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<document id="BC003177E575F7E876E920E3C73999F8" ID-CLB-Dataset="31287" ID-DOI="10.11646/zootaxa.4358.2.1" ID-GBIF-Dataset="d226f633-d454-4e32-b12f-6d00c30bddfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1067996" ID-ZooBank="8861BCC0-022F-4803-98E8-D28B90F666E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511949109352" checkinUser="plazi" docAuthor="Dow, Rory A., Stokvis, Frank &amp; Ngiam, Robin W. J." docDate="2017" docId="03C3A90C985FFF84FF6A89B7FC7FEDB6" docLanguage="en" docName="zootaxa.4358.2.1.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptogomphus Selys in Borneo 1878" docType="treatment" docVersion="8" lastPageNumber="215" masterDocId="FFFAD1749852FF8AFFFD8823FF9DEF7A" masterDocTitle="Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)" masterLastPageNumber="257" masterPageNumber="201" pageNumber="214" updateTime="1698505057438" updateUser="ExternalLinkService">
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<mods:title id="F7A2DDEA511EE99D0D98FB81345C1E33">Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae)</mods:title>
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<mods:namePart id="BAE0AE381A307E7FCFC05E3CC54FF53F">Dow, Rory A.</mods:namePart>
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<mods:namePart id="C8FEE85CA9D40A8AE8CB4266D07EBE05">Stokvis, Frank</mods:namePart>
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<mods:namePart id="0240C23D7F94CC76FFAB07C3F29DBC68">Ngiam, Robin W. J.</mods:namePart>
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<mods:title id="71DEB8D0B3913A6A963864EC35E0A280">Zootaxa</mods:title>
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<mods:date id="7D76BA24A271434557CFD2FCB06FB650">2017</mods:date>
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<treatment id="03C3A90C985FFF84FF6A89B7FC7FEDB6" ID-DOI="http://doi.org/10.5281/zenodo.5631248" ID-GBIF-Taxon="137580771" ID-Zenodo-Dep="5631248" LSID="urn:lsid:plazi:treatment:03C3A90C985FFF84FF6A89B7FC7FEDB6" httpUri="http://treatment.plazi.org/id/03C3A90C985FFF84FF6A89B7FC7FEDB6" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
<subSubSection id="C3704B91985FFF87FF6A89B7FD33EEB5" pageId="13" pageNumber="214" type="nomenclature">
<paragraph id="8BD5181A985FFF87FF6A89B7FD5EEED4" blockId="13.[151,707,404,463]" box="[151,707,404,430]" pageId="13" pageNumber="214">
<heading id="D09DAF76985FFF87FF6A89B7FD5EEED4" bold="true" box="[151,707,404,430]" fontSize="11" level="1" pageId="13" pageNumber="214" reason="1">
<emphasis id="B91EC408985FFF87FF6A89B7FD5EEED4" bold="true" box="[151,707,404,430]" pageId="13" pageNumber="214">
<taxonomicName id="4C6A6399985FFF87FF6A89B7FED7EED4" ID-CoL="5D3K" authorityName="Selys in Borneo" authorityYear="1878" box="[151,330,404,430]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B91EC408985FFF87FF6A89B7FED7EED4" bold="true" box="[151,330,404,430]" italics="true" pageId="13" pageNumber="214">Leptogomphus</emphasis>
</taxonomicName>
sp. cf
<taxonomicName id="4C6A6399985FFF87FE6789B7FD5EEED4" authority="Laidlaw, 1936" authorityName="Laidlaw" authorityYear="1936" box="[410,707,404,430]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FE6789B7FD93EED4" bold="true" box="[410,526,404,430]" italics="true" pageId="13" pageNumber="214">coomansi</emphasis>
<bibRefCitation id="EFFB65EB985FFF87FDE989B7FD5EEED4" author="Laidlaw" box="[532,707,404,430]" pageId="13" pageNumber="214" refString="Laidlaw, F. F. (1936) On some examples of the genus Leptogomphus (Odon., Gomphidae). Treubia, 15 (3), 267 - 270." type="journal article" year="1936">Laidlaw, 1936</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD5181A985FFF87FF6A8995FD33EEB5" blockId="13.[151,707,404,463]" box="[151,686,438,463]" pageId="13" pageNumber="214">
(
<figureCitation id="1351049F985FFF87FF628995FF78EEB5" box="[159,229,438,463]" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="13" pageNumber="214">Figs 1</figureCitation>
,
<figureCitation id="1351049F985FFF87FF0E8994FE9EEEB5" box="[243,259,439,463]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="13" pageNumber="214">2</figureCitation>
,
<figureCitation id="1351049F985FFF87FEF28994FE82EEB5" box="[271,287,439,463]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="13" pageNumber="214">3</figureCitation>
,
<figureCitation id="1351049F985FFF87FED68994FED4EEB5" box="[299,329,439,463]" captionStart="FIGURES10–13" captionStartId="14.[151,264,1864,1887]" captionTargetBox="[151,1436,772,1843]" captionTargetId="figure@14.[151,1436,772,1843]" captionTargetPageId="14" captionText="FIGURES10–13. Male head frontal: (10) L.coomansi SAR09_10_GOM4; (11) L.coomansi holotype; (12) L.sp. cf coomansi SAB12_GOM4; (13) L.pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068006/files/figure.png" pageId="13" pageNumber="214">12</figureCitation>
,
<figureCitation id="1351049F985FFF87FEA88994FEEEEEB5" box="[341,371,439,463]" captionStart="FIGURES18–23" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[183,1420,198,1847]" captionTargetId="figure@18.[183,1420,198,1847]" captionTargetPageId="18" captionText="FIGURES18–23.Female headdorsal:(18) L.coomansi SAR11_12_GOM48;(19)L.mariaeholotype;(20) L.sp.cf coomansi PomS260; (21) L. pasia holotype; (22) L.sii paratype SAR13_14_GOM44; (23) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068010/files/figure.png" pageId="13" pageNumber="214">20</figureCitation>
,
<figureCitation id="1351049F985FFF87FE828994FE00EEB5" box="[383,413,439,463]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="13" pageNumber="214">30</figureCitation>
,
<figureCitation id="1351049F985FFF87FE548994FE5AEEB5" box="[425,455,439,463]" captionStart="FIGURES 36–41" captionStartId="25.[151,264,1973,1996]" captionTargetBox="[204,1387,183,1958]" captionTargetId="figure@25.[204,1387,183,1958]" captionTargetPageId="25" captionText="FIGURES 36–41. Female head frontal: (36) L. coomansi SAR11_12_GOM48; (37) L. mariae holotype; (38) L. sp. cf coomansi PomS260; (39) L.pasia holotype; (40) L.sii paratype SAR13_14_GOM44; (41) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068018/files/figure.png" pageId="13" pageNumber="214">38</figureCitation>
,
<figureCitation id="1351049F985FFF87FE2E8995FE6FEEB5" box="[467,498,438,463]" captionStart="FIGURES 52–55" captionStartId="31.[151,264,1501,1524]" captionTargetBox="[151,1436,409,1479]" captionTargetId="figure@31.[151,1436,409,1480]" captionTargetPageId="31" captionText="FIGURES 52–55. Male synthorax dorsal: (52) L. coomansi SAR09_10_GOM4; (53) L. coomansi holotype; (54) L. sp. cf coomansi SAB12_GOM4; (55) L. pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068024/files/figure.png" pageId="13" pageNumber="214">54</figureCitation>
,
<figureCitation id="1351049F985FFF87FE038995FD81EEB5" box="[510,540,438,463]" captionStart="FIGURES 60–63" captionStartId="34.[151,264,1601,1624]" captionTargetBox="[156,1432,517,1580]" captionTargetId="figure@34.[156,1432,517,1580]" captionTargetPageId="34" captionText="FIGURES 60–63. Male synthorax lateral: (60) L. coomansi SAR09_10_GOM4; (61) L. sp. cf coomansi SAB12_GOM4; (62) L. schieli holotype; (63) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068028/files/figure.png" pageId="13" pageNumber="214">61</figureCitation>
,
<figureCitation id="1351049F985FFF87FDDA8994FDD8EEB5" box="[551,581,439,463]" captionStart="FIGURES 68–73" captionStartId="36.[151,264,1915,1938]" captionTargetBox="[156,1432,193,1894]" captionTargetId="figure@36.[156,1432,193,1894]" captionTargetPageId="36" captionText="FIGURES 68–73. Female synthorax dorsal: (68) L. coomansi SAR11_12_GOM48; (69) L. mariae holotype; (70) L. sp. cf coomansi PomS260; (71) L.pasia holotype; (72) L.sii paratype SAR13_14_GOM44; (73) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068032/files/figure.png" pageId="13" pageNumber="214">70</figureCitation>
,
<figureCitation id="1351049F985FFF87FDAC8994FDF2EEB5" box="[593,623,439,463]" captionStart="FIGURES 78–83" captionStartId="38.[151,264,1952,1975]" captionTargetBox="[169,1406,182,1934]" captionTargetId="figure@38.[169,1406,182,1934]" captionTargetPageId="38" captionText="FIGURES 78–83. Female synthorax lateral: (78) L. coomansi SAR11_12_GOM48; (79) L. mariae holotype, right hand side, flipped for consistency; (80) L. sp. cf coomansi PomS260; (81) L. pasia holotype; (82) L. sii paratype SAR13_14_GOM44; (83) L.species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068036/files/figure.png" pageId="13" pageNumber="214">80</figureCitation>
,
<figureCitation id="1351049F985FFF87FD868995FD3BEEB5" box="[635,678,438,463]" captionStart="FIGURE 126" captionStartId="48.[151,250,1801,1823]" captionTargetBox="[183,1420,421,1756]" captionTargetId="figure@48.[183,1420,421,1756]" captionTargetPageId="48" captionText="FIGURE 126. Distribution of L. coomansi, confirmed (yellow square) and unconfirmed (blue triangle) records, and L. sp. cf coomansi (black circle)." httpUri="https://zenodo.org/record/1068054/files/figure.png" pageId="13" pageNumber="214">126</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3704B91985FFF87FF6A89DDFD7BEDF5" pageId="13" pageNumber="214" type="reference_group">
<paragraph id="8BD5181A985FFF87FF6A89DDFBA3ED49" blockId="13.[151,1435,510,563]" pageId="13" pageNumber="214">
<taxonomicName id="4C6A6399985FFF87FF6A89DDFEF7ED6F" ID-CoL="3TQ4K" authorityName="van Tol" authorityYear="1990" box="[151,362,510,533]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC408985FFF87FF6A89DDFEF7ED6F" box="[151,362,510,533]" italics="true" pageId="13" pageNumber="214">Leptogomphus pasia</emphasis>
</taxonomicName>
;—van
<bibRefCitation id="EFFB65EB985FFF87FE4889DDFD8EED6E" author="Tol" box="[437,531,510,532]" pageId="13" pageNumber="214" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
: 97, 98 (part, male Danum Valley, Sabah, excluded from type series),
<tableCitation id="C6E82DA1985FFF87FB2189DDFABBED6E" box="[1244,1318,510,532]" captionStart="TABLE 2" captionStartId="3.[151,239,256,279]" captionTargetBox="[159,1375,443,1403]" captionText="TABLE 2. Leptogomphus from Borneo used in the molecular analyses; collectors and more details of location given in material lists under each species. Collection codes and BOLD Process IDs are given along with basic location details. The collection codes can be used to locate the sequences on the BOLD website, and also appear as BOLD Sample IDs there." pageId="13" pageNumber="214">Table 2</tableCitation>
(part, male characters);—?
<bibRefCitation id="EFFB65EB985FFF87FEA28A3EFE67ED49" author="Kalkman" box="[351,506,541,563]" pageId="13" pageNumber="214" refString="Kalkman, V. J. (2005) Some notes on dragonflies observed at the KBFSC, Brunei. Agrion, 9 (1), 13 - 14." type="journal article" year="2005">Kalkman 2005</bibRefCitation>
: 14 (male, Kuala Belalong Field Studies Centre, Brunei).
</paragraph>
<paragraph id="8BD5181A985FFF87FF6A8A79FD7BEDF5" blockId="13.[151,1436,602,655]" pageId="13" pageNumber="214">
<taxonomicName id="4C6A6399985FFF87FF6A8A79FEB1ED0A" ID-CoL="5D3K" authorityName="Selys in Borneo" authorityYear="1878" box="[151,300,602,624]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B91EC408985FFF87FF6A8A79FEB1ED0A" box="[151,300,602,624]" italics="true" pageId="13" pageNumber="214">Leptogomphus</emphasis>
</taxonomicName>
cf
<taxonomicName id="4C6A6399985FFF87FEAF8A78FE14ED0A" ID-CoL="3TQ4K" authorityName="van Tol" authorityYear="1990" box="[338,393,603,624]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="pasia">
<emphasis id="B91EC408985FFF87FEAF8A78FE14ED0A" box="[338,393,603,624]" italics="true" pageId="13" pageNumber="214">pasia</emphasis>
</taxonomicName>
;—van
<bibRefCitation id="EFFB65EB985FFF87FE2B8A79FDAAED0A" author="Tol" box="[470,567,602,624]" pageId="13" pageNumber="214" refString="van Tol, J. (1990) Key to the Malesian species of Leptogomphus Selys, with the description of a new species from Sabah (Odonata, Gomphidae). Tijdschrift voor Entomologie, 133, 97 - 105." type="journal article" year="1990">Tol 1990</bibRefCitation>
: 99, 102,
<figureCitation id="1351049F985FFF87FD5F8A79FC9DED0A" box="[674,768,602,624]" captionStart="FIGURES 4–9" captionStartId="9.[151,264,1713,1736]" captionTargetBox="[151,1436,193,1692]" captionTargetId="figure@9.[151,1436,193,1692]" captionTargetPageId="9" captionText="FIGURES 4–9. Male head dorsal: (4) L. coomansi SAR09_10_GOM4; (5) L. pendleburyi SAR13_14_GOM13; (6) L. pasia SAB12_GOM1; (7) L. sii holotype; (8) L.schieli holotype; (9) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068004/files/figure.png" pageId="13" pageNumber="214">Figs 6, 8</figureCitation>
,
<figureCitation id="1351049F985FFF87FCF38A79FCB4ED0A" box="[782,809,602,624]" captionStart="FIGURES 14–17" captionStartId="16.[151,264,1933,1956]" captionTargetBox="[151,1436,841,1912]" captionTargetId="figure@16.[151,1436,841,1912]" captionTargetPageId="16" captionText="FIGURES 14–17. Male head frontal: (14) L. pasia SAB12_GOM1; (15) L. sii holotype; (16) L. schieli holotype; (17) L. williamsoni SAR11_12_GOM56." httpUri="https://zenodo.org/record/1068008/files/figure.png" pageId="13" pageNumber="214">16</figureCitation>
,
<figureCitation id="1351049F985FFF87FCCA8A79FCE4ED0A" box="[823,889,602,624]" captionStart="FIGURES 32–35" captionStartId="23.[151,264,1357,1380]" captionTargetBox="[151,1436,265,1336]" captionTargetId="figure@23.[151,1436,265,1336]" captionTargetPageId="23" captionText="FIGURES 32–35. Female head dorsal detail; h—occipital horn; p—pit; s—occipital spur; t—tubercle: (32) L pendleburyi SAR16_GOM8; (33) L. schieli paratype SAR15_GOM6; (34) L. williamsoni SAR13_14_GOM51; (35) L. sp. cf williamsoni SAR11_12_GOM59." httpUri="https://zenodo.org/record/1068016/files/figure.png" pageId="13" pageNumber="214">32, 33</figureCitation>
(refers to same male specimen as above);—Dow &amp; Ngiam 2015: 24 (Sungai Sii, upper Baram,
<collectingRegion id="49AED6F8985FFF87FD7D8A5AFD41EDF5" box="[640,732,633,655]" country="Malaysia" name="Sarawak" pageId="13" pageNumber="214">
<collectingCountry id="F37D588A985FFF87FD7D8A5AFD41EDF5" box="[640,732,633,655]" name="Malaysia" pageId="13" pageNumber="214">Sarawak</collectingCountry>
</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3704B91985FFF87FF6A8A9FFBF2EA26" pageId="13" pageNumber="214" type="materials_examined">
<paragraph id="8BD5181A985FFF87FF6A8A9FFDFEEBD2" blockId="13.[151,1437,700,2021]" pageId="13" pageNumber="214">
<emphasis id="B91EC408985FFF87FF6A8A9FFDF1EDAF" bold="true" box="[151,620,700,725]" pageId="13" pageNumber="214">
Material studied.
<collectingRegion id="49AED6F8985FFF87FE858A9FFE79EDAF" box="[376,484,700,725]" country="Malaysia" name="Sarawak" pageId="13" pageNumber="214">
<collectingCountry id="F37D588A985FFF87FE858A9FFE79EDAF" box="[376,484,700,725]" name="Malaysia" pageId="13" pageNumber="214">Sarawak</collectingCountry>
</collectingRegion>
,
<collectingCountry id="F37D588A985FFF87FE0B8A9FFDFAEDAF" box="[502,615,700,725]" name="Malaysia" pageId="13" pageNumber="214">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="9D6CD393985FFF87FD848A9FFD34EDAE" box="[633,681,700,724]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(teneral;
<collectionCode id="ED7B80DF985FFF87FCDC8A9FFCC7EDAF" box="[801,858,700,725]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
15_GOM14), Sungai Selabi, Ulu Mujok,
<collectionCode id="ED7B80DF985FFF87FAB48A9FFA05EDAE" box="[1353,1432,700,724]" pageId="13" pageNumber="214">LEWS</collectionCode>
, Sarikei Division,
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,
<geoCoordinate id="EE5E7EDD985FFF87FE298AFCFDC0ED82" box="[468,605,735,760]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="112.16291">112.16291E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FD918AC3FD73ED82" box="[620,750,735,760]" pageId="13" pageNumber="214" value="2015-08-01">1 viii 2015</date>
, leg. RA. Dow, in collection Dow;
<specimenCount id="9D6CD393985FFF87FB738AC3FB26ED8D" box="[1166,1211,736,760]" pageId="13" pageNumber="214" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FB308AFCFA9BED82" box="[1229,1286,735,760]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
15_GOM20), Sungai Temurok and tributaries inside Lanjak Entimau Wildlife Sanctuary boundary, Ulu Mujok, Sarikei Division,
<geoCoordinate id="EE5E7EDD985FFF87FF6A8B0BFE94EC3A" box="[151,265,808,832]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="1" value="1.70348">1.70348N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FEE88B04FE02EC3A" box="[277,415,807,832]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="112.13513">112.13513E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FE518B0BFDB6EC3A" box="[428,555,807,832]" pageId="13" pageNumber="214" value="2015-08-04">4 viii 2015</date>
, leg. R.A. Dow, in collection R.A. Dow;
<specimenCount id="9D6CD393985FFF87FBF98B0BFBB2EC45" box="[1028,1071,808,832]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FBC28B0BFBE5EC3A" box="[1087,1144,808,832]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
07_8_GOM49), stream in unplanted valley, block E2N, Tubau,
<collectingRegion id="49AED6F8985FFF87FDB88B6FFD3BEC1F" box="[581,678,844,869]" country="Malaysia" name="Sarawak" pageId="13" pageNumber="214">
<collectingCountry id="F37D588A985FFF87FDB88B6FFD3BEC1F" box="[581,678,844,869]" name="Malaysia" pageId="13" pageNumber="214">Sarawak</collectingCountry>
</collectingRegion>
Planted Forest Project, Bintulu Division,
<geoCoordinate id="EE5E7EDD985FFF87FB778B6FFB7DEC1E" box="[1162,1248,844,868]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="55" value="3.113">3.113N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FB108B6FFAC6EC1F" box="[1261,1371,844,869]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="55" value="113.662">113.662E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FA948B6FFF4FECF2" pageId="13" pageNumber="214" value="2008-10-20">20 x 2008</date>
, leg. A. Juhin, in collection Dow;
<specimenCount id="9D6CD393985FFF87FD838B53FD33ECFD" box="[638,686,880,904]" pageId="13" pageNumber="214" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FD3E8B4CFD61ECF2" box="[707,764,879,904]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
15_GOM40), Sungai Lamah, mid Baram, Miri Division,
<geoCoordinate id="EE5E7EDD985FFF87FF6A8BB7FE94ECD7" box="[151,265,916,941]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="1" value="3.53978">3.53978N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FEE88BB7FE02ECD7" box="[277,415,916,941]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="114.22956">114.22956E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FE568BB6FD8EECD7" box="[427,531,916,941]" pageId="13" pageNumber="214" value="2015-02-07">7 ii 2015</date>
, leg. L. Southwell, in collection Dow;
<specimenCount id="9D6CD393985FFF87FC378BB7FC68ECD6" box="[970,1013,916,940]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(teneral;
<collectionCode id="ED7B80DF985FFF87FB9F8BB7FB06ECD7" box="[1122,1179,916,941]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
11_12_GOM25), Long Liau/Sungai Dapoi area, ulu Tinjar, Miri Division, (coordinates not available),
<date id="FFD43EDA985FFF87FBDD8B9BFB39ECAA" box="[1056,1188,952,976]" pageId="13" pageNumber="214" value="2011-05-04" valueMax="2011-05-07" valueMin="2011-05-04">4–7 v 2011</date>
, leg. L. Southwell, in collection Dow;
<specimenCount id="9D6CD393985FFF87FEAB8BFFFE1EEC8E" box="[342,387,988,1012]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FE6E8BFFFE51EC8E" box="[403,460,988,1012]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
09_10_GOM37:
<collectionCode id="ED7B80DF985FFF87FD738BFFFD75EC8F" box="[654,744,988,1013]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
.INS.501357), stream in Lio Mato water catchment area, Lio Mato, Upper Baram, Miri Division,
<geoCoordinate id="EE5E7EDD985FFF87FD958BDCFD23EB62" box="[616,702,1023,1048]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="55" value="3.163">3.163N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FD2B8BDCFCD9EB62" box="[726,836,1023,1048]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="55" value="115.235">115.235E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FCA78C23FC78EB62" box="[858,997,1024,1048]" pageId="13" pageNumber="214" value="2009-10-13">13 x 2009</date>
, leg. R.A. Dow, in
<collectionCode id="ED7B80DF985FFF87FB058C23FAD2EB6D" box="[1272,1359,1024,1047]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
;
<specimenCount id="9D6CD393985FFF87FA9B8C23FA06EB6D" box="[1382,1435,1024,1048]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FF628C07FF45EB47" box="[159,216,1060,1085]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="13" pageNumber="214">SAR</collectionCode>
13_14_GOM50), tributary to Sungai Sii downstream from Camp, Upper Baram, Miri Division,
<geoCoordinate id="EE5E7EDD985FFF87FADB8C07FA05EB46" box="[1318,1432,1060,1084]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="1" value="2.99147">2.99147N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FF6A8C64FE83EB1A" box="[151,286,1095,1120]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="114.90256">114.90256E</geoCoordinate>
(Sungai Sii),
<date id="FFD43EDA985FFF87FE428C6BFDD8EB1A" box="[447,581,1095,1120]" pageId="13" pageNumber="214" value="2014-07-17">17 vii 2014</date>
, leg. R.W.J. Ngiam, to be deposited in
<collectionCode id="ED7B80DF985FFF87FBFA8C6BFBDCEB1A" box="[1031,1089,1096,1120]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="214">ZRC</collectionCode>
;
<specimenCount id="9D6CD393985FFF87FBB38C6BFBE7EB25" box="[1102,1146,1096,1120]" pageId="13" pageNumber="214" type="female">1 ♀</specimenCount>
(PomS260-2014, used in illustrations), steep tributary of Deer Cave River,
<collectionCode id="ED7B80DF985FFF87FD248C4FFCB7EBFE" box="[729,810,1132,1156]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13141" name="Gros Morne National Park" pageId="13" pageNumber="214">GMNP</collectionCode>
, Miri Division,
<geoCoordinate id="EE5E7EDD985FFF87FC1B8C4FFBD6EBFE" box="[998,1099,1132,1156]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="5" value="4.0297">4.0297N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FBA48C4FFB49EBFF" box="[1113,1236,1132,1157]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="5" value="114.8355">114.8355E</geoCoordinate>
,
<date id="FFD43EDA985FFF87FB1E8C4FFAF8EBFE" box="[1251,1381,1132,1157]" pageId="13" pageNumber="214" value="2014-04-22">22 iv 2014</date>
, leg. P.O.M. Steinhoff, in collection Steinhoff.
</paragraph>
<paragraph id="8BD5181A985FFF87FF3A8C97FBF2EA26" blockId="13.[151,1437,700,2021]" pageId="13" pageNumber="214">
<emphasis id="B91EC408985FFF87FF3A8C97FE00EBB7" bold="true" box="[199,413,1204,1229]" pageId="13" pageNumber="214">
<collectingRegion id="49AED6F8985FFF87FF3A8C97FE89EBB7" box="[199,276,1204,1229]" country="Malaysia" name="Sabah" pageId="13" pageNumber="214">Sabah</collectingRegion>
,
<collectingCountry id="F37D588A985FFF87FEDA8C97FE05EBB7" box="[295,408,1204,1229]" name="Malaysia" pageId="13" pageNumber="214">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="9D6CD393985FFF87FE518C97FE7DEBB6" box="[428,480,1204,1228]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
, brooklet crossing West Trail, near Sungai Segama, Danum Valley,
<geoCoordinate id="EE5E7EDD985FFF87FADB8C97FA05EBB7" box="[1318,1432,1204,1229]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="1" value="4.91667">4.91667N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FF6A8CF4FE83EB8A" box="[151,286,1239,1264]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="117.66667">117.66667E</geoCoordinate>
(Danum Valley),
<quantity id="4C92B5FF985FFF87FE138CFBFDA4EB8A" box="[494,569,1240,1264]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="214" unit="m" value="170.0">170 m</quantity>
,
<date id="FFD43EDA985FFF87FDBA8CFBFD2FEB8A" box="[583,690,1240,1264]" pageId="13" pageNumber="214" value="1987-05-02">2 v 1987</date>
, leg. J. van Tol, in
<collectionCode id="ED7B80DF985FFF87FC6E8CFAFC76EB8A" box="[915,1003,1241,1264]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
;
<specimenCount id="9D6CD393985FFF87FC048CFBFBB8EB8A" box="[1017,1061,1240,1264]" pageId="13" pageNumber="214" type="female">1 ♀</specimenCount>
, Sungai Palum Tambun, Danum Valley,
<geoCoordinate id="EE5E7EDD985FFF87FF138CDFFEFDEA6F" box="[238,352,1276,1301]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="1" value="4.91667">4.91667N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FE928CDFFE6AEA6F" box="[367,503,1276,1301]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="1" value="117.66667">117.66667E</geoCoordinate>
(Danum Valley),
<quantity id="4C92B5FF985FFF87FD378CDFFCD2EA6E" box="[714,847,1276,1301]" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="13" pageNumber="214" unit="m" value="155.0" valueMax="160.0" valueMin="150.0">150–160 m</quantity>
,
<date id="FFD43EDA985FFF87FCA28CDFFC7CEA6E" box="[863,993,1276,1301]" pageId="13" pageNumber="214" value="1994-04-23">23 iv 1994</date>
, leg. M. Hämäläinen, in
<collectionCode id="ED7B80DF985FFF87FAF58CDEFAFDEA6E" box="[1288,1376,1277,1300]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
;
<specimenCount id="9D6CD393985FFF87FA938CDFFA01EA6E" box="[1390,1436,1276,1300]" pageId="13" pageNumber="214" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED7B80DF985FFF87FF628D03FF4BEA42" box="[159,214,1312,1336]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14076" name="Society of Amateur Botanists" pageId="13" pageNumber="214">SAB</collectionCode>
12_GOM4;
<collectionCode id="ED7B80DF985FFF87FE988D3CFE23EA42" box="[357,446,1311,1336]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
.INS.507752, used in illustrations), Kimamabang waterfall stream, Inobong, Crocker Range National Park,
<geoCoordinate id="EE5E7EDD985FFF87FE738D67FE55EA27" box="[398,456,1348,1373]" direction="north" orientation="latitude" pageId="13" pageNumber="214" precision="5555" value="5.9">5.9N</geoCoordinate>
,
<geoCoordinate id="EE5E7EDD985FFF87FE2E8D67FDA0EA27" box="[467,573,1348,1373]" direction="east" orientation="longitude" pageId="13" pageNumber="214" precision="55" value="116.117">116.117E</geoCoordinate>
(Inobong),
<date id="FFD43EDA985FFF87FD3F8D67FCA0EA26" box="[706,829,1348,1373]" pageId="13" pageNumber="214" value="2012-09-18">18 ix 2012</date>
, leg. R.A. Dow, in
<collectionCode id="ED7B80DF985FFF87FBE98D66FBF1EA26" box="[1044,1132,1349,1372]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="13" pageNumber="214">RMNH</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3704B91985FFF84FF3A8D44FC7FEDB6" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214" type="description">
<paragraph id="8BD5181A985FFF84FF3A8D44FDA0EFAF" blockId="13.[151,1437,700,2021]" lastBlockId="14.[151,1436,151,717]" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
<emphasis id="B91EC408985FFF87FF3A8D44FE5CEAFA" bold="true" box="[199,449,1383,1408]" pageId="13" pageNumber="214">Descriptive remarks.</emphasis>
Morphologically, males of this taxon differ from
<taxonomicName id="4C6A6399985FFF87FC098D4AFBE3EAFA" authorityName="Laidlaw" authorityYear="1936" box="[1012,1150,1385,1408]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FC098D4AFBE3EAFA" box="[1012,1150,1385,1408]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
only in the presence of a complete antehumeral stripe; other characters fall within the range of variation in
<taxonomicName id="4C6A6399985FFF87FBAA8DAEFB79EADE" authorityName="Laidlaw" authorityYear="1936" box="[1111,1252,1421,1444]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FBAA8DAEFB79EADE" box="[1111,1252,1421,1444]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
. The middorsal stripes are narrower than the average seen in
<taxonomicName id="4C6A6399985FFF87FD598D92FCB2EAB2" authorityName="Laidlaw" authorityYear="1936" box="[676,815,1457,1480]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FD598D92FCB2EAB2" box="[676,815,1457,1480]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
, and in the male illustrated in
<figureCitation id="1351049F985FFF87FB6B8D8CFA99EAB2" box="[1174,1284,1455,1480]" captionStart="FIGURES 52–55" captionStartId="31.[151,264,1501,1524]" captionTargetBox="[151,1436,409,1479]" captionTargetId="figure@31.[151,1436,409,1480]" captionTargetPageId="31" captionText="FIGURES 52–55. Male synthorax dorsal: (52) L. coomansi SAR09_10_GOM4; (53) L. coomansi holotype; (54) L. sp. cf coomansi SAB12_GOM4; (55) L. pendleburyi SAR13_14_GOM13." httpUri="https://zenodo.org/record/1068024/files/figure.png" pageId="13" pageNumber="214">Figure 54</figureCitation>
they are just disconnected from the mesothoracic collar, but this is a condition approached in at least one specimen of
<taxonomicName id="4C6A6399985FFF87FA788DF6FE9EE96A" authorityName="Laidlaw" authorityYear="1936" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FA788DF6FE9EE96A" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
examined (see the variation section under that species). In all males examined the arms of the epiproct are at the least curved end of the spectrum seen in
<taxonomicName id="4C6A6399985FFF87FCF78E3EFC05E94E" authorityName="Laidlaw" authorityYear="1936" box="[778,920,1565,1588]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FCF78E3EFC05E94E" box="[778,920,1565,1588]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
. Females are associated with the males by supposition on the basis of their complete antehumeral stripes and the possession of an entirely black clypeus as in the males (but in
<taxonomicName id="4C6A6399985FFF87FE9B8E46FE6FE906" authorityName="Laidlaw" authorityYear="1936" box="[358,498,1637,1660]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FE9B8E46FE6FE906" box="[358,498,1637,1660]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
females only seen in the
<typeStatus id="54D1A6B8985FFF87FCDE8E47FC18E907" box="[803,901,1636,1661]" pageId="13" pageNumber="214" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C6A6399985FFF87FC4C8E46FBBCE906" box="[945,1057,1637,1660]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B91EC408985FFF87FC4C8E46FBBCE906" box="[945,1057,1637,1660]" italics="true" pageId="13" pageNumber="214">L. mariae</emphasis>
</taxonomicName>
and one other specimen). These females also differ in the structures of the dorsum of the head from
<taxonomicName id="4C6A6399985FFF87FC768EAAFB89E9DA" authorityName="Laidlaw" authorityYear="1936" box="[907,1044,1673,1696]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FC768EAAFB89E9DA" box="[907,1044,1673,1696]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
; they also possess pits, but the pits are orientated parallel to the longitudinal axis of the head (more transverse in definite
<taxonomicName id="4C6A6399985FFF87FBAF8E8EFB44E9BE" authorityName="Laidlaw" authorityYear="1936" box="[1106,1241,1709,1732]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FBAF8E8EFB44E9BE" box="[1106,1241,1709,1732]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
) and run right up to rear of the lateral ocelli (separated from the lateral ocelli in definite
<taxonomicName id="4C6A6399985FFF87FC058EF2FB1BE992" authorityName="Laidlaw" authorityYear="1936" box="[1016,1158,1745,1768]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FC058EF2FB1BE992" box="[1016,1158,1745,1768]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
), and the tubercles are correspondingly narrower (
<figureCitation id="1351049F985FFF87FE318ED7FDBFE876" box="[460,546,1780,1805]" captionStart="FIGURES28–31" captionStartId="22.[151,264,1357,1380]" captionTargetBox="[151,1435,266,1336]" captionTargetId="figure@22.[151,1436,265,1336]" captionTargetPageId="22" captionText="FIGURES28–31. Female head dorsaldetail; h—occipital horn; ot—occipital tube; p—pit; s—occipital spur; t—tubercle;thtubercle horn: (28) L. coomansi SAR11_12_GOM48; (29) L. mariae holotype; (30) L. sp. cf coomansi PomS260; (31) L. species SAR11_12_GOM33." httpUri="https://zenodo.org/record/1068014/files/figure.png" pageId="13" pageNumber="214">Fig. 30</figureCitation>
); occipital horns are not present in any of the specimens examined except the female from the Danum Valley, which has one broken occipital horn. The Danum Valley female is also unusual in that there are small areas at the rear of the pits that are raised up, forming additional small tubercles or very small horns; the sternum of S8 was not checked in this specimen. The other three females listed under this taxon are teneral (so reliable measurements cannot be given); the vulvar scale is not suitable for figuring in any, but appears similar to that it
<taxonomicName id="4C6A6399985FFF87FEA88F8AFE42E8BA" authorityName="Laidlaw" authorityYear="1936" box="[341,479,1961,1984]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985FFF87FEA88F8AFE42E8BA" box="[341,479,1961,1984]" italics="true" pageId="13" pageNumber="214">L. coomansi</emphasis>
</taxonomicName>
, but possibly slightly longer. The sternum of S8 is only exposed in the female from Sungai Lamah, which is the most teneral of the three, and severely distorted; the curious tubercle seen on the sternum of S
<quantity id="4C92B5FF985CFF84FED088BBFEC7EFCA" box="[301,346,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="14" pageNumber="215" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="4C6A6399985CFF84FE9988BAFE6DEFCA" authorityName="Laidlaw" authorityYear="1936" box="[356,496,153,176]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FE9988BAFE6DEFCA" box="[356,496,153,176]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
might be present but not fully inflated, however this might be an artefact of the folding and distortion of the segment.
</paragraph>
<paragraph id="8BD5181A985CFF84FF3A88C3FC64EE67" blockId="14.[151,1436,151,717]" pageId="14" pageNumber="215">Measurements (mm). Males: Hw 29.5–31.5, abdomen excluding anal appendages 36–39.5, Ax 16–18 Fw, 11– 12 Hw, Px 10–12 both pairs. Females: teneral and not suitable for measuring.</paragraph>
<paragraph id="8BD5181A985CFF84FF3A8904FA9CEE8F" blockId="14.[151,1436,151,717]" pageId="14" pageNumber="215">
<emphasis id="B91EC408985CFF84FF3A8904FE1CEE3A" bold="true" box="[199,385,295,320]" pageId="14" pageNumber="215">Other remarks.</emphasis>
In the molecular results, this taxon is not clearly distinguished from
<taxonomicName id="4C6A6399985CFF84FB7E890AFA96EE3A" authorityName="Laidlaw" authorityYear="1936" box="[1155,1291,297,320]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FB7E890AFA96EE3A" box="[1155,1291,297,320]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
in COI (
<figureCitation id="1351049F985CFF84FA938904FF3BEE1E" captionStart="FIGURE1" captionStartId="4.[151,250,1913,1935]" captionTargetBox="[168,1423,1134,1886]" captionTargetId="figure@4.[151,1436,1127,1891]" captionTargetPageId="4" captionText="FIGURE1. COI gene tree for 29 specimensof Leptogomphusand three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1067998/files/figure.png" pageId="14" pageNumber="215">Fig. 1</figureCitation>
; possibly this is the result of introgression), but is weakly separated from it in ITS (
<figureCitation id="1351049F985CFF84FB7C896FFB55EE1E" box="[1153,1224,332,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1294,1316]" captionTargetBox="[243,1384,200,1237]" captionTargetId="figure@5.[151,1436,193,1273]" captionTargetPageId="5" captionText="FIGURE 2. ITS gene tree for 20 specimens of Leptogomphus and three outgroup taxon.The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068000/files/figure.png" pageId="14" pageNumber="215">Fig. 2</figureCitation>
); in the combined analysis (
<figureCitation id="1351049F985CFF84FEFE894CFED5EEF2" box="[259,328,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,250,1871,1893]" captionTargetBox="[209,1373,710,1811]" captionTargetId="figure@6.[151,1436,700,1850]" captionTargetPageId="6" captionText="FIGURE 3. Phylogenetic reconstruction for 30 specimens of Leptogomphus and three outgroup taxa using the combined COI+ITS dataset. The best Maximum Likelihood tree is shown, with posterior probabilities from the Bayesian Inference analysis also depicted on the branches. Bootstrap values and posterior probabilities are shown if less than 100 or 1.0 respectively. RMNH collection codes are shown for each specimen, as well as the sex of the specimen and an indication of where it was collected." httpUri="https://zenodo.org/record/1068002/files/figure.png" pageId="14" pageNumber="215">Fig. 3</figureCitation>
) it appears as the close sister to
<taxonomicName id="4C6A6399985CFF84FD3B8952FCCCEEF2" authorityName="Laidlaw" authorityYear="1936" box="[710,849,369,392]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FD3B8952FCCCEEF2" box="[710,849,369,392]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
. Since only two samples of
<emphasis id="B91EC408985CFF84FB618952FB2EEEF2" box="[1180,1203,369,392]" italics="true" pageId="14" pageNumber="215">L.</emphasis>
sp. cf
<taxonomicName id="4C6A6399985CFF84FAF88952FAEDEEF2" authorityName="Laidlaw" authorityYear="1936" box="[1285,1392,369,392]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FAF88952FAEDEEF2" box="[1285,1392,369,392]" italics="true" pageId="14" pageNumber="215">coomansi</emphasis>
</taxonomicName>
are included in these analyses it is not possible to have great confidence that the inclusion of more samples would not result in a blurring of the distinction seen in ITS and the combined analysis presented here. Since none of the supposed females are included in the analyses, no light is thrown on their association with the males.
</paragraph>
<paragraph id="8BD5181A985CFF84FF3A89DCFEFBEDFF" blockId="14.[151,1436,151,717]" pageId="14" pageNumber="215">
The status of this taxon is left open here until more material becomes available; an expanded molecular analysis is needed to resolve the issue of its relation to
<taxonomicName id="4C6A6399985CFF84FCFE8A06FC11ED46" authorityName="Laidlaw" authorityYear="1936" box="[771,908,549,572]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FCFE8A06FC11ED46" box="[771,908,549,572]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
. One or both sexes may ultimately prove to be merely uncommon extremes of variation in
<taxonomicName id="4C6A6399985CFF84FD828A6AFC9AED1A" authorityName="Laidlaw" authorityYear="1936" box="[639,775,585,608]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FD828A6AFC9AED1A" box="[639,775,585,608]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
, but may also prove to be a distinct, but recently separated, biological species.
</paragraph>
<paragraph id="8BD5181A985CFF84FF3A8AACFC7FEDB6" blockId="14.[151,1436,151,717]" pageId="14" pageNumber="215">
This taxon is know from
<collectingRegion id="49AED6F8985CFF84FE168AACFDB2EDD2" box="[491,559,655,680]" country="Malaysia" name="Sabah" pageId="14" pageNumber="215">Sabah</collectingRegion>
and
<collectingRegion id="49AED6F8985CFF84FD948AACFD54EDD2" box="[617,713,655,680]" country="Malaysia" name="Sarawak" pageId="14" pageNumber="215">
<collectingCountry id="F37D588A985CFF84FD948AACFD54EDD2" box="[617,713,655,680]" name="Malaysia" pageId="14" pageNumber="215">Sarawak</collectingCountry>
</collectingRegion>
east of the Lupar River (see
<figureCitation id="1351049F985CFF84FBE68AACFBE3EDD2" box="[1051,1150,655,680]" captionStart="FIGURE 126" captionStartId="48.[151,250,1801,1823]" captionTargetBox="[183,1420,421,1756]" captionTargetId="figure@48.[183,1420,421,1756]" captionTargetPageId="48" captionText="FIGURE 126. Distribution of L. coomansi, confirmed (yellow square) and unconfirmed (blue triangle) records, and L. sp. cf coomansi (black circle)." httpUri="https://zenodo.org/record/1068054/files/figure.png" pageId="14" pageNumber="215">Fig. 126</figureCitation>
). In
<collectingRegion id="49AED6F8985CFF84FB498AACFA89EDD2" box="[1204,1300,655,680]" country="Malaysia" name="Sarawak" pageId="14" pageNumber="215">
<collectingCountry id="F37D588A985CFF84FB498AACFA89EDD2" box="[1204,1300,655,680]" name="Malaysia" pageId="14" pageNumber="215">Sarawak</collectingCountry>
</collectingRegion>
it occurs at some of the same locations as
<taxonomicName id="4C6A6399985CFF84FE168A96FDE9EDB6" authorityName="Laidlaw" authorityYear="1936" box="[491,628,693,716]" class="Insecta" family="Gomphidae" genus="Leptogomphus" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="215" phylum="Arthropoda" rank="species" species="coomansi">
<emphasis id="B91EC408985CFF84FE168A96FDE9EDB6" box="[491,628,693,716]" italics="true" pageId="14" pageNumber="215">L. coomansi</emphasis>
</taxonomicName>
, but appears to be much scarcer.
</paragraph>
</subSubSection>
</treatment>
</document>
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