238 lines
40 KiB
XML
238 lines
40 KiB
XML
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<document id="EE2847BB9C44D118884E59E6124BA581" ID-DOI="10.5281/zenodo.198863" ID-GBIF-Dataset="6f286e35-670c-4bb9-920b-604c51cecd9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="198863" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460315829380" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2010" docId="039F5848FF9FFFEAE5F3FC5AFF36C989" docLanguage="en" docName="zt02652p063.pdf" docOrigin="Zootaxa 2652" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Triacanthoneus pacificus Anker, 2010, n. sp." docType="treatment" docVersion="8" lastPageNumber="55" masterDocId="FFA62030FF98FFE2E564FFD8FFC6CE05" masterDocTitle="A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts" masterLastPageNumber="63" masterPageNumber="47" pageNumber="54" updateTime="1698255077161" updateUser="plazi">
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<mods:titleInfo id="AE6E1C731D2B11A2EE37BFA127DA12B7">
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<mods:title id="E6D72797EEB003FF4B15C6E628D5F83C">A new genus and three new species of alpheid shrimps (Crustacea, Decapoda, Caridea) from the tropical American coasts</mods:title>
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</mods:titleInfo>
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<mods:name id="C1604906F58DF9927EA6E75B1921797C" type="personal">
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<mods:roleTerm id="762C7D9B867BE577923571DD5D053163">Author</mods:roleTerm>
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<mods:namePart id="7B4CEF83E3EDDB77CB195745B74A05D8">Anker, Arthur</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7F516EB070116D7F032261B6FF5C034C">text</mods:typeOfResource>
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<mods:titleInfo id="F31A53AAE33D092F8130A667C5D7F1E7">
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<mods:title id="0672FBD187C9E5EA070B74D45152216B">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B27F6F2E33FD5C87DD1C0993A8A75DD3">
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<mods:date id="EA9E100EC7F8F28B77EFE7BEC2DD6F2A">2010</mods:date>
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<mods:detail id="73489A3BB10A4F8458A10731F3C9B664" type="volume">
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<mods:number id="F5E90DA6AE26CC11C352FAF0A8509770">2652</mods:number>
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</mods:detail>
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<mods:start id="560D6603A00D71C22CBE93C0E8599A92">47</mods:start>
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<mods:classification id="C9626C53ED854D3869AD371D4C2E6FEA">journal article</mods:classification>
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<mods:identifier id="D6CEBC68345FE71ADDFCDE088BF48829" type="DOI">10.5281/zenodo.198863</mods:identifier>
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<mods:identifier id="F10BEF346EAACE1FF3F18B2D98B780B6" type="GBIF-Dataset">6f286e35-670c-4bb9-920b-604c51cecd9f</mods:identifier>
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<mods:identifier id="79A0B20E5EF7099C72B7F7AA569515D1" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D101CD9F35F6BCDE1334C47686EAE426" type="Zenodo-Dep">198863</mods:identifier>
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</mods:mods>
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<treatment id="039F5848FF9FFFEAE5F3FC5AFF36C989" ID-DOI="http://doi.org/10.5281/zenodo.6206918" ID-GBIF-Taxon="119401709" ID-Zenodo-Dep="6206918" LSID="urn:lsid:plazi:treatment:039F5848FF9FFFEAE5F3FC5AFF36C989" httpUri="http://treatment.plazi.org/id/039F5848FF9FFFEAE5F3FC5AFF36C989" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
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<subSubSection id="C32CBAD5FF9FFFE5E5F3FC5AFEB1CDBA" pageId="7" pageNumber="54" type="nomenclature">
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<paragraph id="8B89E95EFF9FFFE5E5F3FC5AFDD2CD99" blockId="7.[151,532,898,959]" box="[151,532,898,924]" pageId="7" pageNumber="54">
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<heading id="D0C15E32FF9FFFE5E5F3FC5AFDD2CD99" bold="true" box="[151,532,898,924]" fontSize="11" level="1" pageId="7" pageNumber="54" reason="1">
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<emphasis id="B942354CFF9FFFE5E5F3FC5AFDD2CD99" bold="true" box="[151,532,898,924]" pageId="7" pageNumber="54">
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<taxonomicName id="4C3692DDFF9FFFE5E5F3FC5AFE0CCD99" ID-CoL="85ZFT" box="[151,458,898,924]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="54" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
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<emphasis id="B942354CFF9FFFE5E5F3FC5AFE0CCD99" bold="true" box="[151,458,898,924]" italics="true" pageId="7" pageNumber="54">Triacanthoneus pacificus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2718837FF9FFFE5E4B6FC5AFDD2CD99" box="[466,532,898,924]" pageId="7" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B89E95EFF9FFFE5E5F3FC7DFEB1CDBA" blockId="7.[151,532,898,959]" box="[151,375,933,959]" pageId="7" pageNumber="54">
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(
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<figureCitation id="130DF5DBFF9FFFE5E5FBFC7DFF34CDBA" box="[159,242,933,959]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Figs. 5</figureCitation>
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,
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<figureCitation id="130DF5DBFF9FFFE5E59BFC7DFEC9CDBA" box="[255,271,933,959]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="7" pageNumber="54">6</figureCitation>
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,
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<figureCitation id="130DF5DBFF9FFFE5E478FC7DFEFECDBA" box="[284,312,933,959]" captionStart="FIGURE 10" captionStartId="14.[151,256,1843,1867]" captionTargetBox="[151,1436,493,1821]" captionTargetId="figure@14.[151,1436,493,1822]" captionTargetPageId="14" captionText="FIGURE 10. Life colouration of three species of Triacanthoneus n. gen.: A – C, Triacanthoneus toro n. sp., paratype, ovigerous female from Bocas del Toro, Caribbean coast of Panama, USNM 1145376; D – F, Triacanthoneus pacificus n. sp., holotype, ovigerous female from Playa Venao, Pacific coast of Panama, USNM 1145374 [D, E], non-type ovigerous female from Bahía Malaga, Pacific coast of Colombia, UVC [F]; G, Triacanthoneus alacranes n. sp., holotype, ovigerous female from Alacranes Reef, Yucatán, Mexico, UNAM-CNCR 26177 [G by J. Duarte-Gutiérrez, all other photographs by author]." httpUri="https://zenodo.org/record/198873/files/figure.png" pageId="7" pageNumber="54">10</figureCitation>
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D–F)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32CBAD5FF9FFFE5E5F3FC37FB2CCAC1" pageId="7" pageNumber="54" type="materials_examined">
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<paragraph id="8B89E95EFF9FFFE5E5F3FC37FE7DCA51" blockId="7.[151,1437,1007,2004]" pageId="7" pageNumber="54">
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<emphasis id="B942354CFF9FFFE5E5F3FC37FE95CA0C" bold="true" box="[151,339,1007,1033]" pageId="7" pageNumber="54">
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<typeStatus id="548D57FCFF9FFFE5E5F3FC37FF10CA0C" box="[151,214,1007,1033]" pageId="7" pageNumber="54">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="548D57FCFF9FFFE5E438FC28FE17CA0F" box="[348,465,1008,1034]" pageId="7" pageNumber="54" type="holotype">Holotype</typeStatus>
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: ovigerous female (CL
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<quantity id="4CCE44BBFF9FFFE5E794FC28FC97CA0F" box="[752,849,1008,1034]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="54" unit="mm" value="2.8">2.8 mm</quantity>
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),
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<collectionCode id="ED27719BFF9FFFE5E603FC28FC7BCA0F" LSID="urn:lsid:biocol.org:col:34871" box="[871,957,1008,1034]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="54">USNM</collectionCode>
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1145374,
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<collectingCountry id="F321A9CEFF9FFFE5E15AFC28FB67CA0F" box="[1086,1185,1008,1034]" name="Panama" pageId="7" pageNumber="54">Panama</collectingCountry>
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, Pacific coast, Playa Venao, intertidal, exposed at low tide, under rocks in sand, coll. J.A. Baeza (processed by A. Anker),
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<date id="FF88CF9EFF9FFFE5E01AFBCDFEEFCA51" pageId="7" pageNumber="54" value="2007-03-18">18 March 2007</date>
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, fcn 07-090.
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</paragraph>
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<paragraph id="8B89E95EFF9FFFE5E5A2FB87FB2CCAC1" blockId="7.[151,1437,1007,2004]" pageId="7" pageNumber="54">
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<emphasis id="B942354CFF9FFFE5E5A2FB87FD81CA7C" bold="true" box="[198,583,1119,1145]" pageId="7" pageNumber="54">Additional material examined.</emphasis>
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2 ovigerous females (CL 2.9,
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<quantity id="4CCE44BBFF9FFFE5E6D2FBB8FBD0CA7F" box="[950,1046,1120,1146]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="54" unit="mm" value="3.4">3.4 mm</quantity>
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),
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<collectionCode id="ED27719BFF9FFFE5E148FBB8FBABCA7F" box="[1068,1133,1120,1146]" httpUri="http://grbio.org/cool/zh0r-sp5k" name="Coleccion de Anfibios y Reptiles, Universidad del Valle, Cali" pageId="7" pageNumber="54">UVC</collectionCode>
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,
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<collectingCountry id="F321A9CEFF9FFFE5E11EFBB8FB37CA7F" box="[1146,1265,1120,1146]" name="Colombia" pageId="7" pageNumber="54">Colombia</collectingCountry>
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, Pacific coast, Bahía Malaga, Los Negros, sand-rock bottom, depth
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<quantity id="4CCE44BBFF9FFFE5E66BFB5DFC86CA9A" box="[783,832,1157,1183]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="54" unit="m" value="2.0">2 m</quantity>
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,
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<collectionCode id="ED27719BFF9FFFE5E629FB5DFC67CA9A" box="[845,929,1157,1183]" pageId="7" pageNumber="54">ARMS</collectionCode>
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(large plastic basket with shells),
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<date id="FF88CF9EFF9FFFE5E05CFB5DFF13CAC1" pageId="7" pageNumber="54" value="2009-04-25">25 April 2009</date>
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, coll. J.F. Lazarus-Agudelo et al. (processed by A. Anker), fcn COL-00135, COL-00136.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32CBAD5FF9FFFEAE5A2FB17FEFFCB87" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54" type="description">
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<paragraph id="8B89E95EFF9FFFE5E5A2FB17FAFCCBEA" blockId="7.[151,1437,1007,2004]" pageId="7" pageNumber="54">
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<emphasis id="B942354CFF9FFFE5E5A2FB17FE99CAEC" bold="true" box="[198,351,1231,1257]" pageId="7" pageNumber="54">Description.</emphasis>
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Small-sized alpheid shrimp. Carapace with one strong sharp mediodorsal tooth situated distinctly posterior to carapace mid-length and two strong sharp dorsolateral teeth situated ventral and anterior to mediodorsal tooth, in post-hepatic area; stiff seta present on mid-dorsal line anterior to mediodorsal tooth (
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<figureCitation id="130DF5DBFF9FFFE5E5FBFA98FF23CB5F" box="[159,229,1344,1370]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
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B, C). Rostrum straight, more than 1.7 times as long as broad at base; lateral margins slightly concave; tip overreaching mid-length of second segment of antennular peduncle; ventral margin with strong, anteroventrally directed subapical dent; rostral carina well developed, reaching posteriorly to mediodorsal tooth, with one stiff seta approximately at carapace mid-length; orbital teeth obscure, present as small bumps lateral to eye (
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<figureCitation id="130DF5DBFF9FFFE5E45AFA0DFE45CBEA" box="[318,387,1493,1519]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
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A–E). Pterygostomial angle not protruding anteriorly, broadly rounded (
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<figureCitation id="130DF5DBFF9FFFE5E1B4FA0DFAD0CBEA" box="[1232,1302,1493,1519]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
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B).
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</paragraph>
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<paragraph id="8B89E95EFF9FFFE5E5A1FA22FE7AC8CA" blockId="7.[151,1437,1007,2004]" pageId="7" pageNumber="54">
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Pleura of first to fourth abdominal somites rounded posteroventrally; fifth pleuron with posteroventral angle subacute. Telson narrow, almost four times as long as wide in proximal third, tapering distally; dorsal surface with two pairs of strong spiniform setae situated at some distance from lateral margins, at about 0.5 and 0.75 telson length, respectively; posterior margin straight, each posterolateral angle with two pairs of spiniform setae, lateral being 0.7 times as long as mesial, and two long plumose setae between mesial spiniform setae (
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<figureCitation id="130DF5DBFF9FFFE5E43FF96DFE67C8CA" box="[347,417,1717,1743]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
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J).
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</paragraph>
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<paragraph id="8B89E95EFF9FFFE5E5A1F902FB0BC91F" blockId="7.[151,1437,1007,2004]" pageId="7" pageNumber="54">
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Eyes partly exposed in dorsal and lateral views; cornea somewhat reduced, well pigmented, occupying anterolateral portion of eyestalk, anteromesial margin with small blunt process (
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<figureCitation id="130DF5DBFF9FFFE5E15BF8D8FB42C91F" box="[1087,1156,1792,1818]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
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A, B).
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</paragraph>
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<paragraph id="8B89E95EFF9FFFEAE5A1F8FDFBB8CED2" blockId="7.[151,1437,1007,2004]" lastBlockId="8.[151,1437,152,1932]" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
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Antennular peduncle stout, with second segment distinctly longer than wide; first segment with distally acute stylocerite reaching to 0.4–0.5 length of second segment; ventromesial carina of first segment with anteriorly directed tooth; lateral flagellum biramous, fused portion composed of at least three segments; accessory ramus composed of four or so partly fused segments with groups of aesthetascs (
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<figureCitation id="130DF5DBFF9FFFE5E061F84DFA89C9AA" box="[1285,1359,1941,1967]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="7" pageNumber="54">Fig. 5</figureCitation>
|
|||
|
A, B). Antenna with stout basicerite bearing sharp distoventral tooth; scaphocerite reaching or almost reaching to end of antennular peduncle, ovate; blade broad, with anterior margin convex, reaching slightly beyond strong distolateral tooth; carpocerite barely reaching 0.5 length of scaphocerite (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE694FF65FBF3CED2" box="[1008,1077,189,215]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="8" pageNumber="55">Fig. 5</figureCitation>
|
|||
|
A, B).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FF3AFCC5CF69" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
Mouthparts not dissected, appearing typical for genus in external view. Third maxilliped with coxa bearing rounded lateral plate above mastigobranch; antepenultimate segment elongate, flattened; penultimate segment elongate, slender; ultimate segment tapering distally, ending in corneous tip, without robust setiform setae on apex; arthrobranch well developed (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE7C7FE8AFD24CF69" box="[675,738,338,364]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="8" pageNumber="55">Fig 5</figureCitation>
|
|||
|
F).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FEA0FE4DCCB9" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
Chelipeds very unequal in size and asymmetrical in shape, carried folded (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE12DFEA0FB57CF97" box="[1097,1169,376,402]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
A, D). Major cheliped enlarged, elongate, slender; ischium rather long, slender, ventrolateral surface with three stout spiniform setae; merus much longer than ischium, slender, smooth, ventrally flattened; carpus much shorter than merus, subcylindrical, distally broadening, smooth; chela subcylindrical, smooth; fingers shorter than palm, compressed, cutting edges evenly serrated, with around 10 subtriangular teeth (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE128FDD5FB52CC22" box="[1100,1172,525,551]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
A–C). Minor cheliped not particularly enlarged, slender; ischium long, ventrolateral surface with three stout spiniform setae; merus slightly longer than ischium, somewhat compressed, smooth; carpus shorter than merus, subcylindrical, slightly widening distally, smooth; chela simple, smooth, with fingers longer than palm, finger cutting edges straight (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE59BFD7AFE82CCB9" box="[255,324,674,700]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
D, E).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FD10FD98CA79" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
Second pereiopod slender; ischium as long as merus, with three spiniform setae on ventrolateral surface; carpus five-segmented, first segment longest, as long as sum of four other segments, ratio of carpal segments (from proximal to distal) more or less equal to: 6: 1.5: 1: 1.5: 2; chela simple, longer than most-distal carpal article (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE595FCE0FEF1CD57" box="[241,311,824,850]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
F, G). Third pereiopod very slender, compressed; ischium elongate, with two spaced spiniform setae on ventrolateral surface; merus elongate, about 10 times as long as wide, smooth; carpus as long as merus; propodus with two slender spiniform setae on ventral margin and one stouter and longer distal spiniform seta adjacent to dactylus; dactylus about 0.45 propodus length, simple, conical, very slender, gradually curving distally, furnished with setae on dorsal margin (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE6B6FC15FBDBCDE2" box="[978,1053,973,999]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
H, I). Fourth pereiopod almost identical to third pereiopod. Fifth pereiopod longer and more slender than third or fourth pereiopods; ischium unarmed; carpus slightly longer than merus, propodus with one small spiniform seta on ventral margin and at least five rows of slender serrulate setae on distolateral surface, most-distal row with longest setae reaching well beyond dactylar base (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE4BCFBBAFDDBCA79" box="[472,541,1122,1148]" captionStart="FIGURE 6" captionStartId="10.[151,255,1552,1576]" captionTargetBox="[152,1435,185,1514]" captionTargetId="figure@10.[151,1436,156,1531]" captionTargetPageId="10" captionText="FIGURE 6. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, left (major) cheliped, lateral; B, same, carpus and chela, mesial; C, same, chela fingers; D, right (minor) cheliped, lateral; E, same, chela, mesial; F, second pereiopod, lateral; G, same, ischium; H, third pereiopod, lateral; I, same, distal propodus and dactylus; J, fifth pereiopod, lateral; K, same, propodus and dactylus, mesial." httpUri="https://zenodo.org/record/198869/files/figure.png" pageId="8" pageNumber="55">Fig. 6</figureCitation>
|
|||
|
J, K).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FB50FC7FCB17" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
Second pleopod (of both ovigerous specimens) with appendix interna and appendix masculina, latter exceeding than appendix interna, with stiff apical and subapical setae (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE685FB75FBEFCAC2" box="[993,1065,1197,1223]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="8" pageNumber="55">Fig. 5</figureCitation>
|
|||
|
G). Uropod with lateral lobe of protopod ending in small subacute tooth; exopod and endopod slender; exopod with sinuous diaeresis and blunt distolateral tooth adjacent to robust spiniform seta (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE651FB20FCBCCB17" box="[821,890,1272,1298]" captionStart="FIGURE 5" captionStartId="9.[151,255,1873,1897]" captionTargetBox="[154,1436,210,1851]" captionTargetId="figure@9.[151,1436,197,1852]" captionTargetPageId="9" captionText="FIGURE 5. Triacanthoneus pacificus n. gen., n. sp., holotype, ovigerous female (CL 2.8 mm) from Playa Venao, Pacific coast of Panama, USNM 1145374: A, frontal region, dorsal; B, carapace and frontal appendages, lateral; C, carapace, dorsal; D, mediodorsal tooth of carapace, lateral; E, distal rostrum, lateral; F, third maxilliped, lateral; G, second pleopod, appendix masculina and appendix interna; H, uropod, dorsal; I, same, posterior region of exopod; J, telson, dorsal." httpUri="https://zenodo.org/record/198868/files/figure.png" pageId="8" pageNumber="55">Fig. 5</figureCitation>
|
|||
|
H, I).
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FAC5FC9ECB32" blockId="8.[151,1437,152,1932]" box="[197,856,1309,1335]" pageId="8" pageNumber="55">Gill-exopod formula as described for genus (see above).</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A1FA9AFEFFCB87" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
Colour in life semitransparent-white, sometimes with pale orange tinge; gonads or eggs yellow-orange (
|
|||
|
<figureCitation id="130DF5DBFF90FFEAE5FBFAB0FF35CB87" box="[159,243,1384,1410]" captionStart="FIGURE 10" captionStartId="14.[151,256,1843,1867]" captionTargetBox="[151,1436,493,1821]" captionTargetId="figure@14.[151,1436,493,1822]" captionTargetPageId="14" captionText="FIGURE 10. Life colouration of three species of Triacanthoneus n. gen.: A – C, Triacanthoneus toro n. sp., paratype, ovigerous female from Bocas del Toro, Caribbean coast of Panama, USNM 1145376; D – F, Triacanthoneus pacificus n. sp., holotype, ovigerous female from Playa Venao, Pacific coast of Panama, USNM 1145374 [D, E], non-type ovigerous female from Bahía Malaga, Pacific coast of Colombia, UVC [F]; G, Triacanthoneus alacranes n. sp., holotype, ovigerous female from Alacranes Reef, Yucatán, Mexico, UNAM-CNCR 26177 [G by J. Duarte-Gutiérrez, all other photographs by author]." httpUri="https://zenodo.org/record/198873/files/figure.png" pageId="8" pageNumber="55">Fig. 10</figureCitation>
|
|||
|
D–F).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C32CBAD5FF90FFEAE5A2FA55FCE7C839" pageId="8" pageNumber="55" type="etymology">
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A2FA55FE89CBC9" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
<emphasis id="B942354CFF90FFEAE5A2FA55FE95CBA2" bold="true" box="[198,339,1421,1447]" pageId="8" pageNumber="55">Etymology.</emphasis>
|
|||
|
Name refers to the Pacific Ocean; the two other species of the new genus are confined to the Atlantic Ocean.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A2FA0FFCE7C839" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
<emphasis id="B942354CFF90FFEAE5A2FA0FFEE8CBF4" bold="true" box="[198,302,1495,1521]" pageId="8" pageNumber="55">Habitat.</emphasis>
|
|||
|
The Panamanian
|
|||
|
<typeStatus id="548D57FCFF90FFEAE764FA00FDF5CBF7" box="[512,563,1496,1522]" pageId="8" pageNumber="55">type</typeStatus>
|
|||
|
specimen was collected in the mixed sand-mud-rock intertidal at low tide, on sand under large rocks; the Colombian specimen was extracted from an improvised ARMS (large basket with shells) placed on a rock-sand bottom at about
|
|||
|
<specimenCount id="9D3022D7FF90FFEAE788F9FAFCE7C839" box="[748,801,1570,1596]" pageId="8" pageNumber="55" type="generic">
|
|||
|
<quantity id="4CCE44BBFF90FFEAE788F9FAFCDBC839" box="[748,797,1570,1596]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="55" unit="m" value="2.0">2 m</quantity>
|
|||
|
.
|
|||
|
</specimenCount>
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C32CBAD5FF90FFEAE5A2F99FFCF2C867" box="[198,820,1607,1634]" pageId="8" pageNumber="55" type="materials_examined">
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A2F99FFCF2C867" blockId="8.[151,1437,152,1932]" box="[198,820,1607,1634]" pageId="8" pageNumber="55">
|
|||
|
<emphasis id="B942354CFF90FFEAE5A2F99FFEAAC864" bold="true" box="[198,364,1607,1633]" pageId="8" pageNumber="55">
|
|||
|
<typeStatus id="548D57FCFF90FFEAE5A2F99FFEC2C864" box="[198,260,1607,1633]" pageId="8" pageNumber="55">Type</typeStatus>
|
|||
|
locality.
|
|||
|
</emphasis>
|
|||
|
Playa Venao, Pacific coast of
|
|||
|
<collectingCountry id="F321A9CEFF90FFEAE7ABF990FCF6C867" box="[719,816,1608,1634]" name="Panama" pageId="8" pageNumber="55">Panama</collectingCountry>
|
|||
|
.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C32CBAD5FF90FFEAE5A2F9B5FE76C8A9" pageId="8" pageNumber="55" type="distribution">
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A2F9B5FE76C8A9" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
<emphasis id="B942354CFF90FFEAE5A2F9B5FEA1C882" bold="true" box="[198,359,1645,1671]" pageId="8" pageNumber="55">Distribution.</emphasis>
|
|||
|
Presently known only from Playa Venao,
|
|||
|
<collectingCountry id="F321A9CEFF90FFEAE614F9B5FC16C882" box="[880,976,1645,1671]" name="Panama" pageId="8" pageNumber="55">Panama</collectingCountry>
|
|||
|
, and Bahía Malaga,
|
|||
|
<collectingCountry id="F321A9CEFF90FFEAE1B4F9B5FA81C882" box="[1232,1351,1645,1671]" name="Colombia" pageId="8" pageNumber="55">Colombia</collectingCountry>
|
|||
|
, in the tropical Eastern Pacific.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C32CBAD5FF90FFEAE5A2F96FFF36C989" pageId="8" pageNumber="55" type="discussion">
|
|||
|
<paragraph id="8B89E95EFF90FFEAE5A2F96FFF36C989" blockId="8.[151,1437,152,1932]" pageId="8" pageNumber="55">
|
|||
|
<emphasis id="B942354CFF90FFEAE5A2F96FFE86C8D4" bold="true" box="[198,320,1719,1745]" pageId="8" pageNumber="55">Remarks.</emphasis>
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE428F960FDBEC8D4" box="[332,632,1720,1745]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE428F960FDBEC8D4" box="[332,632,1720,1745]" italics="true" pageId="8" pageNumber="55">Triacanthoneus pacificus</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE7E0F96FFD0DC8D4" box="[644,715,1719,1745]" pageId="8" pageNumber="55" rank="species">
|
|||
|
<emphasis id="B942354CFF90FFEAE7E0F96FFD0DC8D4" bold="true" box="[644,715,1719,1745]" pageId="8" pageNumber="55">n. sp.</emphasis>
|
|||
|
</taxonomicNameLabel>
|
|||
|
differs from
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE610F960FC04C8D4" box="[884,962,1720,1745]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE610F960FC04C8D4" box="[884,962,1720,1745]" italics="true" pageId="8" pageNumber="55">T. toro</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE6B5F96FFBDEC8D4" box="[977,1048,1719,1745]" pageId="8" pageNumber="55" rank="species">
|
|||
|
<emphasis id="B942354CFF90FFEAE6B5F96FFBDEC8D4" bold="true" box="[977,1048,1719,1745]" pageId="8" pageNumber="55">n. sp.</emphasis>
|
|||
|
</taxonomicNameLabel>
|
|||
|
and
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE13FF960FB29C8D4" box="[1115,1263,1720,1745]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="alacranes" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE13FF960FB29C8D4" box="[1115,1263,1720,1745]" italics="true" pageId="8" pageNumber="55">T. alacranes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE198F96FFA85C8D4" box="[1276,1347,1719,1745]" pageId="8" pageNumber="55" rank="species">
|
|||
|
<emphasis id="B942354CFF90FFEAE198F96FFA85C8D4" bold="true" box="[1276,1347,1719,1745]" pageId="8" pageNumber="55">n. sp.</emphasis>
|
|||
|
</taxonomicNameLabel>
|
|||
|
by the distinctly more robust major chela; and the more ventrally directed subapical dent on the ventral margin of the rostrum; specifically from
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE482F8DBFDF0C919" box="[486,566,1795,1820]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="toro" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE482F8DBFDF0C919" box="[486,566,1795,1820]" italics="true" pageId="8" pageNumber="55">T. toro</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE725F8DAFD4EC919" box="[577,648,1794,1820]" pageId="8" pageNumber="55" rank="species">
|
|||
|
<emphasis id="B942354CFF90FFEAE725F8DAFD4EC919" bold="true" box="[577,648,1794,1820]" pageId="8" pageNumber="55">n. sp.</emphasis>
|
|||
|
</taxonomicNameLabel>
|
|||
|
by the more posterior position of the mediodorsal tooth on the carapace. It specifically differs from
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE739F8F0FD32C944" box="[605,756,1832,1857]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="alacranes" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE739F8F0FD32C944" box="[605,756,1832,1857]" italics="true" pageId="8" pageNumber="55">T. alacranes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE798F8F0FCF8C947" box="[764,830,1832,1858]" pageId="8" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
by the absence of orbital teeth, the more anterior position of the dorsolateral teeth of the carapace, and several other features (see under
|
|||
|
<taxonomicName id="4C3692DDFF90FFEAE1DDF895FA89C963" box="[1209,1359,1869,1894]" class="Malacostraca" family="Alpheidae" genus="Triacanthoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="55" phylum="Arthropoda" rank="species" species="alacranes" status="sp. nov.">
|
|||
|
<emphasis id="B942354CFF90FFEAE1DDF895FA89C963" box="[1209,1359,1869,1894]" italics="true" pageId="8" pageNumber="55">T. alacranes</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
<taxonomicNameLabel id="A2718837FF90FFEAE033F895FA5AC962" box="[1367,1436,1869,1895]" pageId="8" pageNumber="55" rank="species">
|
|||
|
<emphasis id="B942354CFF90FFEAE033F895FA5AC962" bold="true" box="[1367,1436,1869,1895]" pageId="8" pageNumber="55">n. sp.</emphasis>
|
|||
|
</taxonomicNameLabel>
|
|||
|
below).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|