEB3F835" bold="true" box="[151,270,1921,1943]" pageId="9" pageNumber="510">FIGURE 2</emphasis>
. Maximum likelihood phylogenetic tree rooted with 
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<emphasis id="B96FEA92FFA87D68B15DF82DFC68F834" box="[815,981,1921,1943]" italics="true" pageId="9" pageNumber="510">H. masirahensis</emphasis>
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(outgroups not shown). Black dots on the nodes indicate ML bootstrap values ≥ 70 and BI posterior probabilities ≥ 0.95. Mean ages and 95% highest posterior density intervals of the nodes estimated with BEAST dating analysis are respectively indicated above and below the nodes. Clades are colored according to their geographic location marked on the map depicted in Fig. 1.
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<figureCitation id="13202A05FFAB7D6BB351FF6AFE1CFF47" box="[291,417,197,228]" captionStart="FIGURE 3" captionStartId="10.[151,250,1921,1943]" captionText="FIGURE 3. Parsimony networks corresponding to RAG 1, MC 1 R and c-mos nDNA sequence variation. Lines represent a mutational step, circles haplotypes and dots unsampled haplotypes. The size of circles is proportional to the number of individuals. Samples from the same species are similarly colored but with different tones for different geographic locations. For correspondences of sample and location codes see Appendix I." pageId="10" pageNumber="511">Figure 3</figureCitation>
RAG1
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S3260 a S5234 b S3260 b S3419 a S3419 b 
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<paragraph id="8BA43680FFAB7D6BB672F9A6FBB8F9B1" blockId="10.[365,1237,1366,1868]" box="[1024,1029,1545,1554]" pageId="10" pageNumber="511">S</paragraph>
<paragraph id="8BA43680FFAB7D6BB677FA52FBA4F9A5" blockId="10.[365,1237,1366,1868]" box="[1029,1049,1533,1542]" pageId="10" pageNumber="511">3419</paragraph>
<paragraph id="8BA43680FFAB7D6BB677F9A6FB9DF9B1" blockId="10.[365,1237,1366,1868]" box="[1029,1056,1545,1554]" pageId="10" pageNumber="511">3422 a</paragraph>
<paragraph id="8BA43680FFAB7D6BB66EFA52FB9CF9A5" blockId="10.[365,1237,1366,1868]" box="[1052,1057,1533,1542]" pageId="10" pageNumber="511">b</paragraph>
<paragraph id="8BA43680FFAB7D6BB63AF9A6FBF0F9B1" blockId="10.[365,1237,1366,1868]" box="[1096,1101,1545,1554]" pageId="10" pageNumber="511">S</paragraph>
<paragraph id="8BA43680FFAB7D6BB63EFA52FBECF9A5" blockId="10.[365,1237,1366,1868]" box="[1100,1105,1533,1542]" pageId="10" pageNumber="511">S</paragraph>
<paragraph id="8BA43680FFAB7D6BB63FF9A6FBDBF9B1" blockId="10.[365,1237,1366,1868]" box="[1101,1126,1545,1554]" pageId="10" pageNumber="511">10576</paragraph>
<paragraph id="8BA43680FFAB7D6BB623FA52FBD8F9A5" blockId="10.[365,1237,1366,1868]" box="[1105,1125,1533,1542]" pageId="10" pageNumber="511">5648</paragraph>
<paragraph id="8BA43680FFAB7D6BB61AF9A6FBD1F9B1" blockId="10.[365,1237,1366,1868]" box="[1128,1132,1545,1554]" pageId="10" pageNumber="511">a</paragraph>
<paragraph id="8BA43680FFAB7D6BB61AFA52FBD0F9A5" blockId="10.[365,1237,1366,1868]" box="[1128,1133,1533,1542]" pageId="10" pageNumber="511">b</paragraph>
<paragraph id="8BA43680FFAB7D6BB3B4F9A1FCC1F9B6" blockId="10.[365,1237,1366,1868]" box="[454,892,1548,1559]" pageId="10" pageNumber="511">S7676 b S2831 b</paragraph>
<paragraph id="8BA43680FFAB7D6BB029F9BCFD73F9BF" blockId="10.[365,1237,1366,1868]" box="[603,718,1555,1564]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB029F9BCFDD7F9BF" box="[603,618,1555,1564]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
119 b S7909 b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3B4F9B5FCC6F982" blockId="10.[365,1237,1366,1868]" box="[454,891,1560,1571]" pageId="10" pageNumber="511">S7924 b S3562 a</paragraph>
<paragraph id="8BA43680FFAB7D6BB672F9BAFBD0F9BD" blockId="10.[365,1237,1366,1868]" box="[1024,1133,1557,1566]" pageId="10" pageNumber="511">
S3807 a 
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b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB02EF9B0FBD1F989" blockId="10.[365,1237,1366,1868]" box="[604,1132,1567,1581]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB02EF9B0FDD6F98B" box="[604,619,1567,1576]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
136 a S7924 a S3562 b S4209 a 
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a
</paragraph>
<paragraph id="8BA43680FFAB7D6BB029F984FBD0F9EE" blockId="10.[365,1237,1366,1868]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB029F984FDD7F997" box="[603,618,1579,1588]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
136 b S7929 a S5305 a S4209 b 
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b 
<collectingCountry id="F30C7610FFAB7D6BB013F999FDCDF99C" box="[609,624,1590,1599]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
81 a S7929 b S5305 b S5004 a 
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a 
<collectingCountry id="F30C7610FFAB7D6BB012F9EDFDD2F9E8" box="[608,623,1602,1611]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
81 b S7966 a S5189 a S5004 b 
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b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB013F9E1FBD1F9FA" blockId="10.[365,1237,1366,1868]" box="[609,1132,1614,1628]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB013F9E1FDCDF9F4" box="[609,624,1614,1623]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
85 a S5189 b S5060 a 
<accessionNumber id="9448AB63FFAB7D6BB63AF9FFFBDBF9FA" box="[1096,1126,1616,1625]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10630" pageId="10" pageNumber="511" type="EnaNcbi">S10630</accessionNumber>
a
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3B7F9F5FBD0F96C" blockId="10.[365,1237,1366,1868]" pageId="10" pageNumber="511">
S7966 b 
<collectingCountry id="F30C7610FFAB7D6BB012F9F5FDD2F9C0" box="[608,623,1626,1635]" name="Austria" pageId="10" pageNumber="511">AO</collectingCountry>
85 b S5060 b 
<accessionNumber id="9448AB63FFAB7D6BB635F9F3FBD8F9C6" box="[1095,1125,1628,1637]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10630" pageId="10" pageNumber="511" type="EnaNcbi">S10630</accessionNumber>
b JS75 a JS5 b S5097 a 
<accessionNumber id="9448AB63FFAB7D6BB63AF9C8FBDBF9D3" box="[1096,1126,1639,1648]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10631" pageId="10" pageNumber="511" type="EnaNcbi">S10631</accessionNumber>
a JS75 b JS6 b S5097 b 
<accessionNumber id="9448AB63FFAB7D6BB63AF9DCFBDBF9DF" box="[1096,1126,1651,1660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10687" pageId="10" pageNumber="511" type="EnaNcbi">S10687</accessionNumber>
a S7091 a JS8 a S3257 a S3314 a S5142 a 
<accessionNumber id="9448AB63FFAB7D6BB635F9D0FBD8F92B" box="[1095,1125,1663,1672]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10687" pageId="10" pageNumber="511" type="EnaNcbi">S10687</accessionNumber>
b S7091 b JS8 b S3257 b S3314 b S5142 b 
<accessionNumber id="9448AB63FFAB7D6BB63AF924FBDBF937" box="[1096,1126,1675,1684]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10745" pageId="10" pageNumber="511" type="EnaNcbi">S10745</accessionNumber>
a S7871 a S3260 a S3351 a S5234 a 
<accessionNumber id="9448AB63FFAB7D6BB635F939FBD8F93C" box="[1095,1125,1686,1695]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10745" pageId="10" pageNumber="511" type="EnaNcbi">S10745</accessionNumber>
b S7893 a S3260 b S3351 b S5234 b 24Hhomo a S7893 b S3264 a S3398 a S5430 a 24Hhomo b S3281 a S3399 a S5624 a SPM001511 a S3281 b S3399 b S5624 b SPM001511 b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB30AF94AFE14F950" blockId="10.[365,1237,1366,1868]" box="[376,425,1765,1779]" pageId="10" pageNumber="511">
<emphasis id="B96FEA92FFAB7D6BB30AF94AFE14F950" bold="true" box="[376,425,1765,1779]" pageId="10" pageNumber="511">Legend</emphasis>
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3FBF8AEFB1EF8A4" blockId="10.[365,1237,1366,1868]" box="[393,1187,1790,1805]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB3FBF8AEFE14F8AE" box="[393,425,1793,1805]" name="Oman" pageId="10" pageNumber="511">Oman</collectingCountry>
S3422 b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3FBF8BEFCECF881" blockId="10.[365,1237,1366,1868]" box="[393,849,1809,1826]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB3FBF8BEFE12F8BE" box="[393,431,1809,1821]" name="Yemen" pageId="10" pageNumber="511">Yemen</collectingCountry>
(mainland) S3398 b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3FBF88FFCECF88D" blockId="10.[365,1237,1366,1868]" box="[393,849,1824,1838]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB3FBF88FFE12F88F" box="[393,431,1824,1836]" name="Yemen" pageId="10" pageNumber="511">Yemen</collectingCountry>
(Darsa Island) S5430 b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB63EF885FBD0F890" blockId="10.[365,1237,1366,1868]" box="[1100,1133,1834,1843]" pageId="10" pageNumber="511">S3807 b</paragraph>
<paragraph id="8BA43680FFAB7D6BB3FBF89FFCECF89A" blockId="10.[365,1237,1366,1868]" box="[393,849,1840,1852]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB3FBF89FFE12F89F" box="[393,431,1840,1852]" name="Yemen" pageId="10" pageNumber="511">Yemen</collectingCountry>
(Samha Island) 
<accessionNumber id="9448AB63FFAB7D6BB15EF89FFCF7F89A" box="[812,842,1840,1849]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S10631" pageId="10" pageNumber="511" type="EnaNcbi">S10631</accessionNumber>
b
</paragraph>
<paragraph id="8BA43680FFAB7D6BB3FBF8EFFDB5F8EF" blockId="10.[365,1237,1366,1868]" box="[393,520,1856,1868]" pageId="10" pageNumber="511">
<collectingCountry id="F30C7610FFAB7D6BB3FBF8EFFE12F8EF" box="[393,431,1856,1868]" name="Yemen" pageId="10" pageNumber="511">Yemen</collectingCountry>
(
<collectingCountry id="F30C7610FFAB7D6BB3C4F8EFFE62F8EF" box="[438,479,1856,1868]" name="Yemen" pageId="10" pageNumber="511">Socotra</collectingCountry>
Island)
</paragraph>
</subSubSection>
<caption id="DF646608FFAB7D6BB2E5F82EFCACF850" pageId="10" pageNumber="511">
<paragraph id="8BA43680FFAB7D6BB2E5F82EFCACF850" blockId="10.[151,1436,1921,2035]" pageId="10" pageNumber="511">
<emphasis id="B96FEA92FFAB7D6BB2E5F82EFEADF835" bold="true" box="[151,272,1921,1943]" pageId="10" pageNumber="511">FIGURE 3</emphasis>
. Parsimony networks corresponding to RAG1, MC1R and c-mos nDNA sequence variation. Lines represent a mutational step, circles haplotypes and dots unsampled haplotypes. The size of circles is proportional to the number of individuals. Samples from the same species are similarly colored but with different tones for different geographic locations. For correspondences of sample and location codes see Appendix I.
</paragraph>
</caption>
<caption id="DF646608FFAA7D6AB2E5FAA5FB0DFAFE" httpUri="https://zenodo.org/record/228116/files/figure.png" pageId="11" pageNumber="512" targetBox="[196,1375,263,1264]" targetPageId="11">
<paragraph id="8BA43680FFAA7D6AB2E5FAA5FB0DFAFE" blockId="11.[151,1436,1290,1373]" pageId="11" pageNumber="512">
<emphasis id="B96FEA92FFAA7D6AB2E5FAA5FEB1FABC" bold="true" box="[151,268,1290,1312]" pageId="11" pageNumber="512">FIGURE 4</emphasis>
. Size and shape differences between 
<taxonomicName id="4C1B4D03FFAA7D6AB0F4FAA5FCD5FA83" box="[646,872,1290,1312]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAA7D6AB0F4FAA5FCD5FA83" box="[646,872,1290,1312]" italics="true" pageId="11" pageNumber="512">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAA7D6AB11CFAA5FC02FABC" box="[878,959,1290,1311]" pageId="11" pageNumber="512" rank="species">
<emphasis id="B96FEA92FFAA7D6AB11CFAA5FC02FABC" bold="true" box="[878,959,1290,1311]" pageId="11" pageNumber="512">sp. nov.</emphasis>
</taxonomicNameLabel>
(blue lines and squares) from mainland Arabia and 
<taxonomicName id="4C1B4D03FFAA7D6AB2B1FA85FED9FA9D" box="[195,356,1320,1343]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFAA7D6AB2B1FA85FED9FA9D" box="[195,356,1320,1343]" italics="true" pageId="11" pageNumber="512">H. homoeolepis</emphasis>
</taxonomicName>
from Socotra (red lines and rectangles). The total contribution of the first three components of the PCA analysis to explain the total morphological variation is also given. See material and methods for details.
</paragraph>
</caption>
<subSubSection id="C301650BFFAA7D6AB2B5FA25FBFBF839" pageId="11" pageNumber="512" type="reference_group">
<paragraph id="8BA43680FFAA7D6AB2B5FA25FCE1F93D" blockId="11.[151,1436,1418,2019]" pageId="11" pageNumber="512">
<taxonomicName id="4C1B4D03FFAA7D6AB2B5FA25FE7CFA01" box="[199,449,1418,1443]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAA7D6AB2B5FA25FE7CFA01" box="[199,449,1418,1443]" italics="true" pageId="11" pageNumber="512">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAA7D6AB3BBFA25FD9EFA00" box="[457,547,1418,1443]" pageId="11" pageNumber="512" rank="species">
<emphasis id="B96FEA92FFAA7D6AB3BBFA25FD9EFA00" bold="true" box="[457,547,1418,1443]" pageId="11" pageNumber="512">sp. nov.</emphasis>
</taxonomicNameLabel>
is clearly differentiated from sympatric or geographically close populations of 
<taxonomicName id="4C1B4D03FFAA7D6AB2E5FA00FDDDFA65" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[151,608,1453,1478]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="hajarensis">
<emphasis id="B96FEA92FFAA7D6AB2E5FA00FE92FA65" box="[151,303,1453,1478]" italics="true" pageId="11" pageNumber="512">H. hajarensis</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB34AFA01FDDDFA65" author="Carranza" box="[312,608,1453,1478]" pageId="11" pageNumber="512" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C1B4D03FFAA7D6AB01EFA00FB97FA65" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[620,1066,1453,1478]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="alkiyumii">
<emphasis id="B96FEA92FFAA7D6AB01EFA00FD44FA65" box="[620,761,1453,1478]" italics="true" pageId="11" pageNumber="512">H. alkiyumii</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB170FA01FB97FA65" author="Carranza" box="[770,1066,1453,1478]" pageId="11" pageNumber="512" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold, 2012</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C1B4D03FFAA7D6AB645FA00FF6FFA49" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="festivus">
<emphasis id="B96FEA92FFAA7D6AB645FA00FB12FA65" box="[1079,1199,1454,1478]" italics="true" pageId="11" pageNumber="512">H. festivus</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB6CAFA01FF6FFA49" author="Carranza" pageId="11" pageNumber="512" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold, 2012</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C1B4D03FFAA7D6AB37FFA7CFD8EFA49" authority="Heyden, 1827" authorityName="Heyden" authorityYear="1827" box="[269,563,1490,1515]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B96FEA92FFAA7D6AB37FFA7CFE32FA48" box="[269,399,1490,1515]" italics="true" pageId="11" pageNumber="512">H. robustus</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB3E7FA7DFD8EFA49" author="Heyden" box="[405,563,1490,1515]" pageId="11" pageNumber="512" refString="Heyden, C. H. G. von (1827) Atlas zu der Reise im nordlichen Afrika von Eduard Ruppel. Reptilien. Bronner, Frankfurt am Main, 24 pp." type="book" year="1827">Heyden, 1827</bibRefCitation>
</taxonomicName>
by its smaller adult body size and absence of enlarged tubercles anywhere on the body (all four species with maximum SVL larger than 
<quantity id="4CE39B65FFAA7D6AB13DFA5AFC04F9AD" box="[847,953,1525,1550]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="11" pageNumber="512" unit="mm" value="34.6">34.6 mm</quantity>
, and with conical or strongly keeled and extensive tubercles on the body, nape hind legs and tail); from 
<taxonomicName id="4C1B4D03FFAA7D6AB124F9B4FB35F991" authority="Arnold, 1980" authorityName="Arnold" authorityYear="1980" box="[854,1160,1562,1587]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="lemurinus">
<emphasis id="B96FEA92FFAA7D6AB124F9B4FC54F990" box="[854,1001,1562,1587]" italics="true" pageId="11" pageNumber="512">H. lemurinus</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB182F9B5FB35F991" author="Arnold" box="[1008,1160,1562,1587]" pageId="11" pageNumber="512" refString="Arnold, E. N. (1980) The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. Journal of Oman Studies Special Report, 2, 273 - 332." type="journal article" year="1980">Arnold, 1980</bibRefCitation>
</taxonomicName>
, and from 
<taxonomicName id="4C1B4D03FFAA7D6AB772F9B4FE85F9F5" authority="Ruppell, 1835" authorityName="Ruppell" authorityYear="1835" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="flaviviridis">
<emphasis id="B96FEA92FFAA7D6AB772F9B4FA21F990" box="[1280,1436,1562,1587]" italics="true" pageId="11" pageNumber="512">H. flaviviridis</emphasis>
<bibRefCitation id="EF8A4B71FFAA7D6AB2E5F992FE85F9F5" author="Ruppell" box="[151,312,1597,1622]" pageId="11" pageNumber="512" refString="Ruppell, E. (1835) Neue Wirbelthiere zu der Fauna von Abyssinien gehorig. Vol. 3. Amphibien. S. Schmerber, Frankfurt am Main, 18 pp." type="book" year="1835">Rüppell, 1835</bibRefCitation>
</taxonomicName>
by its smaller adult body size (maximum SVL larger than 
<quantity id="4CE39B65FFAA7D6AB199F992FBEFF9F5" box="[1003,1106,1597,1622]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="11" pageNumber="512" unit="mm" value="34.6">34.6 mm</quantity>
in both species), absence of femoral pores (4–14 femoral pores on the underside of each thigh in 
<taxonomicName id="4C1B4D03FFAA7D6AB1D2F9CCFB81F9D8" box="[928,1084,1634,1659]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="flaviviridis">
<emphasis id="B96FEA92FFAA7D6AB1D2F9CCFB81F9D8" box="[928,1084,1634,1659]" italics="true" pageId="11" pageNumber="512">H. flaviviridis</emphasis>
</taxonomicName>
), and fewer lamellae under the first toe of pes (counts higher than four or five in both species).
</paragraph>
<paragraph id="8BA43680FFAA7D6AB2B5F905FBFBF839" blockId="11.[151,1436,1418,2019]" pageId="11" pageNumber="512">
<emphasis id="B96FEA92FFAA7D6AB2B5F905FD23F960" bold="true" box="[199,670,1706,1731]" pageId="11" pageNumber="512">Genetic and phylogeographic remarks.</emphasis>
The phylogenetic analyses by 
<bibRefCitation id="EF8A4B71FFAA7D6AB670F905FA89F961" author="Carranza" box="[1026,1332,1706,1731]" pageId="11" pageNumber="512" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold (2012)</bibRefCitation>
, Gómez- Díaz 
<emphasis id="B96FEA92FFAA7D6AB2A6F960FEB3F945" box="[212,270,1741,1766]" italics="true" pageId="11" pageNumber="512">et al.</emphasis>
(2012), and 
<bibRefCitation id="EF8A4B71FFAA7D6AB3D1F962FD3DF945" box="[419,640,1741,1766]" pageId="11" pageNumber="512" refString="Smid, J., Carranza, S., Kratochvil, L., Gvozdik, V., Nasher, A. K. &amp; Moravec, J. (2013 a) Out of Arabia: A complex biogeographic history of multiple vicariance and dispersal events in the gecko genus Hemidactylus (Reptilia: Gekkonidae). PloS one, 8 (5), e 64018. http: // dx. doi. org / 10.1371 / journal. pone. 0064018" type="journal article">
Šmíd 
<emphasis id="B96FEA92FFAA7D6AB394F960FDA7F945" box="[486,538,1741,1766]" italics="true" pageId="11" pageNumber="512">et al</emphasis>
. (2013a)
</bibRefCitation>
clearly indicate that 
<taxonomicName id="4C1B4D03FFAA7D6AB100F960FC52F945" box="[882,1007,1743,1766]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAA7D6AB100F960FC52F945" box="[882,1007,1743,1766]" italics="true" pageId="11" pageNumber="512">H. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAA7D6AB185F962FBEEF945" box="[1015,1107,1741,1766]" pageId="11" pageNumber="512" rank="species">
<emphasis id="B96FEA92FFAA7D6AB185F962FBEEF945" bold="true" box="[1015,1107,1741,1766]" pageId="11" pageNumber="512">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="4C1B4D03FFAA7D6AB6FEF960FA83F945" box="[1164,1342,1741,1766]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="512" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFAA7D6AB6FEF960FA83F945" box="[1164,1342,1741,1766]" italics="true" pageId="11" pageNumber="512">H. homoeolepis</emphasis>
</taxonomicName>
are two genetically well differentiated sister taxa. The phylogenetic analyses performed in this study support the hypothesis that these species are reciprocally monophyletic (
<figureCitation id="13202A05FFAA7D6AB0BFF8BAFCAEF88D" box="[717,787,1813,1838]" captionStart="FIGURE 2" captionStartId="9.[151,250,1921,1943]" captionText="FIGURE 2. Maximum likelihood phylogenetic tree rooted with H. masirahensis and H. inexpectatus (outgroups not shown). Black dots on the nodes indicate ML bootstrap values ≥ 70 and BI posterior probabilities ≥ 0.95. Mean ages and 95 % highest posterior density intervals of the nodes estimated with BEAST dating analysis are respectively indicated above and below the nodes. Clades are colored according to their geographic location marked on the map depicted in Fig. 1." pageId="11" pageNumber="512">Fig. 2</figureCitation>
) and present a high level of genetic divergence between them in the mitochondrial markers: 
<emphasis id="B96FEA92FFAA7D6AB030F894FDE4F8F1" box="[578,601,1851,1874]" italics="true" pageId="11" pageNumber="512">p-</emphasis>
distance (12S, cyt 
<emphasis id="B96FEA92FFAA7D6AB14AF895FCFBF8F0" box="[824,838,1850,1875]" italics="true" pageId="11" pageNumber="512">b</emphasis>
, ND4) = 6.5 ± 1.1, 11.0 ± 1.5, and 11.5 ± 1.3%, respectively (Table 1). In addition, the network analyses also depict no haplotype sharing between the two species in two of the three nuclear markers and only one haplotype shared in c-mos (
<figureCitation id="13202A05FFAA7D6AB181F82DFB85F839" box="[1011,1080,1922,1947]" captionStart="FIGURE 3" captionStartId="10.[151,250,1921,1943]" captionText="FIGURE 3. Parsimony networks corresponding to RAG 1, MC 1 R and c-mos nDNA sequence variation. Lines represent a mutational step, circles haplotypes and dots unsampled haplotypes. The size of circles is proportional to the number of individuals. Samples from the same species are similarly colored but with different tones for different geographic locations. For correspondences of sample and location codes see Appendix I." pageId="11" pageNumber="512">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C301650BFFAA7D6FB2B5F80AFE16FC6B" lastPageId="14" lastPageNumber="515" pageId="11" pageNumber="512" type="description">
<paragraph id="8BA43680FFAA7D6AB2B5F80AFD63F841" blockId="11.[151,1436,1418,2019]" pageId="11" pageNumber="512">
<emphasis id="B96FEA92FFAA7D6AB2B5F80AFDABF81D" bold="true" box="[199,534,1957,1982]" pageId="11" pageNumber="512">
Description of the 
<typeStatus id="54A08822FFAA7D6AB3DBF80AFDAFF81D" box="[425,530,1957,1982]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
.
</emphasis>
NHMUK2013.901. Data on 10 morphometric, and five meristic variables (see Material and Methods) are provided in Appendix II.
</paragraph>
<caption id="DF646608FFAD7D6DB2E5F8A6FCD4F856" httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="12" pageNumber="513" targetBox="[220,1356,204,1774]" targetPageId="12">
<paragraph id="8BA43680FFAD7D6DB2E5F8A6FCD4F856" blockId="12.[151,1437,1801,2037]" pageId="12" pageNumber="513">
<emphasis id="B96FEA92FFAD7D6DB2E5F8A6FEB3F8BD" bold="true" box="[151,270,1801,1823]" pageId="12" pageNumber="513">FIGURE 5</emphasis>
. Type localities, general dorsal views and details of 
<taxonomicName id="4C1B4D03FFAD7D6DB15BF8A6FBB1F8BC" box="[809,1036,1801,1823]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAD7D6DB15BF8A6FBB1F8BC" box="[809,1036,1801,1823]" italics="true" pageId="12" pageNumber="513">Hemidactylus minutus</emphasis>
</taxonomicName>
<emphasis id="B96FEA92FFAD7D6DB666F8A6FBD8F8BD" bold="true" box="[1044,1125,1801,1822]" pageId="12" pageNumber="513">
<taxonomicNameLabel id="A25C57E9FFAD7D6DB666F8A6FBD8F8BD" box="[1044,1125,1801,1822]" pageId="12" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from mainland Arabia and 
<taxonomicName id="4C1B4D03FFAD7D6DB7F1F8A5FEAAF89E" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFAD7D6DB7F1F8A5FEAAF89E" italics="true" pageId="12" pageNumber="513">H. homoeolepis</emphasis>
</taxonomicName>
from Socotra. A) and C) type locality (Asylah, Oman) of 
<taxonomicName id="4C1B4D03FFAD7D6DB105F886FC55F89D" box="[887,1000,1833,1854]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAD7D6DB105F886FC55F89D" box="[887,1000,1833,1854]" italics="true" pageId="12" pageNumber="513">H. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAD7D6DB183F887FBFEF89E" box="[1009,1091,1832,1853]" pageId="12" pageNumber="513" rank="species">
<emphasis id="B96FEA92FFAD7D6DB183F887FBFEF89E" bold="true" box="[1009,1091,1832,1853]" pageId="12" pageNumber="513">sp. nov.</emphasis>
</taxonomicNameLabel>
; B) dorsal view of 
<taxonomicName id="4C1B4D03FFAD7D6DB762F888FF5AF8FF" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAD7D6DB762F888FF5AF8FF" italics="true" pageId="12" pageNumber="513">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAD7D6DB29CF8E8FE82F8FF" box="[238,319,1863,1884]" pageId="12" pageNumber="513" rank="species">
<emphasis id="B96FEA92FFAD7D6DB29CF8E8FE82F8FF" bold="true" box="[238,319,1863,1884]" pageId="12" pageNumber="513">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype (voucher code: NHMUK2013.901; sample code: S7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1st and 4th toes of the holotype; H) dorsal view of one of the two 
<taxonomicName id="4C1B4D03FFAD7D6DB7F1F8C9FEAAF839" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFAD7D6DB7F1F8C9FEAAF839" italics="true" pageId="12" pageNumber="513">H. homoeolepis</emphasis>
</taxonomicName>
syntypes (voucher code: original BMNH81.7.22.6 and after the II World War BMNH1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of 
<taxonomicName id="4C1B4D03FFAD7D6DB1D8F80CFBF7F81B" box="[938,1098,1954,1976]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="513" phylum="Chordata" rank="species" species="homoeolepis">
<emphasis id="B96FEA92FFAD7D6DB1D8F80CFBF7F81B" box="[938,1098,1954,1976]" italics="true" pageId="12" pageNumber="513">H. homoeolepis</emphasis>
</taxonomicName>
, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1st and 4th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen.
</paragraph>
</caption>
<paragraph id="8BA43680FFAC7D6CB2E5FB9AFC92FB49" blockId="13.[151,1437,1077,1258]" pageId="13" pageNumber="514">
<emphasis id="B96FEA92FFAC7D6CB2E5FB9AFEB0FBE8" bold="true" box="[151,269,1077,1099]" pageId="13" pageNumber="514">FIGURE 6</emphasis>
. General dorsal views and details of two possibly undescribed new species from mainland Arabia. A) dorsal view of the female 
<taxonomicName id="4C1B4D03FFAC7D6CB353FBFBFE11FBC9" box="[289,428,1108,1130]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="genus">
<emphasis id="B96FEA92FFAC7D6CB353FBFBFE11FBC9" box="[289,428,1108,1130]" italics="true" pageId="13" pageNumber="514">Hemidactylus</emphasis>
</taxonomicName>
sp. 
<specimenCount id="9D1DFD09FFAC7D6CB3A7FBFBFDE9FBC9" box="[469,596,1108,1130]" pageId="13" pageNumber="514" type="generic">12 specimen</specimenCount>
from Shaqra, 
<collectingCountry id="F30C7610FFAC7D6CB091FBFAFC94FBC9" box="[739,809,1109,1130]" name="Yemen" pageId="13" pageNumber="514">Yemen</collectingCountry>
(voucher code: BMNH1953.1.6.99); B), C), D) and E) details of the head, posterior dorsal part with large tubercles, absence of preanal pores, and lamellae of the 1st and 4th toes of the same specimen, respectively; F) dorsal view of the juvenile 
<taxonomicName id="4C1B4D03FFAC7D6CB0CBFB3BFCF9FB09" box="[697,836,1172,1194]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="514" phylum="Chordata" rank="genus">
<emphasis id="B96FEA92FFAC7D6CB0CBFB3BFCF9FB09" box="[697,836,1172,1194]" italics="true" pageId="13" pageNumber="514">Hemidactylus</emphasis>
</taxonomicName>
sp. 
<specimenCount id="9D1DFD09FFAC7D6CB11FFB3BFC51FB09" box="[877,1004,1172,1194]" pageId="13" pageNumber="514" type="generic">13 specimen</specimenCount>
from Khiyat, 
<collectingCountry id="F30C7610FFAC7D6CB608FB3BFB42FB09" box="[1146,1279,1172,1194]" name="Saudi Arabia" pageId="13" pageNumber="514">Saudi Arabia</collectingCountry>
(voucher code: BMNH1992.168); G), H), I), and J) details of the head, posterior dorsal part with no tubercles, absence of preanal pores, and lamellae of the 1st and 4th toes of the same specimen, respectively.
</paragraph>
<paragraph id="8BA43680FFAC7D6CB2B5FAB9FCC5F878" blockId="13.[151,1436,1302,2011]" pageId="13" pageNumber="514">
Adult male, small (SVL 
<quantity id="4CE39B65FFAC7D6CB3ACFAB8FDFAFA8C" box="[478,583,1303,1327]" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="13" pageNumber="514" unit="mm" value="32.8">32.8 mm</quantity>
), body distinctly depressed (
<figureCitation id="13202A05FFAC7D6CB1FBFAB9FC70FA8C" box="[905,973,1302,1327]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
B). Right thigh muscle partially removed for tissue sample; small patch of skin missing on the dorsal surface of the right posterior limb (Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB2E5FAF1FF44FAD4" box="[151,249,1374,1399]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99996" pageId="13" pageNumber="514" type="EnaNcbi">M99996</accessionNumber>
-– 
<accessionNumber id="9448AB63FFAC7D6CB37DFAF1FECCFAD4" box="[271,369,1374,1399]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99997" pageId="13" pageNumber="514" type="EnaNcbi">M99997</accessionNumber>
). Absence of enlarged tubercles; homogeneous small, granular, and generally round, not imbricate dorsal scaling on head, body, limbs, and tail (
<figureCitation id="13202A05FFAC7D6CB166FA2DFCE4FA38" box="[788,857,1410,1435]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
B; Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB667FA2CFBC4FA38" box="[1045,1145,1411,1435]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99994" pageId="13" pageNumber="514" type="EnaNcbi">M99994</accessionNumber>
). Heterogeneous, roughly hexagonal ventral scales; gular and anteriormost ventral scales small, juxtaposed or slightly imbricate, increasing in size, height, and degree of imbrication towards the pelvic area; scales on the sides of the body with dark spot in the middle (Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB3BAFA41FD91F9A4" box="[456,556,1518,1543]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99995" pageId="13" pageNumber="514" type="EnaNcbi">M99995</accessionNumber>
). Supralabials 9/9, infralabials 8/8 (Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB60AFA40FB64F9A4" box="[1144,1241,1519,1543]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99998" pageId="13" pageNumber="514" type="EnaNcbi">M99998</accessionNumber>
– 
<accessionNumber id="9448AB63FFAC7D6CB69AFA40FAF4F9A4" box="[1256,1353,1519,1543]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99999" pageId="13" pageNumber="514" type="EnaNcbi">M99999</accessionNumber>
). Head narrow (HW/SVL=0.2), pointed and elongated (HW/SVL=0.3). Round nostrils not protuberant, defined by rostral (entering broadly into lower nostril borders on both sides), first supralabial and three supranasals in contact with upper borders. Rostral approximately square with a median notch above. Inner supranasals in contact with rostral and separated from each other by one vertical column of two polygonal scales. 
<collectingCountry id="F30C7610FFAC7D6CB660F9D1FBF5F934" box="[1042,1096,1662,1687]" name="China" pageId="13" pageNumber="514">Chin</collectingCountry>
shield rhomboid, followed by two pairs of postmentals. Anterior postmentals large and very elongated, in wide contact behind mental, both in contact with the first and second infralabials; posterior postmentals smaller, both in contact with the second infralabials (Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB3B8F945FD93F8A0" box="[458,558,1770,1795]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99995" pageId="13" pageNumber="514" type="EnaNcbi">M99995</accessionNumber>
). Eye relatively large (ED/HL=0.3); ear opening oval to round (
<figureCitation id="13202A05FFAC7D6CB745F945FA3DF8A0" box="[1335,1408,1770,1795]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
D; Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB340F8A0FE2EF884" box="[306,403,1807,1831]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99998" pageId="13" pageNumber="514" type="EnaNcbi">M99998</accessionNumber>
– 
<accessionNumber id="9448AB63FFAC7D6CB3D0F8A0FDBEF884" box="[418,515,1807,1831]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99999" pageId="13" pageNumber="514" type="EnaNcbi">M99999</accessionNumber>
). Parietal and temporal region covered with scales of similar size and shape to others parts of the head. Lamellae under the first toe 5/5 (
<figureCitation id="13202A05FFAC7D6CB16AF89DFCE7F8E8" box="[792,858,1842,1867]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
G); lamellae under the fourth toe 8/8 (
<figureCitation id="13202A05FFAC7D6CB771F89DFAFBF8E8" box="[1283,1350,1842,1867]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
G). Tail complete, longer than body (TL/SVL=1.1); the continuous row of enlarged tile-like subcaudal scales begins about a quarter of the tail length behind the vent (
<figureCitation id="13202A05FFAC7D6CB0FEF8D5FD6DF830" box="[652,720,1914,1939]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
E; Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB1FEF8D5FC4DF830" box="[908,1008,1914,1939]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99996" pageId="13" pageNumber="514" type="EnaNcbi">M99996</accessionNumber>
). Hemipenial swellings very obvious, with two visible elliptical openings below the vent; six pre-anal pores, PAP, in a V-shaped single row, no femoral pores or enlarged femoral scales (
<figureCitation id="13202A05FFAC7D6CB062F86DFDEFF878" box="[528,594,1986,2011]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="13" pageNumber="514">Fig. 5</figureCitation>
F; Morphobank 
<accessionNumber id="9448AB63FFAC7D6CB17AF86CFCD1F878" box="[776,876,1987,2011]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99997" pageId="13" pageNumber="514" type="EnaNcbi">M99997</accessionNumber>
).
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5FF38FAD0FF76" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
Color in life much richer than in the fixed specimen; very pale magenta-ochre, almost transparent dorsally, whiteyellowish on tail (
<figureCitation id="13202A05FFAF7D6FB32EFF13FE21FF76" box="[348,412,188,213]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="14" pageNumber="515">Fig. 5</figureCitation>
B). Iris in life colorful, golden with brown venations and an orange central vertical stripe.
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5FF70FCBAFE2B" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
Coloration in alcohol pale grey-buff or buff; a broad dark stripe from the nostril, through the eye, on to cheek and interrupted above ear; dorsal part of body, head, and limbs with irregular dark spots and streaks; belly pale (Morphobank 
<accessionNumber id="9448AB63FFAF7D6FB34BFE87FE21FEE3" box="[313,412,296,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99994" pageId="14" pageNumber="515" type="EnaNcbi">M99994</accessionNumber>
, 
<accessionNumber id="9448AB63FFAF7D6FB3DAFE87FDB5FEE3" box="[424,520,296,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99998" pageId="14" pageNumber="515" type="EnaNcbi">M99998</accessionNumber>
– 
<accessionNumber id="9448AB63FFAF7D6FB065FE87FDCAFEE3" box="[535,631,296,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99999" pageId="14" pageNumber="515" type="EnaNcbi">M99999</accessionNumber>
, 
<accessionNumber id="9448AB63FFAF7D6FB0F6FE88FD58FEE3" box="[644,741,295,320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99995" pageId="14" pageNumber="515" type="EnaNcbi">M99995</accessionNumber>
, respectively). Tail with ten dark bands, being narrower than pale intervening areas and increasing in intensity distally; ventral surface of tail pale with the three most distal dorsal bands extending on to it (Morphobank 
<accessionNumber id="9448AB63FFAF7D6FB0E5FEC0FD46FE2B" box="[663,763,367,392]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M99996" pageId="14" pageNumber="515" type="EnaNcbi">M99996</accessionNumber>
).
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5FE3BFD8CFC23" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
<emphasis id="B96FEA92FFAF7D6FB2B5FE3BFE86FE0E" bold="true" box="[199,315,404,429]" pageId="14" pageNumber="515">Variation.</emphasis>
Data for 10 morphometric, and five meristic characters (see Material and Methods) for all four 
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, NHMUK2013.902, NHMUK2013.903, NMP 
<date id="FFA51040FFAF7D6FB166FE18FC89FE73" box="[788,820,439,464]" pageId="14" pageNumber="515">
6 
<accessionNumber id="9448AB63FFAF7D6FB152FE18FC89FE73" box="[800,820,439,464]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="14" pageNumber="515" type="EnaNcbi">V</accessionNumber>
</date>
74885 and ONHM3716, are provided in Appendix II. All the specimens are very similar to each other, varying slightly in size related measurements, number of supralabials, infralabials and lamellae under the fourth toe (see Appendix II). Tails of 
<typeStatus id="54A08822FFAF7D6FB622FDAFFB01FDBB" box="[1104,1212,512,536]" pageId="14" pageNumber="515" type="paratype">paratypes</typeStatus>
NHMUK2013.902, and NHMUK2013.903 broken at the base, and of NMP 
<date id="FFA51040FFAF7D6FB17FFD8BFC93FD9E" box="[781,814,548,573]" pageId="14" pageNumber="515">
6 
<accessionNumber id="9448AB63FFAF7D6FB168FD8BFC93FD9E" box="[794,814,548,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="14" pageNumber="515" type="EnaNcbi">V</accessionNumber>
</date>
74885, and ONHM3716 almost at the base, but pieces are preserved intact together with the specimens (e.g. Morphobank M100008, M100006). A piece of tail of 
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NHMUK2013.903 and left forearms of 
<typeStatus id="54A08822FFAF7D6FB0B4FDC2FC8CFD26" box="[710,817,621,645]" pageId="14" pageNumber="515" type="paratype">paratypes</typeStatus>
NMP 
<date id="FFA51040FFAF7D6FB10AFDC3FC24FD26" box="[888,921,620,645]" pageId="14" pageNumber="515">
6 
<accessionNumber id="9448AB63FFAF7D6FB1F7FDC3FC24FD26" box="[901,921,620,645]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="14" pageNumber="515" type="EnaNcbi">V</accessionNumber>
</date>
74885 and ONHM3716 removed for genetic analyses. 
<typeStatus id="54A08822FFAF7D6FB37BFD3FFECBFD0B" box="[265,374,656,680]" pageId="14" pageNumber="515" type="paratype">Paratypes</typeStatus>
NHMUK2013.902 and NHMUK2013.903 with ventral side of tail irregularly stippled or blotched with undefined banding. 
<typeStatus id="54A08822FFAF7D6FB053FD1AFD3EFD6E" box="[545,643,693,717]" pageId="14" pageNumber="515" type="paratype">Paratype</typeStatus>
ONHM3716 with regenerated tail. 
<typeStatus id="54A08822FFAF7D6FB653FD1AFB33FD6E" box="[1057,1166,693,717]" pageId="14" pageNumber="515" type="paratype">Paratypes</typeStatus>
NHMUK2013.902 and NHMUK2013.903 with lateral dark stripe from the nostril continuous on to neck (Morphobank M100010, M100016). Tail of 
<typeStatus id="54A08822FFAF7D6FB306FD52FE69FCB6" box="[372,468,765,789]" pageId="14" pageNumber="515" type="paratype">paratype</typeStatus>
NMP 
<date id="FFA51040FFAF7D6FB06FFD53FD83FCB6" box="[541,574,764,789]" pageId="14" pageNumber="515">
6 
<accessionNumber id="9448AB63FFAF7D6FB058FD53FD83FCB6" box="[554,574,764,789]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="14" pageNumber="515" type="EnaNcbi">V</accessionNumber>
</date>
74885 with the four most distal dorsal bands extending on to ventral side (Morphobank M323685). Anterior right postmental of 
<typeStatus id="54A08822FFAF7D6FB140FC8FFC2FFC9B" box="[818,914,800,824]" pageId="14" pageNumber="515" type="paratype">paratype</typeStatus>
NMP 
<date id="FFA51040FFAF7D6FB192FCB0FBBCFC9B" box="[992,1025,799,824]" pageId="14" pageNumber="515">
6 
<accessionNumber id="9448AB63FFAF7D6FB19FFCB0FBBCFC9B" box="[1005,1025,799,824]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V74885" pageId="14" pageNumber="515" type="EnaNcbi">V</accessionNumber>
</date>
74885 only in contact with first infralabial; posterior left postmental of 
<typeStatus id="54A08822FFAF7D6FB03EFCEAFD10FCFE" box="[588,685,837,861]" pageId="14" pageNumber="515" type="paratype">paratype</typeStatus>
NHMUK 
<quantity id="4CE39B65FFAF7D6FB16CFCEBFC1FFCFE" box="[798,930,836,861]" metricMagnitude="1" metricUnit="m" metricValue="5.11531362" pageId="14" pageNumber="515" unit="in" value="2013.903">2013.903 in</quantity>
contact with the second and third infralabials (Morphobank M100040, M100013).
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5FC23FE16FC6B" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
Main coloration very similar to the 
<typeStatus id="54A08822FFAF7D6FB013FC23FD7BFC06" box="[609,710,908,933]" pageId="14" pageNumber="515" type="holotype">holotype</typeStatus>
, with NHMUK2013.902 and NHMUK2013.903 
<typeStatus id="54A08822FFAF7D6FB684FC22FADFFC06" box="[1270,1378,909,933]" pageId="14" pageNumber="515" type="paratype">paratypes</typeStatus>
with darker spots and streaks.
</paragraph>
</subSubSection>
<subSubSection id="C301650BFFAF7D6FB2B5FC7BFDDBF9F0" pageId="14" pageNumber="515" type="distribution">
<paragraph id="8BA43680FFAF7D6FB2B5FC7BFD07FB4B" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
<emphasis id="B96FEA92FFAF7D6FB2B5FC7BFEDDFC4E" bold="true" box="[199,352,980,1005]" pageId="14" pageNumber="515">Distribution.</emphasis>
Distributed along the Arabian Sea coast, from northeastern 
<collectingCountry id="F30C7610FFAF7D6FB652FC7BFBD8FC4F" box="[1056,1125,980,1004]" name="Oman" pageId="14" pageNumber="515">Oman</collectingCountry>
to extreme eastern 
<collectingCountry id="F30C7610FFAF7D6FB73CFC7AFA21FC4F" box="[1358,1436,981,1004]" name="Yemen" pageId="14" pageNumber="515">Yemen</collectingCountry>
(
<figureCitation id="13202A05FFAF7D6FB2EDFC58FF5EFBB3" box="[159,227,1015,1040]" captionStart="FIGURE 1" captionStartId="4.[151,250,1741,1763]" captionTargetBox="[160,1430,237,1715]" captionTargetId="figure@4.[153,1435,231,1719]" captionTargetPageId="4" captionText="FIGURE 1. Distribution map of the samples used in this study. Color dots indicate specimens included in the molecular analyses and listed in Appendix I. Color stars indicate voucher specimens examined and listed in Appendix II, most of which were included in the morphological analyses. The specimens’ location colors match the colors used in the following figures. Localities of the undescribed specimens Hemidactylus sp. 12 (BMNH 1953.1.6.99 from Shaqra, Yemen) and Hemidactylus sp. 13 (BMNH 1992.168 from Khiyat, Saudi Arabia) are also shown in the map." httpUri="https://zenodo.org/record/228115/files/figure.png" pageId="14" pageNumber="515">Fig. 1</figureCitation>
; Appendix I, II). In Dhofar it is found more than 
<quantity id="4CE39B65FFAF7D6FB17CFC57FCE8FBB3" box="[782,853,1015,1040]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="14" pageNumber="515" unit="km" value="70.0">70 km</quantity>
inland, as far as Thumrait. Although the population of 
<emphasis id="B96FEA92FFAF7D6FB2C9FBB3FDB3FB96" box="[187,526,1052,1077]" italics="true" pageId="14" pageNumber="515">
<taxonomicName id="4C1B4D03FFAF7D6FB2C9FBB3FEE8FB96" box="[187,341,1052,1077]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
“ 
<taxonomicName id="4C1B4D03FFAF7D6FB31BFBB3FDBBFB96" box="[361,518,1052,1077]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="homoeolepis">homoeolepis</taxonomicName>
”
</emphasis>
from the Hasikaya Island in the Hallaniyat Archipelago may belong to this species, a detailed genetic and morphological analysis is needed to assess whether this population is 
<taxonomicName id="4C1B4D03FFAF7D6FB68BFBEEFAC9FBFB" box="[1273,1396,1089,1112]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAF7D6FB68BFBEEFAC9FBFB" box="[1273,1396,1089,1112]" italics="true" pageId="14" pageNumber="515">H. minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAF7D6FB708FB90FF75FBDE" pageId="14" pageNumber="515" rank="species">
<emphasis id="B96FEA92FFAF7D6FB708FB90FF75FBDE" bold="true" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
, 
<taxonomicName id="4C1B4D03FFAF7D6FB2ABFBCAFE7AFBDE" box="[217,455,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="paucituberculatus">
<emphasis id="B96FEA92FFAF7D6FB2ABFBCAFE7AFBDE" box="[217,455,1124,1149]" italics="true" pageId="14" pageNumber="515">H. paucituberculatus</emphasis>
</taxonomicName>
, (also present in coastal Dhofar) or a new species. Previous reports of 
<emphasis id="B96FEA92FFAF7D6FB770FBCBFEF9FB03" italics="true" pageId="14" pageNumber="515">
<taxonomicName id="4C1B4D03FFAF7D6FB770FBCBFA21FBDE" box="[1282,1436,1124,1149]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
“ 
<taxonomicName id="4C1B4D03FFAF7D6FB2EDFB28FE81FB03" box="[159,316,1159,1184]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="homoeolepis">homoeolepis</taxonomicName>
”
</emphasis>
from Masirah Island and from Jazirat Hamar an Nafur Island have been recently assigned to 
<taxonomicName id="4C1B4D03FFAF7D6FB7F3FB26FE96FB66" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="masirahensis">
<emphasis id="B96FEA92FFAF7D6FB7F3FB26FE96FB66" italics="true" pageId="14" pageNumber="515">H. masirahensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1B4D03FFAF7D6FB312FB02FDB3FB67" box="[352,526,1197,1220]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="inexpectatus">
<emphasis id="B96FEA92FFAF7D6FB312FB02FDB3FB67" box="[352,526,1197,1220]" italics="true" pageId="14" pageNumber="515">H. inexpectatus</emphasis>
</taxonomicName>
, respectively (
<bibRefCitation id="EF8A4B71FFAF7D6FB0C3FB03FBD4FB66" author="Carranza" box="[689,1129,1196,1221]" pageId="14" pageNumber="515" refString="Carranza, S., Arnold, E. N. (2012) A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species. Zootaxa, 3378, 1 - 95." type="journal article" year="2012">Carranza &amp; Arnold 2012; pers. observ.</bibRefCitation>
). 
<taxonomicName id="4C1B4D03FFAF7D6FB609FB03FAC9FB67" box="[1147,1396,1196,1221]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAF7D6FB609FB03FAC9FB67" box="[1147,1396,1196,1221]" italics="true" pageId="14" pageNumber="515">Hemidactylus minutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25C57E9FFAF7D6FB708FB03FF75FB4B" pageId="14" pageNumber="515" rank="species">
<emphasis id="B96FEA92FFAF7D6FB708FB03FF75FB4B" bold="true" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
can be considered nearly endemic to 
<collectingCountry id="F30C7610FFAF7D6FB01CFB7FFD0BFB4B" box="[622,694,1232,1256]" name="Oman" pageId="14" pageNumber="515">Oman</collectingCountry>
.
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5FB5BFC6BF98C" blockId="14.[151,1437,151,1619]" pageId="14" pageNumber="515">
<emphasis id="B96FEA92FFAF7D6FB2B5FB5BFE3AFAAE" bold="true" box="[199,391,1268,1293]" pageId="14" pageNumber="515">Natural history.</emphasis>
The new species is a ground dwelling strictly nocturnal gecko, usually found in dry places with stony, gravely or even sandy substrates with rocky outcrops (
<figureCitation id="13202A05FFAF7D6FB1FAFAB8FC71FA93" box="[904,972,1303,1328]" captionStart="FIGURE 5" captionStartId="12.[151,250,1801,1823]" captionTargetBox="[220,1356,204,1774]" captionTargetId="figure@12.[219,1371,193,1775]" captionTargetPageId="12" captionText="FIGURE 5. Type localities, general dorsal views and details of Hemidactylus minutus sp. nov. from mainland Arabia and H. homoeolepis from Socotra. A) and C) type locality (Asylah, Oman) of H. minutus sp. nov.; B) dorsal view of Hemidactylus minutus sp. nov. holotype (voucher code: NHMUK 2013.901; sample code: S 7676); D), E), F) and G) details of the head, tail underside pattern, preanal pores, and lamellae of the 1 st and 4 th toes of the holotype; H) dorsal view of one of the two H. homoeolepis syntypes (voucher code: original BMNH 81.7.22.6 and after the II World War BMNH 1946.9.6.99), and I) of an unvouchered specimen (photograph by Edoardo Razzetti); J) habitat type of H. homoeolepis, generally dry places with rocky substrate; K), M) and N) details of the head, preanal pores, and lamellae of the 1 st and 4 th toes of the syntype, respectively; L) detail of the pattern of underside the tail of and unvouchered specimen." httpUri="https://zenodo.org/record/228117/files/figure.png" pageId="14" pageNumber="515">Fig. 5</figureCitation>
A, C). It is abundant in many parts of its distribution range. Observations of specimens of 
<taxonomicName id="4C1B4D03FFAF7D6FB0B0FA92FC83FAF7" box="[706,830,1341,1364]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAF7D6FB0B0FA92FC83FAF7" box="[706,830,1341,1364]" italics="true" pageId="14" pageNumber="515">H. minutus</emphasis>
</taxonomicName>
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<emphasis id="B96FEA92FFAF7D6FB137FA93FC22FAF6" bold="true" box="[837,927,1340,1365]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
carried out by 
<bibRefCitation id="EF8A4B71FFAF7D6FB639FA93FB50FAF7" author="Arnold" box="[1099,1261,1340,1365]" pageId="14" pageNumber="515" refString="Arnold, E. N. (1980) The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. Journal of Oman Studies Special Report, 2, 273 - 332." type="journal article" year="1980">Arnold (1980)</bibRefCitation>
at Wadi Ayoun and Thumrait indicated that almost all the specimens (63/64) were first sighted either on the ground (38/64) or lower than 
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from it (25/64), and only one was found on a rock above this height. At Wadi Sayq, eighteen individuals reported also by 
<bibRefCitation id="EF8A4B71FFAF7D6FB3ABFA08FDC7FA63" author="Arnold" box="[473,634,1447,1472]" pageId="14" pageNumber="515" refString="Arnold, E. N. (1980) The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. Journal of Oman Studies Special Report, 2, 273 - 332." type="journal article" year="1980">Arnold (1980)</bibRefCitation>
were at heights of between 
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and 
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on rock faces, but this may have been because the ground here was covered by dense vegetation following the monsoon. 
<taxonomicName id="4C1B4D03FFAF7D6FB6D1FA61FA21FA44" box="[1187,1436,1486,1511]" class="Reptilia" family="Gekkonidae" genus="Hemidacytlus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="515" phylum="Chordata" rank="species" species="minutus" status="sp. nov.">
<emphasis id="B96FEA92FFAF7D6FB6D1FA61FA21FA44" box="[1187,1436,1486,1511]" italics="true" pageId="14" pageNumber="515">Hemidacytlus minutus</emphasis>
</taxonomicName>
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<emphasis id="B96FEA92FFAF7D6FB2E5FA5DFF4FF9A8" bold="true" box="[151,242,1522,1547]" pageId="14" pageNumber="515">sp. nov.</emphasis>
</taxonomicNameLabel>
is very agile, often proceeding in a series of leaps when pursued. Gravid females, each carrying a single egg, have been recorded in late September at Khawr Sawli (
<bibRefCitation id="EF8A4B71FFAF7D6FB144F9B9FC77F98C" author="Arnold" box="[822,970,1558,1583]" pageId="14" pageNumber="515" refString="Arnold, E. N. (1980) The scientific results of the Oman flora and fauna survey 1977 (Dhofar). The reptiles and amphibians of Dhofar, southern Arabia. Journal of Oman Studies Special Report, 2, 273 - 332." type="journal article" year="1980">Arnold 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8BA43680FFAF7D6FB2B5F995FDDBF9F0" blockId="14.[151,1437,151,1619]" box="[199,614,1594,1619]" pageId="14" pageNumber="515">
<emphasis id="B96FEA92FFAF7D6FB2B5F995FE06F9F0" bold="true" box="[199,443,1594,1619]" pageId="14" pageNumber="515">Conservation status.</emphasis>
Not evaluated.
</paragraph>
</subSubSection>
</treatment>
</document>