424 lines
46 KiB
XML
424 lines
46 KiB
XML
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<document id="39221FC238F0980A4435F50F10F5849F" ID-DOI="10.1080/002229301300323910" ID-ISSN="1464-5262" ID-Zenodo-Dep="5277129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630022732653" checkinUser="felipe" docAuthor="Gussmann, S. M. V. & Scholtz, C. H." docDate="2001" docId="DA651E6CFF83FB33FE403586FE60FEB6" docLanguage="en" docName="JNatHist.35.7.1013-1084.pdf" docOrigin="Journal of Natural History 35 (7)" docSource="http://www.tandfonline.com/doi/abs/10.1080/002229301300323910" docStyle="DocumentStyle:B6EF65DE5B4D2965D4D236BA736D172F.2:JNatHist.2000-2002.journal_article.2cover" docStyleId="B6EF65DE5B4D2965D4D236BA736D172F" docStyleName="JNatHist.2000-2002.journal_article.2cover" docStyleVersion="2" docTitle="Bolbaoeer nikolajevi Gussmann & Scholtz 2001, sp. nov." docType="treatment" docVersion="4" lastPageNumber="1059" masterDocId="265C6614FFADFB02FFDA3151FFDBFFCD" masterDocTitle="Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species" masterLastPageNumber="1084" masterPageNumber="1013" pageNumber="1057" updateTime="1699137266390" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="33507900C1D9D8C34F1CB47D85D1609F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5BD292186F0CB510C5CF77A2422EE36F">
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<mods:title id="B579A0CA30927CF95B9AD77465C98FE9">Systematic revision of the Afrotropical genus BolbaOEer Vulcano, Martinez and Pereira (Coleoptera: Scarabaeoidea: Bolboceratidae: Bolboceratinae), with descriptions of eight new species</mods:title>
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</mods:titleInfo>
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<mods:name id="FE7818E0D697BDEB82DC8D2F7F791B23" type="personal">
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</mods:role>
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<mods:namePart id="47C3273DFCDDF840A6C77763BA25F4DE">Gussmann, S. M. V.</mods:namePart>
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</mods:name>
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<mods:name id="45969F47BE9FBAF2F750247378025506" type="personal">
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<mods:roleTerm id="315B31D9B61BACF7D2DAB2B65B952F97">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8318481C5F8A464D37D1AE14571765A4">Scholtz, C. H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="690BD6F29642138324A5A41957585F81">text</mods:typeOfResource>
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<mods:relatedItem id="2885BC72F9FB9B8CE0B76E30B13BEAB7" type="host">
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<mods:titleInfo id="39AF588B20FE6A276E40C188A9D8A9A5">
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<mods:title id="6C54627D80324E4865EEA58585DE4675">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="653BB51D24FF2353B508EFC1E59674AE">
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<mods:date id="0AD2C1F4FDF569FB378333B5DE0A417B">2001</mods:date>
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<mods:detail id="C73C99797043FDC72AC0768115087EB7" type="pubDate">
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<mods:number id="7D093CDFE15C04AEA0C2E15762F3A949">2001-07-31</mods:number>
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</mods:detail>
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<mods:detail id="AB273C45AD6DCE3D6F229F8D22AD8775" type="volume">
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<mods:number id="83CD7A42BC6EB829901C0F5764FBE011">35</mods:number>
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</mods:detail>
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<mods:detail id="A96E4C1388F456618F5D6EA3B23A17E5" type="issue">
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<mods:number id="DC0413EDF51A0CE8678737B3E2443685">7</mods:number>
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</mods:detail>
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<mods:extent id="2038E12CBB1E020C288B734AD7D6D07D" unit="page">
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<mods:start id="D1249874A083CFD8DA4960A499DB98A8">1013</mods:start>
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<mods:end id="9CDCA3EFD3DDA3BC521CF58A5A6DE6E4">1084</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="A2DB3240069D8BD61D3DE327787056AA">
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<mods:url id="CB0698DCE5728957352EFE8A6E13BC25">http://www.tandfonline.com/doi/abs/10.1080/002229301300323910</mods:url>
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</mods:location>
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<mods:classification id="F2F584D38B219FEEEE8925E7EBA612C3">journal article</mods:classification>
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<mods:identifier id="BE9543486AEA823075E8FE0A1382B475" type="DOI">10.1080/002229301300323910</mods:identifier>
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<mods:identifier id="B0729121F8D3AB6739CD2D8FE8FC1D0E" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="AD290FA60773AF4CD3B50DCB89A5B96D" type="Zenodo-Dep">5277129</mods:identifier>
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</mods:mods>
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<treatment id="DA651E6CFF83FB33FE403586FE60FEB6" ID-DOI="http://doi.org/10.5281/zenodo.5279278" ID-Zenodo-Dep="5279278" LSID="urn:lsid:plazi:treatment:DA651E6CFF83FB33FE403586FE60FEB6" httpUri="http://treatment.plazi.org/id/DA651E6CFF83FB33FE403586FE60FEB6" lastPageId="49" lastPageNumber="1059" pageId="46" pageNumber="1057">
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<subSubSection id="1AD6FCF1FF83FB2CFE403586FD01FAC2" pageId="46" pageNumber="1057" type="nomenclature">
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<paragraph id="5273AF7AFF83FB2CFE403586FD0CFB22" blockId="46.[408,730,1239,1296]" box="[410,727,1239,1263]" pageId="46" pageNumber="1057">
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<heading id="093B1816FF83FB2CFE403586FD0CFB22" box="[410,727,1239,1263]" centered="true" fontSize="10" level="2" pageId="46" pageNumber="1057" reason="3">
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<taxonomicName id="95CCD4F9FF83FB2CFE403586FDAFFB22" authority="Gussmann & Scholtz, 2001" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[410,628,1239,1263]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="1057" phylum="Arthropoda" rank="species" species="nikolajevi" status="sp. nov.">
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<emphasis id="60B87368FF83FB2CFE403586FDAFFB22" bold="true" box="[410,628,1239,1263]" italics="true" pageId="46" pageNumber="1057">BolbaOEer nikolajevi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7B8BCE13FF83FB2CFDA73589FD0CFB22" box="[637,727,1240,1263]" pageId="46" pageNumber="1057" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="5273AF7AFF83FB2CFE4235A6FD01FAC2" blockId="46.[408,730,1239,1296]" box="[408,730,1271,1296]" pageId="46" pageNumber="1057">
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(®gures 2c,
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<quantity id="9534029FFF83FB2CFDC135A6FDEAFAC2" box="[539,561,1271,1295]" metricMagnitude="3" metricUnit="kg" metricValue="3.0" pageId="46" pageNumber="1057" unit="t" value="3.0">3t</quantity>
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±w, 17a±g, 18)
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</paragraph>
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</subSubSection>
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<subSubSection id="1AD6FCF1FF83FB33FF863410FE60FEB6" lastPageId="49" lastPageNumber="1060" pageId="46" pageNumber="1057" type="description">
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<paragraph id="5273AF7AFF83FB2CFF863410FEC3FA95" blockId="46.[92,1047,1344,1687]" box="[92,280,1344,1368]" pageId="46" pageNumber="1057">
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<emphasis id="60B87368FF83FB2CFF863410FEC3FA95" box="[92,280,1344,1368]" italics="true" pageId="46" pageNumber="1057">Description male</emphasis>
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</paragraph>
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<paragraph id="5273AF7AFF83FB2DFF5E3431FF2CFCD4" blockId="46.[92,1047,1344,1687]" lastBlockId="47.[92,1047,132,794]" lastPageId="47" lastPageNumber="1058" pageId="46" pageNumber="1057">
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<emphasis id="60B87368FF83FB2CFF5E3431FED7FAB5" box="[132,268,1376,1400]" italics="true" pageId="46" pageNumber="1057">Body length</emphasis>
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<quantity id="9534029FFF83FB2CFEC23431FE6EFABA" box="[280,437,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.21" metricValueMin="1.9899999999999998" pageId="46" pageNumber="1057" unit="mm" value="21.0" valueMax="22.1" valueMin="19.9">19.9±22.1 mm</quantity>
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(
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<specimenCount id="44CA64F3FF83FB2CFE173431FD82FAB5" box="[461,601,1376,1400]" pageId="46" pageNumber="1057" type="generic">5 specimens</specimenCount>
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).
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<emphasis id="60B87368FF83FB2CFDAF3431FD1BFAB5" box="[629,704,1376,1400]" italics="true" pageId="46" pageNumber="1057">Colour</emphasis>
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light brown to dark brown.
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<emphasis id="60B87368FF83FB2CFF8634D1FE55FA55" box="[92,398,1408,1432]" italics="true" pageId="46" pageNumber="1057">Outer margin of mandible</emphasis>
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feebly sinuate (®gure 2c).
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<emphasis id="60B87368FF83FB2CFD0F34D1FCF6FA55" box="[725,813,1408,1432]" italics="true" pageId="46" pageNumber="1057">Clypeus</emphasis>
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with bituberculate, feebly arcuate transverse posterior carina; tubercles in line with antennal insertions (®gure 2c); posterior carina as long as or shorter than anterior carina.
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<emphasis id="60B87368FF83FB2CFC563490FC11FA15" box="[908,970,1473,1496]" italics="true" pageId="46" pageNumber="1057">Frons</emphasis>
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feebly depressed posterior to posterior clypeal carina.
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<emphasis id="60B87368FF83FB2CFDA4348EFCCCFA3A" box="[638,791,1503,1527]" italics="true" pageId="46" pageNumber="1057">Antennal club</emphasis>
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with glabrous area of basal segment about one-half of exposed surface.
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<emphasis id="60B87368FF83FB2CFD703751FCCCF9DA" box="[682,791,1536,1559]" italics="true" pageId="46" pageNumber="1057">Pronotum</emphasis>
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with anterior margin medially raised, in major males in dorsal view with bisinuate curvature and slightly projecting over frons (®gure 2c), in minor males in dorsal view with simple curvature and not projecting over frons; with one outer and one inner pair of equally long, slightly inward curved horns along posterior margin in major males (®gure
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<quantity id="9534029FFF83FB2CFC65372FFC0DF95B" box="[959,982,1662,1686]" metricMagnitude="3" metricUnit="kg" metricValue="3.0" pageId="46" pageNumber="1057" unit="t" value="3.0">3t</quantity>
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), but protrusions more approximated, shorter and eventually reduced to small tubercles in minor males (®gure 3u, v); disc sometimes with scattered ®ne punctures but always with large, fairly densely spaced punctures, these increasingly conūent and more densely spaced towards sides and along anterior margin; punctures decreasing in size and density and increasingly interspersed by ®ne punctures usually in between and always on pronotal protrusions.
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<taxonomicName id="95CCD4F9FF82FB2DFDFC3075FD49FEF1" box="[550,658,292,316]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="47" pageNumber="1058" phylum="Arthropoda" rank="genus">
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<emphasis id="60B87368FF82FB2DFDFC3075FD49FEF1" box="[550,658,292,316]" italics="true" pageId="47" pageNumber="1058">Scutellum</emphasis>
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</taxonomicName>
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with surface coarsely punctate.
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<emphasis id="60B87368FF82FB2DFF863015FF74FE91" box="[92,175,324,348]" italics="true" pageId="47" pageNumber="1058">Elytron</emphasis>
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with strial punctures separate d by three and a half to ®ve puncture diameters; intervals with approximately four to ®ve punctures between two striae.
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<emphasis id="60B87368FF82FB2DFC663035FBCCFEB1" box="[956,1047,356,380]" italics="true" pageId="47" pageNumber="1058">Protibia</emphasis>
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®ve-dentate.
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<emphasis id="60B87368FF82FB2DFF2F30D2FE4EFE56" box="[245,405,387,411]" italics="true" pageId="47" pageNumber="1058">Protibial spur</emphasis>
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more or less as long as ®fth tarsomere, acuminate.
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<emphasis id="60B87368FF82FB2DFF8630F2FED3FE76" box="[92,264,419,443]" italics="true" pageId="47" pageNumber="1058">Metatrochanter</emphasis>
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of unmodi®ed shape; with very dense, long setation except for an asetose area along anterior margin where metatrochanter is attached to metafemur.
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<emphasis id="60B87368FF82FB2DFF8630B2FF0CFE36" box="[92,215,483,507]" italics="true" pageId="47" pageNumber="1058">Metafemur</emphasis>
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in ventral view in posterior third with line of densely spaced, moderately long and long setae; with dense, long setae in basal area and basal third along posterior margin; with fairly dense, long setae in anterior third; remaining areas with sparse, long setation.
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<emphasis id="60B87368FF82FB2DFE4E3313FDDAFD97" box="[404,513,578,602]" italics="true" pageId="47" pageNumber="1058">Metatibia</emphasis>
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in lateral view with subapical carina bilobed; with unmodi®ed spurs.
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<emphasis id="60B87368FF82FB2DFEB23333FE0DFDB7" box="[360,470,610,634]" italics="true" pageId="47" pageNumber="1058">Underside</emphasis>
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with posterior margin of sternite 4 medially feebly and roundedly produced (®gure 17b) and with dense setation; posterior margin of sternite 5 broadly triangularly emarginate (®gure 17b); posterior margin of sternite 6 medially deeply divided as in ®gure 17b; pygidium unmodi®ed (®gure 17a).
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<emphasis id="60B87368FF82FB2DFF8633B3FF1AFD37" box="[92,193,738,762]" italics="true" pageId="47" pageNumber="1058">Genitalia</emphasis>
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with aedeagus as in ®gure 17c±e; genital capsule apically with moderately long setation.
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</paragraph>
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<paragraph id="5273AF7AFF82FB2DFF863212FEF6FC97" blockId="47.[92,1047,834,1692]" box="[92,301,834,858]" pageId="47" pageNumber="1058">
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<emphasis id="60B87368FF82FB2DFF863212FEF6FC97" box="[92,301,834,858]" italics="true" pageId="47" pageNumber="1058">Description female</emphasis>
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</paragraph>
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<paragraph id="5273AF7AFF82FB2DFF5E3233FE26FA7A" blockId="47.[92,1047,834,1692]" pageId="47" pageNumber="1058">
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<emphasis id="60B87368FF82FB2DFF5E3233FEDCFCB7" box="[132,263,866,890]" italics="true" pageId="47" pageNumber="1058">Body length</emphasis>
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<quantity id="9534029FFF82FB2DFED53233FE77FCB4" box="[271,428,866,890]" metricMagnitude="-2" metricUnit="m" metricValue="1.9799999999999998" metricValueMax="2.04" metricValueMin="1.92" pageId="47" pageNumber="1058" unit="mm" value="19.799999999999997" valueMax="20.4" valueMin="19.2">19.2±20.4 mm</quantity>
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(
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<specimenCount id="44CA64F3FF82FB2DFE653233FD9DFCB7" box="[447,582,866,890]" pageId="47" pageNumber="1058" type="generic">7 specimens</specimenCount>
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).
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<emphasis id="60B87368FF82FB2DFD873233FD73FCB7" box="[605,680,866,890]" italics="true" pageId="47" pageNumber="1058">Colour</emphasis>
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as in male.
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<emphasis id="60B87368FF82FB2DFCEB3233FF65FC57" italics="true" pageId="47" pageNumber="1058">Outer margin female mandible</emphasis>
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rounded.
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<emphasis id="60B87368FF82FB2DFEEE32D2FEA9FC57" box="[308,370,899,922]" italics="true" pageId="47" pageNumber="1058">Frons</emphasis>
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with distinctly raised bituberculate, transverse carina; position of carina variable, from in line with to mostly slightly posterior to posterolateral genal angle; carina straight and distinctly shorter than, usually half length of anterior clypeal carina.
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<emphasis id="60B87368FF82FB2DFEB632B0FDDEFC34" box="[364,517,993,1017]" italics="true" pageId="47" pageNumber="1058">Antennal club</emphasis>
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as in male.
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<emphasis id="60B87368FF82FB2DFD4832B3FD24FC34" box="[658,767,994,1017]" italics="true" pageId="47" pageNumber="1058">Pronotum</emphasis>
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with anterior margin as in minor males; with transverse, feebly carinate swelling at median portion of disc; carina in dorsal view more or less straight, in frontal view more or less straight but laterally arcuate (®gure 3w); swelling either without or with very few ®ne punctures, but always with some large punctures; with large, mostly conūent punctures anterior, lateral and posterior to carinate swelling.
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<emphasis id="60B87368FF82FB2DFD7B35D1FC4FFB55" box="[673,916,1152,1176]" italics="true" pageId="47" pageNumber="1058">
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<taxonomicName id="95CCD4F9FF82FB2DFD7B35D1FCD5FB55" box="[673,782,1152,1176]" class="Trilobita" family="Styginidae" genus="Scutellum" kingdom="Animalia" order="Corynexochida" pageId="47" pageNumber="1058" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
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and elytron
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</emphasis>
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as in male.
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<emphasis id="60B87368FF82FB2DFF8635F1FE55FB75" box="[92,398,1184,1208]" italics="true" pageId="47" pageNumber="1058">Protibia and protibial spur</emphasis>
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as in male.
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<emphasis id="60B87368FF82FB2DFDF235F1FD0FFB75" box="[552,724,1184,1208]" italics="true" pageId="47" pageNumber="1058">Metatrochanter</emphasis>
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of unmodi®ed shape; with sparse, long setation in basal third and along posterior margin; surface otherwise asetose.
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<emphasis id="60B87368FF82FB2DFF1B35B1FEE7FB35" box="[193,316,1248,1272]" italics="true" pageId="47" pageNumber="1058">Metafemur</emphasis>
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in ventral view in posterior third with line of fairly dense, moderately long and long setae; with fairly dense and long setae in anterior third and apical area; with fairly dense, moderately long setae along posterior margin; remaining areas with few isolated setae.
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<emphasis id="60B87368FF82FB2DFDFB3411FD55FA95" box="[545,654,1344,1368]" italics="true" pageId="47" pageNumber="1058">Metatibia</emphasis>
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as in male.
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|
<emphasis id="60B87368FF82FB2DFCCF3411FC58FA95" box="[789,899,1344,1368]" italics="true" pageId="47" pageNumber="1058">Underside</emphasis>
|
|||
|
|
with sternite 5 postero-medially feebly but discernibly swollen (®gure 17f); posterior margin of sternite 6 and pygidium medially in ventral view somewhat pointed (®gure
|
|||
|
|
<quantity id="9534029FFF82FB2DFC0634D1FBDEFA55" box="[988,1029,1408,1432]" metricMagnitude="-2" metricUnit="kg" metricValue="1.7" pageId="47" pageNumber="1058" unit="g" value="17.0">17g</quantity>
|
|||
|
|
), otherwise unmodi®ed (®gure 17f, g).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="5273AF7AFF82FB2DFF5E3491FC10FA1A" blockId="47.[92,1047,834,1692]" box="[132,971,1472,1496]" pageId="47" pageNumber="1058">
|
|||
|
|
<emphasis id="60B87368FF82FB2DFF5E3491FED3FA15" box="[132,264,1472,1496]" italics="true" pageId="47" pageNumber="1058">Distribution</emphasis>
|
|||
|
|
(®gure 18). West Africa, from
|
|||
|
|
<collectingCountry id="2ADBEFEAFF82FB2DFD583491FD03FA15" box="[642,728,1472,1496]" name="Senegal" pageId="47" pageNumber="1058">Senegal</collectingCountry>
|
|||
|
|
eastward to
|
|||
|
|
<collectingCountry id="2ADBEFEAFF82FB2DFCB53491FC18FA15" box="[879,963,1472,1496]" name="Nigeria" pageId="47" pageNumber="1058">Nigeria</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="5273AF7AFF82FB32FF5E34B1FDA1FC57" blockId="47.[92,1047,834,1692]" lastBlockId="48.[92,1048,834,1686]" lastPageId="48" lastPageNumber="1059" pageId="47" pageNumber="1058">
|
|||
|
|
<emphasis id="60B87368FF82FB2DFF5E34B1FE4EFA3A" box="[132,405,1503,1527]" italics="true" pageId="47" pageNumber="1058">Type material examined</emphasis>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFE7D34B1FBCAF9DA" collectingDate="1954-07" collectionCode="BMNH" collectorName="W. E. S. Merrett' & W. E. S. Merrett" country="Nigeria" latitude="12.0" location="Kano Dist." longLatPrecision="776" longitude="9.0" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Kano" typeStatus="holotype">
|
|||
|
|
<typeStatus id="8D7711D8FF82FB2DFE7D34B1FDC1FA3A" box="[423,538,1503,1527]" pageId="47" pageNumber="1058" type="holotype">HOLOTYPE</typeStatus>
|
|||
|
|
l[diss.]:`
|
|||
|
|
<collectingRegion id="90086198FF82FB2DFD47348EFCC4FA3A" box="[669,799,1503,1527]" country="Nigeria" name="Kano" pageId="47" pageNumber="1058">Kano Dist.</collectingRegion>
|
|||
|
|
[
|
|||
|
|
<geoCoordinate id="37F8C9BDFF82FB2DFCE9348EFC5EFA3A" box="[819,901,1503,1527]" degrees="12.00" direction="north" orientation="latitude" pageId="47" pageNumber="1058" precision="555" value="12.0">12.00N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="37F8C9BDFF82FB2DFC55348EFC05FA3A" box="[911,990,1503,1527]" degrees="09.00" direction="east" orientation="longitude" pageId="47" pageNumber="1058" precision="555" value="9.0">09.00E</geoCoordinate>
|
|||
|
|
], N.
|
|||
|
|
<collectingCountry id="2ADBEFEAFF82FB2DFF8634AEFF6BF9DA" box="[92,176,1535,1559]" name="Nigeria" pageId="47" pageNumber="1058">Nigeria</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<date id="267289BAFF82FB2DFF1E34AEFEEEF9DA" box="[196,309,1535,1559]" pageId="47" pageNumber="1058" value="1954-07">
|
|||
|
|
<collectingDate id="36367052FF82FB2DFF1E34AEFEEEF9DA" box="[196,309,1535,1559]" pageId="47" pageNumber="1058" value="1954-07">July 1954</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="FF39CAACFF82FB2DFE9034AEFDD8F9DA" box="[330,515,1535,1559]" pageId="47" pageNumber="1058">W.E.S. Merrett’</collectorName>
|
|||
|
|
<emphasis id="60B87368FF82FB2DFDDC34AEFDCDF9DA" bold="true" box="[518,534,1535,1565]" pageId="47" pageNumber="1058">/`</emphasis>
|
|||
|
|
<collectorName id="FF39CAACFF82FB2DFDCC34AEFD11F9DA" box="[534,714,1535,1559]" pageId="47" pageNumber="1058">W.E.S. Merrett</collectorName>
|
|||
|
|
B.M. 1962-270.’ (
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFC713751FBD8F9DA" box="[939,1027,1536,1559]" pageId="47" pageNumber="1058">BMNH</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFF86374EFDD9F9FA" box="[92,514,1567,1591]" collectingDate="1954-07" collectionCode="BMNH" collectorName="W. E. S. Merrett' & W. E. S. Merrett" country="Nigeria" latitude="12.0" location="Kano Dist." longLatPrecision="776" longitude="9.0" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Kano" typeStatus="allotype">
|
|||
|
|
<typeStatus id="8D7711D8FF82FB2DFF86374EFF1CF9FA" box="[92,199,1567,1591]" pageId="47" pageNumber="1058" type="allotype">ALLOTYPE</typeStatus>
|
|||
|
|
m: same data (
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFEAE3771FE17F9FA" box="[372,460,1568,1591]" pageId="47" pageNumber="1058">BMNH</collectionCode>
|
|||
|
|
); 10
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFDD0374EFBD4F9FA" box="[522,1039,1567,1591]" collectingDate="1954-07" collectionCode="TMSA" collectorName="W. E. S. Merrett' & W. E. S. Merrett" country="Nigeria" elevation="1" latitude="12.0" location="Kano Dist." longLatPrecision="776" longitude="9.0" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Kano" typeStatus="paratype">
|
|||
|
|
<typeStatus id="8D7711D8FF82FB2DFDD0374EFDA6F9FA" box="[522,637,1567,1591]" pageId="47" pageNumber="1058" type="paratype">PARATYPES</typeStatus>
|
|||
|
|
: 1 l[diss.],
|
|||
|
|
<quantity id="9534029FFF82FB2DFCDE374EFCC4F9FA" box="[772,799,1567,1591]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">
|
|||
|
|
<elevation id="D9E14849FF82FB2DFCDE374EFCC4F9FA" box="[772,799,1567,1591]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">1m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
, same data (
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFC6F374EFBDCF9FA" box="[949,1031,1567,1591]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="47" pageNumber="1058">TMSA</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFF86376EFD0DF99A" box="[92,726,1599,1623]" collectingDate="1953-07" collectionCode="BMNH" collectorName="W. E. S. Merrett' & W. E. S. Merrett" country="Nigeria" elevation="1" latitude="12.0" location="Kano Dist." longLatPrecision="776" longitude="9.0" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Kano" typeStatus="paratype">
|
|||
|
|
1 l[diss.],
|
|||
|
|
<quantity id="9534029FFF82FB2DFF01376EFF2DF99A" box="[219,246,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">
|
|||
|
|
<elevation id="D9E14849FF82FB2DFF01376EFF2DF99A" box="[219,246,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">1m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
: same data but:`
|
|||
|
|
<date id="267289BAFF82FB2DFE16376EFD9EF99A" box="[460,581,1599,1623]" pageId="47" pageNumber="1058" value="1953-07">
|
|||
|
|
<collectingDate id="36367052FF82FB2DFE16376EFD9EF99A" box="[460,581,1599,1623]" pageId="47" pageNumber="1058" value="1953-07">July 1953</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
...’ (
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFDAA3711FD13F99A" box="[624,712,1600,1623]" pageId="47" pageNumber="1058">BMNH</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFD32376EFEB1F9BA" collectingDate="1945-08" collectionCode="BMNH" collectorName="W. E. S. Merrett' & W. E. S. Merrett" country="Nigeria" elevation="1" latitude="12.0" location="Kano Dist." longLatPrecision="776" longitude="9.0" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Kano" typeStatus="paratype">
|
|||
|
|
1 l[diss.], same data but:`
|
|||
|
|
<date id="267289BAFF82FB2DFFB8370EFF05F9BA" box="[98,222,1631,1655]" pageId="47" pageNumber="1058" value="1945-08">
|
|||
|
|
<collectingDate id="36367052FF82FB2DFFB8370EFF05F9BA" box="[98,222,1631,1655]" pageId="47" pageNumber="1058" value="1945-08">Aug. 1945</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
...’ (
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFEDE3731FE87F9BA" box="[260,348,1632,1655]" pageId="47" pageNumber="1058">BMNH</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB2DFEAD370EFC39F95B" collectionCode="MNHN" collectorName="E. Laglaize & Museum Paris & Coll. R. Oberthur'" country="Senegal" latitude="12.44" location="Casamance" longLatPrecision="776" longitude="-15.33" pageId="47" pageNumber="1058" specimenCount="1" stateProvince="Sedhiou" typeStatus="paratype">
|
|||
|
|
1, 2mm,`[
|
|||
|
|
<collectingCountry id="2ADBEFEAFF82FB2DFE34370EFD9FF9BA" box="[494,580,1631,1655]" name="Senegal" pageId="47" pageNumber="1058">Senegal</collectingCountry>
|
|||
|
|
],
|
|||
|
|
<location id="5713F9A1FF82FB2DFD85370EFD05F9BA" LSID="urn:lsid:plazi:treatment:DA651E6CFF83FB33FE403586FE60FEB6:5713F9A1FF82FB2DFD85370EFD05F9BA" box="[607,734,1631,1655]" country="Senegal" latitude="12.44" longLatPrecision="776" longitude="-15.33" name="Casamance" pageId="47" pageNumber="1058" stateProvince="Sedhiou">Casamance</location>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="90086198FF82FB2DFD37370EFC92F9BA" box="[749,841,1631,1655]" country="Senegal" name="Sedhiou" pageId="47" pageNumber="1058">Sedhiou</collectingRegion>
|
|||
|
|
[SeÂdhiou,
|
|||
|
|
<geoCoordinate id="37F8C9BDFF82FB2DFC1E370EFBCCF9BA" box="[964,1047,1631,1655]" degrees="12.44" direction="north" orientation="latitude" pageId="47" pageNumber="1058" precision="555" value="12.44">12.44N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="37F8C9BDFF82FB2DFF86372FFF69F95B" box="[92,178,1662,1686]" degrees="15.33" direction="west" orientation="longitude" pageId="47" pageNumber="1058" precision="555" value="-15.33">15.33W</geoCoordinate>
|
|||
|
|
],
|
|||
|
|
<collectorName id="FF39CAACFF82FB2DFF08372EFE8EF95B" box="[210,341,1662,1686]" pageId="47" pageNumber="1058">E. Laglaize</collectorName>
|
|||
|
|
1891’
|
|||
|
|
<emphasis id="60B87368FF82FB2DFE47372FFE77F95B" bold="true" box="[413,428,1662,1692]" pageId="47" pageNumber="1058">/`</emphasis>
|
|||
|
|
<collectorName id="FF39CAACFF82FB2DFE77372FFD8DF95B" box="[429,598,1662,1686]" pageId="47" pageNumber="1058">Museum Paris</collectorName>
|
|||
|
|
ex
|
|||
|
|
<collectorName id="FF39CAACFF82FB2DFD5C372FFC84F95B" box="[646,863,1662,1686]" pageId="47" pageNumber="1058">Coll. R. Oberthur’</collectorName>
|
|||
|
|
(
|
|||
|
|
<collectionCode id="34DD37BFFF82FB2DFCA2372EFC0FF95B" box="[888,980,1663,1686]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="47" pageNumber="1058">MNHN</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF82FB32FC2E372FFE92FCB7" collectionCode="BMNH" collectorName="J. Risbec. & Pres. & Com & Inst. Ent. B. M." country="Senegal" elevation="1" lastPageId="48" lastPageNumber="1059" latitude="14.42" location="Bambey" longLatPrecision="773" longitude="-16.28" pageId="47" pageNumber="1058" specimenCount="1" typeStatus="paratype">
|
|||
|
|
<quantity id="9534029FFF82FB2DFC2E372FFBD4F95B" box="[1012,1039,1662,1686]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">
|
|||
|
|
<elevation id="D9E14849FF82FB2DFC2E372FFBD4F95B" box="[1012,1039,1662,1686]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="1058" unit="m" value="1.0">1m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,`
|
|||
|
|
<collectingCountry id="2ADBEFEAFF9DFB32FFB83213FF63FC97" box="[98,184,834,858]" name="Senegal" pageId="48" pageNumber="1059">Senegal</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<location id="5713F9A1FF9DFB32FF123213FEFFFC97" LSID="urn:lsid:plazi:treatment:DA651E6CFF83FB33FE403586FE60FEB6:5713F9A1FF9DFB32FF123213FEFFFC97" box="[200,292,834,858]" country="Senegal" latitude="14.42" longLatPrecision="773" longitude="-16.28" name="Bambey" pageId="48" pageNumber="1059">Bambey</location>
|
|||
|
|
[
|
|||
|
|
<geoCoordinate id="37F8C9BDFF9DFB32FEEF3213FE5CFC94" box="[309,391,834,858]" degrees="14.42" direction="north" orientation="latitude" pageId="48" pageNumber="1059" precision="555" value="14.42">14.42N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="37F8C9BDFF9DFB32FE553213FE3EFC94" box="[399,485,834,858]" degrees="16.28" direction="west" orientation="longitude" pageId="48" pageNumber="1059" precision="555" value="-16.28">16.28W</geoCoordinate>
|
|||
|
|
], 1943,
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FD9E3213FD71FC97" box="[580,682,834,858]" pageId="48" pageNumber="1059">J.Risbec.</collectorName>
|
|||
|
|
’
|
|||
|
|
<emphasis id="60B87368FF9DFB32FD6B3213FD1AFC97" bold="true" box="[689,705,834,864]" pageId="48" pageNumber="1059">/`</emphasis>
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FD1B3213FD22FC94" box="[705,761,834,858]" pageId="48" pageNumber="1059">Pres.</collectorName>
|
|||
|
|
by
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FCFE3213FC87FC97" box="[804,860,834,858]" pageId="48" pageNumber="1059">Com</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FCB03213FBCDFC94" box="[874,1046,834,857]" pageId="48" pageNumber="1059">Inst. Ent. B.M.</collectorName>
|
|||
|
|
1948-525.’ (
|
|||
|
|
<collectionCode id="34DD37BFFF9DFB32FF393233FEE0FCB4" box="[227,315,866,889]" pageId="48" pageNumber="1059">BMNH</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation id="E2A4A527FF9DFB32FE823233FDADFC57" collectionCode="MRAC" collectorName="Coll. Mus. Tervuren & P. C. Fernandez" country="Burkina Faso" elevation="1" latitude="12.22" location="Ougadougo" longLatPrecision="776" longitude="-1.31" pageId="48" pageNumber="1059" specimenCount="1" typeStatus="paratype">
|
|||
|
|
<quantity id="9534029FFF9DFB32FE823233FEA8FCB7" box="[344,371,866,890]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="1059" unit="m" value="1.0">
|
|||
|
|
<elevation id="D9E14849FF9DFB32FE823233FEA8FCB7" box="[344,371,866,890]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="48" pageNumber="1059" unit="m" value="1.0">1m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,`
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FE503233FDA9FCB4" box="[394,626,866,890]" pageId="48" pageNumber="1059">Coll. Mus. Tervuren</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="2ADBEFEAFF9DFB32FD593233FCD5FCB7" box="[643,782,866,890]" name="Burkina Faso" pageId="48" pageNumber="1059">Haute-Volta</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<location id="5713F9A1FF9DFB32FCC53233FC79FCB7" LSID="urn:lsid:plazi:treatment:DA651E6CFF83FB33FE403586FE60FEB6:5713F9A1FF9DFB32FCC53233FC79FCB7" box="[799,930,866,890]" country="Burkina Faso" latitude="12.22" longLatPrecision="776" longitude="-1.31" name="Ougadougo" pageId="48" pageNumber="1059">Ougadougo</location>
|
|||
|
|
u [
|
|||
|
|
<geoCoordinate id="37F8C9BDFF9DFB32FC1E3233FBCCFCB4" box="[964,1047,866,890]" degrees="12.22" direction="north" orientation="latitude" pageId="48" pageNumber="1059" precision="555" value="12.22">12.22N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="37F8C9BDFF9DFB32FF8632D3FF69FC54" box="[92,178,898,922]" degrees="01.31" direction="west" orientation="longitude" pageId="48" pageNumber="1059" precision="555" value="-1.31">01.31W</geoCoordinate>
|
|||
|
|
] (leg.
|
|||
|
|
<collectorName id="FF39CAACFF9DFB32FED932D3FE70FC57" box="[259,427,898,922]" pageId="48" pageNumber="1059">P.C.Fernandez</collectorName>
|
|||
|
|
) X-70’ (
|
|||
|
|
<collectionCode id="34DD37BFFF9DFB32FDCA32D3FDBDFC57" box="[528,614,898,922]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="48" pageNumber="1059" type="Museum">MRAC</collectionCode>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="06B3FFF2FF9DFB32FF7C3386FC16FD21" ID-DOI="http://doi.org/10.5281/zenodo.5277161" ID-Zenodo-Dep="5277161" box="[166,973,726,749]" httpUri="https://zenodo.org/record/5277161/files/figure.png" pageId="48" pageNumber="1059" startId="48.[166,182,727,748]" targetBox="[95,1045,133,698]" targetPageId="48">
|
|||
|
|
<paragraph id="5273AF7AFF9DFB32FF7C3386FC16FD21" blockId="48.[166,973,726,749]" box="[166,973,726,749]" pageId="48" pageNumber="1059">
|
|||
|
|
FIG. 18. Distributional records:
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FE253386FD00FD20" authorityName="Gussmann & Scholtz" authorityYear="2001" baseAuthorityName="Gussmann & Scholtz" baseAuthorityYear="2001" box="[511,731,727,749]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="bidenticollis">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FE253386FD00FD20" box="[511,731,727,749]" italics="true" pageId="48" pageNumber="1059">BolbaOEer bidenticollis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="60B87368FF9DFB32FD303387FD25FD21" bold="true" box="[746,766,726,748]" pageId="48" pageNumber="1059">_</emphasis>
|
|||
|
|
);
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FCCD3389FC4EFD20" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[791,917,727,749]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FCCD3389FC4EFD20" box="[791,917,727,749]" italics="true" pageId="48" pageNumber="1059">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="60B87368FF9DFB32FC7F3387FC62FD21" bold="true" box="[933,953,726,748]" pageId="48" pageNumber="1059">E</emphasis>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="5273AF7AFF9DFB32FF5E32F3FDF5FC14" blockId="48.[92,1048,834,1686]" pageId="48" pageNumber="1059">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FF5E32F3FF20FC77" box="[132,251,930,954]" italics="true" pageId="48" pageNumber="1059">Etymology</emphasis>
|
|||
|
|
. The species is named after Dr Georgy Nikolajev in recognition of his revisional work on
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FEBE3290FE11FC14" box="[356,458,961,985]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FEBE3290FE11FC14" box="[356,458,961,985]" italics="true" pageId="48" pageNumber="1059">BolbaOEer</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in 1982.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="5273AF7AFF9DFB32FF5E32B0FBCCFAD5" blockId="48.[92,1048,834,1686]" pageId="48" pageNumber="1059">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FF5E32B0FF23FC34" box="[132,248,993,1017]" italics="true" pageId="48" pageNumber="1059">Comments</emphasis>
|
|||
|
|
. Males of this new species are easily identi®ed by the following combination of character states: always with four, equally long protrusions along posterior pronotal margin (®gure
|
|||
|
|
<quantity id="9534029FFF9DFB32FE0D3570FE36FBF4" box="[471,493,1057,1081]" metricMagnitude="3" metricUnit="kg" metricValue="3.0" pageId="48" pageNumber="1059" unit="t" value="3.0">3t</quantity>
|
|||
|
|
±v); posterior margin of sternite 4 medially feebly and roundedly produced (®gure 17b) and with dense setation; posterior margin of sternite 5 broadly triangularly incised (®gure 17b); sternite 6 medially deeply divided and bilobed (®gure 17b), lobes apically densely setose; pygidium evenly rounded, unmodi®ed (®gure 17a); and distinct aedeagus with relatively broad parameres, their tips acute (®gure 17c±e). A number of character states shared with or similar to
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FC243590FF6DFB35" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="princeps">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FC243590FF6DFB35" italics="true" pageId="48" pageNumber="1059">B. princeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FF2E35B0FE94FB35" box="[244,335,1249,1272]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="gigas">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FF2E35B0FE94FB35" box="[244,335,1249,1272]" italics="true" pageId="48" pageNumber="1059">B. gigas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
prompted us to place
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FD8235B0FD3EFB35" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[600,741,1248,1272]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FD8235B0FD3EFB35" box="[600,741,1248,1272]" italics="true" pageId="48" pageNumber="1059">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in the
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FCE035B0FC6CFB35" box="[826,951,1249,1272]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="princeps">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FCE035B0FC6CFB35" box="[826,951,1249,1272]" italics="true" pageId="48" pageNumber="1059">B. princeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
speciesgroup, as already explained above under the discussion of the various species-groups.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="5273AF7AFF9DFB33FF5E3471FE60FEB6" blockId="48.[92,1048,834,1686]" lastBlockId="49.[92,1048,132,380]" lastPageId="49" lastPageNumber="1060" pageId="48" pageNumber="1059">
|
|||
|
|
We are con®dent that the female specimens designated above as
|
|||
|
|
<typeStatus id="8D7711D8FF9DFB32FC5C3471FC3AFAF5" box="[902,993,1312,1336]" pageId="48" pageNumber="1059" type="allotype">allotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<typeStatus id="8D7711D8FF9DFB32FF863411FF11FA95" box="[92,202,1344,1368]" pageId="48" pageNumber="1059" type="paratype">paratypes</typeStatus>
|
|||
|
|
are correctly associated with the males because most of them were collected together in male
|
|||
|
|
<emphasis id="60B87368FF9DFB32FECF3431FEC5FAB3" bold="true" box="[277,286,1376,1406]" pageId="48" pageNumber="1059">/</emphasis>
|
|||
|
|
female combinations. Male
|
|||
|
|
<emphasis id="60B87368FF9DFB32FD893431FD87FAB3" bold="true" box="[595,604,1376,1406]" pageId="48" pageNumber="1059">/</emphasis>
|
|||
|
|
female combinations of
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FCB53430FC38FAB5" box="[879,995,1376,1400]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="bremeri">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FCB53430FC38FAB5" box="[879,995,1376,1400]" italics="true" pageId="48" pageNumber="1059">B. bremeri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FF8634D0FF0CFA55" box="[92,215,1409,1432]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="princeps">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FF8634D0FF0CFA55" box="[92,215,1409,1432]" italics="true" pageId="48" pageNumber="1059">B. princeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
were, incidentally, also collected from the
|
|||
|
|
<typeStatus id="8D7711D8FF9DFB32FD1834D1FCFDFA55" box="[706,806,1408,1432]" pageId="48" pageNumber="1059" type="holotype">holotype</typeStatus>
|
|||
|
|
locality of
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FC7234D0FF59FA75" authorityName="Gussmann & Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FC7234D0FF59FA75" italics="true" pageId="48" pageNumber="1059">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, with all of them sharing identical label data. But because the male
|
|||
|
|
<emphasis id="60B87368FF9DFB32FC1934F1FC17FA73" bold="true" box="[963,972,1440,1470]" pageId="48" pageNumber="1059">/</emphasis>
|
|||
|
|
female associations for
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FEC63490FE48FA15" box="[284,403,1472,1496]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="bremeri">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FEC63490FE48FA15" box="[284,403,1472,1496]" italics="true" pageId="48" pageNumber="1059">B. bremeri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FE0F3490FD88FA15" box="[469,595,1473,1496]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="princeps">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FE0F3490FD88FA15" box="[469,595,1473,1496]" italics="true" pageId="48" pageNumber="1059">B. princeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are already clearly manifested from a number of other localities not shared by
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FDE734B1FD10FA3A" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[573,715,1503,1527]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FDE734B1FD10FA3A" box="[573,715,1503,1527]" italics="true" pageId="48" pageNumber="1059">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, a possible mix up is out of the question. Females of
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FEA63751FDDCF9DA" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[380,519,1535,1559]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FEA63751FDDCF9DA" box="[380,519,1535,1559]" italics="true" pageId="48" pageNumber="1059">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be identi®ed in ventral view by the feeble but discernible postero-median swelling on sternite 5 and the somewhat pointed posterior margins of the otherwise unmodi®ed sternite 6 and pygidium. Although females of members of the
|
|||
|
|
<taxonomicName id="95CCD4F9FF9DFB32FE763731FD95F9BA" box="[428,590,1631,1655]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="1059" phylum="Arthropoda" rank="species" species="abyssinicus">
|
|||
|
|
<emphasis id="60B87368FF9DFB32FE763731FD95F9BA" box="[428,590,1631,1655]" italics="true" pageId="48" pageNumber="1059">B. abyssinicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species-group also display a more or less pronounced swelling on sternite 5, they are immediately distinguishable from
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FF8631D4FF3DFF51" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[92,230,132,156]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FF8631D4FF3DFF51" box="[92,230,132,156]" italics="true" pageId="49" pageNumber="1060">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by their strongly modi®ed pygidium (e.g. compare ®gures 17f and 24h). Furthermore, at least with regard to females of
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FD4231F4FCDCFF71" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[664,775,164,188]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="sasakii">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FD4231F4FCDCFF71" box="[664,775,164,188]" italics="true" pageId="49" pageNumber="1060">B. sasakii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FCC031F4FC67FF71" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[794,956,164,188]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="namibiensis">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FCC031F4FC67FF71" box="[794,956,164,188]" italics="true" pageId="49" pageNumber="1060">B. namibiensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FC2431F4FF1DFF11" authorityName="Gussmann & Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="sebastiani">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FC2431F4FF1DFF11" italics="true" pageId="49" pageNumber="1060">B. sebastiani</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the pronotal punctation consists of small, deep and densely spaced punctures anterior to transverse carinate swelling as opposed to large, shallow and densely spaced punctures in
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FE8B3054FE3BFED1" authorityName="Gussmann & Scholtz" authorityYear="2001" box="[337,480,260,284]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FE8B3054FE3BFED1" box="[337,480,260,284]" italics="true" pageId="49" pageNumber="1060">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Lastly the
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FDAF3054FCCEFED1" box="[629,789,260,284]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="abyssinicus">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FDAF3054FCCEFED1" box="[629,789,260,284]" italics="true" pageId="49" pageNumber="1060">B. abyssinicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species-group and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FC243054FF1EFEF1" authorityName="Gussmann & Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FC243054FF1EFEF1" italics="true" pageId="49" pageNumber="1060">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are clearly allopatric (compare ®gure 18 with 25). DiOEerences between
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FC243074FF1EFE91" authorityName="Gussmann & Scholtz" authorityYear="2001" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="nikolajevi">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FC243074FF1EFE91" italics="true" pageId="49" pageNumber="1060">B. nikolajevi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
females and females of the closely related
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FD103014FC93FE91" box="[714,840,325,348]" class="Insecta" family="Bolboceratidae" genus="Bolboceras" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="princeps">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FD103014FC93FE91" box="[714,840,325,348]" italics="true" pageId="49" pageNumber="1060">B. princeps</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="95CCD4F9FF9CFB33FC503014FC33FE91" box="[906,1000,325,348]" class="Insecta" family="Bolboceratidae" genus="Bolbaoeer" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="1060" phylum="Arthropoda" rank="species" species="gigas">
|
|||
|
|
<emphasis id="60B87368FF9CFB33FC503014FC33FE91" box="[906,1000,325,348]" italics="true" pageId="49" pageNumber="1060">B. gigas</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are discussed under the latter two.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|