treatments-xml/data/03/C4/87/03C487CBEEE93B8EFCEFFE55FC47FE74.xml

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<mods:title id="CECB908B10120F2B889AA5E3B1BC2140">Untold diversity: the astonishing species richness of the Notodelphyidae (Copepoda: Cyclopoida), a family of symbiotic copepods associated with ascidians (Tunicata)</mods:title>
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<mods:namePart id="E0BB5F07DC2C0B39027E8E9A605EFE93">Kim, Il-Hoi</mods:namePart>
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<mods:affiliation id="C19CF5F6FDD2AE150132266AFF2EA3D7">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<mods:namePart id="384BD1DCCB23C5C475B74DA324158AE3">Boxshall, Geoff A.</mods:namePart>
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<mods:affiliation id="05D73331FD8F695355F19875922C2C1A">Korea Institute of Coastal Ecology, Inc., 802 - ho, 302 - dong, 397 Seokcheon-ro, Ojeong-gu, Bucheon, Gyeonggi-do 14449, Republic of Korea ® ihkim @ gwnu. ac. kr; https: // orcid. org / 0000 - 0002 - 7332 - 0043</mods:affiliation>
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<treatment id="03C487CBEEE93B8EFCEFFE55FC47FE74" ID-DOI="http://doi.org/10.5281/zenodo.6422111" ID-GBIF-Taxon="194475055" ID-Zenodo-Dep="6422111" LSID="urn:lsid:plazi:treatment:03C487CBEEE93B8EFCEFFE55FC47FE74" httpUri="http://treatment.plazi.org/id/03C487CBEEE93B8EFCEFFE55FC47FE74" lastPageId="443" lastPageNumber="444" pageId="440" pageNumber="441" scope_class="Copepoda" scope_family="Notodelphyidae" scope_order="Cyclopoida" scope_subPhylum="Tunicata">
<subSubSection id="C3776556EEE93B8DFCEFFE55FB52FE74" box="[810,1175,410,437]" pageId="440" pageNumber="441" type="nomenclature">
<paragraph id="8BD236DDEEE93B8DFCEFFE55FB52FE74" blockId="440.[810,1175,410,471]" box="[810,1175,410,437]" pageId="440" pageNumber="441">
<heading id="D09A81B1EEE93B8DFCEFFE55FB52FE74" box="[810,1175,410,437]" fontSize="11" level="2" pageId="440" pageNumber="441" reason="2">
<emphasis id="B919EACFEEE93B8DFCEFFE55FB52FE74" bold="true" box="[810,1175,410,437]" pageId="440" pageNumber="441">
<taxonomicName id="4C6D4D5EEEE93B8DFCEFFE55FBF7FE74" ID-CoL="7ZG3Q" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[810,1074,410,437]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="440" pageNumber="441" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEE93B8DFCEFFE55FBF7FE74" bold="true" box="[810,1074,410,437]" italics="true" pageId="440" pageNumber="441">Bysone brevicaudatus</emphasis>
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<taxonomicNameLabel id="A22A57B4EEE93B8DFBFFFE55FB52FE74" box="[1082,1175,411,437]" pageId="440" pageNumber="441" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BD236DDEEE93B8DFCEFFE73FC18FE16" blockId="440.[810,1175,410,471]" box="[810,989,445,471]" pageId="440" pageNumber="441">
(
<figureCitation id="13562A58EEE93B8DFCF7FE73FC5BFE16" box="[818,926,445,471]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Figs. 292</figureCitation>
, 
<figureCitation id="13562A58EEE93B8DFC6CFE73FC11FE16" box="[937,980,445,471]" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="440" pageNumber="441">293</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEE93B8DFCEFFDCFFC23FD07" pageId="440" pageNumber="441" type="materials_examined">
<paragraph id="8BD236DDEEE93B8DFCEFFDCFFC23FD07" blockId="440.[810,1452,513,2070]" pageId="440" pageNumber="441">
<emphasis id="B919EACFEEE93B8DFCEFFDCFFC1FFDDA" bold="true" box="[810,986,513,539]" pageId="440" pageNumber="441">Type material</emphasis>
. 
<materialsCitation id="3B053C80EEE93B8DFC37FDCFFC23FD07" ID-GBIF-Occurrence="3748446423" collectedFrom="from Stolonica multitestis Monniot C., Monniot F., Griffiths &amp; Schleyer, 2001" collectingDate="1995-11-17" collectorName="Monniot" country="Mozambique" location="Matemo" pageId="440" pageNumber="441" specimenCode="MNHN-IU-2014-21355" specimenCount="1" specimenCount-female="1" stateProvince="Matemo" typeStatus="holotype">
<typeStatus id="54D6887FEEE93B8DFC37FDCFFB9FFDDA" box="[1010,1114,513,539]" pageId="440" pageNumber="441" type="holotype">Holotype</typeStatus>
<specimenCount id="9D6BFD54EEE93B8DFBAFFDCCFBBAFDDD" box="[1130,1151,514,540]" pageId="440" pageNumber="441" type="female">♀</specimenCount>
(dissected and mounted on a slide, 
<specimenCode id="DBCB9EA6EEE93B8DFC09FDEDFB1FFDFF" box="[972,1242,547,574]" pageId="440" pageNumber="441">MNHN-IU-2014-21355</specimenCode>
) 
<collectedFrom id="38717520EEE93B8DFB32FDEDFCA7FD40" pageId="440" pageNumber="441">
from 
<taxonomicName id="4C6D4D5EEEE93B8DFA87FDEDFCA7FD40" authority="Monniot C., Monniot F., Griffiths &amp; Schleyer, 2001" authorityName="Monniot C., Monniot F., Griffiths &amp; Schleyer" authorityYear="2001" class="Ascidiacea" family="Styelidae" genus="Stolonica" kingdom="Animalia" order="Pleurogona" pageId="440" pageNumber="441" phylum="Chordata" rank="species" species="multitestis">
<emphasis id="B919EACFEEE93B8DFA87FDEDFC58FD9E" italics="true" pageId="440" pageNumber="441">Stolonica multitestis</emphasis>
Monniot C., Monniot F., Griffiths &amp; Schleyer, 2001
</taxonomicName>
</collectedFrom>
(MNHN-IT-2008-8032 = MNHN A1/RIT/25), south of 
<collectingRegion id="49A9F83FEEE93B8DFC48FD44FC28FD65" box="[909,1005,650,676]" pageId="440" pageNumber="441">Matemo</collectingRegion>
, Ibo I., 
<collectingCountry id="F37A764DEEE93B8DFB80FD47FB19FD62" box="[1093,1244,649,675]" name="Mozambique" pageId="440" pageNumber="441">Mozambique</collectingCountry>
, 
<collectorName id="2698530BEEE93B8DFB2DFD47FA8DFD62" box="[1256,1352,649,675]" pageId="440" pageNumber="441">Monniot</collectorName>
coll., 
<date id="FFD3101DEEE93B8DFA4AFD47FC27FD07" pageId="440" pageNumber="441" value="1995-11-17">
<collectingDate id="EF97E9F5EEE93B8DFA4AFD47FC27FD07" pageId="440" pageNumber="441" value="1995-11-17">17 November 1995</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEE93B8DFC9AFD03FBA6FCEA" pageId="440" pageNumber="441" type="etymology">
<paragraph id="8BD236DDEEE93B8DFC9AFD03FBA6FCEA" blockId="440.[810,1452,513,2070]" pageId="440" pageNumber="441">
<emphasis id="B919EACFEEE93B8DFC9AFD03FC25FD26" bold="true" box="[863,992,717,743]" pageId="440" pageNumber="441">Etymology</emphasis>
. The specific name is derived from the Latin 
<emphasis id="B919EACFEEE93B8DFCB7FD21FC66FCC8" box="[882,931,751,777]" italics="true" pageId="440" pageNumber="441">brev</emphasis>
(=short) and 
<emphasis id="B919EACFEEE93B8DFB88FD21FB46FCC8" box="[1101,1155,751,777]" italics="true" pageId="440" pageNumber="441">caud</emphasis>
(=tail) and refers to the relatively stout caudal rami.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEE93B8EFC9AFCFDFEBBFAE8" lastPageId="443" lastPageNumber="444" pageId="440" pageNumber="441" type="description">
<paragraph id="8BD236DDEEE93B8DFC9AFCFDFC72FA8A" blockId="440.[810,1452,513,2070]" pageId="440" pageNumber="441">
<emphasis id="B919EACFEEE93B8DFC9AFCFDFBAAFC8C" bold="true" box="[863,1135,819,845]" pageId="440" pageNumber="441">Descriptionoffemale</emphasis>
. Body (
<figureCitation id="13562A58EEE93B8DFB1FFCFDFA92FC8C" box="[1242,1367,819,845]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292A</figureCitation>
) stout, curved ventrally. Prosome 1.40 mm long: dorsal cephalic shield clearly defined; first to fourth pedigerous somites discernible by constrictions; second and third pedigerous somites each with trace of tergite. Third and fourth pedigerous somites forming brood pouch. Free urosome (
<figureCitation id="13562A58EEE93B8DFCF7FC31FC69FBD8" box="[818,940,1023,1049]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292B</figureCitation>
) 5-segmented: genital somite well defined ventrally from preceding somite, but dorsally confluent. Anal somite about 1.5 times longer than preceding abdominal somite. Caudalramus (
<figureCitation id="13562A58EEE93B8DFB14FBABFA88FBBE" box="[1233,1357,1125,1151]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292C</figureCitation>
) nearly rectangular, about 2.7 times longerthan wide (148×54 μm) and aslongas anal somite: armedwith 6 smallsetae (outer proximal, ventral, and 4 distal); all setae small, less than half of width of ramus at base; outer proximal and ventral setae located at 38 and 62% of ramus length, respectively.
</paragraph>
<paragraph id="8BD236DDEEE93B8DFC9AFA9AFAAFF925" blockId="440.[810,1452,513,2070]" pageId="440" pageNumber="441">
Rostrum (
<figureCitation id="13562A58EEE93B8DFC1BFA9DFBA7FAAC" box="[990,1122,1363,1389]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292D</figureCitation>
) 95×118 μm, indistinctly articulated at base tapering distally towards rounded apex. Antennule (
<figureCitation id="13562A58EEE93B8DFC37FA59FBA0FA73" box="[1010,1125,1431,1458]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292E</figureCitation>
) 270 μmlong, 6-segmented; armatureformula 3, 14, 7+aesthetasc, 3+aesthetasc, 2+aesthetasc, and 9+2 aesthetascs; setae naked and short; distal seta on fourth and fifth segment annulated at base. Antenna (
<figureCitation id="13562A58EEE93B8DFC5FF9D1FBC8F9FB" box="[922,1037,1567,1594]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292F</figureCitation>
) 4-segmented; coxa, basis, and first endopodal segment unarmed; compound distal endopodal segment about 3.1 times longer than wide (75×24 μm) and 1.3 times longerthan firstendopodal segment: armed with 1 subdistal and 3 distal setae, plus strongly curved terminal claw, about 0.65 times as long as segment.
</paragraph>
<paragraph id="8BD236DDEEE93B8EFC9AF925FEF1FE3B" blockId="440.[810,1452,513,2070]" lastBlockId="443.[136,778,173,982]" lastPageId="443" lastPageNumber="444" pageId="440" pageNumber="441">
Labrum (
<figureCitation id="13562A58EEE93B8DFC0DF925FBF8F8C7" box="[968,1085,1771,1798]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292G</figureCitation>
) with concave posterior margin, pointed posterolateral corners, and broad posteromedian lobe bearing 2 denticles at apex. Mandible (
<figureCitation id="13562A58EEE93B8DFAE9F8FEFA66F88B" box="[1324,1443,1840,1866]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="440" pageNumber="441">Fig. 292H</figureCitation>
) withbroadcoxalgnathobasebearing 5 teethand 1 spinulelike subsidiary tooth distally at base of distalmost tooth: basis with 1 setaon medial margin; exopod as long as endopod, armed with 2 long proximal and 2 short distal setae: endopodarmedwith 2 and 5 setaeon firstand second segments, respectively; middle seta on second segment markedly longer than other 4 setae. Maxillule (
<figureCitation id="13562A58EEEA3B8EFD5DFF63FCC7FF06" box="[664,770,173,199]" captionStart="FIGURE 292" captionStartId="441.[136,245,1907,1933]" captionTargetBox="[246,1343,216,1845]" captionTargetId="figure-18@441.[197,1391,185,1870]" captionTargetPageId="441" captionText="FIGURE 292. Bysone brevicaudatus sp. nov., female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="443" pageNumber="444">Fig. 292I</figureCitation>
) with 9 setae on arthrite, 1 thin seta on coxal endite, 2 unequal setaeon epipodite, 3 unequal setae onmedial margin of basis, 3 setae distally on exopod and 2 setae distally on endopod. Maxilla (
<figureCitation id="13562A58EEEA3B8EFE2DFEFBFDA5FE8E" box="[488,608,309,335]" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="443" pageNumber="444">Fig. 293A</figureCitation>
) 5-segmented; syncoxa with 8 setae arranged as 3, 1, 2, and 2; basis with robust claw plus 1 seta; endopod with 1, 1, and 3 setae on first to third segments, respectively. Maxilliped (
<figureCitation id="13562A58EEEA3B8EFD19FE55FF03FE16" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="443" pageNumber="444">Fig. 293B</figureCitation>
) unsegmented, armed with 8 medial setae and 1 long outer seta.
</paragraph>
<caption id="DF126655EEE83B8CFF4DF8BDFEEEF813" ID-DOI="http://doi.org/10.5281/zenodo.4591879" ID-Zenodo-Dep="4591879" httpUri="https://zenodo.org/record/4591879/files/figure.png" pageId="441" pageNumber="442" startId="441.[136,245,1907,1933]" targetBox="[246,1343,216,1845]" targetPageId="441">
<paragraph id="8BD236DDEEE83B8CFF4DF8BDFEEEF813" blockId="441.[136,1451,1907,2002]" pageId="441" pageNumber="442">
<emphasis id="B919EACFEEE83B8CFF4DF8BDFEEAF84F" bold="true" box="[136,303,1907,1934]" pageId="441" pageNumber="442">FIGURE 292.</emphasis>
<taxonomicName id="4C6D4D5EEEE83B8CFEF2F8BAFDEAF84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[311,559,1908,1934]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="441" pageNumber="442" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEE83B8CFEF2F8BAFDEAF84F" box="[311,559,1908,1934]" italics="true" pageId="441" pageNumber="442">Bysone brevicaudatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEE83B8CFDF2F8BAFD56F84F" bold="true" box="[567,659,1908,1934]" pageId="441" pageNumber="442">
<taxonomicNameLabel id="A22A57B4EEE83B8CFDF2F8BAFD56F84F" box="[567,659,1908,1934]" pageId="441" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, habitus, right; B, urosome, dorsal; C, right caudal ramus, dorsal; D, rostrum; E, antennule; F, antenna; G, labrum; H, mandible; I, maxillule. Scale bars: A, 0.2 mm; B, 0.1 mm; C–I, 0.05 mm.
</paragraph>
</caption>
<caption id="DF126655EEEB3B8FFF4DF8BDFD2CF871" ID-DOI="http://doi.org/10.5281/zenodo.4591881" ID-Zenodo-Dep="4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="442" pageNumber="443" startId="442.[136,245,1907,1933]" targetBox="[179,1399,274,1789]" targetPageId="442">
<paragraph id="8BD236DDEEEB3B8FFF4DF8BDFD2CF871" blockId="442.[136,1452,1907,1968]" pageId="442" pageNumber="443">
<emphasis id="B919EACFEEEB3B8FFF4DF8BDFEE8F84F" bold="true" box="[136,301,1907,1934]" pageId="442" pageNumber="443">FIGURE 293.</emphasis>
<taxonomicName id="4C6D4D5EEEEB3B8FFEF1F8BAFDEFF84F" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[308,554,1908,1934]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="442" pageNumber="443" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEEB3B8FFEF1F8BAFDEFF84F" box="[308,554,1908,1934]" italics="true" pageId="442" pageNumber="443">Bysone brevicaudatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEEB3B8FFDF4F8BAFD4FF84F" bold="true" box="[561,650,1908,1934]" pageId="442" pageNumber="443">
<taxonomicNameLabel id="A22A57B4EEEB3B8FFDF4F8BAFD4FF84F" box="[561,650,1908,1934]" pageId="442" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm.
</paragraph>
</caption>
<paragraph id="8BD236DDEEEA3B8EFF78FDCCFDA4FC14" blockId="443.[136,778,173,982]" pageId="443" pageNumber="444">
Leg 1 (
<figureCitation id="13562A58EEEA3B8EFEDDFDCFFE55FDDA" box="[280,400,513,539]" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="443" pageNumber="444">Fig. 293C</figureCitation>
) small, with 2-segmented rami; second segments of both rami compound, with traces of articulation; inner coxal seta pinnate; outer seta on basis much larger than that of legs 2–4; first exopodal segment lacking inner seta. Legs 2–4 each with 3-segmented exopod and 2-segmented endopod (
<figureCitation id="13562A58EEEA3B8EFDE1FD65FD7AFD07" box="[548,703,683,710]" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="443" pageNumber="444">Fig. 293D, E</figureCitation>
); both rami elongate; endopod extending to distal border of second exopodal segment; second endopodal segment about 3 times longer than first in legs 2 and 3, about 4 times longer in leg 4. Inner coxal seta present in legs 2–4, but that of leg 4 minute. Outer and distal setae on exopod of leg 2-4 short, naked, and bluntly tipped. Inner setae on exopod and all setae on endopod of leg 4 small and naked. Armature formula for legs 1–4 as follows:
</paragraph>
<paragraph id="8BD236DDEEEA3B8EFED2FC31FCC0FBBE" pageId="443" pageNumber="444">
<table id="F96DC47DEEEAC5CAFF4DFC31FCC2FB41" box="[136,775,1023,1152]" gridcols="5" gridrows="4" inLine="true" pageId="443" pageNumber="444">
<tr id="355D349FEEEAC5CAFF4DFC31FCC2FBD8" box="[136,775,1023,1049]" gridrow="0" pageId="443" pageNumber="444" rowspan-0="1">
<th id="768C5DE3EEEAC5CAFED2FC31FE94FBD8" box="[279,337,1023,1049]" gridcol="1" gridrow="0" pageId="443" pageNumber="444">Coxa</th>
<th id="768C5DE3EEEAC5CAFEAFFC31FE62FBD8" box="[362,423,1023,1049]" gridcol="2" gridrow="0" pageId="443" pageNumber="444">Basis</th>
<th id="768C5DE3EEEAC5CAFE07FC31FDA8FBD8" box="[450,621,1023,1049]" gridcol="3" gridrow="0" pageId="443" pageNumber="444">Exopod</th>
<th id="768C5DE3EEEAC5CAFD4BFC31FCC2FBD8" box="[654,775,1023,1049]" gridcol="4" gridrow="0" pageId="443" pageNumber="444">Endopod</th>
</tr>
<tr id="355D349FEEEAC5CAFF4DFBEFFCC2FBFD" box="[136,775,1057,1084]" gridrow="1" pageId="443" pageNumber="444">
<th id="768C5DE3EEEAC5CAFF4DFBEFFEC6FBFD" box="[136,259,1057,1084]" gridcol="0" gridrow="1" pageId="443" pageNumber="444">Leg 1</th>
<td id="768C5DE3EEEAC5CAFED2FBEFFE94FBFD" box="[279,337,1057,1084]" gridcol="1" gridrow="1" pageId="443" pageNumber="444">0-1</td>
<td id="768C5DE3EEEAC5CAFEAFFBEFFE62FBFD" box="[362,423,1057,1084]" gridcol="2" gridrow="1" pageId="443" pageNumber="444">1-0</td>
<td id="768C5DE3EEEAC5CAFE07FBEFFDA8FBFD" box="[450,621,1057,1084]" gridcol="3" gridrow="1" pageId="443" pageNumber="444">I-0; IV, I, 3</td>
<td id="768C5DE3EEEAC5CAFD4BFBEFFCC2FBFD" box="[654,775,1057,1084]" gridcol="4" gridrow="1" pageId="443" pageNumber="444">0-1; 1, 2, 4</td>
</tr>
<tr id="355D349FEEEAC5CAFF4DFB8DFCC2FB9F" box="[136,775,1091,1118]" gridrow="2" pageId="443" pageNumber="444">
<th id="768C5DE3EEEAC5CAFF4DFB8DFEC6FB9F" box="[136,259,1091,1118]" gridcol="0" gridrow="2" pageId="443" pageNumber="444">Legs 2 &amp; 3</th>
<td id="768C5DE3EEEAC5CAFED2FB8DFE94FB9F" box="[279,337,1091,1118]" gridcol="1" gridrow="2" pageId="443" pageNumber="444">0-1</td>
<td id="768C5DE3EEEAC5CAFEAFFB8DFE62FB9F" box="[362,423,1091,1118]" gridcol="2" gridrow="2" pageId="443" pageNumber="444">1-0</td>
<td id="768C5DE3EEEAC5CAFE07FB8DFDA8FB9F" box="[450,621,1091,1118]" gridcol="3" gridrow="2" pageId="443" pageNumber="444">1-1; 1-1; 3, 1, 5</td>
<td id="768C5DE3EEEAC5CAFD4BFB8DFCC2FB9F" box="[654,775,1091,1118]" gridcol="4" gridrow="2" pageId="443" pageNumber="444">0-1; 1, 2, 5</td>
</tr>
<tr id="355D349FEEEAC5CAFF4DFBABFCC2FB41" box="[136,775,1125,1152]" gridrow="3" pageId="443" pageNumber="444">
<th id="768C5DE3EEEAC5CAFF4DFBABFEC6FB41" box="[136,259,1125,1152]" gridcol="0" gridrow="3" pageId="443" pageNumber="444">Leg 4</th>
<td id="768C5DE3EEEAC5CAFED2FBABFE94FB41" box="[279,337,1125,1152]" gridcol="1" gridrow="3" pageId="443" pageNumber="444">0-1</td>
<td id="768C5DE3EEEAC5CAFEAFFBABFE62FB41" box="[362,423,1125,1152]" gridcol="2" gridrow="3" pageId="443" pageNumber="444">1-0</td>
<td id="768C5DE3EEEAC5CAFE07FBABFDA8FB41" box="[450,621,1125,1152]" gridcol="3" gridrow="3" pageId="443" pageNumber="444">1-1; 1-1; 2, 1, 5</td>
<td id="768C5DE3EEEAC5CAFD4BFBABFCC2FB41" box="[654,775,1125,1152]" gridcol="4" gridrow="3" pageId="443" pageNumber="444">0-1; 1, 2, 4</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD236DDEEEA3B8EFF78FB64FED4FAC6" blockId="443.[136,778,1193,2070]" pageId="443" pageNumber="444">
Leg 5 (
<figureCitation id="13562A58EEEA3B8EFED6FB67FE46FB02" box="[275,387,1193,1219]" captionStart="FIGURE 293" captionStartId="442.[136,245,1907,1933]" captionTargetBox="[179,1399,274,1789]" captionTargetId="figure-20@442.[136,1451,236,1819]" captionTargetPageId="442" captionText="FIGURE 293. Bysone brevicaudatus sp. nov., female. A, maxilla; B, maxilliped; C, leg 1; D, leg 2; E, leg 4; F, leg 5. Scale bars: A–C, 0.05 mm; D, E, 0.1 mm; F, 0.02 mm." figureDoi="http://doi.org/10.5281/zenodo.4591881" httpUri="https://zenodo.org/record/4591881/files/figure.png" pageId="443" pageNumber="444">Fig. 293F</figureCitation>
) represented by small ventrodistal lobe bearing 2 small exopodal setae distally; protopodal seta lacking.
</paragraph>
<paragraph id="8BD236DDEEEA3B8EFF78FAC1FEBBFAE8" blockId="443.[136,778,1193,2070]" box="[189,382,1295,1322]" pageId="443" pageNumber="444">
<emphasis id="B919EACFEEEA3B8EFF78FAC1FF3FFAE8" bold="true" box="[189,250,1295,1321]" pageId="443" pageNumber="444">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3776556EEEA3B8EFF78FAFFFC47FE74" pageId="443" pageNumber="444" type="discussion">
<paragraph id="8BD236DDEEEA3B8EFF78FAFFFE15F871" blockId="443.[136,778,1193,2070]" pageId="443" pageNumber="444">
<emphasis id="B919EACFEEEA3B8EFF78FAFFFEEFFA8A" bold="true" box="[189,298,1329,1355]" pageId="443" pageNumber="444">Remarks</emphasis>
. The genus 
<taxonomicName id="4C6D4D5EEEEA3B8EFE00FAFCFDD3FA8A" authorityName="Stock &amp; Humes" authorityYear="1970" box="[453,534,1330,1355]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEEEA3B8EFE00FAFCFDD3FA8A" box="[453,534,1330,1355]" italics="true" pageId="443" pageNumber="444">Bysone</emphasis>
</taxonomicName>
has comprised only the 
<typeStatus id="54D6887FEEEA3B8EFF73FA9AFF23FAAF" box="[182,230,1364,1390]" pageId="443" pageNumber="444">type</typeStatus>
species, 
<taxonomicName id="4C6D4D5EEEEA3B8EFE91FA9AFE3EFAAC" authorityName="Stock &amp; Humes" authorityYear="1970" box="[340,507,1363,1390]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFE91FA9AFEA0FAAC" box="[340,357,1364,1389]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFEBDFA9DFE3EFAAC" box="[376,507,1363,1389]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
since it was originally established based on material collected in 
<collectingCountry id="F37A764DEEEA3B8EFD47FABBFCCCFA4E" box="[642,777,1397,1423]" name="Madagascar" pageId="443" pageNumber="444">Madagascar</collectingCountry>
(
<bibRefCitation id="EFFC4B2CEEEA3B8EFF55FA59FE53FA73" author="Stock, J. H. &amp; Humes, A. G." box="[144,406,1431,1458]" journalOrPublisher="Zoologischer Anzeiger" pageId="443" pageNumber="444" pagination="193 - 212" part="184" refId="ref331227" refString="Stock, J. H. &amp; Humes, A. G. (1970) On four new notodelphyid copepods, associated with an octocoral, Parerythropodium fulvum (Forskal), in Madagascar. Zoologischer Anzeiger, 184, 193 - 212." title="On four new notodelphyid copepods, associated with an octocoral, Parerythropodium fulvum (Forskal)" type="journal article" volumeTitle="Madagascar" year="1970">Stock &amp; Humes, 1970</bibRefCitation>
). 
<taxonomicName id="4C6D4D5EEEEA3B8EFE69FA56FD61FA70" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" box="[428,676,1431,1457]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEEA3B8EFE69FA56FD61FA70" box="[428,676,1431,1457]" italics="true" pageId="443" pageNumber="444">Bysone brevicaudatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEEA3B8EFD6BFA56FCCFFA73" bold="true" box="[686,778,1432,1458]" pageId="443" pageNumber="444">
<taxonomicNameLabel id="A22A57B4EEEA3B8EFD6BFA56FCCFFA73" box="[686,778,1432,1458]" pageId="443" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares important character states with the 
<typeStatus id="54D6887FEEEA3B8EFDB8FA74FD68FA15" box="[637,685,1466,1492]" pageId="443" pageNumber="444">type</typeStatus>
species including: a 6-segmented antennule, 4 setae on the mandibular exopod, 2 and 5 setae, respectively, on the first and second segments of mandibular endopod, a 2- segmented exopod of leg 1, a similar form of maxilliped, and a similar form of the mandibular gnathobase which bears a spinule-like distal subsidiary denticle and has no proximal seta. These similarities are sufficient to confirm the placement of the new species in 
<taxonomicName id="4C6D4D5EEEEA3B8EFD81F904FD50F922" authorityName="Stock &amp; Humes" authorityYear="1970" box="[580,661,1738,1763]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="genus">
<emphasis id="B919EACFEEEA3B8EFD81F904FD50F922" box="[580,661,1738,1763]" italics="true" pageId="443" pageNumber="444">Bysone</emphasis>
</taxonomicName>
. The two species can be distinguished by reference to the maxilla, which has a stong claw plus a seta on the basis and the endopod is 3-segmented in the new species. In contrast, 
<taxonomicName id="4C6D4D5EEEEA3B8EFF4DF89CFEEEF8AD" authorityName="Stock &amp; Humes" authorityYear="1970" box="[136,299,1874,1900]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFF4DF89CFF5CF8AA" box="[136,153,1874,1899]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFF6DF89CFEEEF8AD" box="[168,299,1874,1900]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
has the maxillary basis armed with only 2 setae and the endopod is 2-segmented. These differences in the maxilla are significant.
</paragraph>
<paragraph id="8BD236DDEEEA3B8EFF78F876FC47FE74" blockId="443.[136,778,1193,2070]" lastBlockId="443.[810,1452,173,437]" pageId="443" pageNumber="444">
There are other differences between these two species: the maxilliped is armed with 9 (8+1) setae in 
<taxonomicName id="4C6D4D5EEEEA3B8EFD37F814FEE3F7D7" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEEA3B8EFD37F814FCC6F832" box="[754,771,2010,2035]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFF4DF832FEE3F7D7" box="[136,294,2044,2070]" italics="true" pageId="443" pageNumber="444">brevicaudatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEEA3B8EFEEBF832FE4CF7D7" bold="true" box="[302,393,2044,2070]" pageId="443" pageNumber="444">
<taxonomicNameLabel id="A22A57B4EEEA3B8EFEEBF832FE4CF7D7" box="[302,393,2044,2070]" pageId="443" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(vs. 5+1 setaein 
<taxonomicName id="4C6D4D5EEEEA3B8EFD93F832FD3CF7D7" authorityName="Stock &amp; Humes" authorityYear="1970" box="[598,761,2044,2070]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFD93F832FDA2F7D4" box="[598,615,2044,2069]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFDB3F832FD3CF7D7" box="[630,761,2044,2070]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
), the maxillulary arthrite is armed with 9 setae in 
<taxonomicName id="4C6D4D5EEEEA3B8EFA56FF60FC0DFF28" authority="Kim &amp; Boxshall, 2020" authorityName="Kim &amp; Boxshall" authorityYear="2020" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="brevicaudatus" status="sp. nov.">
<emphasis id="B919EACFEEEA3B8EFA56FF60FA61FF06" box="[1427,1444,174,199]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFCEFFF01FC0DFF28" box="[810,968,207,233]" italics="true" pageId="443" pageNumber="444">brevicaudatus</emphasis>
</taxonomicName>
<emphasis id="B919EACFEEEA3B8EFC0BFF01FBEDFF28" bold="true" box="[974,1064,207,233]" pageId="443" pageNumber="444">
<taxonomicNameLabel id="A22A57B4EEEA3B8EFC0BFF01FBEDFF28" box="[974,1064,207,233]" pageId="443" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(vs. 5 setaein 
<taxonomicName id="4C6D4D5EEEEA3B8EFB15FF1EFAB7FF28" authorityName="Stock &amp; Humes" authorityYear="1970" box="[1232,1394,207,233]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFB15FF1EFB24FF28" box="[1232,1249,208,233]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFB2AFF01FAB7FF28" box="[1263,1394,207,233]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
), the basis of leg 1 bears an outer seta (vs. seta absent in 
<taxonomicName id="4C6D4D5EEEEA3B8EFA56FF3CFC68FEEC" authorityName="Stock &amp; Humes" authorityYear="1970" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFA56FF3CFA61FECA" box="[1427,1444,242,267]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFCEFFEDDFC68FEEC" box="[810,941,275,301]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
), the second endopodal segment of leg 1 is armed with 7 setae (vs. 6 setae), and leg 5 is short, lobate (vs. digitiform in 
<taxonomicName id="4C6D4D5EEEEA3B8EFBCBFE96FB7FFEB0" authorityName="Stock &amp; Humes" authorityYear="1970" box="[1038,1210,343,369]" class="Copepoda" family="Notodelphyidae" genus="Bysone" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cyclopoida" pageId="443" pageNumber="444" phylum="Arthropoda" rank="species" species="operculatus">
<emphasis id="B919EACFEEEA3B8EFBCBFE96FBDAFEB0" box="[1038,1055,344,369]" italics="true" pageId="443" pageNumber="444">B</emphasis>
. 
<emphasis id="B919EACFEEEA3B8EFBF2FE99FB7FFEB0" box="[1079,1210,343,369]" italics="true" pageId="443" pageNumber="444">operculatus</emphasis>
</taxonomicName>
). These differences are sufficient to support the establishment of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>