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<document id="9E8B09EF95D5F745A1E068DF0930F62B" ID-CLB-Dataset="30140" ID-DOI="10.11646/zootaxa.4415.3.6" ID-GBIF-Dataset="f7dfd7e8-238c-432a-9eec-0929f785f747" ID-ISSN="1175-5326" ID-Zenodo-Dep="1242185" ID-ZooBank="30CF1B69-020C-43E8-B8F0-C30B41E04FE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1525600684827" checkinUser="plazi" docAuthor="Coelho, Beatriz W. T. &amp; Engel, Michael S." docDate="2018" docId="C55BAA03FFAC7B53FF47FAAF358AFDD9" docLanguage="en" docName="zootaxa.4415.3.6.imf" docOrigin="Zootaxa 4415 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ruizantheda colombiana Coelho &amp; Engel 2018, n. sp." docType="treatment" docVersion="9" lastPageNumber="519" masterDocId="3962D27BFFA87B55FFD0FFBB3400FFB9" masterDocTitle="A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)" masterLastPageNumber="532" masterPageNumber="513" originalUpdateDomain="Boston" originalUpdateTime="1545217612389" originalUpdateUser="gglab" pageNumber="517" updateTime="1698541935543" updateUser="ExternalLinkService">
<mods:mods id="BD557049792B2052F7894AE7671DE25C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87841729A2B50F344CF1EE6A94A1856B">
<mods:title id="B260085ACFEF2FA2BB63AF8750C61B53">A new group of species within the bee genus Ruizantheda, with a revised key to the males of the genus (Hymenoptera: Halictidae: Caenohalictini)</mods:title>
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<mods:name id="24AAF98D4DFF19191D88434A07266EC6" type="personal">
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<mods:namePart id="B3F36A007EF373F2EC5FE916088AF292">Coelho, Beatriz W. T.</mods:namePart>
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<mods:name id="DC9A910B9A6031BCEB86B605F1939512" type="personal">
<mods:role id="5749D3A819FCB4C12BD8ED2AE822FCB9">
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<mods:namePart id="BAE048314E82CE5ED067DCC559A38C1F">Engel, Michael S.</mods:namePart>
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<mods:typeOfResource id="37EA97A737C3400F65A7D0DF5B2DDACD">text</mods:typeOfResource>
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<mods:title id="ADF949E448D4373BCB7DED1A623EA253">Zootaxa</mods:title>
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<mods:part id="FDC956DE08FAD8C5BA30C5963A43CC2B">
<mods:date id="3D4E67FB75162EAE3C1D9DDEACCE9237">2018</mods:date>
<mods:detail id="19EE8C64CD215E334EEA51D836045CD2" type="volume">
<mods:number id="CCC6A8B34BF7AEF4A1BFAC7CA20F99DD">4415</mods:number>
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<mods:classification id="E70025E38AE9B04289FA50F9672E9C2E">journal article</mods:classification>
<mods:identifier id="770ECFC2D855EBC87926AE3454DDA92C" type="CLB-Dataset">30140</mods:identifier>
<mods:identifier id="02F82CBCC6A42C2CA4A84AEF4F2118C1" type="DOI">10.11646/zootaxa.4415.3.6</mods:identifier>
<mods:identifier id="914AAD2E10E456A7690E54ECAD4FCE3E" type="GBIF-Dataset">f7dfd7e8-238c-432a-9eec-0929f785f747</mods:identifier>
<mods:identifier id="C3DB5A2E0E8F4C7B0996462EFB79E3D6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="596C66462337E431E55042BE94A79C77" type="Zenodo-Dep">1242185</mods:identifier>
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<treatment id="C55BAA03FFAC7B53FF47FAAF358AFDD9" ID-DOI="http://doi.org/10.5281/zenodo.6489463" ID-GBIF-Taxon="143811443" ID-Zenodo-Dep="6489463" LSID="urn:lsid:plazi:treatment:C55BAA03FFAC7B53FF47FAAF358AFDD9" httpUri="http://treatment.plazi.org/id/C55BAA03FFAC7B53FF47FAAF358AFDD9" lastPageId="6" lastPageNumber="519" pageId="4" pageNumber="517">
<subSubSection id="05E8489EFFAC7B51FF47FAAF360EFAF6" pageId="4" pageNumber="517" type="nomenclature">
<paragraph id="4D4D1B15FFAC7B51FF47FAAF360FFA97" blockId="4.[151,527,1300,1359]" box="[151,527,1300,1326]" pageId="4" pageNumber="517">
<heading id="1605AC79FFAC7B51FF47FAAF360FFA97" bold="true" box="[151,527,1300,1326]" fontSize="11" level="1" pageId="4" pageNumber="517" reason="1">
<emphasis id="7F86C707FFAC7B51FF47FAAF360FFA97" bold="true" box="[151,527,1300,1326]" pageId="4" pageNumber="517">
<taxonomicName id="8AF26096FFAC7B51FF47FAAF35C5FA97" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[151,453,1300,1326]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="517" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="7F86C707FFAC7B51FF47FAAF35C5FA97" bold="true" box="[151,453,1300,1326]" italics="true" pageId="4" pageNumber="517">Ruizantheda colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFAC7B51FE1CFAAF360FFA97" box="[460,527,1300,1326]" pageId="4" pageNumber="517" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4D4D1B15FFAC7B51FF47FA8D360EFAF6" blockId="4.[151,527,1300,1359]" box="[151,526,1334,1359]" pageId="4" pageNumber="517">
(
<figureCitation id="D5C90790FFAC7B51FF4EFA8D350BFAF6" box="[158,267,1334,1359]" captionStart="FIGURES3–8" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES3–8. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="4" pageNumber="517">Figures 4</figureCitation>
,
<figureCitation id="D5C90790FFAC7B51FEC7FA8C355EFAF6" box="[279,350,1335,1359]" captionStart="FIGURES 11–13" captionStartId="7.[151,264,1902,1925]" captionTargetBox="[225,1360,951,1880]" captionTargetId="figure@7.[219,1367,949,1881]" captionTargetPageId="7" captionText="FIGURES 11–13. Male legs. 11–12, mesofemur and mesotrochanter: 11, Ruizantheda colombiana n. sp.; 12, R. venezuelana n. sp. 13, hind leg of R. colombiana n. sp." httpUri="https://zenodo.org/record/1242193/files/figure.png" pageId="4" pageNumber="517">11, 13</figureCitation>
,
<figureCitation id="D5C90790FFAC7B51FEBAFA8C3589FAF6" box="[362,393,1335,1359]" captionStart="FIGURES 26–27" captionStartId="12.[151,264,1992,2015]" captionTargetBox="[259,1316,1494,1970]" captionTargetId="figure@12.[259,1328,1494,1971]" captionTargetPageId="12" captionText="FIGURES 26–27. Male.26, habitus, lateral view, Ruizanthedanigra n. sp. 27, S5,R. colombianan. sp." httpUri="https://zenodo.org/record/1242199/files/figure.png" pageId="4" pageNumber="517">27</figureCitation>
,
<figureCitation id="D5C90790FFAC7B51FE44FA8D35DDFAF6" box="[404,477,1334,1359]" captionStart="FIGURES 34–37" captionStartId="15.[151,264,1703,1726]" captionTargetBox="[225,1363,542,1683]" captionTargetId="figure@15.[224,1363,541,1683]" captionTargetPageId="15" captionText="FIGURES 34–37. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 34–35, R. colombiana n. sp.; 36–37, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="4" pageNumber="517">34–35</figureCitation>
,
<figureCitation id="D5C90790FFAC7B51FE38FA8C3606FAF6" box="[488,518,1335,1359]" captionStart="FIGURES 42–44" captionStartId="16.[151,264,1426,1449]" captionTargetBox="[241,1331,260,1411]" captionTargetId="figure@16.[240,1346,251,1423]" captionTargetPageId="16" captionText="FIGURES 42–44. Male. 42–43, genital capsule, ventral view; 44, gonostylus, ventral view. 42, Ruizantheda nigra n. sp.; 43, R.venezuelana n. sp.; 44, R. colombianan. sp." httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="4" pageNumber="517">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFAC7B51FF47FAC530DAF9BA" pageId="4" pageNumber="517" type="diagnosis">
<paragraph id="4D4D1B15FFAC7B51FF47FAC530DAF9BA" blockId="4.[151,1437,1406,2008]" pageId="4" pageNumber="517">
<emphasis id="7F86C707FFAC7B51FF47FAC5350AFA2E" bold="true" box="[151,266,1406,1431]" pageId="4" pageNumber="517">Diagnosis</emphasis>
. Metatibia with subapical region concave ventrally (
<figureCitation id="D5C90790FFAC7B51FC5BFAC537E3FA2E" box="[907,995,1406,1431]" captionStart="FIGURES 11–13" captionStartId="7.[151,264,1902,1925]" captionTargetBox="[225,1360,951,1880]" captionTargetId="figure@7.[219,1367,949,1881]" captionTargetPageId="7" captionText="FIGURES 11–13. Male legs. 11–12, mesofemur and mesotrochanter: 11, Ruizantheda colombiana n. sp.; 12, R. venezuelana n. sp. 13, hind leg of R. colombiana n. sp." httpUri="https://zenodo.org/record/1242193/files/figure.png" pageId="4" pageNumber="517">Fig. 13</figureCitation>
) and
<emphasis id="7F86C707FFAC7B51FBE0FAC53052FA2E" bold="true" box="[1072,1106,1406,1431]" pageId="4" pageNumber="517">rvl</emphasis>
with projection formed by sclerotized area that partially borders a translucent and flat surface (
<figureCitation id="D5C90790FFAC7B51FC4FFA1837FEFA02" box="[927,1022,1443,1468]" captionStart="FIGURES 34–37" captionStartId="15.[151,264,1703,1726]" captionTargetBox="[225,1363,542,1683]" captionTargetId="figure@15.[224,1363,541,1683]" captionTargetPageId="15" captionText="FIGURES 34–37. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 34–35, R. colombiana n. sp.; 36–37, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="4" pageNumber="517">Figs. 34</figureCitation>
,
<figureCitation id="D5C90790FFAC7B51FBDCFA18302AFA02" box="[1036,1066,1443,1467]" captionStart="FIGURES 42–44" captionStartId="16.[151,264,1426,1449]" captionTargetBox="[241,1331,260,1411]" captionTargetId="figure@16.[240,1346,251,1423]" captionTargetPageId="16" captionText="FIGURES 42–44. Male. 42–43, genital capsule, ventral view; 44, gonostylus, ventral view. 42, Ruizantheda nigra n. sp.; 43, R.venezuelana n. sp.; 44, R. colombianan. sp." httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="4" pageNumber="517">44</figureCitation>
) are distinct autapomorphies for this species. Other diagnostic characters include: mesotrochanter with strong pointed keel ventro-proximally (
<figureCitation id="D5C90790FFAC7B51FABFFA7D34B4F9BA" captionStart="FIGURES 11–13" captionStartId="7.[151,264,1902,1925]" captionTargetBox="[225,1360,951,1880]" captionTargetId="figure@7.[219,1367,949,1881]" captionTargetPageId="7" captionText="FIGURES 11–13. Male legs. 11–12, mesofemur and mesotrochanter: 11, Ruizantheda colombiana n. sp.; 12, R. venezuelana n. sp. 13, hind leg of R. colombiana n. sp." httpUri="https://zenodo.org/record/1242193/files/figure.png" pageId="4" pageNumber="517">Fig. 11</figureCitation>
); S5 with minute U-shaped gradulus (
<figureCitation id="D5C90790FFAC7B51FD8EFA5036B2F9BA" box="[606,690,1515,1540]" captionStart="FIGURES 26–27" captionStartId="12.[151,264,1992,2015]" captionTargetBox="[259,1316,1494,1970]" captionTargetId="figure@12.[259,1328,1494,1971]" captionTargetPageId="12" captionText="FIGURES 26–27. Male.26, habitus, lateral view, Ruizanthedanigra n. sp. 27, S5,R. colombianan. sp." httpUri="https://zenodo.org/record/1242199/files/figure.png" pageId="4" pageNumber="517">Fig. 27</figureCitation>
); and basal region of volsella L-shaped (
<figureCitation id="D5C90790FFAC7B51FBA9FA5030CDF9BA" box="[1145,1229,1515,1540]" captionStart="FIGURES 34–37" captionStartId="15.[151,264,1703,1726]" captionTargetBox="[225,1363,542,1683]" captionTargetId="figure@15.[224,1363,541,1683]" captionTargetPageId="15" captionText="FIGURES 34–37. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 34–35, R. colombiana n. sp.; 36–37, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="4" pageNumber="517">Fig. 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFAC7B53FF17F9B5363AFE31" lastPageId="6" lastPageNumber="519" pageId="4" pageNumber="517" type="description">
<paragraph id="4D4D1B15FFAC7B51FF17F9B534F4F9F2" blockId="4.[151,1437,1406,2008]" pageId="4" pageNumber="517">
<emphasis id="7F86C707FFAC7B51FF17F9B53551F99E" bold="true" box="[199,337,1550,1575]" pageId="4" pageNumber="517">Description</emphasis>
. Male:
<emphasis id="7F86C707FFAC7B51FE77F9B435D4F99F" box="[423,468,1551,1574]" italics="true" pageId="4" pageNumber="517">Size</emphasis>
. Male. Length approximately 11.0 mm. Forewing length 8.0 mm. Intertegular distance
<quantity id="8A0AB6F0FFAC7B51FF47F98834F0F9F2" box="[151,240,1587,1611]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="517" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="4D4D1B15FFAC7B51FF17F9ED34E6F946" blockId="4.[151,1437,1406,2008]" pageId="4" pageNumber="517">
<emphasis id="7F86C707FFAC7B51FF17F9ED3515F9D6" box="[199,277,1622,1647]" italics="true" pageId="4" pageNumber="517">Colour</emphasis>
. Head and mesosoma bright green, with blue highlights on clypeus, pronotum, and propodeum. About apical one third of clypeus with transverse yellow band. Mandible dark brown with reddish apex. Labral basal area yellow, labral distal process brown. Legs brown, pro- and metacoxae, pro- and mesofemora with faint green-blue highlights. Metasoma brown, discs of T1 and basal bands of T2–4 with blue-green highlights. Pygidial plate brown.
</paragraph>
<paragraph id="4D4D1B15FFAC7B50FF17F8B035C5FEA4" blockId="4.[151,1437,1406,2008]" lastBlockId="5.[151,1437,151,752]" lastPageId="5" lastPageNumber="518" pageId="4" pageNumber="517">
<emphasis id="7F86C707FFAC7B51FF17F8B0354BF89D" box="[199,331,1803,1828]" italics="true" pageId="4" pageNumber="517">Pubescence</emphasis>
. Compound eyes with brownish setae, approximately as long as ocellar diameter. Face with long, plumose, whitish setae, less-branched brownish setae on vertex, dark setae on clypeus. Lower paraocular area with strip of short, plumose, white setae along inner margin of compound eye. Mesosoma with brownish-yellow setae, except anterior margin of mesoscutum with white setae. Legs with whitish setae, except inner surface of tibiae with yellowish-white setae, golden on tarsi, and some blackish setae on outer surface of metatibia. Discs of T2–5 with semi-erect to semi-decumbent setae of variable length. Posterior marginal zone of T2 with many short, simple, appressed, whitish setae. Apicolateral lobes of S4 with tuft of branched setae and 7–9 stout, simple, recurved setae on margin, and the remainder with scattered few setae. S5 with conspicuous triangular patch of short, decumbent setae under minute U-shaped gradulus, setae becoming slightly longer toward apical margin; apical margin with setae simple and branched.
</paragraph>
<paragraph id="4D4D1B15FFAD7B50FF17FE9335B2FD49" blockId="5.[151,1437,151,752]" pageId="5" pageNumber="518">
<emphasis id="7F86C707FFAD7B50FF17FE93352CFE86" box="[199,300,296,319]" italics="true" pageId="5" pageNumber="518">Structure</emphasis>
. Head about as long as wide (
<figureCitation id="D5C90790FFAD7B50FD50FE9C36C6FEF9" box="[640,710,295,320]" captionStart="FIGURES3–8" captionStartId="4.[151,264,1172,1195]" captionTargetBox="[229,1362,387,1142]" captionTargetId="figure@4.[220,1366,375,1150]" captionTargetPageId="4" captionText="FIGURES3–8. Male heads. 3, Ruizantheda nigra n. sp; 4, R.colombianan. sp.; 5, R.venezuelana n. sp.; 6, R. baeri (Vachal); 7, R. pilosa n. sp.; 8, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242189/files/figure.png" pageId="5" pageNumber="518">Fig. 4</figureCitation>
). Upper to lower interocular distance ratio about 1.2. Malar area linear. Mandible simple. Labral distal process in the form of a minute inverted triangle. Epistomal sulcus forming obtuse angle and slightly rounded. Clypeus almost entirely projecting below lower orbital tangent. Ocellocular area slightly concave. Vertex expanded behind ocelli. Frons slightly depressed. Preoccipital ridge rounded. Profemur swollen. Mesotrochanter with strong pointed keel ventro-proximally (
<figureCitation id="D5C90790FFAD7B50FC0DFE0C3036FE69" box="[989,1078,439,464]" captionStart="FIGURES 11–13" captionStartId="7.[151,264,1902,1925]" captionTargetBox="[225,1360,951,1880]" captionTargetId="figure@7.[219,1367,949,1881]" captionTargetPageId="7" captionText="FIGURES 11–13. Male legs. 11–12, mesofemur and mesotrochanter: 11, Ruizantheda colombiana n. sp.; 12, R. venezuelana n. sp. 13, hind leg of R. colombiana n. sp." httpUri="https://zenodo.org/record/1242193/files/figure.png" pageId="5" pageNumber="518">Fig. 11</figureCitation>
); mesofemur greatly swollen, ventrally flattened; mesotibia slightly swollen with flat, minutely ridged ventral area and small distal tooth on outer surface; mesobasitarsus less than two-thirds length of mesotibia, unmodified. Metatibia with subapical region concave ventrally (
<figureCitation id="D5C90790FFAD7B50FEBBFD9F35BDFD85" box="[363,445,548,573]" captionStart="FIGURES 11–13" captionStartId="7.[151,264,1902,1925]" captionTargetBox="[225,1360,951,1880]" captionTargetId="figure@7.[219,1367,949,1881]" captionTargetPageId="7" captionText="FIGURES 11–13. Male legs. 11–12, mesofemur and mesotrochanter: 11, Ruizantheda colombiana n. sp.; 12, R. venezuelana n. sp. 13, hind leg of R. colombiana n. sp." httpUri="https://zenodo.org/record/1242193/files/figure.png" pageId="5" pageNumber="518">Fig. 13</figureCitation>
). S4 shortened medially, largely hidden under S3, with transverse median depression and apical margin shallowly emarginate. Apical margin of S5 very shallowly emarginate, with minute U-shaped gradulus (
<figureCitation id="D5C90790FFAD7B50FED7FDD7355BFD3D" box="[263,347,620,645]" captionStart="FIGURES 26–27" captionStartId="12.[151,264,1992,2015]" captionTargetBox="[259,1316,1494,1970]" captionTargetId="figure@12.[259,1328,1494,1971]" captionTargetPageId="12" captionText="FIGURES 26–27. Male.26, habitus, lateral view, Ruizanthedanigra n. sp. 27, S5,R. colombianan. sp." httpUri="https://zenodo.org/record/1242199/files/figure.png" pageId="5" pageNumber="518">Fig. 27</figureCitation>
). Gradulus of S6 interrupted medially. S7 with median apical process angulate, short and without setae. S8 with median apical process broad and trapezoidal in shape, apical margin bearing lateral, membranous lobes, separated by a short and sclerotized surface (as Fig. 32), lobe medially rounded. Pygidial plate oval, very slightly truncate apically.
</paragraph>
<caption id="198D4B9DFFAD7B50FF47FAC730E4FA2B" box="[151,1252,1404,1427]" httpUri="https://zenodo.org/record/1242191/files/figure.png" pageId="5" pageNumber="518" startId="5.[151,264,1404,1427]" targetBox="[230,1366,810,1375]" targetPageId="5">
<paragraph id="4D4D1B15FFAD7B50FF47FAC730E4FA2B" blockId="5.[151,1252,1404,1427]" box="[151,1252,1404,1427]" pageId="5" pageNumber="518">
<emphasis id="7F86C707FFAD7B50FF47FAC73540FA28" bold="true" box="[151,320,1404,1427]" pageId="5" pageNumber="518">FIGURES 9–10</emphasis>
. 9, head of
<taxonomicName id="8AF26096FFAD7B50FE68FAC7367AFA2B" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[440,634,1404,1426]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="7F86C707FFAD7B50FE68FAC7367AFA2B" box="[440,634,1404,1426]" italics="true" pageId="5" pageNumber="518">Ruizantheda pilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFAD7B50FD51FAC636BAFA2B" box="[641,698,1405,1426]" pageId="5" pageNumber="518" rank="species">
<emphasis id="7F86C707FFAD7B50FD51FAC636B3FA2B" bold="true" box="[641,691,1405,1426]" pageId="5" pageNumber="518">n. sp</emphasis>
.
</taxonomicNameLabel>
, female. 10, metasomal terga of
<taxonomicName id="8AF26096FFAD7B50FBD6FAC63060FA2B" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[1030,1120,1404,1426]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="518" phylum="Arthropoda" rank="species" species="pilosa" status="sp. nov.">
<emphasis id="7F86C707FFAD7B50FBD6FAC63015FA2B" box="[1030,1045,1405,1426]" italics="true" pageId="5" pageNumber="518">R</emphasis>
.
<emphasis id="7F86C707FFAD7B50FBF2FAC73060FA2B" box="[1058,1120,1404,1426]" italics="true" pageId="5" pageNumber="518">pilosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFAD7B50FBB6FAC6309FFA2B" box="[1126,1183,1405,1426]" pageId="5" pageNumber="518" rank="species">
<emphasis id="7F86C707FFAD7B50FBB6FAC63099FA2B" bold="true" box="[1126,1177,1405,1426]" pageId="5" pageNumber="518">n. sp</emphasis>
.
</taxonomicNameLabel>
, male.
</paragraph>
</caption>
<paragraph id="4D4D1B15FFAD7B50FF17FA043772F8DD" blockId="5.[151,1437,1471,2036]" pageId="5" pageNumber="518">
<emphasis id="7F86C707FFAD7B50FF17FA043531FA61" box="[199,305,1471,1496]" italics="true" pageId="5" pageNumber="518">Genitalia</emphasis>
(
<figureCitation id="D5C90790FFAD7B50FE91FA0435CBFA61" box="[321,459,1471,1496]" captionStart="FIGURES 34–37" captionStartId="15.[151,264,1703,1726]" captionTargetBox="[225,1363,542,1683]" captionTargetId="figure@15.[224,1363,541,1683]" captionTargetPageId="15" captionText="FIGURES 34–37. Male genital capsule. 34, 36 ventral view; 35, 37 dorsal view. 34–35, R. colombiana n. sp.; 36–37, R. gaullei (Vachal)." httpUri="https://zenodo.org/record/1242203/files/figure.png" pageId="5" pageNumber="518">Figs. 34–35</figureCitation>
,
<figureCitation id="D5C90790FFAD7B50FE07FA0435F7FA6E" box="[471,503,1471,1495]" captionStart="FIGURES 42–44" captionStartId="16.[151,264,1426,1449]" captionTargetBox="[241,1331,260,1411]" captionTargetId="figure@16.[240,1346,251,1423]" captionTargetPageId="16" captionText="FIGURES 42–44. Male. 42–43, genital capsule, ventral view; 44, gonostylus, ventral view. 42, Ruizantheda nigra n. sp.; 43, R.venezuelana n. sp.; 44, R. colombianan. sp." httpUri="https://zenodo.org/record/1242205/files/figure.png" pageId="5" pageNumber="518">44</figureCitation>
). Basal region of gonobase with W-shaped in ventral view. Gonocoxae with outer margins divergent and convex near apex; inner dorsal margins progressively divergent, at about midpoint becoming slightly curved and much more divergent. Basal region of gonostylus with
<emphasis id="7F86C707FFAD7B50FBBCF9BC308EF999" bold="true" box="[1132,1166,1543,1568]" pageId="5" pageNumber="518">rvl</emphasis>
sclerotized, long, more than one-half length of gonocoxite and rounded apically, with simple setae at extremity, overlapping those of opposing gonostylus; with median projection formed by sclerotized area that partially borders a translucent and flat surface; inner lobe present.
<emphasis id="7F86C707FFAD7B50FE06F9C83606F935" bold="true" box="[470,518,1651,1676]" pageId="5" pageNumber="518">Mgl</emphasis>
crescent-shaped with outer surface expanded medially, and with scattered short setae.
<emphasis id="7F86C707FFAD7B50FF0CF92C350FF909" bold="true" box="[220,271,1687,1712]" pageId="5" pageNumber="518">Ogp</emphasis>
large, terminating before apex of
<emphasis id="7F86C707FFAD7B50FD59F92C36B6F909" bold="true" box="[649,694,1687,1712]" pageId="5" pageNumber="518">mgl</emphasis>
, with many long and plumose setae arising at juncture of
<emphasis id="7F86C707FFAD7B50FAE8F92C3164F909" bold="true" box="[1336,1380,1687,1712]" pageId="5" pageNumber="518">ogp</emphasis>
with
<emphasis id="7F86C707FFAD7B50FF47F90034C4F96D" bold="true" box="[151,196,1723,1748]" pageId="5" pageNumber="518">mgl</emphasis>
; outer margin less sclerotized, extending to ventral region and laterally forming an inflated area with cuticular wrinkles; clump of median, erect, plumose setae at base of
<emphasis id="7F86C707FFAD7B50FCFCF9643757F941" bold="true" box="[812,855,1759,1784]" pageId="5" pageNumber="518">ogp</emphasis>
. Penis valve strongly arched in lateral view; strong, main, central dorsal ridge; apex pointed; outer lateral expansion bilobed, forming an obtuse angle between two sections, and without projection posteriorly; prong very narrow, extending past volsella posteriorly. Volsella medio-apical margin strongly emarginate; basal region L-shaped.
</paragraph>
<paragraph id="4D4D1B15FFAD7B53FF17F8D4363AFE31" blockId="5.[151,1437,1471,2036]" lastBlockId="6.[151,1437,151,608]" lastPageId="6" lastPageNumber="519" pageId="5" pageNumber="518">
<emphasis id="7F86C707FFAD7B50FF17F8D43531F831" box="[199,305,1903,1928]" italics="true" pageId="5" pageNumber="518">Sculpture</emphasis>
. Upper paraocular area and frons densely punctate, with fine, contiguous punctures and scattered setal bases intermixed. Lower paraocular area with coarse punctures separated by 1–3 PD. Supraclypeal area sparsely punctured medially, becoming more densely punctured laterally. Disc of clypeus with coarse punctures separated by 1–3 PD. Face with surface between punctures microreticulate, weaker on clypeus. Mesoscutum with setal bases separated by 3–8 PD centrally, surface slightly roughened. Mesoscutellum with a few coarse setal bases centrally, becoming more densely punctured posteriorly and laterally (2–4 PD), surface with appearance bright. Pre-episternum with setal bases separated by 2–4 PD. Mesepisternum with setal bases separated by 2–6 PD. Metapostnotum with narrowly angulate triangular area defined by microareolate sculpture and a fine stria in the middle of metapostnotum, from the base to the apex. Surface between setal bases reticulate throughout mesosoma. Discs of T2–3 with fine punctures separated by 2–3 PD, sparsely punctate on apical margins, surfaces reticulate, slightly finer than remainder of body.
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFAE7B53FF17FE2F3092FE14" box="[199,1170,404,429]" pageId="6" pageNumber="519" type="distribution">
<paragraph id="4D4D1B15FFAE7B53FF17FE2F3092FE14" blockId="6.[151,1437,151,608]" box="[199,1170,404,429]" pageId="6" pageNumber="519">
<emphasis id="7F86C707FFAE7B53FF17FE2F3560FE14" bold="true" box="[199,352,404,429]" pageId="6" pageNumber="519">Distribution.</emphasis>
<taxonomicName id="8AF26096FFAE7B53FEB7FE2F367FFE14" authority="Coelho &amp; Engel, 2018" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[359,639,404,429]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="519" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="7F86C707FFAE7B53FEB7FE2F367FFE14" box="[359,639,404,429]" italics="true" pageId="6" pageNumber="519">Ruizantheda colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="64B57A7CFFAE7B53FD55FE2F36C4FE14" box="[645,708,404,429]" pageId="6" pageNumber="519" rank="species">
<emphasis id="7F86C707FFAE7B53FD55FE2F36C4FE14" bold="true" box="[645,708,404,429]" pageId="6" pageNumber="519">n. sp.</emphasis>
</taxonomicNameLabel>
is presently known only from
<collectingCountry id="35E55B85FFAE7B53FBCCFE2F308EFE14" box="[1052,1166,404,429]" name="Colombia" pageId="6" pageNumber="519">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFAE7B53FF17FE0C3087FDA1" pageId="6" pageNumber="519" type="materials_examined">
<paragraph id="4D4D1B15FFAE7B53FF17FE0C3087FDA1" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="519">
<materialsCitation id="FD9A1148FFAE7B53FF17FE0C3087FDA1" ID-GBIF-Occurrence="1949259678" collectingDate="1977-02-07" collectionCode="SEMC" collectorName="M. Breed &amp; C. D. Michener" country="Colombia" elevation="3300" location="above Tenerife" pageId="6" pageNumber="519" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="7F86C707FFAE7B53FF17FE0C356FFE69" bold="true" box="[199,367,439,464]" pageId="6" pageNumber="519">
<typeStatus id="9249A5B7FFAE7B53FF17FE0C3502FE69" box="[199,258,439,464]" pageId="6" pageNumber="519">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="9249A5B7FFAE7B53FEAEFE0C35E6FE69" box="[382,486,439,464]" pageId="6" pageNumber="519" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="2BE383D0FFAE7B53FDE7FE0C3688FE69" box="[567,648,439,464]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="6" pageNumber="519">
<emphasis id="7F86C707FFAE7B53FDE7FE0C3688FE69" bold="true" box="[567,648,439,464]" pageId="6" pageNumber="519">SEMC</emphasis>
</collectionCode>
), “
<collectingCountry id="35E55B85FFAE7B53FD7BFE033747FE69" box="[683,839,440,464]" name="Colombia" pageId="6" pageNumber="519">COLOMBIA</collectingCountry>
: Prov. Valle\ above
<location id="482D4DCEFFAE7B53FBFAFE0C3086FE69" LSID="urn:lsid:plazi:treatment:C55BAA03FFAC7B53FF47FAAF358AFDD9:482D4DCEFFAE7B53FBFAFE0C3086FE69" box="[1066,1158,439,464]" country="Colombia" name="Tenerife" pageId="6" pageNumber="519">Tenerife</location>
\
<collectingDate id="2908C43DFFAE7B53FB48FE033122FE69" box="[1176,1314,439,464]" pageId="6" pageNumber="519" value="1977-02-07">
<date id="394C3DD5FFAE7B53FB48FE033122FE69" box="[1176,1314,439,464]" pageId="6" pageNumber="519" value="1977-02-07">7 Feb. 1977</date>
</collectingDate>
.
<elevation id="C6DFFC26FFAE7B53FAFFFE033188FE76" box="[1327,1416,440,464]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="519" unit="m" value="3300.0">
<quantity id="8A0AB6F0FFAE7B53FAFFFE033188FE76" box="[1327,1416,440,464]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="6" pageNumber="519" unit="m" value="3300.0">3300 m</quantity>
</elevation>
. \
<collectorName id="E0077EC3FFAE7B53FF47FE663505FE4C" box="[151,261,476,501]" pageId="6" pageNumber="519">M. Breed</collectorName>
,
<collectorName id="E0077EC3FFAE7B53FEC2FE6735BEFE4C" box="[274,446,476,501]" pageId="6" pageNumber="519">C.D. Michener</collectorName>
” and “
<typeStatus id="9249A5B7FFAE7B53FDDBFE6736A1FE4D" box="[523,673,476,500]" pageId="6" pageNumber="519" type="holotype">HOLOTYPE</typeStatus>
\
<taxonomicName id="8AF26096FFAE7B53FD6BFE6737D3FE4C" authorityName="Coelho &amp; Engel" authorityYear="2018" box="[699,979,476,501]" class="Insecta" family="Halictidae" genus="Ruizantheda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="519" phylum="Arthropoda" rank="species" species="colombiana">Ruizantheda colombiana</taxonomicName>
\
<location id="482D4DCEFFAE7B53FC0BFE67302BFE4C" LSID="urn:lsid:plazi:treatment:C55BAA03FFAC7B53FF47FAAF358AFDD9:482D4DCEFFAE7B53FC0BFE67302BFE4C" box="[987,1067,476,501]" country="Colombia" county="Specimen" municipality="Engel" name="Coelho" pageId="6" pageNumber="519">Coelho</location>
&amp;
<collectingMunicipality id="AD29816FFFAE7B53FB82FE673096FE4C" box="[1106,1174,476,501]" pageId="6" pageNumber="519">Engel</collectingMunicipality>
” [red label].
<collectingCounty id="A42C6399FFAE7B53FAFEFE67319CFE4C" box="[1326,1436,476,501]" pageId="6" pageNumber="519">Specimen</collectingCounty>
lacking antennae, and with head, left metatibia, metatarsus, and metasoma glued to board.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="05E8489EFFAE7B53FF17FD9F358AFDD9" pageId="6" pageNumber="519" type="etymology">
<paragraph id="4D4D1B15FFAE7B53FF17FD9F358AFDD9" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="519">
<emphasis id="7F86C707FFAE7B53FF17FD9F354EFD84" bold="true" box="[199,334,548,573]" pageId="6" pageNumber="519">Etymology.</emphasis>
The specific epithet refers to the occurrence of the species in
<collectingCountry id="35E55B85FFAE7B53FBDFFD9F3080FD84" box="[1039,1152,548,573]" name="Colombia" pageId="6" pageNumber="519">Colombia</collectingCountry>
, one of the northernmost records for the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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