treatments-xml/data/03/93/2D/03932D70B62AC37EFF7B0DDCA55ADB87.xml

<document id="65FCDE121095FDC651AD90B3B1BB98DE" ID-DOI="10.11646/zootaxa.5026.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10100344" ID-ZooBank="07E0C922-6B0F-4916-85C2-AD95146A8F1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979906524" checkinUser="plazi" docAuthor="Neita-Moreno, Jhon C." docDate="2021" docId="03932D70B62AC37EFF7B0DDCA55ADB87" docLanguage="en" docName="zootaxa.5026.1.1.pdf" docOrigin="Zootaxa 5026 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala rogerpauli Moore, Branham, &amp; Cave 2018" docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="FFAA5508B623C377FFEC0C06A02ADC12" masterDocTitle="A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia" masterLastPageNumber="58" masterPageNumber="1" pageNumber="10" updateTime="1699603307102" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A1730425A518C47EECBF53DB50E059F2">A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia</mods:title>
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<mods:namePart id="B9EA81C60F3F0055E434C41E72EE0971">Neita-Moreno, Jhon C.</mods:namePart>
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<mods:date id="35BA86FA920462350A429ACC20E1568D">2021</mods:date>
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<subSubSection id="C320CFEDB62AC37EFF7B0DDCA372DDE6" box="[151,856,474,501]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8B859C66B62AC37EFF7B0DDCA372DDE6" blockId="9.[151,856,474,537]" box="[151,856,474,501]" pageId="9" pageNumber="10">
<heading id="D0CD2B0AB62AC37EFF7B0DDCA372DDE6" bold="true" box="[151,856,474,501]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<taxonomicName id="4C3AE7E5B62AC37EFF7B0DDCA372DDE6" authority="Moore, Branham, &amp; Cave, 2018" authorityName="Moore, Branham, &amp; Cave" authorityYear="2018" box="[151,856,474,501]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rogerpauli">
<emphasis id="B94E4074B62AC37EFF7B0DDCA372DDE6" bold="true" box="[151,856,474,501]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF7B0DDCA1EADDE7" bold="true" box="[151,448,474,501]" italics="true" pageId="9" pageNumber="10">Cyclocephala rogerpauli</emphasis>
Moore, Branham, &amp; Cave, 2018
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB62AC37EFF7B0DF9A2B7DE0B" box="[151,669,510,537]" pageId="9" pageNumber="10" type="description">
<paragraph id="8B859C66B62AC37EFF7B0DF9A2B7DE0B" blockId="9.[151,856,474,537]" box="[151,669,510,537]" pageId="9" pageNumber="10">
(
<figureCitation id="130180E3B62AC37EFF730DF9A135DE0B" box="[159,287,511,537]" captionStart="FIGURE 1" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,345,1699]" captionTargetId="figure-72@4.[151,1436,345,1699]" captionTargetPageId="4" captionText="FIGURE 1A–F. Cyclocephala cribrata species group. A, Cyclocephala marqueti male; B, Cyclocephala marqueti female; C, Cyclocephala proxima male; D, Cyclocephala proxima female; E, Cyclocephala rogerpauli male (holotype); F, Cyclocephala rogerpauli female. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100346" httpUri="https://zenodo.org/record/10100346/files/figure.png" pageId="9" pageNumber="10">Figs. 1E–F</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFEC60DF9A176DE0B" box="[298,348,511,537]" captionStart="FIGURE 19" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[182,1400,188,1910]" captionTargetId="figure-29@30.[175,1411,181,1919]" captionTargetPageId="30" captionText="FIGURE 19A–H. Metatibiae (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C. Cyclocephala rogerpauli; D, Cyclocephala dalensi; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala rufa; H, Cyclocephala arhuacana. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10100392" httpUri="https://zenodo.org/record/10100392/files/figure.png" pageId="9" pageNumber="10">19C</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFE840DF9A1B0DE0B" box="[360,410,511,537]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="9" pageNumber="10">20C</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFE490DF9A1FDDE0B" box="[421,471,511,537]" captionStart="FIGURE 21" captionStartId="32.[151,250,1894,1918]" captionTargetBox="[161,1435,158,1858]" captionTargetId="figure-29@32.[151,1436,151,1870]" captionTargetPageId="32" captionText="FIGURE 21A–D.Wing. Cyclocephala cribrata species group. A, Cyclocephala marqueti; B, C, D, Cyclocephala proxima." figureDoi="http://doi.org/10.5281/zenodo.10100396" httpUri="https://zenodo.org/record/10100396/files/figure.png" pageId="9" pageNumber="10">21C</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFE0F0DF9A21BDE0B" box="[483,561,511,537]" captionStart="FIGURE 23" captionStartId="34.[151,250,1841,1865]" captionTargetBox="[157,1408,191,1807]" captionTargetId="figure-29@34.[151,1436,181,1817]" captionTargetPageId="34" captionText="FIGURE 23A–D. Wing. Cyclocephala ligyrina species group. A, Cyclocephala arhuacana; B, C, D, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100400" httpUri="https://zenodo.org/record/10100400/files/figure.png" pageId="9" pageNumber="10">23E–F</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFDD00DF9A247DE0B" box="[572,621,511,537]" captionStart="FIGURE 29" captionStartId="43.[151,250,1627,1651]" captionTargetBox="[163,1424,191,1592]" captionTargetId="figure-15@43.[151,1436,181,1604]" captionTargetPageId="43" captionText="FIGURE 29A–F. Parameres and phallobases, Cyclocephala ligyrina species-group. A, Cyclocephala kaszabi parameres, caudal view; B, Cyclocephala kaszabi phallobase, lateral view; C, Cyclocephala dalensi parameres, caudal view; D, Cyclocephala dalensi phallobase, lateral view; E, Cyclocephala rufa parameres, caudal view; F, Cyclocephala rufa phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100413" httpUri="https://zenodo.org/record/10100413/files/figure.png" pageId="9" pageNumber="10">29E</figureCitation>
; 
<figureCitation id="130180E3B62AC37EFD940DF8A2BFDE0B" box="[632,661,510,537]" captionStart="FIGURE 35" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[274,1311,181,1912]" captionTargetId="figure-15@49.[265,1322,181,1919]" captionTargetPageId="49" captionText="FIGURE 35A–G. Cyclocephala marqueti, third instar. A, Labium, dorsal view; B, right maxilla, dorsal view; C, apex of mala showing unci; D, stridulatory teeth of maxilla; E, prothoracic claw, lateral view; F, mesothoracic claw, lateral view; G, metathoracic claw, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100422" httpUri="https://zenodo.org/record/10100422/files/figure.png" pageId="9" pageNumber="10">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB62AC37EFF7B0E40A2CDDEDF" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="8B859C66B62AC37EFF7B0E40A2CDDEDF" blockId="9.[151,1437,582,1941]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF7B0E40A120DE72" bold="true" box="[151,266,582,608]" pageId="9" pageNumber="10">Diagnosis</emphasis>
. 
<taxonomicName id="4C3AE7E5B62AC37EFEF40E41A21ADE73" authorityName="Moore, Branham, &amp; Cave" authorityYear="2018" box="[280,560,583,609]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rogerpauli">
<emphasis id="B94E4074B62AC37EFEF40E41A21ADE73" box="[280,560,583,609]" italics="true" pageId="9" pageNumber="10">Cyclocephala rogerpauli</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C3AE7E5B62AC37EFD2C0E41A36EDE72" box="[704,836,583,609]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="marqueti">
C. 
<emphasis id="B94E4074B62AC37EFD0C0E41A36EDE72" box="[736,836,583,608]" italics="true" pageId="9" pageNumber="10">marqueti</emphasis>
</taxonomicName>
because the inner edge of the metatibia has a longitudinal keel (
<figureCitation id="130180E3B62AC37EFEFF0E6DA15DDE97" box="[275,375,619,645]" captionStart="FIGURE 19" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[182,1400,188,1910]" captionTargetId="figure-29@30.[175,1411,181,1919]" captionTargetPageId="30" captionText="FIGURE 19A–H. Metatibiae (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C. Cyclocephala rogerpauli; D, Cyclocephala dalensi; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala rufa; H, Cyclocephala arhuacana. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10100392" httpUri="https://zenodo.org/record/10100392/files/figure.png" pageId="9" pageNumber="10">Fig. 19C</figureCitation>
) and the surface of pygidium has denser punctures (
<figureCitation id="130180E3B62AC37EFC510E6DA40EDE97" box="[957,1060,619,645]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="9" pageNumber="10">Fig. 24C</figureCitation>
). 
<taxonomicName id="4C3AE7E5B62AC37EFBDA0E6DA564DE97" authorityName="Moore, Branham, &amp; Cave" authorityYear="2018" box="[1078,1358,619,645]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rogerpauli">
<emphasis id="B94E4074B62AC37EFBDA0E6DA564DE97" box="[1078,1358,619,645]" italics="true" pageId="9" pageNumber="10">Cyclocephala rogerpauli</emphasis>
</taxonomicName>
can be distinguished by the dense punctures on the surface of the metasternite. The female epipleuron is weakly enlarged at the level of the first abdominal sternite (
<figureCitation id="130180E3B62AC37EFD980EB5A2F1DEDF" box="[628,731,691,717]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="9" pageNumber="10">Fig. 20C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB62AC37EFF2B0ED0A1D8DA3F" pageId="9" pageNumber="10" type="description">
<paragraph id="8B859C66B62AC37EFF2B0ED0A18AD947" blockId="9.[151,1437,582,1941]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF2B0ED0A147DEE2" bold="true" box="[199,365,726,752]" pageId="9" pageNumber="10">Redescription</emphasis>
. 
<emphasis id="B94E4074B62AC37EFE900ED1A193DEE3" bold="true" box="[380,441,727,753]" pageId="9" pageNumber="10">Male</emphasis>
(
<figureCitation id="130180E3B62AC37EFE250ED1A234DEE3" box="[457,542,727,753]" captionStart="FIGURE 1" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,345,1699]" captionTargetId="figure-72@4.[151,1436,345,1699]" captionTargetPageId="4" captionText="FIGURE 1A–F. Cyclocephala cribrata species group. A, Cyclocephala marqueti male; B, Cyclocephala marqueti female; C, Cyclocephala proxima male; D, Cyclocephala proxima female; E, Cyclocephala rogerpauli male (holotype); F, Cyclocephala rogerpauli female. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100346" httpUri="https://zenodo.org/record/10100346/files/figure.png" pageId="9" pageNumber="10">Fig. 1E</figureCitation>
). Length 25.0–26.0 mm; width 12.5–13.0 mm. Color black. 
<emphasis id="B94E4074B62AC37EFB250ED1A538DEE3" bold="true" box="[1225,1298,727,753]" pageId="9" pageNumber="10">Head:</emphasis>
Frons moderately to moderately densely punctate (rarely sparsely), punctures mostly moderately large (rarely small). Clypeus with surface similar to frons, apical half often with only a few, small punctures; apex convexly rounded, margined, usually with weak angle at center, weakly reflexed. Mandibles apically rounded, internal face with a groove; labium densely setose, paraglossa undeveloped, apex widely notched; maxilla with galea developed, with 7 teeth, lacinia sclerotized and developed toward base of teeth. Interocular width equals 4.0 transverse eye diameters.Antenna with 10 antennomeres, club with a few long setae and slightly longer than antennomeres 2–7. 
<emphasis id="B94E4074B62AC37EFB680FA9A52CDFDB" bold="true" box="[1156,1286,943,969]" pageId="9" pageNumber="10">Pronotum:</emphasis>
Surface with punctures moderately dense, moderately large to large (especially on sides), ocellate, deep. Base lacking marginal bead. 
<emphasis id="B94E4074B62AC37EFF360FF1A11AD803" bold="true" box="[218,304,1015,1041]" pageId="9" pageNumber="10">Elytra:</emphasis>
Surface with punctate rows, punctures of striae and intervals smilar to those of pronotum; entire surface often coarsely and transversely wrinkled obscuring rows of punctures. 
<emphasis id="B94E4074B62AC37EFC0F081CA475D826" bold="true" box="[995,1119,1050,1076]" pageId="9" pageNumber="10">Pygidium:</emphasis>
Surface moderately densely to more usually sparsely punctate; punctures moderate in size to more usually moderately large, ocellate, setigerous; setae short, spine like, tawny (
<figureCitation id="130180E3B62AC37EFE1C0865A272D86F" box="[496,600,1123,1149]" captionStart="FIGURE 24" captionStartId="35.[151,250,1840,1864]" captionTargetBox="[152,1423,181,1807]" captionTargetId="figure-15@35.[151,1436,181,1816]" captionTargetPageId="35" captionText="FIGURE 24A–H. Pygidia (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala santandereana (holotype); F, Cyclocephala dalensi; G, Cyclocephala kaszabi; H, Cyclocephala ligyrina." figureDoi="http://doi.org/10.5281/zenodo.10100402" httpUri="https://zenodo.org/record/10100402/files/figure.png" pageId="9" pageNumber="10">Fig. 24C</figureCitation>
). In lateral view, surface in male evenly convex, weakly convex to nearly flat in female. 
<emphasis id="B94E4074B62AC37EFED60881A151D8B3" bold="true" box="[314,379,1159,1185]" pageId="9" pageNumber="10">Legs:</emphasis>
Protibia tridentate, basal tooth slightly removed from others. Protarsus enlarged (
<figureCitation id="130180E3B62AC37EFAE70881A54AD8B3" box="[1291,1376,1159,1185]" captionStart="FIGURE 1" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,345,1699]" captionTargetId="figure-72@4.[151,1436,345,1699]" captionTargetPageId="4" captionText="FIGURE 1A–F. Cyclocephala cribrata species group. A, Cyclocephala marqueti male; B, Cyclocephala marqueti female; C, Cyclocephala proxima male; D, Cyclocephala proxima female; E, Cyclocephala rogerpauli male (holotype); F, Cyclocephala rogerpauli female. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100346" httpUri="https://zenodo.org/record/10100346/files/figure.png" pageId="9" pageNumber="10">Fig. 1E</figureCitation>
), tarsomeres 2–4 successively gradually larger, fifth large, curved, with small angulation at base on ventral side; median claw at base with small tooth, claw large, curved, apex cleft. Metatarsus subequal in length to metatibia. The inner edge of the metatibia has a longitudinal keel (
<figureCitation id="130180E3B62AC37EFD7608F5A32BD91F" box="[666,769,1267,1293]" captionStart="FIGURE 19" captionStartId="30.[151,250,1943,1967]" captionTargetBox="[182,1400,188,1910]" captionTargetId="figure-29@30.[175,1411,181,1919]" captionTargetPageId="30" captionText="FIGURE 19A–H. Metatibiae (dorsal view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C. Cyclocephala rogerpauli; D, Cyclocephala dalensi; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala rufa; H, Cyclocephala arhuacana. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10100392" httpUri="https://zenodo.org/record/10100392/files/figure.png" pageId="9" pageNumber="10">Fig. 19C</figureCitation>
). 
<emphasis id="B94E4074B62AC37EFCFA08F5A345D91F" bold="true" box="[790,879,1267,1293]" pageId="9" pageNumber="10">Venter:</emphasis>
Prosternal process moderate in length, columnar, apex obliquely flattened into a transverse oval with anterior 2/3 to 4/5 raised into convex “button”. 
<emphasis id="B94E4074B62AC37EFB0F0910A571D922" bold="true" box="[1251,1371,1302,1328]" pageId="9" pageNumber="10">Genitalia:</emphasis>
Parameres as in 
<figureCitation id="130180E3B62AC37EFEF0093DA1B6D947" box="[284,412,1339,1365]" captionStart="FIGURE 26" captionStartId="40.[151,250,1885,1909]" captionTargetBox="[161,1424,192,1849]" captionTargetId="figure-29@40.[151,1436,181,1862]" captionTargetPageId="40" captionText="FIGURE 26A–F. Parameres and phallobases, Cyclocephala cribrata species group. A, Cyclocephala marqueti parameres, caudal view; B, Cyclocephala marqueti phallobase, lateral view; C, Cyclocephala proxima parameres, caudal view; D, Cyclocephala proxima phallobase, lateral view; E, Cyclocephala rogerpauli parameres, caudal view; F, Cyclocephala rogerpauli phallobase, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100406" httpUri="https://zenodo.org/record/10100406/files/figure.png" pageId="9" pageNumber="10">Fig. 26E–F</figureCitation>
.
</paragraph>
<paragraph id="8B859C66B62AC37EFF2B0959A1D8DA3F" blockId="9.[151,1437,582,1941]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF2B0959A134D96B" bold="true" box="[199,286,1375,1401]" pageId="9" pageNumber="10">Female</emphasis>
(
<figureCitation id="130180E3B62AC37EFEC20959A1A9D96B" box="[302,387,1375,1401]" captionStart="FIGURE 1" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,345,1699]" captionTargetId="figure-72@4.[151,1436,345,1699]" captionTargetPageId="4" captionText="FIGURE 1A–F. Cyclocephala cribrata species group. A, Cyclocephala marqueti male; B, Cyclocephala marqueti female; C, Cyclocephala proxima male; D, Cyclocephala proxima female; E, Cyclocephala rogerpauli male (holotype); F, Cyclocephala rogerpauli female. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100346" httpUri="https://zenodo.org/record/10100346/files/figure.png" pageId="9" pageNumber="10">Fig. 1F</figureCitation>
). Length 24.0– 
<quantity id="4CC23183B62AC37EFDC30958A2BCD96B" box="[559,662,1374,1401]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" pageId="9" pageNumber="10" unit="mm" value="25.9">25.9 mm</quantity>
; width 12.0– 
<quantity id="4CC23183B62AC37EFCC60959A3A5D96B" box="[810,911,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="9" pageNumber="10" unit="mm" value="12.1">12.1 mm</quantity>
. Similar to male, punctures on pronotum similar. Epipleuron (ventral view, 
<figureCitation id="130180E3B62AC37EFE0B0985A261D98F" box="[487,587,1411,1437]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="9" pageNumber="10">Fig. 20C</figureCitation>
) slightly enlarged at level of first abdominal sternite. In dorsal view, lateral margin slightly enlarged just before middle (
<figureCitation id="130180E3B62AC37EFD6309A1A2DDD9D3" box="[655,759,1447,1473]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="9" pageNumber="10">Fig. 20C</figureCitation>
); in lateral view, ventral margin of epipleuron without tooth or angulation (
<figureCitation id="130180E3B62AC37EFEAF09CDA184D9F7" box="[323,430,1483,1509]" captionStart="FIGURE 20" captionStartId="31.[151,250,1913,1937]" captionTargetBox="[196,1389,161,1875]" captionTargetId="figure-15@31.[185,1402,151,1890]" captionTargetPageId="31" captionText="FIGURE 20A–G. Female epipleura (ventral view). A, Cyclocephala marqueti; B, Cyclocephala proxima; C, Cyclocephala rogerpauli; D, Cyclocephala brevipennis; E, Cyclocephala kaszabi; F, Cyclocephala ligyrina; G, Cyclocephala arhuacana." figureDoi="http://doi.org/10.5281/zenodo.10100394" httpUri="https://zenodo.org/record/10100394/files/figure.png" pageId="9" pageNumber="10">Fig. 20C</figureCitation>
). Abdominal ventrite VIII entire, not emarginate at center. Pygidium slightly concave, punctures sparse. Protarsus simple (
<figureCitation id="130180E3B62AC37EFDC709E9A2AADA1B" box="[555,640,1519,1545]" captionStart="FIGURE 1" captionStartId="4.[151,250,1722,1746]" captionTargetBox="[151,1436,345,1699]" captionTargetId="figure-72@4.[151,1436,345,1699]" captionTargetPageId="4" captionText="FIGURE 1A–F. Cyclocephala cribrata species group. A, Cyclocephala marqueti male; B, Cyclocephala marqueti female; C, Cyclocephala proxima male; D, Cyclocephala proxima female; E, Cyclocephala rogerpauli male (holotype); F, Cyclocephala rogerpauli female. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10100346" httpUri="https://zenodo.org/record/10100346/files/figure.png" pageId="9" pageNumber="10">Fig. 1F</figureCitation>
). Gonocoxite larger than gonocoxal sternite, gonocoxite at center with a membranous area (Fig. 32E).
</paragraph>
</subSubSection>
<subSubSection id="C320CFEDB62AC37EFF2B0A30A55ADB87" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B859C66B62AC37EFF2B0A30A493DA43" blockId="9.[151,1437,582,1941]" box="[199,1209,1590,1617]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF2B0A30A173DA42" bold="true" box="[199,345,1590,1616]" pageId="9" pageNumber="10">Distribution</emphasis>
. 
<taxonomicName id="4C3AE7E5B62AC37EFE8B0A31A255DA43" authorityName="Moore, Branham, &amp; Cave" authorityYear="2018" box="[359,639,1591,1617]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="rogerpauli">
<emphasis id="B94E4074B62AC37EFE8B0A31A255DA43" box="[359,639,1591,1617]" italics="true" pageId="9" pageNumber="10">Cyclocephala rogerpauli</emphasis>
</taxonomicName>
is known only from 
<collectingCountry id="F32DDCF6B62AC37EFC850A31A3FDDA43" box="[873,983,1591,1617]" name="Colombia" pageId="9" pageNumber="10">Colombia</collectingCountry>
(
<bibRefCitation id="EFABE197B62AC37EFC0A0A31A486DA43" author="Dechambre, R. P." box="[998,1196,1591,1617]" pageId="9" pageNumber="10" pagination="1 - 24" refId="ref22901" refString="Dechambre, R. P. (1999) Vingt nouvelles especes et une nouvelle sous-espece de Cyclocephala Burmeister, 1847. Les Coleopteres du Monde, 14 (Supplement 1), 1 - 24." type="journal article" year="1999">Dechambre 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8B859C66B62AC37EFF2B0A5CA377DA8B" blockId="9.[151,1437,582,1941]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF2B0A5CA17BDA66" bold="true" box="[199,337,1626,1652]" pageId="9" pageNumber="10">Life history</emphasis>
. Adults are attracted to lights at night. The adults are also attracted to the flowers of 
<taxonomicName id="4C3AE7E5B62AC37EFB1F0A5DA094DA8A" class="Liliopsida" family="Arecaceae" genus="Attalea" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="butyracea">
<emphasis id="B94E4074B62AC37EFB1F0A5DA094DA8A" italics="true" pageId="9" pageNumber="10">Attalea butyracea</emphasis>
</taxonomicName>
(Mutis 
<emphasis id="B94E4074B62AC37EFEFB0A79A11ADA8A" box="[279,304,1663,1688]" italics="true" pageId="9" pageNumber="10">ex</emphasis>
L.f.) Wess. Boer (
<taxonomicName id="4C3AE7E5B62AC37EFE100A79A25EDA8B" box="[508,628,1663,1689]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
) (
<bibRefCitation id="EFABE197B62AC37EFD660A79A37ADA8B" author="Dechambre, R. P." box="[650,848,1663,1689]" pageId="9" pageNumber="10" pagination="13 - 27" refId="ref22871" refString="Dechambre, R. P. (1997) Revision des Cyclocephala du groupe cribrata Burmeister (Coleoptera, Dynastidae). Coleopteres, 3, 13 - 27." type="journal article" year="1997">Dechambre 1997</bibRefCitation>
).
</paragraph>
<paragraph id="8B859C66B62AC37EFF2B0AA4A43EDAAF" blockId="9.[151,1437,582,1941]" box="[199,1044,1698,1725]" pageId="9" pageNumber="10">
<emphasis id="B94E4074B62AC37EFF2B0AA4A1A0DAAF" bold="true" box="[199,394,1698,1725]" pageId="9" pageNumber="10">Locality records</emphasis>
(
<figureCitation id="130180E3B62AC37EFE750AA5A1C7DAAF" box="[409,493,1698,1725]" captionStart="FIGURE 35" captionStartId="49.[151,250,1943,1967]" captionTargetBox="[274,1311,181,1912]" captionTargetId="figure-15@49.[265,1322,181,1919]" captionTargetPageId="49" captionText="FIGURE 35A–G. Cyclocephala marqueti, third instar. A, Labium, dorsal view; B, right maxilla, dorsal view; C, apex of mala showing unci; D, stridulatory teeth of maxilla; E, prothoracic claw, lateral view; F, mesothoracic claw, lateral view; G, metathoracic claw, lateral view." figureDoi="http://doi.org/10.5281/zenodo.10100422" httpUri="https://zenodo.org/record/10100422/files/figure.png" pageId="9" pageNumber="10">Fig. 35</figureCitation>
). 
<materialsCitation id="3B52963BB62AC37EFDED0AA5A43ADAAF" box="[513,1040,1699,1725]" collectionCode="IAvH, MNHN" pageId="9" pageNumber="10" specimenCount="7">
<specimenCount id="9D3C57EFB62AC37EFDED0AA5A2A1DAAF" box="[513,651,1699,1725]" pageId="9" pageNumber="10" type="generic">7 specimens</specimenCount>
examined from 
<collectionCode id="ED2B04A3B62AC37EFCAF0AA5A3ABDAAF" box="[835,897,1699,1725]" pageId="9" pageNumber="10">IAvH</collectionCode>
and 
<collectionCode id="ED2B04A3B62AC37EFC5B0AA5A43ADAAF" box="[951,1040,1699,1725]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B859C66B62AC37EFF2B0AC1A55ADB87" blockId="9.[151,1437,582,1941]" pageId="9" pageNumber="10">
“ 
<materialsCitation id="3B52963BB62AC37EFF3D0AC1A547DB87" collectingDate="1995-07-17" collectingDateMax="2019-06-04" collectingDateMin="1995-07-17" collectionCode="MNHN" country="Colombia" county="San Onofre" elevation="84347" latitude="11.192611" location="Palm" municipality="Sucre" pageId="9" pageNumber="10" specimenCode="EC10190" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Magdalena" typeStatus="holotype">
San Onofre (Sucre) / 
<collectingCountry id="F32DDCF6B62AC37EFE290AC1A219DAF3" box="[453,563,1735,1761]" name="Colombia" pageId="9" pageNumber="10">Colombia</collectingCountry>
/ 
<date id="FF84BAA6B62AC37EFDA60AC1A33EDAF3" box="[586,788,1734,1761]" pageId="9" pageNumber="10" value="1995-07-17" valueMax="1995-07-24" valueMin="1995-07-17">
<collectingDate id="EFC0434EB62AC37EFDA60AC1A33EDAF3" box="[586,788,1734,1761]" pageId="9" pageNumber="10" value="1995-07-17" valueMax="1995-07-24" valueMin="1995-07-17">17-24 juillet 1995</collectingDate>
</date>
/ 
<taxonomicName id="4C3AE7E5B62AC37EFCC60AC1A3D9DAF3" box="[810,1011,1735,1761]" class="Liliopsida" family="Arecaceae" genus="Attalea" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="butyracea">
<emphasis id="B94E4074B62AC37EFCC60AC1A3D9DAF3" box="[810,1011,1735,1761]" italics="true" pageId="9" pageNumber="10">Attalea butyracea</emphasis>
</taxonomicName>
/ Palm 3/1 / 
<typeStatus id="548122C4B62AC37EFB690AC1A4C7DAF3" box="[1157,1261,1735,1761]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
/ 
<taxonomicName id="4C3AE7E5B62AC37EFAE80AC1A0FBDB17" authority="Neita-Moreno, 2021" authorityName="Neita-Moreno" authorityYear="2021" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">Cyclocephala nigra</taxonomicName>
<taxonomicNameLabel id="A27DFD0FB62AC37EFF3B0AEDA138DB17" box="[215,274,1771,1797]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
<typeStatus id="548122C4B62AC37EFEF40AEDA186DB17" box="[280,428,1771,1797]" pageId="9" pageNumber="10" type="holotype">HOLOTYPE</typeStatus>
/ R. P. DECHAMBRE det. 1998” 
<collectionCode id="ED2B04A3B62AC37EFCC50AEDA355DB17" box="[809,895,1771,1797]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
<specimenCode id="DB9C341DB62AC37EFC690AEDA3D8DB17" box="[901,1010,1771,1797]" collectionCode="MNHN" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">EC10190</specimenCode>
. “ 
<collectingCounty id="62E4E4EAB62AC37EFBEA0AEDA4ADDB17" box="[1030,1159,1771,1797]" pageId="9" pageNumber="10">San Onofre</collectingCounty>
(
<collectingMunicipality id="6BE1061CB62AC37EFB7A0AEDA4F3DB17" box="[1174,1241,1771,1797]" pageId="9" pageNumber="10">Sucre</collectingMunicipality>
) / 
<collectingCountry id="F32DDCF6B62AC37EFB180AEDA548DB17" box="[1268,1378,1771,1797]" name="Colombia" pageId="9" pageNumber="10">Colombia</collectingCountry>
/ 
<date id="FF84BAA6B62AC37EFA9B0AEDA110DB3B" pageId="9" pageNumber="10" value="1995-07-17" valueMax="1995-07-24" valueMin="1995-07-17">
<collectingDate id="EFC0434EB62AC37EFA9B0AEDA110DB3B" pageId="9" pageNumber="10" value="1995-07-17" valueMax="1995-07-24" valueMin="1995-07-17">17- 24 juillet 1995</collectingDate>
</date>
/ 
<taxonomicName id="4C3AE7E5B62AC37EFEA20B09A23CDB3B" box="[334,534,1807,1833]" class="Liliopsida" family="Arecaceae" genus="Attalea" kingdom="Plantae" order="Arecales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="butyracea">
<emphasis id="B94E4074B62AC37EFEA20B09A23CDB3B" box="[334,534,1807,1833]" italics="true" pageId="9" pageNumber="10">Attalea butyracea</emphasis>
</taxonomicName>
/ 
<location id="8EE5CABDB62AC37EFDC80B09A274DB3B" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFDC80B09A274DB3B" box="[548,606,1807,1833]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Palm" pageId="9" pageNumber="10" stateProvince="Magdalena">Palm</location>
3/1 / 
<typeStatus id="548122C4B62AC37EFD710B09A32FDB3B" box="[669,773,1807,1833]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
/ 
<taxonomicName id="4C3AE7E5B62AC37EFCF60B09A3D9DB3B" authority="Neita-Moreno, 2021" authorityName="Neita-Moreno" authorityYear="2021" box="[794,1011,1807,1833]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nigra" status="sp. nov.">Cyclocephala nigra</taxonomicName>
<taxonomicNameLabel id="A27DFD0FB62AC37EFC150B09A41EDB3B" box="[1017,1076,1807,1833]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
PARATIPES / R. P. DECHAM- BRE det. 1998” 
<collectionCode id="ED2B04A3B62AC37EFEBF0B35A183DB5F" box="[339,425,1843,1869]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="10">MNHN</collectionCode>
(
<specimenCount id="9D3C57EFB62AC37EFE570B35A22FDB5F" box="[443,517,1843,1869]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
and 
<specimenCount id="9D3C57EFB62AC37EFDD10B35A285DB5C" box="[573,687,1843,1870]" pageId="9" pageNumber="10" type="female">4 females</specimenCount>
). “ 
<collectingCountry id="F32DDCF6B62AC37EFD200B35A36ADB5F" box="[716,832,1843,1869]" name="Colombia" pageId="9" pageNumber="10">Colombia</collectingCountry>
, 
<collectingRegion id="49FE5284B62AC37EFCA00B35A3E1DB5F" box="[844,971,1843,1869]" country="Colombia" name="Magdalena" pageId="9" pageNumber="10">Magdalena</collectingRegion>
, 
<location id="8EE5CABDB62AC37EFC3B0B35A43FDB5F" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFC3B0B35A43FDB5F" box="[983,1045,1843,1869]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Santa" pageId="9" pageNumber="10" stateProvince="Magdalena">Santa</location>
/ 
<location id="8EE5CABDB62AC37EFBC10B35A459DB5F" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFBC10B35A459DB5F" box="[1069,1139,1843,1869]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Marta" pageId="9" pageNumber="10" stateProvince="Magdalena">Marta</location>
, 
<location id="8EE5CABDB62AC37EFB920B35A5B7DB5F" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFB920B35A5B7DB5F" box="[1150,1437,1843,1869]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Reserva Biologica Caoba" pageId="9" pageNumber="10" stateProvince="Magdalena">Reserva Biológica Caoba</location>
/ Corrg. Bonda, Vda. Theran, / 
<location id="8EE5CABDB62AC37EFDFC0B51A2DEDB63" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFDFC0B51A2DEDB63" box="[528,756,1879,1905]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Bosque Intervenido" pageId="9" pageNumber="10" stateProvince="Magdalena">Bosque Intervenido</location>
, 
<location id="8EE5CABDB62AC37EFCE80B51A419DB63" LSID="urn:lsid:plazi:treatment:03932D70B62AC37EFF7B0DDCA55ADB87:8EE5CABDB62AC37EFCE80B51A419DB63" box="[772,1075,1879,1905]" country="Colombia" county="San Onofre" latitude="11.192611" municipality="Sucre" name="Tropical Alternohidrigrico" pageId="9" pageNumber="10" stateProvince="Magdalena">Tropical Alternohidrígrico</location>
, 
<geoCoordinate id="EE0EFAA1B62AC37EFBAF0B51A4CCDB63" box="[1091,1254,1879,1905]" degrees="11" direction="north" minutes="11" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="33.4" value="11.192611">11º11´33.4´´N</geoCoordinate>
; 74º/0´24.4´´W. WGS 
<quantity id="4CC23183B62AC37EFF340B7DA163DB87" box="[216,329,1915,1941]" metricMagnitude="4" metricUnit="m" metricValue="8.4347" pageId="9" pageNumber="10" unit="m" value="84347.0">
<elevation id="00177B55B62AC37EFF340B7DA163DB87" box="[216,329,1915,1941]" metricMagnitude="4" metricUnit="m" metricValue="8.4347" pageId="9" pageNumber="10" unit="m" value="84347.0">84, 347 m</elevation>
</quantity>
. Trampa / de Luz Negra, Manual, 
<date id="FF84BAA6B62AC37EFD250B7DA31CDB87" box="[713,822,1915,1941]" pageId="9" pageNumber="10" value="2019-06-04">
<collectingDate id="EFC0434EB62AC37EFD250B7DA31CDB87" box="[713,822,1915,1941]" pageId="9" pageNumber="10" value="2019-06-04">4.vi.2019</collectingDate>
</date>
, / Jhon 
<collectingRegion id="49FE5284B62AC37EFC600B7DA3E6DB87" box="[908,972,1915,1941]" country="Colombia" name="Cesar" pageId="9" pageNumber="10">César</collectingRegion>
Neita Moreno” [
<specimenCount id="9D3C57EFB62AC37EFB650B7CA4BCDB87" box="[1161,1174,1914,1941]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
-IAvH-E-216386]
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>