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<document id="11979B26BD45EFE1CCB190BD2678D913" ID-CLB-Dataset="46358" ID-DOI="10.11646/zootaxa.3768.3.1" ID-GBIF-Dataset="0dd03941-3a6b-49e3-8431-5169527af5ab" ID-ISSN="1175-5326" ID-Zenodo-Dep="252362" ID-ZooBank="AE4CFE9E-2AB6-4099-8F50-C49310808060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460388819172" checkinUser="plazi" docAuthor="Veijalainen, Anu, Broad, Gavin R. &amp; Sääksjärvi, Ilari E." docDate="2014" docId="03F926428F14B14F30BBFB1A56F23C5B" docLanguage="en" docName="zt03768p252.pdf" docOrigin="Zootaxa 3768 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orthocentrus pentagonum Broad, sp. nov." docType="treatment" docVersion="10" lastPageNumber="210" masterDocId="FFC05E3A8F1CB146302CFFD8556D395F" masterDocTitle="Twenty seven new species of Orthocentrus (Hymenoptera: Ichneumonidae; Orthocentrinae) with a key to the Neotropical species of the genus" masterLastPageNumber="252" masterPageNumber="201" pageNumber="209" updateTime="1698297975084" updateUser="ExternalLinkService">
<mods:mods id="D7175F77E8690DC8BD6CEB9448655BE4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C70064F8751BEE418B91BC1CDE6FE474">
<mods:title id="08B1DEE3630F0B8B8E87EF5E7EB05DDB">Twenty seven new species of Orthocentrus (Hymenoptera: Ichneumonidae; Orthocentrinae) with a key to the Neotropical species of the genus</mods:title>
</mods:titleInfo>
<mods:name id="A31D515FDD9A281B40CD217FD83D30E8" type="personal">
<mods:role id="208059F8268F1003F15F4ACED68571F3">
<mods:roleTerm id="E5A3D40E2E3E660B1D64FEA288B0FDA1">Author</mods:roleTerm>
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<mods:namePart id="8D70A1FA860C8E9AB694FFF8A6F756E8">Veijalainen, Anu</mods:namePart>
</mods:name>
<mods:name id="1B00EC14DE412CCCF023BC09074B5EC8" type="personal">
<mods:role id="3D7BF6D2EB0A1062E9174F3D71D0CC81">
<mods:roleTerm id="3258C9A85FC2E561B05415450CF6A34D">Author</mods:roleTerm>
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<mods:namePart id="0BEBC066AAF97090ABCC6930083CC0CF">Broad, Gavin R.</mods:namePart>
</mods:name>
<mods:name id="434CC93F40E8A626618F26C643E705D3" type="personal">
<mods:role id="85F281AD8D2B725EFEC0B28A732E890A">
<mods:roleTerm id="A0A39CE617FA9DC0BD95CA259DBAB9CB">Author</mods:roleTerm>
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<mods:namePart id="4291FE8D0647D15078F0231AF46D5721">Sääksjärvi, Ilari E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="14B7BCDCDF6460F95140EA02A4C3A5A3">text</mods:typeOfResource>
<mods:relatedItem id="4ADBCD746E2B2CDE43A288BFB5625CD7" type="host">
<mods:titleInfo id="64C37877A6B89C75F98AB658FD8C5B27">
<mods:title id="4F85CDC1B543707FE459B766F8944D9F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="77FD4043BD9CEB56EDABE25E8D8E1F04">
<mods:date id="A8CF3BE65E57991CA255EF04E2D33002">2014</mods:date>
<mods:detail id="BF88026EBDB3D3BEC33F54E5FC181066" type="volume">
<mods:number id="5DBC5C53C1C6F8E494AC009C3125AFBD">3768</mods:number>
</mods:detail>
<mods:detail id="BC39B27418732A34198FF4408275C1C3" type="issue">
<mods:number id="AFA6F627DF3C5A86A5FAFB4870546176">3</mods:number>
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<mods:extent id="BF2F550F0DF45CF1D8E076B04033D3C6" unit="page">
<mods:start id="E71816A242DAA4CC1A996A438B46A91F">201</mods:start>
<mods:end id="4241296B31B1680D32E5604A88B56362">252</mods:end>
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<mods:classification id="5F0FD657C2CEDF9F667A6CF9C4F133F4">journal article</mods:classification>
<mods:identifier id="753DC8A1E23D7397D1369770DEE45C02" type="CLB-Dataset">46358</mods:identifier>
<mods:identifier id="0B7D192F1A68BB6B5F8E321A73193723" type="DOI">10.11646/zootaxa.3768.3.1</mods:identifier>
<mods:identifier id="DCFD547258EE5C86DEE438C39422FEE6" type="GBIF-Dataset">0dd03941-3a6b-49e3-8431-5169527af5ab</mods:identifier>
<mods:identifier id="F82C821EA795FBE519BD9EB30EB5E413" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="21540E463677A996DA1F939D3F6C3097" type="Zenodo-Dep">252362</mods:identifier>
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<treatment id="03F926428F14B14F30BBFB1A56F23C5B" ID-DOI="http://doi.org/10.5281/zenodo.6134668" ID-GBIF-Taxon="119416152" ID-Zenodo-Dep="6134668" LSID="urn:lsid:plazi:treatment:03F926428F14B14F30BBFB1A56F23C5B" httpUri="http://treatment.plazi.org/id/03F926428F14B14F30BBFB1A56F23C5B" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209">
<subSubSection id="C34AC4DF8F14B14F30BBFB1A51373A6B" lastPageId="9" lastPageNumber="210" pageId="8" pageNumber="209" type="nomenclature">
<paragraph id="8BEF97548F14B14E30BBFB1A57FA3D82" blockId="8.[151,663,1218,1278]" box="[151,663,1218,1245]" pageId="8" pageNumber="209">
<heading id="D0A720388F14B14E30BBFB1A57FA3D82" bold="true" box="[151,663,1218,1245]" fontSize="11" level="1" pageId="8" pageNumber="209" reason="1">
<emphasis id="B9244B468F14B14E30BBFB1A57FA3D82" bold="true" box="[151,663,1218,1245]" pageId="8" pageNumber="209">
<taxonomicName id="4C50ECD78F14B14E30BBFB1A57413D82" authority="Broad" authorityName="Broad" box="[151,556,1218,1245]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="pentagonum" status="sp. nov.">
<emphasis id="B9244B468F14B14E30BBFB1A54BB3D83" bold="true" box="[151,470,1218,1244]" italics="true" pageId="8" pageNumber="209">Orthocentrus pentagonum</emphasis>
Broad
</taxonomicName>
,
<taxonomicNameLabel id="A217F63D8F14B14E3216FB1B57FA3D82" box="[570,663,1219,1245]" pageId="8" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF97548F14B14E30BBFB3D54983DA2" blockId="8.[151,663,1218,1278]" box="[151,501,1253,1278]" pageId="8" pageNumber="209">
<figureCitation id="136B8BD18F14B14E30BBFB3D55B13DA1" box="[151,220,1253,1278]" captionStart="FIGURE 1" captionStartId="39.[151,250,1973,1995]" captionTargetBox="[164,1421,187,1947]" captionTargetId="figure@39.[164,1423,187,1949]" captionTargetPageId="39" captionText="FIGURE 1. Lateral views: Orthocentrus shieldsi species-group (in part). A: anguillae, B: pentagonum, C: concrispus, D: hispidus, E: quercus, F: shieldsi. Scale 0.5 mm." httpUri="https://zenodo.org/record/252363/files/figure.png" pageId="8" pageNumber="209">Figs 1</figureCitation>
B, 6 D, 8 D, 11 D, 13 D.
</paragraph>
<paragraph id="8BEF97548F14B14E30BBFAFF54863C61" blockId="8.[151,1436,1318,2027]" box="[151,491,1318,1343]" pageId="8" pageNumber="209">
Fore wing length
<quantity id="4CA83AB18F14B14E3171FAFF548A3C61" box="[349,487,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="8" pageNumber="209" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
.
</paragraph>
<paragraph id="8BEF97548F14B14E30EBFA92578E3C3C" blockId="8.[151,1436,1318,2027]" box="[199,739,1354,1379]" pageId="8" pageNumber="209">Whole body, especially head, laterally flattened.</paragraph>
<paragraph id="8BEF97548F14B14E30EBFAB7516B3F94" blockId="8.[151,1436,1318,2027]" pageId="8" pageNumber="209">Face medially 1.0x wider than high; head entirely smooth and polished except for small coriaceous area on edge of antennal shelf, face with shallow punctures, eyes not setose, dorsal ridge of face inbetween antennal sockets without a median prominence; face profile very straight except just before antennal sockets impressed, edge of clypeus very straight, antennal sockets on a high shelf; malar space with narrow, very shallow, slightly curved sulcus, represented by discrete line, not microsculptured; maxillary palp short, reaching to fore coxa. In dorsal view, head posteriorly deeply concave, temples distinct, posterior ocelli distant from eye by 0.7x ocellar diameter, anterior ocellus separated from eye by 1.5x ocellar diameter, lacking ocellar-ocular grooves. Minimum distance between antennal sockets very narrow, sockets almost touching each other; antenna thick, curled, with 20– 21 (n=47) transverse and similar-sized flagellomeres which not gradually shortening apically; basal flagellomere 1.1x as wide as high and about 1/4 of the length of scape; scape parallel-sided.</paragraph>
<paragraph id="8BEF97548F14B14E30EBF90F515C3E20" blockId="8.[151,1436,1318,2027]" pageId="8" pageNumber="209">Mesosoma smooth and polished except pronotum slightly coriaceous antero-dorsally, postero-ventral corner of pronotum with short striae or vague rugae, propodeum with coriaceous microsculpture, especially antero-medially; mesoscutum lacking notauli; in profile, scutellum not particularly high, metapleuron not convex; propodeum with complete posterior transverse carina, median longitudinal and lateral longitudinal carinae weakly present posterior to spiracle, median longitudinal carinae faint or lacking anteriorly, spiracles small.</paragraph>
<paragraph id="8BEF97548F14B14E30EBF853548B3EB5" blockId="8.[151,1436,1318,2027]" pageId="8" pageNumber="209">Legs all slightly flattened, broad; coxae and femora polished, tibiae and tarsi coriaceous-granulate; hind coxa 1.1x as long as first tergite, hind femur 2.6x as long as high, hind tibia 3.0–3.8x as long as apically wide; tibiae dorsally with spine-like setae.</paragraph>
<paragraph id="8BEF97548F15B14F30EBFF4F54553843" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">Wings somewhat narrow, cells thus comparatively long and narrow; fore wing with areolet closed but 3rs-m sometimes weak, areolet relatively large, longer than high, anterior side about same length as vein Rs+2r, 2rs-m shorter than 3rs-m, 2m-cu meeting areolet at apical 0.6–0.7, vein Rs straight; hind wing vein cu-a intercepted below middle.</paragraph>
<paragraph id="8BEF97548F15B14F30EBFEFE56AC3B02" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">First tergite long elongate, 2.0x as long as apically wide, in dorsal view, slightly wider at spiracles; coriaceous, polished apically, with weak median longitudinal carinae, with transverse impressions originating at about middle or slightly behind middle of tergite, sloping posteriorly, not meeting centrally. Second tergite 1.3–1.6x times as long as apically wide; coriaceous with weak to fairly strong striae, polished posteriorly, transverse impressions originating at about middle or slightly posterior to middle of tergite, sloping anteriorly and posteriorly, not meeting clearly centrally, weakly delimiting central rounded rhombic area; basal thyridia rounded-rectangular, contrastingly coloured. Remaining tergites smooth and polished, lacking thyridia; third tergite sometimes with coriaceous microsculpture antero-medially. Ovipositor straight, thin, without notch; ovipositor sheath bluntly pointed, with setae longer than sheath width and strongly curved, backwards pointing.</paragraph>
<paragraph id="8BEF97548F15B14F30EBFDB0562A3BFA" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">Body setose except eyes, pronotum, mesopleuron, metapleuron; setae on propodeum, basal tergites and posterior sides of coxae very few, scattered on apical tergites.</paragraph>
<paragraph id="8BEF97548F15B14F30EBFD7751373A6B" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">Dark brown to blackish brown except mouthparts creamy, sternites creamy to light yellow or brownish creamy, fore and mid legs largely yellow, hind trochanter and trochantellus dull orange, antenna with scape, pedicel and variable number of basal flagellomeres dull yellow/orange, scape infuscate over entire dorsal side. Clypeus apically and sometimes dorsal ridge of upper face between antennae narrowly orange.</paragraph>
</subSubSection>
<subSubSection id="C34AC4DF8F15B14F30EBFCE654E83A08" box="[199,389,830,855]" pageId="9" pageNumber="210" type="description">
<paragraph id="8BEF97548F15B14F30EBFCE654E83A08" blockId="9.[151,1436,151,1284]" box="[199,389,830,855]" pageId="9" pageNumber="210">
<emphasis id="B9244B468F15B14F30EBFCE6546C3A08" box="[199,257,830,855]" italics="true" pageId="9" pageNumber="210">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34AC4DF8F15B14F30EBFCBA503C3A24" box="[199,1361,866,891]" pageId="9" pageNumber="210" type="biology_ecology">
<paragraph id="8BEF97548F15B14F30EBFCBA503C3A24" blockId="9.[151,1436,151,1284]" box="[199,1361,866,891]" pageId="9" pageNumber="210">
<emphasis id="B9244B468F15B14F30EBFCBA544A3A24" bold="true" box="[199,295,866,891]" pageId="9" pageNumber="210">Biology.</emphasis>
Hosts unknown. All individuals were collected from high-altitude Costa Rican
<taxonomicName id="4C50ECD78F15B14F3489FCBB506E3A25" box="[1189,1283,867,890]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="251" phylum="Arthropoda" rank="species" species="quercus">
<emphasis id="B9244B468F15B14F3489FCBB506E3A25" box="[1189,1283,867,890]" italics="true" pageId="9" pageNumber="210">Quercus</emphasis>
</taxonomicName>
forest.
</paragraph>
</subSubSection>
<subSubSection id="C34AC4DF8F15B14F30EBFC5E513D3AC0" box="[199,1104,902,927]" pageId="9" pageNumber="210" type="etymology">
<paragraph id="8BEF97548F15B14F30EBFC5E513D3AC0" blockId="9.[151,1436,151,1284]" box="[199,1104,902,927]" pageId="9" pageNumber="210">
<emphasis id="B9244B468F15B14F30EBFC5E54233AC0" bold="true" box="[199,334,902,927]" pageId="9" pageNumber="210">Etymology.</emphasis>
Named after the elongate, pentagonal areolet (a noun in apposition).
</paragraph>
</subSubSection>
<subSubSection id="C34AC4DF8F15B14F30EBFC72513E3D71" pageId="9" pageNumber="210" type="discussion">
<paragraph id="8BEF97548F15B14F30EBFC72513E3D71" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">
<emphasis id="B9244B468F15B14F30EBFC7254223A9C" bold="true" box="[199,335,938,963]" pageId="9" pageNumber="210">Comments.</emphasis>
Very similar to
<taxonomicName id="4C50ECD78F15B14F3238FC7357D93A9D" box="[532,692,939,962]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="concrispus">
<emphasis id="B9244B468F15B14F3238FC7357D93A9D" box="[532,692,939,962]" italics="true" pageId="9" pageNumber="210">O. concrispus</emphasis>
</taxonomicName>
but differs most prominently in the strongly sculptured second tergite (matt and longitudinally striate versus predominantly unsculptured and shiny in
<taxonomicName id="4C50ECD78F15B14F34A1FC1650413AB9" box="[1165,1324,974,998]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="concrispus">
<emphasis id="B9244B468F15B14F34A1FC1650413AB9" box="[1165,1324,974,998]" italics="true" pageId="9" pageNumber="210">O. concrispus</emphasis>
</taxonomicName>
) but also differs in the larger fore wing areolet, the smaller number of antennal flagellomeres, smaller ocelli, darker dorsal surface of the scape, and the more strongly sculptured propodeum and second tergite.
</paragraph>
</subSubSection>
<subSubSection id="C34AC4DF8F15B14F30EBFBE156F23C5B" pageId="9" pageNumber="210" type="materials_examined">
<paragraph id="8BEF97548F15B14F30EBFBE156243D2A" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">
<emphasis id="B9244B468F15B14F30EBFBE154D93D0D" bold="true" box="[199,436,1081,1106]" pageId="9" pageNumber="210">Material examined.</emphasis>
<typeStatus id="54EB29F68F15B14F3193FBE1574A3D0D" box="[447,551,1081,1106]" pageId="9" pageNumber="210" type="holotype">Holotype</typeStatus>
female: ‘
<collectingCountry id="F347D7C48F15B14F32BAFBE156763D0D" box="[662,795,1081,1106]" name="Costa Rica" pageId="9" pageNumber="210">Costa Rica</collectingCountry>
, Sn José Pv. [San José Province], Cerro de la Muerte,
<taxonomicName id="4C50ECD78F15B14F30BBFB8555993D2A" box="[151,244,1117,1141]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="251" phylum="Arthropoda" rank="species" species="quercus">Quercus</taxonomicName>
forest,
<quantity id="4CA83AB18F15B14F3165FB8554CD3D2B" box="[329,416,1117,1141]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="210" unit="m" value="2700.0">2700 m</quantity>
,
<date id="FFEEB1948F15B14F3180FB8554953D2A" box="[428,504,1117,1141]" pageId="9" pageNumber="210" value="1988-07">VII.88</date>
, Gauld &amp; Mitchell’ (
<collectionCode id="ED410F918F15B14F32CFFB8556503D2A" box="[739,829,1117,1141]" pageId="9" pageNumber="210">BMNH</collectionCode>
).
</paragraph>
<paragraph id="8BEF97548F15B14F30EBFB5956F23C5B" blockId="9.[151,1436,151,1284]" pageId="9" pageNumber="210">
<typeStatus id="54EB29F68F15B14F30EBFB5954553DC6" box="[199,312,1153,1177]" pageId="9" pageNumber="210" type="paratype">Paratypes</typeStatus>
: 9 ♀
<collectingCountry id="F347D7C48F15B14F3156FB5954943DC6" box="[378,505,1152,1177]" name="Costa Rica" pageId="9" pageNumber="210">Costa Rica</collectingCountry>
, San José Pv,
<quantity id="4CA83AB18F15B14F32B3FB5957853DC6" box="[671,744,1152,1177]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="210" unit="km" value="20.0">20 km</quantity>
S Empalme, Cerro de la Muerte,
<quantity id="4CA83AB18F15B14F345CFB5951A43DC7" box="[1136,1225,1153,1177]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="210" unit="m" value="2800.0">2800 m</quantity>
,
<date id="FFEEB1948F15B14F34FBFB5950603DC6" box="[1239,1293,1153,1177]" pageId="9" pageNumber="210" value="1988-05">V.88</date>
, Gauld; 4 ♀
<collectingCountry id="F347D7C48F15B14F30BBFB7D547E3DE2" box="[151,275,1188,1213]" name="Costa Rica" pageId="9" pageNumber="210">Costa Rica</collectingCountry>
, San José Pv, Cerro de la Muerte,
<taxonomicName id="4C50ECD78F15B14F32A2FB7D57863DE2" box="[654,747,1189,1213]" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="251" phylum="Arthropoda" rank="species" species="quercus">Quercus</taxonomicName>
forest,
<quantity id="4CA83AB18F15B14F3313FB7D56F83DE3" box="[831,917,1189,1213]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="210" unit="m" value="2700.0">2700 m</quantity>
,
<date id="FFEEB1948F15B14F338CFB7D56BA3DE2" box="[928,983,1189,1213]" pageId="9" pageNumber="210" value="1988-05">V.88</date>
, Gauld &amp; Mitchell; 24 ♀ as previous but collection date
<date id="FFEEB1948F15B14F316DFB1054EC3DBF" box="[321,385,1224,1248]" pageId="9" pageNumber="210" value="1988-06">VI.88</date>
(all
<collectionCode id="ED410F918F15B14F319FFB1057613DBF" box="[435,524,1224,1248]" pageId="9" pageNumber="210">BMNH</collectionCode>
); 9 ♀ as previous but collection date
<date id="FFEEB1948F15B14F338BFB10569D3DBF" box="[935,1008,1224,1248]" pageId="9" pageNumber="210" value="1988-07">VII.88</date>
(4
<collectionCode id="ED410F918F15B14F3439FB1151083DBF" box="[1045,1125,1225,1248]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="9" pageNumber="210">ZMUT</collectionCode>
, 5
<collectionCode id="ED410F918F15B14F34A8FB1051B73DBF" LSID="urn:lsid:biocol.org:col:34871" box="[1156,1242,1224,1248]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="210">USNM</collectionCode>
); 1 ♀
<collectingCountry id="F347D7C48F15B14F3530FB1050F53DBF" box="[1308,1432,1223,1248]" name="Costa Rica" pageId="9" pageNumber="210">Costa Rica</collectingCountry>
,
<quantity id="4CA83AB18F15B14F30BBFB3455833C5C" box="[151,238,1260,1284]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="210" unit="m" value="2800.0">2800 m</quantity>
, San José Pv,
<quantity id="4CA83AB18F15B14F31A0FB3454BF3C5B" box="[396,466,1259,1284]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="210" unit="km" value="20.0">20 km</quantity>
S Empalme,
<date id="FFEEB1948F15B14F3245FB3457C13C5B" box="[617,684,1260,1284]" pageId="9" pageNumber="210" value="1988-09">IX.88</date>
, I.D. Gauld (
<collectionCode id="ED410F918F15B14F3312FB3456F93C5B" box="[830,916,1260,1284]" httpUri="http://grbio.org/cool/9aq9-rt4h" name="University of Tokyo, Department of Zoology" pageId="9" pageNumber="210">ZMUT</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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