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213 lines
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<document id="B48A2B70535E7EAA069BB7D17083E665" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="6C7F2A5F71CD3C4798E11829B5224AB1" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus contiro Guignot 1952" docType="treatment" docVersion="5" lastPageNumber="194" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="192" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="9054B90368A302531DFE860BDD0E8101">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="6ABF105F611B3FBE7FBEE6594B1BAFA3">Bistroem, Olof</mods:namePart>
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<mods:namePart id="BAE6D8B2DC8AA5F0D8619EF1B7931F33">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="D9368477274C439AFFA878CDBEB830E7">Bergsten, Johannes</mods:namePart>
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<mods:title id="B83B8649A6E1CA6CCFEA8E4F4E763156">ZooKeys</mods:title>
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<mods:date id="2C7C816A960427739E56ECE85BB3DE00">2015</mods:date>
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<mods:number id="AB9971E3E726802F2A2032E705C50129">542</mods:number>
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<mods:classification id="4183007872AACCA7C694890D339B9F09">journal article</mods:classification>
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<treatment id="6C7F2A5F71CD3C4798E11829B5224AB1" ID-GBIF-Taxon="127862798" LSID="urn:lsid:plazi:treatment:6C7F2A5F71CD3C4798E11829B5224AB1" httpUri="http://treatment.plazi.org/id/6C7F2A5F71CD3C4798E11829B5224AB1" lastPageId="193" lastPageNumber="194" pageId="191" pageNumber="192">
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<subSubSection id="1C1598499A8F8F58F7F3FF11C8C18C80" pageId="191" pageNumber="192" type="multiple">
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<paragraph id="3E18363EECA3929928660B87946E9A24" pageId="191" pageNumber="192">
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<pageBreakToken id="43E948BF3598811CD1694C483A80C55D" pageId="191" pageNumber="192" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Dytiscidae
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</paragraph>
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</subSubSection>
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<subSubSection id="EC1227E7F5EDD89DDC04434C43C155F4" pageId="191" pageNumber="192" type="nomenclature">
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<paragraph id="37AC1EC16F6CB4121C3019E0B2108A98" pageId="191" pageNumber="192">
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<taxonomicName id="5DB74EBAE68E33AF810FFA7260D1BC2C" ID-CoL="3RM6F" authority="Guignot, 1952" authorityName="Guignot" authorityYear="1952" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro Guignot, 1952</taxonomicName>
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Figs 158-159, 341-342, 487-488, 570
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</paragraph>
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</subSubSection>
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<subSubSection id="C01509BBD490E16B8BB5CEDA27E9BE09" pageId="191" pageNumber="192" type="reference_group">
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<paragraph id="46039BDFE102D6DEC72E89986962D0EE" pageId="191" pageNumber="192">
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<taxonomicName id="8FC459EDC43233B53FA7A05A05B16777" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
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<bibRefCitation id="4ADCE9EB4CAB97D168AECFC116775DDB" author="Guignot, F" journalOrPublisher="Bulletin Institut royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 7" title="Description de nouveaux dytiscides africains." volume="28" year="1952 d">Guignot 1952d</bibRefCitation>
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: 5 (original description, faunistics);
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<bibRefCitation id="9D0A299546FE28273764F3B548D5954C" author="Guignot, F" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="257" pageNumber="258" pagination="233 - 237" title="Hydrocanthares captures a la lumiere dans le Haut-Katanga." volume="47" year="1953 b">Guignot 1953b</bibRefCitation>
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: 234 (faunistics);
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<bibRefCitation id="CB98B10B86F4452939A6546B2F9E1F0E" author="Omer-Cooper, Joyce" journalOrPublisher="Memoirs of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="1 - 90" title="Dytiscidae (Coleoptera) collected in the Transvaal (Rhodes University Expedition 1948)." volume="5" year="1957">Omer-Cooper 1957</bibRefCitation>
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: 10 (discussion, description);
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<bibRefCitation id="EC6068DE836EDB96F7D7F12CEAF24525" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
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: 37, 38, 39, 40 (description, faunistics, biology);
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<bibRefCitation id="5D6676373E665531C0B062E7C4652FB9" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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: 550, 554 (description, discussion, faunistics);
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<bibRefCitation id="45F21BBD6DB501A0277E374EBD85F1CB" author="Guignot, F" journalOrPublisher="Resultats Scientifiques de l'Exploration Hydrobiologique des Lacs Kivu, Edouard et Albert (1952 - 1954)" pageId="258" pageNumber="259" pagination="133 - 169" title="Haliplidae et Dytiscidae (Coleoptera Hydrocanthares)." volume="3" year="1959 d">Guignot 1959d</bibRefCitation>
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: 161 (discussion);
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<bibRefCitation id="CD39DC2E9EA9A835D67A90271FC6695B" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="258" pageNumber="259" pagination="231 - 239" title="XXIV. ColeopteraDytiscoidea et Gyrinoidea (Supplement). Le Parc National du Niokolo Koba. 2." volume="62" year="1961 b">Guignot 1961b</bibRefCitation>
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: 238 (discussion);
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<bibRefCitation id="93A0B3A8B17367B2E2357BD40AB4B807" author="Omer-Cooper, Joyce" journalOrPublisher="Annals of the Cape Provincial Museums" pageId="261" pageNumber="262" pagination="292 - 296" title="The distribution of the Cape Province Dytiscidae and speculations as to their possible origins." volume="2" year="1962">Omer-Cooper 1962</bibRefCitation>
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: 295 (faunistics);
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<bibRefCitation id="3B8C7CAA496948FE0B10BC0F3D805914" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
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: 76, 77, 78 (description, discussion, faunistics);
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<bibRefCitation id="0ECCD5CD0F1929CB99D3EFD198DD19F6" author="Bilardo, A" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="254" pageNumber="255" pagination="93 - 130" title="Recoltes de coleopteres aquatiques Haliplidae et Dytiscidae dans le Gabon et la Cote d'Ivoire." volume="56" year="1978">Bilardo and Pederzani 1978</bibRefCitation>
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: 119 (faunistics, description);
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<bibRefCitation id="0D961D25E134E439C451557F16327A47" author="Medler, JT" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="259" pageNumber="260" pagination="1 - 919" title="Insects of Nigeria - Check list and bibliography." volume="30" year="1980">Medler 1980</bibRefCitation>
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: 155 (faunistics, list);
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<bibRefCitation id="F41556645148D862C7219A3631253830" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civivico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="441 - 461" title="Contributo alla conoscenza dei Dytiscidae del Cameroun (Coleoptera)." volume="123" year="1982 a">Bilardo 1982a</bibRefCitation>
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: 447 (given as
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<taxonomicName id="038B10A409533CE3216C6B5C00720E4A" genus="Laccophylus" lsidName="Laccophylus" pageId="191" pageNumber="192" rank="genus">Laccophylus</taxonomicName>
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, description, faunistics);
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<bibRefCitation id="F3D046835AA31DAF235C0A9EA6EA1110" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali del Museo Civivico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="250 - 254" title="Guignotusovatus sp. n. e Copelatusvirungaensis sp. n. del Virunga National Park (Zaire) (ColeopteraDytiscidae)." volume="123" year="1982 b">1982b</bibRefCitation>
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: 251 (faunistics);
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<bibRefCitation id="78FE0D9C13BE91C35186D59DAD4B264E" author="Pederzani, F" journalOrPublisher="REDIA" pageId="261" pageNumber="262" pagination="65 - 88" title="Dytiscidae (Coleoptera) collected in the People's Republic of Congo by Mr. Giovanni Onore in the years 1977 - 1980." volume="65" year="1982">Pederzani and Rocchi 1982</bibRefCitation>
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: 72 (faunistics);
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<bibRefCitation id="AB38863BCD6E47759A5D69531372CACE" author="Pederzani, F" journalOrPublisher="Atti dell'Accademia Roveretana degli Agiati, Contribute della Classe di Scienze Matematiche, Fisiche e Naturali" pageId="261" pageNumber="262" pagination="103 - 118" title="Dytiscidae collected in Zambia and description of Hyphydrusfluviatilis sp. n. and Hyphydruszambiensis sp. n. (ColeopteraDytiscidae)." volume="26" year="1988">Pederzani 1988</bibRefCitation>
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: 107 (faunistics, biology);
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<bibRefCitation id="7994A364E253AF1641B686C292DE684A" author="Nilsson, AN" journalOrPublisher="Entomologica Fennica" pageId="260" pageNumber="261" pagination="57 - 94" title="Taxonomy, distribution and habitats of the Dytiscidae (Coleoptera) of Ethiopia." volume="4" year="1993">Nilsson and Persson 1993</bibRefCitation>
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: 80: (faunistics);
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<bibRefCitation id="723E58301849DC99586C5EB04F287EAA" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="215 - 236" title="Haliplidae e Dytiscidae (Coleoptera) del Gabon (parte quinta)." volume="140" year="1999">Bilardo and Rocchi 1999</bibRefCitation>
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: 232, 234 (faunistics);
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<bibRefCitation id="50A4C7818FC054CCD2BD3EA6662E4813" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 242 (catalogue, faunistics);
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<bibRefCitation id="B73C1B7A83ECC97628BAA816178BB664" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="147 - 182" title="Haliplidae e Dytiscidae (Coleoptera) del Gabon. (Parte quarta)." volume="143" year="2002">Bilardo and Rocchi 2002</bibRefCitation>
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: 174 (list, faunistics);
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<bibRefCitation id="21DE599EB8CD69752DC5C6B700660F27" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 210 (catalogue, faunistics). [Comment: in the original description
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<bibRefCitation id="5DABC8ADB408A9C70C2C7B1F2895AAFD" author="Guignot, F" journalOrPublisher="Bulletin Institut royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 7" title="Description de nouveaux dytiscides africains." volume="28" year="1952 d">Guignot (1952d</bibRefCitation>
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: 6) also distinguishes a separate morph and gives it the name
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<taxonomicName id="138FBD2C1D8FAC9291CC6E7EDE179D77" aberration="nigrovirgatus" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" infraspecific-rank="ab." kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="aberration" species="contiro">Laccophilus contiro ab. nigrovirgatus</taxonomicName>
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. This name is, however, infrasubspecific and has no value in nomenclature.]
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</paragraph>
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</subSubSection>
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<subSubSection id="E499E67DDEE6D922D9BFB824B0C336FB" pageId="191" pageNumber="192" type="type locality">
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<paragraph id="517C1A44C6810C65B749DA79296CA6D3" pageId="191" pageNumber="192">Type locality.</paragraph>
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<paragraph id="067E6891AF7405A1A148FD76D760B1A5" pageId="191" pageNumber="192">Ethiopia: Mt Chilalu.</paragraph>
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</subSubSection>
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<subSubSection id="7C410E3A79E288AA4D71F996EFE2C5BB" pageId="191" pageNumber="192" type="type material">
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<paragraph id="CAAF9D6E3F960266C888E911BB4D8867" pageId="191" pageNumber="192">Type material studied</paragraph>
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<paragraph id="384A904E3DF60E5A98CEBAA9DAD1E56A" pageId="191" pageNumber="192">
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(4 exs.). Holotype: male: "Abessinia 7,000 ft. Mt. Chilalu 8.xi. 1926 J. Omer-Cooper / male symbol / Type" (MNHN). - Paratypes: "Abyssinia Mount
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<normalizedToken id="3CF6C78AC2546007FF82DCD2069BA57E" originalValue="Chillálo">Chillalo</normalizedToken>
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ponds 7,000-8,000 ft. 8-9.xi. 1926 J. Omer-Cooper / female symbol / Allotype /
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<taxonomicName id="6B07E92D66CEB123A09054B0BB88FDC9" genus="Algae" lsidName="Algae contiro" pageId="191" pageNumber="192" rank="species" species="contiro">contiro</taxonomicName>
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" (1 ex. MNHN); "Abyssinia 7000 ft. Mt Chilalu 8.xi. 1926 J. Omer-Cooper / male symbol / Paratype" (1 ex. MNHN; habitus in Fig. 487). - Uganda: "Uganda Kampala K 11? IX. 1929 H. E. Hopkins / female symbol / paratype" (1 ex. MNHN; habitus in Fig. 488).
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</paragraph>
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</subSubSection>
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<subSubSection id="CD1588E6A18E94CFE37AFC8C88430C3A" lastPageId="192" lastPageNumber="193" pageId="191" pageNumber="192" type="materials_examined">
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<paragraph id="775443DC57AF84336B23AEB554481098" pageId="191" pageNumber="192">Additional material studied</paragraph>
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<paragraph id="F952BD8482179F1418E1796714F7B6C0" lastPageId="192" lastPageNumber="193" pageId="191" pageNumber="192">
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(162 exs.). Sudan: "W. Nile IX. 1929" (2 exs. AMGS); "Gallery Forest Jebel Marra
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<geoCoordinate id="D7AC9A3208048CB8992112E9FB503648" direction="north" orientation="latitude" precision="925" value="12.916667">12°55'N</geoCoordinate>
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,
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<geoCoordinate id="41FD95874B0BC39D626D0081D0D17EEB" direction="east" orientation="longitude" precision="925" value="24.133333">24°08'E</geoCoordinate>
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, 7. 1984 Ruse P. light trap" (8 exs. BMNH, 1 ex. MZH). - Ethiopia: "Suc-Suci Lake Zwai 5,500 ft 12.11. 1926 JOC." (1 ex. AMGS); "7000 ft Mt. Chilalu 8.11. 1926 JOC." (5 exs. AMGS); "West Marsh Lake Zwai 5,500 ft 2-3.11. 1926 JOC." (3 exs. AMGS); "March N of Lake Zwai ca 5,500 ft. 4.11. 1926" (1 ex. AMGS); "Kattere River Lake Zwai 6,000 ft 5.11. 1926 JOC." (1 ex. AMGS); "Water Hole N of Makki River 6,000 ft 28.11. 1926 JOC." (1 ex. AMGS); "C Abyssinia, Abesata Wajju, Bull Bullo 6.3. 1915 Lovacs leg." (1 ex. BMNH); "Belleta Forest 13-14.6. 1963 Linnavuori" (1 ex. MZH). - Sierra Leone: "Freetown 1945/Walton G.A." (1 ex. BMNH). - Nigeria: "NC St. Malumfashi 26-30.7. 1973 Linnavuori leg."(2 exs. MZH); "W St. Ife 7-8.7. 1973 Linnavuori leg." (1 ex. MZH); "Stream crossing Kaduna rd. nr Zaria 4.4. 1963 JOC." (2 exs. AMGS); "Zaria 1969 Brancucci" (1 ex. NHMB); "Stream & reservoir Jos 10.4. 1963 JOC." (3 exs. AMGS); "Stream nr Bakori en rte Katsina 5.4.1963 JOC." (1 ex. AMGS); "Stream nr Zaria 4.4. 1963 JOC" (1 ex. AMGS); "Ponds in
|
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|
|
<pageBreakToken id="B75F2385964FFAB5BF5D07949F85F69A" pageId="192" pageNumber="193" start="start">dry</pageBreakToken>
|
|||
|
|
stream bed Kontagora 5.4. 1963" (2 exs. AMGS); "Kontagora stream 3.4. 1963 JOC." (6 exs. AMGS). - Cameroon: "20 km NW Ban-Gante Forest, savannah at river, at light 15.1. 1978 /
|
|||
|
|
<normalizedToken id="0F918C489F7F41A9B496824E1C774C68" originalValue="Gärdenfors">Gaerdenfors</normalizedToken>
|
|||
|
|
, Hall & Samuelsson" (1 ex. MZLU); "Kamerun int. Satsche 10-14.5. 1909 Riggenbach" (5 exs. ZMHB); "Koza 1.7.1974"(1 ex. NHMB). - Uganda: "Prov.
|
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|
|
<normalizedToken id="79D506A9F143190B8694EA0E642848FE" originalValue="d’Ounyoro">d'Ounyoro</normalizedToken>
|
|||
|
|
, Albert Nyanza S-E, Riv. Mousisi, Alluaud 1909 / Type" (1 ex. MNHN;
|
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|
|
<normalizedToken id="85FE02C8E8E362871DBB52CCBE4EDF2A" originalValue="”type”">"type"</normalizedToken>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="5F133B910F39D2761F91C9C7FEF88177" aberration="nigrovirgatus" genus="Algae" infraspecific-rank="ab." lsidName="Algae ab subsp. nigrovirgatus" pageId="192" pageNumber="193" rank="aberration" species="ab" subSpecies="nigrovirgatus">ab. nigrovirgatus</taxonomicName>
|
|||
|
|
); "Kampala 13.7. 1929 G.H.E. Hopkins" (1 ex. AMGS); same but "15.9. 1929" (1 ex. AMGS); same but "28.2. 1929" (1 ex. AMGS); same but 28.11. 1929" (1 ex. BMNH); same but "29.11. 1929" (1 ex. BMNH); same but "21.2. 1930" (1 ex. BMNH); same but "5.12. 1929" (1 ex. MZH); same but "21.8. 1929" (1 ex. BMNH); same but 9. 1929/Paratype" (1 ex. MNHN; not type material); same but "Kitante Swamp 26.9. 1969" (1 ex. BMNH); "Madi 5. 1927 G.D.H. Carpenter" (1 ex. AMS). - Kenya: "Nyeri 2.12. 1989
|
|||
|
|
<normalizedToken id="C489842A4B0799D8F98D0042DBAEAF07" originalValue="Jäch”">Jaech"</normalizedToken>
|
|||
|
|
(1 ex. NMW); "Ngong Forestry Sta., 13.4, 1968 Spangler" (7 exs. USNM, 2 exs. MZH); "Nairobi 3.11. 1967 / Reichert collector" (1 ex. USNM). - Tanzania: "?stream Mbeya-Tunduma rd. 18.10. 1948 JOC." (3 exs. AMGS); "Mpemba stream Mbeya-Tunduma 16.X. 1948" (2 exs. AMGS); "D.O.A. W Ruangwa 1.9. 1911, 1850 m Sumpf H. Meyer S.G." (2 exs. ZMHB). - Zambia: "Mbesuma Ranch (Isoka) 9-10.12. 2004 Werner & Smrz" (2 exs. NHRS); "Watergreen Farm Chongwe Valley 60 km E Lusaka 4.8. 1986 Pederzani /
|
|||
|
|
<taxonomicName id="6F76096511C2181D88B9C046D746C77E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="192" pageNumber="193" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
Guignot det. Pederzani" (1 ex. CSR); "Kapiri Mpushi env. 13.12. 2002 Kantner" (6 exs. NHMB, 2 exs. MZH). - Malawi: "Swamp
|
|||
|
|
<normalizedToken id="43AAB4CE91B668A4A65A32B84A515EDB" originalValue="Dally’s">Dally's</normalizedToken>
|
|||
|
|
Hotel nr Ft Johnstone 23.8. 1948" (6 exs. AMGS); "Swamp Hawkes Bay 25.9. 1948" (2 exs. AMGS); "Dambo below Livingstonia lake shore 21.10. 1948" (15 exs. AMGS);
|
|||
|
|
<normalizedToken id="C4FE4F420EDC4E6DA97910EFF3002B40" originalValue="“Dally’s">"Dally's</normalizedToken>
|
|||
|
|
18.12. 1946 R.H. Lowe" (2 exs. BMNH); "Ft. Johnston
|
|||
|
|
<normalizedToken id="3518BE336F7B8AF7AB35029C6FCD7FE3" originalValue="Dally’s">Dally's</normalizedToken>
|
|||
|
|
swamp nr L. Nyasa 7.6. 1946" (1 ex. BMNH); "Selima env., 4.1. 2002 Kantner" (1 ex. NHMB); "Selima env., 60 km E Lilongwe 5-8.1. 2002 Kantner" (1 ex. NHMB); "Dedza env., 85 km SE Lilongwe 7-13.1. 2002 Kantner" (1 ex. NHMB). - Namibia: "East Caprivi Katima Mulilo
|
|||
|
|
<geoCoordinate id="75CC6866B42C67D0C70B08F859783DE4" direction="south" orientation="latitude" precision="925" value="-17.483334">17°29'S-</geoCoordinate>
|
|||
|
|
<geoCoordinate id="8DCCD6AAB6EE62B1E179714006BD565C" direction="east" orientation="longitude" precision="925" value="24.283333">24°17'E</geoCoordinate>
|
|||
|
|
, 3-8.3. 1992 Uhlig leg., lux" (8 exs. ZMHB, 2 exs. MZH, 1 ex. NMNW); same but "Deckert leg." (1 ex. ZMHB). - Zimbabwe: "Mashunald Salisbury" (1 ex. SAMC). - South Africa: "Gauteng, RAU system, Germiston Lake 12.8. 1997 Reavell" (1 ex. AMGS); "Natal Umlazi R. 19.9. 1962" (1 ex. AMGS); "Kw. Natal S, Port Shepstone 20 km W, 2.2. 2000 Halada leg." (1 ex. NMW); "Kw. Natal, Barringtonia Swamp For., Amotikela Nat. Res., 24.2. 1991 Reavell" (1 ex. AMGS); "Natal, Durban, Stamford Hill Umgeni
|
|||
|
|
<normalizedToken id="105B777694C73D24F7BC4F3426481847" originalValue="Trägårdh">Traegardh</normalizedToken>
|
|||
|
|
leg. /
|
|||
|
|
<taxonomicName id="6CD86153157A07F72310CF2127476FF4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="192" pageNumber="193" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
Guign. det. Omer-Cooper" (1 ex. MZLU); "ECPR. St. Johns 16.2. 1956 JOC. /
|
|||
|
|
<taxonomicName id="33B0BDB82F82EDE26AA1056A02FB399A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="192" pageNumber="193" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
Guign. det. Omer-Cooper" (6 exs. AMGS); "ECPr. 6 km S of port of St Johns, outside Silaka Nature Res., pond, S31°38.862, E29°30.551, alt. 90 m 26-27.1. 2005 Bergsten leg." (4 exs. NHRS); "EC, East London, Gorncie Park, coastal pond 18.3. 1955" (2 exs. AMGS); "EC., Pond on Hwy 344 at Adelaide 17.5. 2005 Challet" (3 exs. CGC, 1 ex. MZH); "EC, Groot R., Humansdorp 19.2. 1947 JOC." (11 exs. AMGS).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="2D5841EAF7890B936711958ABC635E73" lastPageId="193" lastPageNumber="194" pageId="192" pageNumber="193" type="diagnosis">
|
|||
|
|
<paragraph id="78DEEFB4C3D50CD8672EF4A04A6BE0A1" pageId="192" pageNumber="193">Diagnosis.</paragraph>
|
|||
|
|
<paragraph id="4A1BBCF06F9BC0BAB69D19DEA4BC6C31" lastPageId="193" lastPageNumber="194" pageId="192" pageNumber="193">
|
|||
|
|
Dorsal colour pattern of
|
|||
|
|
<taxonomicName id="385317E5F097654C8E351DBA63753D0A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="192" pageNumber="193" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
is variable. Extremes are represented by specimens with almost totally confluent dark, longitudinal lines of elytra (dark lines
|
|||
|
|
<pageBreakToken id="057282D0E80CCD05A4EF475EB413D3C4" pageId="193" pageNumber="194" start="start">only</pageBreakToken>
|
|||
|
|
discernible posterior to middle) or specimens with separate dark, longitudinal lines. Fortunately penis is quite characteristic, the apex being
|
|||
|
|
<normalizedToken id="60B3477F06084689F4EF74C95A9179B8" originalValue="“harpoon-shaped”">"harpoon-shaped"</normalizedToken>
|
|||
|
|
, and this character distinguishes
|
|||
|
|
<taxonomicName id="EA0EB1E979D385F62BF30FAE417591E9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="193" pageNumber="194" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
from all other
|
|||
|
|
<taxonomicName id="AF2D49B5FD0CF33C336C96B7E905D556" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="193" pageNumber="194" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
|
|||
|
|
species. Morphological variation is still quite extensive and further study will reveal if
|
|||
|
|
<taxonomicName id="FBE291DE5555FCDE589D7EFA03C3E242" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus contiro" order="Coleoptera" pageId="193" pageNumber="194" phylum="Arthropoda" rank="species" species="contiro">Laccophilus contiro</taxonomicName>
|
|||
|
|
must be split up in different species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="5DC5998CC0BE7644F7FEE6A5708F09FC" pageId="193" pageNumber="194" type="description">
|
|||
|
|
<paragraph id="233BDD56946A9271845C81D2372585B1" pageId="193" pageNumber="194">Description.</paragraph>
|
|||
|
|
<paragraph id="BA4EAAE07DEFA1ADFE08DAD12FA79EA5" pageId="193" pageNumber="194">Body length 3.5-3.9 mm, width 1.9-2.1 mm. Elytral colour pattern variable; separate, dark, longitudinal areas almost absent because merged to larger dark areas, or elytral colour pattern consists of distinct longitudinal, dark areas, which sometimes are reduced (Figs 487-488).</paragraph>
|
|||
|
|
<paragraph id="59B7FE65F42B43BBEED070ED44D3A7A7" pageId="193" pageNumber="194">Head: Pale ferrugineous to ferrugineous; no distinct colour pattern. Impunctate, except at eyes; with fine, dense and irregular punctures. Area of punctures extended towards middle of head-disc but they are not connected. Submat, finely microsculptured. Reticulation double but difference in delimitation of size classes very small; in part hardly discernible. Large meshes, when discernible, contain 2-5 fine meshes.</paragraph>
|
|||
|
|
<paragraph id="400C843318D4FAA0598051EB514EF200" pageId="193" pageNumber="194">Pronotum: Pale ferrugineous to ferrugineous, frontally and basally in middle with a distinct blackish to dark ferrugineous area. Submat, finely microsculptured. Reticulation quite distinctly double; large meshes contain generally 2-5 small meshes. Almost impunctate. At margins, except basally in middle, with fine, sparse and scattered punctures.</paragraph>
|
|||
|
|
<paragraph id="FE4ACBC6CA2FD130B89BD65EBE3116DC" pageId="193" pageNumber="194">Elytra: Colour pattern variable. Dark elytral lines, to a variable degree discernible; sometimes distinct and sometimes almost absent (Figs 487-488). Submat, finely microsculptured. Reticulation indistinctly double. Large meshes almost absent because strongly reduced (delimiting lines of meshes have often vanished); sometimes indistinctly visible as fine fragments in frontal parts of elytra. Almost impunctate; very fine, sparse punctures form a discal, dorsolateral and lateral row of punctures.</paragraph>
|
|||
|
|
<paragraph id="C5FDBA99DDA808925F2E603810DEB22C" pageId="193" pageNumber="194">Ventral aspect: Dark ferrugineous to ferrugineous, prosternum paler, pale ferrugineous. Almost impunctate. Rather shiny; scattered very fine, in part indistinct microsculpture may be discerned. Metacoxal plates with approximately 10 shallow and transversely placed furrows. Base of abdomen with curved striae. Prosternal process rather slender, apex somewhat extended and pointed. Apical ventrite with a lateral, sharp knob (Fig. 158).</paragraph>
|
|||
|
|
<paragraph id="C7F493AD603071BFE8AD1080E531D3F5" pageId="193" pageNumber="194">Legs: Pro- and mesotarsus somewhat extended, provided with distinct suckers.</paragraph>
|
|||
|
|
<paragraph id="F09128BDF97B57ECA4E8A94C1103AD1F" pageId="193" pageNumber="194">Male genitalia: Penis in lateral aspect quite broad; from base to apex slightly curved. Extreme apex broad, only slightly extended (Figs 341-342).</paragraph>
|
|||
|
|
<paragraph id="8EA40A2CCF0A20EA2D1B10E2EE023CA7" pageId="193" pageNumber="194">Female: Pro- and mesotarsus slender, somewhat extended. Apical ventrite lacks knob (Fig. 159).</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="B1E5CAA6840B087D8F062B1606D80257" pageId="193" pageNumber="194" type="distribution">
|
|||
|
|
<paragraph id="4DF4AB7633139BA7BB1FFDD4A4ADFBE2" pageId="193" pageNumber="194">Distribution.</paragraph>
|
|||
|
|
<paragraph id="6EFD46887E9FD8E022DC55E7F9FC7180" pageId="193" pageNumber="194">
|
|||
|
|
Sudan, Ethiopia, Sierra Leone, Nigeria, Cameroon, Uganda, Kenya, Tanzania, Zambia, Malawi, Namibia, Zimbabwe, South Africa (Fig. 570). Additional country records are Zaire, Rwanda-Burundi (
|
|||
|
|
<bibRefCitation id="0CE20B2D15781A4DB40A60EDE7B256F8" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
|
|||
|
|
), the Ivory Coast (
|
|||
|
|
<bibRefCitation id="4A01A28847DF2EE49E39F18199B24620" author="Bilardo, A" journalOrPublisher="Memorie della Societa Entomologica Italiana" pageId="254" pageNumber="255" pagination="93 - 130" title="Recoltes de coleopteres aquatiques Haliplidae et Dytiscidae dans le Gabon et la Cote d'Ivoire." volume="56" year="1978">Bilardo and Pederzani 1978</bibRefCitation>
|
|||
|
|
), Congo (Brazzaville) (
|
|||
|
|
<bibRefCitation id="2A8B90AF8A9547828F436299779C57CD" author="Pederzani, F" journalOrPublisher="REDIA" pageId="261" pageNumber="262" pagination="65 - 88" title="Dytiscidae (Coleoptera) collected in the People's Republic of Congo by Mr. Giovanni Onore in the years 1977 - 1980." volume="65" year="1982">Pederzani and Rocchi 1982</bibRefCitation>
|
|||
|
|
) and Gabon (
|
|||
|
|
<bibRefCitation id="1172A78904B29CFD53DC4F1B26F6603F" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="215 - 236" title="Haliplidae e Dytiscidae (Coleoptera) del Gabon (parte quinta)." volume="140" year="1999">Bilardo and Rocchi 1999</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="8FE420578550F4200940929F9B9F53F9" pageId="193" pageNumber="194" type="collecting circumstances">
|
|||
|
|
<paragraph id="546670F67E1364A64CF0486F739BDA12" pageId="193" pageNumber="194">Collecting circumstances.</paragraph>
|
|||
|
|
<paragraph id="467431D1DF7308201781BDAF1B3250F9" pageId="193" pageNumber="194">Label data gives limited information on ecological preferences; collected in pools, streams and swamps and sometimes at light. Literature data are considered rather poor.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|