treatments-xml/data/03/F3/87/03F387B2FFDB5317CD245096FC8BA424.xml

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<mods:title id="007F4E5B2E235B2D47B72C950708804B">Revision of some types of North American aleocharines (Coleoptera: Staphylinidae: Aleocharinae), with synonymic notes</mods:title>
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<taxonomicName id="4C5A4D27FFDB5317CD245096FDE9A1E1" ID-CoL="8VRN8" authority="Pace, 1982" authorityName="Pace" authorityYear="1982" box="[264,561,964,990]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CD245096FDE9A1E1" bold="true" box="[264,561,964,990]" pageId="91" pageNumber="92">
<emphasis id="B92EEAB6FFDB5317CD245096FE7EA1E1" bold="true" box="[264,422,964,990]" italics="true" pageId="91" pageNumber="92">Emmelostiba</emphasis>
<bibRefCitation id="EFCB4B55FFDB5317CD815096FDE9A1E1" author="Pace, R." box="[429,561,964,990]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace, 1982</bibRefCitation>
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<treatmentCitation id="0AFB10B5FFDB5317CD24575DFD93A61A" author="Pace, R." box="[264,587,1038,1062]" page="446" pageId="91" pageNumber="92" year="1982">
<taxonomicName id="4C5A4D27FFDB5317CD24575DFD93A61A" ID-CoL="8VRN8" authority="Pace, 1982: 446" authorityName="Pace" authorityPageNumber="446" authorityYear="1982" box="[264,587,1038,1062]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CD24575DFE4BA619" box="[264,403,1039,1062]" italics="true" pageId="91" pageNumber="92">Emmelostiba</emphasis>
<bibRefCitation id="EFCB4B55FFDB5317CDB6575CFD93A61A" author="Pace, R." box="[410,587,1038,1061]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace, 1982: 446</bibRefCitation>
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(
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species: 
<taxonomicName id="4C5A4D27FFDB5317CEDC575DFBBCA61A" ID-CoL="39H4B" authority="Pace, 1982" authorityName="Pace" authorityYear="1982" box="[752,1124,1038,1062]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="besucheti">
<emphasis id="B92EEAB6FFDB5317CEDC575DFC3EA619" box="[752,998,1039,1062]" italics="true" pageId="91" pageNumber="92">Emmelostiba besucheti</emphasis>
<bibRefCitation id="EFCB4B55FFDB5317CFC1575CFBBCA61A" author="Pace, R." box="[1005,1124,1038,1061]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace, 1982</bibRefCitation>
</taxonomicName>
, by original designation).
</paragraph>
<paragraph id="8BE536A4FFDB5317CD24571DFACCA6B9" blockId="91.[264,1322,1038,1158]" pageId="91" pageNumber="92">
<treatmentCitationGroup id="AB4A118AFFDB5317CD24571DFCC0A65A" box="[264,792,1102,1126]" pageId="91" pageNumber="92">
<taxonomicName id="4C5A4D27FFDB5317CD24571DFE2DA65A" ID-CoL="8XPGW" authority="Lohse" authorityName="Lohse" authorityYear="1990" box="[264,501,1102,1126]" class="Insecta" family="Staphylinidae" genus="Pseudousipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CD24571DFE74A659" box="[264,428,1103,1126]" italics="true" pageId="91" pageNumber="92">Pseudousipalia</emphasis>
Lohse
</taxonomicName>
<emphasis id="B92EEAB6FFDB5317CDD0571DFDC9A659" box="[508,529,1103,1126]" italics="true" pageId="91" pageNumber="92">in</emphasis>
<treatmentCitation id="0AFB10B5FFDB5317CE34571CFCC0A65A" author="Lohse, G. A. &amp; Klimaszewski, J. &amp; Smetana, A." box="[536,792,1102,1126]" page="149" pageId="91" pageNumber="92" year="1990">
<bibRefCitation id="EFCB4B55FFDB5317CE34571CFCC0A65A" author="Lohse, G. A. &amp; Klimaszewski, J. &amp; Smetana, A." box="[536,792,1102,1126]" pageId="91" pageNumber="92" pagination="121 - 202" refId="ref79361" refString="Lohse, G. A., Klimaszewski, J. &amp; Smetana, A. (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 44 (2), 121 - 202." type="journal article" year="1990">
Lohse 
<emphasis id="B92EEAB6FFDB5317CE4D571DFD4FA659" box="[609,663,1103,1126]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
, 1990: 149
</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
(
<typeStatus id="54E18806FFDB5317CF0B571CFC8EA65A" box="[807,854,1102,1125]" pageId="91" pageNumber="92">type</typeStatus>
species: 
<taxonomicName id="4C5A4D27FFDB5317CFED571DFAF2A65A" ID-CoL="6WHDX" authority="Lohse" authorityName=": Lohse" authorityYear="1991" box="[961,1322,1102,1126]" class="Insecta" family="Staphylinidae" genus="Pseudousipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="microptera">
<emphasis id="B92EEAB6FFDB5317CFED571DFB3AA659" box="[961,1250,1103,1126]" italics="true" pageId="91" pageNumber="92">Pseudousipalia microptera</emphasis>
Lohse
</taxonomicName>
<emphasis id="B92EEAB6FFDB5317CD14573DFE95A6B9" box="[312,333,1135,1158]" italics="true" pageId="91" pageNumber="92">in</emphasis>
<bibRefCitation id="EFCB4B55FFDB5317CD78573CFDCEA6BA" author="Lohse, G. A. &amp; Klimaszewski, J. &amp; Smetana, A." box="[340,534,1134,1158]" pageId="91" pageNumber="92" pagination="121 - 202" refId="ref79361" refString="Lohse, G. A., Klimaszewski, J. &amp; Smetana, A. (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 44 (2), 121 - 202." type="journal article" year="1990">
Lohse 
<emphasis id="B92EEAB6FFDB5317CDB7573DFE09A6B9" box="[411,465,1135,1158]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
, 1990
</bibRefCitation>
, by original designation; in subtribe 
<taxonomicName id="4C5A4D27FFDB5317CFB0573CFB47A6BA" authority="Lohse, 1969" authorityName="Lohse" authorityYear="1969" box="[924,1183,1134,1157]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="subTribe" subTribe="Callicerina">
Callicerina 
<bibRefCitation id="EFCB4B55FFDB5317C834573CFB47A6BA" author="Lohse, G. A." box="[1048,1183,1134,1157]" pageId="91" pageNumber="92" pagination="169 - 175" refId="ref78926" refString="Lohse, G. A. (1969) Vorschlage zur Anderung der Aleocharinensystematik (Coleoptera: Staphylinidae). In: Bericht uber die 10. Wanderversammlung deutscher Entomologen, 15. bis 19. September 1965 in Dresden. Deutsche Akademie der Landwirtschaftswissenschaften zu Berlin, Berlin, pp. 169 - 175." type="book chapter" year="1969">Lohse, 1969</bibRefCitation>
</taxonomicName>
), 
<emphasis id="B92EEAB6FFDB5317C89E573CFACCA6B9" bold="true" box="[1202,1300,1134,1158]" pageId="91" pageNumber="92">syn. nov.</emphasis>
</paragraph>
<paragraph id="8BE536A4FFDB5317CD2457F3FD92A687" blockId="91.[264,586,1185,1208]" box="[264,586,1185,1208]" pageId="91" pageNumber="92">(Other references are omitted)</paragraph>
</subSubSection>
<subSubSection id="C340652FFFDB5317CD2457BBFD84A73C" box="[264,604,1257,1283]" pageId="91" pageNumber="92" type="diagnosis">
<paragraph id="8BE536A4FFDB5317CD2457BBFD84A73C" blockId="91.[264,1323,1257,1563]" box="[264,604,1257,1283]" pageId="91" pageNumber="92">
<emphasis id="B92EEAB6FFDB5317CD2457BBFE50A73C" bold="true" box="[264,392,1257,1283]" pageId="91" pageNumber="92">Diagnosis.</emphasis>
See 
<bibRefCitation id="EFCB4B55FFDB5317CDE457BBFD81A73C" author="Pace, R." box="[456,601,1257,1283]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace (1982)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340652FFFDB5317CD145643FC8BA424" pageId="91" pageNumber="92" type="discussion">
<paragraph id="8BE536A4FFDB5317CD145643FC8BA424" blockId="91.[264,1323,1257,1563]" pageId="91" pageNumber="92">
<emphasis id="B92EEAB6FFDB5317CD145643FE1BA714" bold="true" box="[312,451,1297,1323]" pageId="91" pageNumber="92">Discussion.</emphasis>
The 
<typeStatus id="54E18806FFDB5317CE255643FDE3A714" box="[521,571,1297,1323]" pageId="91" pageNumber="92">type</typeStatus>
species of 
<taxonomicName id="4C5A4D27FFDB5317CEEC5643FCADA715" authorityName="Lohse" authorityYear="1990" box="[704,885,1297,1322]" class="Insecta" family="Staphylinidae" genus="Pseudousipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CEEC5643FCADA715" box="[704,885,1297,1322]" italics="true" pageId="91" pageNumber="92">Pseudousipalia</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5A4D27FFDB5317CF835643FB90A715" authorityName="Pace" authorityYear="1982" box="[943,1096,1297,1322]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CF835643FB90A715" box="[943,1096,1297,1322]" italics="true" pageId="91" pageNumber="92">Emmelostiba</emphasis>
</taxonomicName>
are similar in their mouthparts, particularly the shape of the ligula which is split into two separate lobes (
<figureCitation id="13612A21FFDB5317C8D7566BFEFDA744" captionStart="FIGURES 10–14" captionStartId="15.[264,383,1761,1785]" captionTargetBox="[306,1280,627,1718]" captionTargetId="figure-163@15.[294,1292,612,1734]" captionTargetPageId="15" captionText="FIGURES 10–14. Genitalia of Oxypoda grandipennis (Casey) (male from Armagh, Québec (10– 13); and female from 10 km S Ninilchik, Alaska (14)). 10 — median lobe, parameral view; 11 — apex of median lobe, parameral view; 12 — median lobe, lateral view; 13 — apex of median lobe, lateral view; 14 — spermatheca. Scale bar 0.1 mm (14), 0.2 mm (11, 13), 0.4 mm (10, 12)." figureDoi="http://doi.org/10.5281/zenodo.5019797" httpUri="https://zenodo.org/record/5019797/files/figure.png" pageId="91" pageNumber="92">
Fig. 
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in 
<bibRefCitation id="EFCB4B55FFDB5317CD785633FE0DA744" author="Pace, R." box="[340,469,1377,1403]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace 1982</bibRefCitation>
); in their pronotal setation pattern (
<figureCitation id="13612A21FFDB5317CFA15633FC3CA744" box="[909,996,1377,1403]" captionStart="FIGURES 10–14" captionStartId="15.[264,383,1761,1785]" captionTargetBox="[306,1280,627,1718]" captionTargetId="figure-163@15.[294,1292,612,1734]" captionTargetPageId="15" captionText="FIGURES 10–14. Genitalia of Oxypoda grandipennis (Casey) (male from Armagh, Québec (10– 13); and female from 10 km S Ninilchik, Alaska (14)). 10 — median lobe, parameral view; 11 — apex of median lobe, parameral view; 12 — median lobe, lateral view; 13 — apex of median lobe, lateral view; 14 — spermatheca. Scale bar 0.1 mm (14), 0.2 mm (11, 13), 0.4 mm (10, 12)." figureDoi="http://doi.org/10.5281/zenodo.5019797" httpUri="https://zenodo.org/record/5019797/files/figure.png" pageId="91" pageNumber="92">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="EFCB4B55FFDB5317C83F5633FB4DA744" author="Pace, R." box="[1043,1173,1377,1403]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace 1982</bibRefCitation>
); and in the distinct shape of the aedeagus, particularly the deeply split apex of the median lobe (
<figureCitation id="13612A21FFDB5317C8D756DBFEFDA7F4" captionStart="FIGURES 15–18" captionStartId="18.[264,382,1069,1093]" captionTargetBox="[279,1307,335,1056]" captionTargetId="figure-287@18.[269,1318,321,1068]" captionTargetPageId="18" captionText="FIGURES 15–18. Genitalia of Oxypoda amica Casey (male, lectotype of O. amica (15–17); and female from 2.5 km N Lawrence (18)). 15 — median lobe, parameral view; 16 — median lobe with details of retracted internal sac, lateral view; 17 — median lobe with details of retracted internal sac, abparameral view; 18 — spermatheca. Scale bar 0.1 mm (18), 0.2 mm (15–17)." figureDoi="http://doi.org/10.5281/zenodo.5019800" httpUri="https://zenodo.org/record/5019800/files/figure.png" pageId="91" pageNumber="92">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="EFCB4B55FFDB5317CD7656E3FE07A7F4" author="Pace, R." box="[346,479,1457,1483]" pageId="91" pageNumber="92" pagination="443 - 450" refId="ref80432" refString="Pace, R. (1982) Oncosomechusa besucheti n. gen., n. sp. dell'India ed Emmelostiba besucheti n. gen., n. sp. del Libano (Coleoptera, Staphylinidae). Revue suisse de Zoologie, 89 (2), 443 - 450." type="journal article" year="1982">Pace 1982</bibRefCitation>
; 
<figureCitation id="13612A21FFDB5317CDDE56E3FD95A7F4" box="[498,589,1457,1483]" captionStart="FIGURES 52–55" captionStartId="48.[264,382,1473,1497]" captionTargetBox="[279,1295,618,1436]" captionTargetId="figure-199@48.[273,1315,609,1446]" captionTargetPageId="48" captionText="FIGURES 52–55. Genitalia of Atheta fenyesi Bernhauer (male, lectotype of At. querula Casey (52–54); and female, paralectotype of At. fenyesi (55)). 52 — median lobe, parameral view; 53 — median lobe, lateral view; 54 — median lobe, abparameral view; 55 — spermatheca. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5019822" httpUri="https://zenodo.org/record/5019822/files/figure.png" pageId="91" pageNumber="92">
Fig. 
<quantity id="4CA29B41FFDB5317CE1C56E3FD95A7F4" box="[560,589,1457,1483]" metricMagnitude="0" metricUnit="m" metricValue="1.3716" pageId="91" pageNumber="92" unit="in" value="54.0">54</quantity>
</figureCitation>
in 
<bibRefCitation id="EFCB4B55FFDB5317CEAE56E3FCB0A7F4" author="Lohse, G. A. &amp; Klimaszewski, J. &amp; Smetana, A." box="[642,872,1457,1483]" pageId="91" pageNumber="92" pagination="121 - 202" refId="ref79361" refString="Lohse, G. A., Klimaszewski, J. &amp; Smetana, A. (1990) Revision of Arctic Aleocharinae of North America (Coleoptera: Staphylinidae). The Coleopterists Bulletin, 44 (2), 121 - 202." type="journal article" year="1990">
Lohse 
<emphasis id="B92EEAB6FFDB5317CEF456E3FCC3A7F5" box="[728,795,1457,1482]" italics="true" pageId="91" pageNumber="92">et al.</emphasis>
1990
</bibRefCitation>
). Based on this similarity, I place 
<taxonomicName id="4C5A4D27FFDB5317CD24568BFE64A7CD" authorityName="Lohse" authorityYear="1990" box="[264,444,1497,1522]" class="Insecta" family="Staphylinidae" genus="Pseudousipalia" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CD24568BFE64A7CD" box="[264,444,1497,1522]" italics="true" pageId="91" pageNumber="92">Pseudousipalia</emphasis>
</taxonomicName>
in synonymy with 
<taxonomicName id="4C5A4D27FFDB5317CE88568BFCE5A7CD" authorityName="Pace" authorityYear="1982" box="[676,829,1497,1522]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CE88568BFCE5A7CD" box="[676,829,1497,1522]" italics="true" pageId="91" pageNumber="92">Emmelostiba</emphasis>
</taxonomicName>
. The genus 
<taxonomicName id="4C5A4D27FFDB5317CFFB568BFBA8A7CD" authorityName="Pace" authorityYear="1982" box="[983,1136,1497,1522]" class="Insecta" family="Staphylinidae" genus="Emmelostiba" kingdom="Animalia" order="Coleoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAB6FFDB5317CFFB568BFBA8A7CD" box="[983,1136,1497,1522]" italics="true" pageId="91" pageNumber="92">Emmelostiba</emphasis>
</taxonomicName>
was previously known from the Palaearctic and Oriental Regions.
</paragraph>
</subSubSection>
</treatment>
</document>