340 lines
49 KiB
XML
340 lines
49 KiB
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<document id="7447E0CDB25CD0186D261FF13FF9DA25" ID-DOI="10.11646/zootaxa.5323.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="8209415" ID-ZooBank="C022FD72-9C54-4A89-A880-3D99F52C8499" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1691006370328" checkinUser="plazi" docAuthor="Tan, Ming Kai, Salvador, Jewel Anne G., Sabang, Aira Mae M., Bahoy, Daphne Cayle M., Nuñeza, Olga Macas & Robillard, Tony" docDate="2023" docId="03B587EEE750676C0CFEC52F2903F818" docLanguage="en" docName="zootaxa.5323.3.1.pdf" docOrigin="Zootaxa 5323 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5323.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ectatoderus dubius Tan & Salvador & Sabang & Bahoy & Nuñeza & Robillard 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="344" masterDocId="FF8CFF96E77A67470C69C3582A4BFFA1" masterDocTitle="Taxonomy and bioacoustics of little-known Grylloidea crickets (Orthoptera, Ensifera) from Mindanao, Philippines" masterLastPageNumber="348" masterPageNumber="301" pageNumber="343" updateTime="1691007180022" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="AB026776C2B9E4DCD891EBBA3B29EAA2">
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<mods:title id="F884D2FEEF1370EB7BAE39F64645AD0A">Taxonomy and bioacoustics of little-known Grylloidea crickets (Orthoptera, Ensifera) from Mindanao, Philippines</mods:title>
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<mods:namePart id="6A0E2159E18ABD7D62FAA8BE9F5C64BD">Tan, Ming Kai</mods:namePart>
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<mods:affiliation id="067705082C59F87B6CB838AFCA4C3FF7">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore</mods:affiliation>
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<mods:namePart id="DB6787D437C5E37AC10EE2C3D50C3ECD">Salvador, Jewel Anne G.</mods:namePart>
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<mods:affiliation id="06EFF1E6CA9783B2ED659E772F460FA4">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:namePart id="C4109200DA0278BB05F12ED9549DCFF3">Sabang, Aira Mae M.</mods:namePart>
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<mods:affiliation id="C0FEAA378E20E2152819FB2A8B77C165">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:namePart id="E7C06466DD3AF341F0E4C7D868FD0023">Bahoy, Daphne Cayle M.</mods:namePart>
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<mods:affiliation id="DE5D5E5D2E46179A27DC01D7AAA54EE3">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:namePart id="6BECD48CCB468943E13CBC3698B4436B">Nuñeza, Olga Macas</mods:namePart>
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<mods:affiliation id="2747944BF70C4E50454A3CDE9BEA0239">Department of Biological Sciences, College of Science and Mathematics, Mindanao State University-Iligan Institute of Technology, Tibanga, Iligan City 9200, Philippines</mods:affiliation>
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<mods:namePart id="5E98C75A0A25DAFB5611F13190C09BCA">Robillard, Tony</mods:namePart>
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<mods:affiliation id="2B3EB4365D4A9CFC9FEF829AD189E414">Institut de Systématique, Evolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France.</mods:affiliation>
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<mods:typeOfResource id="98911423CCB6B75E60E464D58CB8930D">text</mods:typeOfResource>
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<mods:title id="BB09893534EC0585B1CAEDD56B03D99D">Zootaxa</mods:title>
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<mods:part id="2EFD4C55CC5F117B7E79325238FA0B3D">
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<mods:date id="893934D5B87FAC8FD3AC0F7803EDBC46">2023</mods:date>
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<mods:number id="209107987B84C10B4682814FD2B7AB9E">2023-08-02</mods:number>
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<mods:number id="A71B49F915B915B6935EC369A69928FE">5323</mods:number>
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<mods:number id="509EAD50AC83AAAB42CCBF7A0B84622A">3</mods:number>
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<mods:url id="15DEE4CA455FCB28AC013F175F75D902">http://dx.doi.org/10.11646/zootaxa.5323.3.1</mods:url>
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<mods:classification id="2F5B503F2FE9538706E960B22A99425D">journal article</mods:classification>
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<mods:identifier id="A6534EE455C3AE68AD03FBAB450421D8" type="DOI">10.11646/zootaxa.5323.3.1</mods:identifier>
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<mods:identifier id="643E65E44331CB061A8CEEB002D94169" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6C3053FA201E4EC14CD651EC85382A58" type="Zenodo-Dep">8209415</mods:identifier>
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<mods:identifier id="950053C146A7198E18F413630A34FC25" type="ZooBank">C022FD72-9C54-4A89-A880-3D99F52C8499</mods:identifier>
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</mods:mods>
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<treatment id="03B587EEE750676C0CFEC52F2903F818" LSID="urn:lsid:plazi:treatment:03B587EEE750676C0CFEC52F2903F818" httpUri="http://treatment.plazi.org/id/03B587EEE750676C0CFEC52F2903F818" lastPageId="43" lastPageNumber="344" pageId="42" pageNumber="343">
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<subSubSection id="C3066573E750676D0CFEC52F28F2F930" box="[151,697,1655,1682]" pageId="42" pageNumber="343" type="nomenclature">
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<paragraph id="8BA336F8E750676D0CFEC52F28F2F930" blockId="42.[151,697,1655,1717]" box="[151,697,1655,1682]" pageId="42" pageNumber="343">
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<heading id="D0EB8194E750676D0CFEC52F28F2F930" bold="true" box="[151,697,1655,1682]" fontSize="11" level="1" pageId="42" pageNumber="343" reason="1">
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<emphasis id="B968EAEAE750676D0CFEC52F28F2F930" bold="true" box="[151,697,1655,1682]" pageId="42" pageNumber="343">
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<taxonomicName id="4C1C4D7BE750676D0CFEC52F2BF3F930" authority="Tan" authorityName="Tan & Salvador & Sabang & Bahoy & Nuñeza & Robillard" authorityYear="2023" box="[151,440,1655,1682]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="species" species="dubius">
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<emphasis id="B968EAEAE750676D0CFEC52F2BC9F933" bold="true" box="[151,386,1655,1682]" italics="true" pageId="42" pageNumber="343">Ectatoderus dubius</emphasis>
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Tan
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</taxonomicName>
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&
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<collectingCountry id="F30B7668E750676D0DB6C52F2804F930" box="[479,591,1655,1681]" name="El Salvador" pageId="42" pageNumber="343">Salvador</collectingCountry>
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,
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<taxonomicNameLabel id="A25B5791E750676D0E32C52F28F2F930" box="[603,697,1655,1681]" pageId="42" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3066573E750676D0CFEC5C32B98F8C8" pageId="42" pageNumber="343" type="description">
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<paragraph id="8BA336F8E750676D0CFEC5C32B6CF914" blockId="42.[151,697,1655,1717]" box="[151,295,1691,1717]" pageId="42" pageNumber="343">
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(
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<figureCitation id="13272A7DE750676D0CF6C5C32B54F914" box="[159,287,1691,1717]" captionStart-0="FIGURE 37" captionStart-1="FIGURE 38" captionStart-2="FIGURE 39" captionStartId-0="42.[151,250,1549,1573]" captionStartId-1="44.[151,250,1943,1967]" captionStartId-2="45.[151,250,1344,1368]" captionTargetBox-0="[151,1436,669,1526]" captionTargetBox-1="[183,1407,183,1918]" captionTargetBox-2="[159,1426,191,1304]" captionTargetId-0="figure-214@42.[151,1436,669,1526]" captionTargetId-1="figure-17@44.[179,1407,181,1919]" captionTargetId-2="figure-17@45.[151,1436,181,1320]" captionTargetPageId-0="42" captionTargetPageId-1="44" captionTargetPageId-2="45" captionText-0="FIGURE 37. Ectatoderus dubius sp. nov. ♁ in its natural environment, a mossy forest near Mt Sumagaya." captionText-1="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." captionText-2="FIGURE 39. Ectatoderus dubius sp. nov. ♁ calling song: oscillograms showing one echeme (A); spectrogram of the same echeme (B); power spectra of a syllable (C) where the grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." figureDoi-0="http://doi.org/10.5281/zenodo.8209507" figureDoi-1="http://doi.org/10.5281/zenodo.8209511" figureDoi-2="http://doi.org/10.5281/zenodo.8209513" httpUri-0="https://zenodo.org/record/8209507/files/figure.png" httpUri-1="https://zenodo.org/record/8209511/files/figure.png" httpUri-2="https://zenodo.org/record/8209513/files/figure.png" pageId="42" pageNumber="343">Figs 37–39</figureCitation>
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)
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</paragraph>
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<paragraph id="8BA336F8E750676D0CFEC5BB2B98F8C8" blockId="42.[151,1437,1763,2042]" pageId="42" pageNumber="343">
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<emphasis id="B968EAEAE750676D0CFEC5BB2BD7F95C" bold="true" box="[151,412,1763,1789]" pageId="42" pageNumber="343">Specimens examined.</emphasis>
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<materialsCitation id="3B743CA5E750676D0DC0C5BB2B98F8C8" collectingDate="2023-02-16" collectionCode="PNM" collectorName="M. K. Tan & Tan, J. A. G & Salvador, A. M & Sabang, A & Gono, D. C. M & K. Grumo" country="Philippines" county="Mindanao" elevation="7" latitude="8.70132" location="Barangay Lunotan" longLatPrecision="1" longitude="125.0098" municipality="Gingoog City" pageId="42" pageNumber="343" specimenCount="1" stateProvince="Misamis Oriental" typeStatus="holotype">
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<emphasis id="B968EAEAE750676D0DC0C5BB2854F95C" bold="true" box="[425,543,1763,1789]" pageId="42" pageNumber="343">
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<typeStatus id="54A7885AE750676D0DC0C5BB2851F95C" box="[425,538,1763,1789]" pageId="42" pageNumber="343">Holotype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="F30B7668E750676D0E45C5BB289AF95C" box="[556,721,1763,1789]" name="Philippines" pageId="42" pageNumber="343">PHILIPPINES</collectingCountry>
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• ♁;
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<collectingCounty id="62C24E74E750676D0F77C5BB29D9F95C" box="[798,914,1763,1789]" pageId="42" pageNumber="343">Mindanao</collectingCounty>
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,
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<collectingRegion id="49D8F81AE750676D0FCAC5BB2E24F95C" box="[931,1135,1763,1789]" country="Philippines" name="Misamis Oriental" pageId="42" pageNumber="343">Misamis Oriental</collectingRegion>
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,
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<collectingMunicipality id="6BC7AC82E750676D08E9C5BB2F6BF95C" box="[1152,1312,1763,1789]" pageId="42" pageNumber="343">Gingoog City</collectingMunicipality>
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,
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<location id="8EC36023E750676D0958C5BC2ABCF883" LSID="urn:lsid:plazi:treatment:03B587EEE750676C0CFEC52F2903F818:8EC36023E750676D0958C5BC2ABCF883" country="Philippines" county="Mindanao" latitude="8.70132" longLatPrecision="1" longitude="125.0098" municipality="Gingoog City" name="Barangay Lunotan" pageId="42" pageNumber="343" stateProvince="Misamis Oriental">Barangay Lunotan</location>
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, near
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<location id="8EC36023E750676D0D53C4502B92F880" LSID="urn:lsid:plazi:treatment:03B587EEE750676C0CFEC52F2903F818:8EC36023E750676D0D53C4502B92F880" box="[314,473,1799,1826]" country="Philippines" county="Mindanao" latitude="8.70132" longLatPrecision="1" longitude="125.0098" municipality="Gingoog City" name="Mt Sumagaya" pageId="42" pageNumber="343" stateProvince="Misamis Oriental">Mt Sumagaya</location>
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;
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<geoCoordinate id="EE28503FE750676D0D8CC45F281CF880" box="[485,599,1799,1825]" degrees="8.70132" direction="north" orientation="latitude" pageId="42" pageNumber="343" precision="1" value="8.70132">N8.70132</geoCoordinate>
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,
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<geoCoordinate id="EE28503FE750676D0E0BC45F28A6F883" box="[610,749,1799,1826]" degrees="125.00980" direction="east" orientation="longitude" pageId="42" pageNumber="343" precision="1" value="125.0098">E125.00980</geoCoordinate>
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, 1197.0±
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<quantity id="4CE49B1DE750676D0F3DC45F29D1F880" box="[852,922,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="7.3" pageId="42" pageNumber="343" unit="m" value="7.3">
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<elevation id="0031D1CBE750676D0F3DC45F29D1F880" box="[852,922,1799,1825]" metricMagnitude="0" metricUnit="m" metricValue="7.3" pageId="42" pageNumber="343" unit="m" value="7.3">7.3 m</elevation>
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</quantity>
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.a.s.l.;
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<date id="FFA21038E750676D0FB4C45F2EECF880" box="[989,1191,1799,1825]" pageId="42" pageNumber="343" value="2023-02-16">
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<collectingDate id="EFE6E9D0E750676D0FB4C45F2EECF880" box="[989,1191,1799,1825]" pageId="42" pageNumber="343" value="2023-02-16">16 February 2023</collectingDate>
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</date>
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, 19h44; mossy forest, on a leaf near the ground; coll.
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<collectorName id="26E9532EE750676D0E69C4742824F8E7" box="[512,623,1836,1862]" pageId="42" pageNumber="343">M.K. Tan</collectorName>
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, J.A.G.
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<collectorName id="26E9532EE750676D0EA7C473293DF8E7" box="[718,886,1835,1862]" pageId="42" pageNumber="343">Salvador, A.M</collectorName>
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.
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<collectorName id="26E9532EE750676D0FEBC47329B2F8E7" box="[898,1017,1835,1862]" pageId="42" pageNumber="343">Sabang, A</collectorName>
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.
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<collectorName id="26E9532EE750676D086FC4732EEFF8E4" box="[1030,1188,1835,1861]" pageId="42" pageNumber="343">Gono, D.C.M</collectorName>
|
|||
|
|
. Bahoy &
|
|||
|
|
<collectorName id="26E9532EE750676D0948C4742FD3F8E4" box="[1313,1432,1835,1862]" pageId="42" pageNumber="343">K. Grumo</collectorName>
|
|||
|
|
; MIN.23.66 (
|
|||
|
|
<collectionCode id="ED0DAE3DE750676D0D4BC4172B29F8C8" box="[290,354,1871,1897]" country="Philippines" httpUri="http://grbio.org/cool/mxaf-c5e2" name="Philippine National Museum" pageId="42" pageNumber="343">PNM</collectionCode>
|
|||
|
|
) (
|
|||
|
|
<figureCitation id="13272A7DE750676D0D10C4172B80F8C8" box="[377,459,1871,1897]" captionStart="FIGURE 37" captionStartId="42.[151,250,1549,1573]" captionTargetBox="[151,1436,669,1526]" captionTargetId="figure-214@42.[151,1436,669,1526]" captionTargetPageId="42" captionText="FIGURE 37. Ectatoderus dubius sp. nov. ♁ in its natural environment, a mossy forest near Mt Sumagaya." figureDoi="http://doi.org/10.5281/zenodo.8209507" httpUri="https://zenodo.org/record/8209507/files/figure.png" pageId="42" pageNumber="343">Fig. 37</figureCitation>
|
|||
|
|
)
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E750676C0CAEC42B282AFEE0" lastPageId="43" lastPageNumber="344" pageId="42" pageNumber="343" type="discussion">
|
|||
|
|
<paragraph id="8BA336F8E750676C0CAEC42B282AFEE0" blockId="42.[151,1437,1763,2042]" lastBlockId="43.[151,1437,151,1977]" lastPageId="43" lastPageNumber="344" pageId="42" pageNumber="343">
|
|||
|
|
<emphasis id="B968EAEAE750676D0CAEC42B2B70F82C" bold="true" box="[199,315,1907,1933]" pageId="42" pageNumber="343">Remarks.</emphasis>
|
|||
|
|
The generic status of this species is very peculiar and this species could potentially be placed under either
|
|||
|
|
<taxonomicName id="4C1C4D7BE750676D0CB6C4CF2B0CF810" box="[223,327,1943,1969]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE750676D0CB6C4CF2B0CF810" box="[223,327,1943,1969]" italics="true" pageId="42" pageNumber="343">Ornebius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
or
|
|||
|
|
<taxonomicName id="4C1C4D7BE750676D0D07C4CF2BBEF810" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[366,501,1943,1969]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE750676D0D07C4CF2BBEF810" box="[366,501,1943,1969]" italics="true" pageId="42" pageNumber="343">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The male
|
|||
|
|
<typeStatus id="54A7885AE750676D0E1FC4CF2893F810" box="[630,728,1943,1969]" pageId="42" pageNumber="343">holotype</typeStatus>
|
|||
|
|
has the TI with a small round inner tympanum and the frontal rostrum about as wide as the scapes; all of which are characteristics of
|
|||
|
|
<taxonomicName id="4C1C4D7BE750676D0FD9C4E32E53F874" box="[944,1048,1979,2005]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE750676D0FD9C4E32E53F874" box="[944,1048,1979,2005]" italics="true" pageId="42" pageNumber="343">Ornebius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C1C4D7BE750676D0826C4E32E9DF874" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[1103,1238,1979,2005]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE750676D0826C4E32E9DF874" box="[1103,1238,1979,2005]" italics="true" pageId="42" pageNumber="343">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The pronotum is strongly produced backwards, leaving only the apex of the tegmen free, which is characteristics of
|
|||
|
|
<taxonomicName id="4C1C4D7BE750676D088AC4872F21F858" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[1251,1386,2015,2041]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="343" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE750676D088AC4872F21F858" box="[1251,1386,2015,2041]" italics="true" pageId="42" pageNumber="343">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
; but the male genitalia are more characteristics of an
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0EAAC3CF29C3FF10" box="[707,904,151,177]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0EAAC3CF2960FF10" box="[707,811,151,177]" italics="true" pageId="43" pageNumber="344">Ornebius</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
, particularly of species from the species group
|
|||
|
|
<emphasis id="B968EAEAE751676C0CFEC3E32B47FF74" box="[151,268,187,213]" italics="true" pageId="43" pageNumber="344">kanetataki</emphasis>
|
|||
|
|
(Matsummura, 1904). Moreover,
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0EE2C3E32959FF74" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[651,786,187,213]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0EE2C3E32959FF74" box="[651,786,187,213]" italics="true" pageId="43" pageNumber="344">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has not been recorded in the
|
|||
|
|
<collectingCountry id="F30B7668E751676C0835C3E32E95FF74" box="[1116,1246,187,213]" name="Philippines" pageId="43" pageNumber="344">Philippines</collectingCountry>
|
|||
|
|
. However, based on the generic diagnosis of
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0DACC3872943FF58" authority="(Ingrisch, 2006)" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[453,776,223,249]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0DACC3872807FF58" box="[453,588,223,249]" italics="true" pageId="43" pageNumber="344">Ectatoderus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF8D4B09E751676C0E33C387294BFF58" author="Ingrisch, S." box="[602,768,223,249]" pageId="43" pageNumber="344" pagination="133 - 227" refId="ref17607" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, we tentatively place this species under
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C08D6C3872F0DFF58" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[1215,1350,223,249]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C08D6C3872F0DFF58" box="[1215,1350,223,249]" italics="true" pageId="43" pageNumber="344">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Future phylogenetic work may reveal that this species may belong to another genus (noting that it is likely that
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C097CC25B2FD7FEBC" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[1301,1436,259,285]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C097CC25B2FD7FEBC" box="[1301,1436,259,285]" italics="true" pageId="43" pageNumber="344">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is not monophyletic, see
|
|||
|
|
<bibRefCitation id="EF8D4B09E751676C0DC7C27F281EFEE0" author="Ingrisch, S." box="[430,597,295,321]" pageId="43" pageNumber="344" pagination="133 - 227" refId="ref17607" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC21229CBFD6C" pageId="43" pageNumber="344" type="diagnosis">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC2122BE8FE70" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC2122B09FEC5" bold="true" box="[199,322,330,356]" pageId="43" pageNumber="344">Diagnosis.</emphasis>
|
|||
|
|
From the
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0DA8C2132803FEC4" baseAuthorityName="Ingrisch" baseAuthorityYear="2006" box="[449,584,331,357]" class="Insecta" family="Mogoplistidae" genus="Ectatoderus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0DA8C2132803FEC4" box="[449,584,331,357]" italics="true" pageId="43" pageNumber="344">Ectatoderus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from Southeast Asia, this new species is characterised by the epiphallus membranous with its lateral valves forming separate lobes; these lobes are weakly sclerotized, with their apices rounded and more sclerotized; the medial valve more strongly sclerotized, curved at the base forming a semi-circle, having its apex pointed.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC28329CBFD6C" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
From
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0D67C2832B3DFE54" box="[270,374,475,501]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0D67C2832B3DFE54" box="[270,374,475,501]" italics="true" pageId="43" pageNumber="344">Ornebius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, this species is characterised by the pronotum very strongly produced posteriorly, covering almost entire FWs. The new species also shares similarities in the pronotum with
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0822C2A72A92FD9C" authority="(Bolivar, 1889)" baseAuthorityName="Bolivar" baseAuthorityYear="1889" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="species" species="cucullatus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0822C2A72F64FDB8" box="[1099,1327,511,537]" italics="true" pageId="43" pageNumber="344">Ornebius cucullatus</emphasis>
|
|||
|
|
(Bolívar, 1889)
|
|||
|
|
</taxonomicName>
|
|||
|
|
from Mayon Volcano (in
|
|||
|
|
<collectingRegion id="49D8F81AE751676C0D93C17B2808FD9C" box="[506,579,547,573]" country="Philippines" name="Albay" pageId="43" pageNumber="344">Albay</collectingRegion>
|
|||
|
|
), in the pronotum extending more posteriorly and nearly surpassing the base of the FIII; but differs by the FWs not extending well beyond pronotum and by the mirror covered by the pronotal dorsal disk, and by the pronotal dorsal disk covering almost entirely of the abdomen. It also shares similarities in the pronotum with
|
|||
|
|
<taxonomicName id="4C1C4D7BE751676C0D05C1D728B6FD08" authority="Ingrisch, 2006" authorityName="Ingrisch" authorityYear="2006" box="[364,765,655,681]" class="Insecta" family="Mogoplistidae" genus="Ornebius" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="344" phylum="Arthropoda" rank="species" species="consternus">
|
|||
|
|
<emphasis id="B968EAEAE751676C0D05C1D7281FFD09" box="[364,596,655,681]" italics="true" pageId="43" pageNumber="344">Ornebius consternus</emphasis>
|
|||
|
|
<bibRefCitation id="EF8D4B09E751676C0E32C1D728B6FD08" author="Ingrisch, S." box="[603,765,655,681]" pageId="43" pageNumber="344" pagination="133 - 227" refId="ref17607" refString="Ingrisch, S. (2006) New taxa and notes on some previously described species of scaly crickets from South East Asia (Orthoptera, Grylloidea, Mogoplistidae, Mogoplistinae). Revue Suisse de Zoologie, 113 (1), 133 - 227. https: // doi. org / 10.5962 / bhl. part. 80345" type="journal article" year="2006">Ingrisch, 2006</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from
|
|||
|
|
<collectingRegion id="49D8F81AE751676C0F28C1D729E2FD08" box="[833,937,655,681]" country="Indonesia" name="Sulawesi Utara" pageId="43" pageNumber="344">Sulawesi</collectingRegion>
|
|||
|
|
, but differs by the male genitalia without the lateral valves sclerotized, and by not forming a tube near the apex.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC18F29C3FD50" box="[199,904,727,753]" pageId="43" pageNumber="344" type="etymology">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC18F29C3FD50" blockId="43.[151,1437,151,1977]" box="[199,904,727,753]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC18F2B05FD50" bold="true" box="[199,334,727,753]" pageId="43" pageNumber="344">Etymology.</emphasis>
|
|||
|
|
The species name refers to peculiar generic status.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC1A22B8EF938" pageId="43" pageNumber="344" type="description">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC1A22AA9FB64" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC1A22B12FCB5" bold="true" box="[199,345,762,788]" pageId="43" pageNumber="344">Description.</emphasis>
|
|||
|
|
Small species for the genus; habitus of male as shown in
|
|||
|
|
<figureCitation id="13272A7DE751676C0FB2C1A32E7BFCB4" box="[987,1072,763,789]" captionStart="FIGURE 37" captionStartId="42.[151,250,1549,1573]" captionTargetBox="[151,1436,669,1526]" captionTargetId="figure-214@42.[151,1436,669,1526]" captionTargetPageId="42" captionText="FIGURE 37. Ectatoderus dubius sp. nov. ♁ in its natural environment, a mossy forest near Mt Sumagaya." figureDoi="http://doi.org/10.5281/zenodo.8209507" httpUri="https://zenodo.org/record/8209507/files/figure.png" pageId="43" pageNumber="344">Figs 37</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13272A7DE751676C0852C1A32E25FCB4" box="[1083,1134,763,789]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">38A</figureCitation>
|
|||
|
|
. Dorsum of head flattened, covered with dark-grey scales (
|
|||
|
|
<figureCitation id="13272A7DE751676C0E69C0472827FC98" box="[512,620,799,825]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38A</figureCitation>
|
|||
|
|
). Frontal rostrum 1.5 times wider than scapes (
|
|||
|
|
<figureCitation id="13272A7DE751676C08E4C0472EB2FC98" box="[1165,1273,799,825]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38A</figureCitation>
|
|||
|
|
). scapes dark-grey (
|
|||
|
|
<figureCitation id="13272A7DE751676C0CB0C01B2B0FFCFC" box="[217,324,835,861]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38A</figureCitation>
|
|||
|
|
); basal antennal segments dark-coloured, thereafter brown with some segments black (
|
|||
|
|
<figureCitation id="13272A7DE751676C0972C01B2FDBFCFC" box="[1307,1424,835,861]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Figs. 38A</figureCitation>
|
|||
|
|
). Frons and mouthparts of red brown (
|
|||
|
|
<figureCitation id="13272A7DE751676C0E5CC03F28D6FC20" box="[565,669,871,897]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38B</figureCitation>
|
|||
|
|
). Maxillary palps dark-grey to black; with apical (= fifth), subapical (= fourth) and third segments of subequal lengths; apical segment pyriform (
|
|||
|
|
<figureCitation id="13272A7DE751676C0866C0D32E32FC04" box="[1039,1145,907,933]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38B</figureCitation>
|
|||
|
|
). Head with lateral parts, including genae, of dark colour, without bands (
|
|||
|
|
<figureCitation id="13272A7DE751676C0EACC0F72964FC68" box="[709,815,943,969]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38B</figureCitation>
|
|||
|
|
). Pronotal disk brown and wholly covered with dark-grey scales, about 1.7 times longer than wide, with anterior margin narrow and straight; lateral margin widening posteriorly; covering entire mirror of FW; posterior margin convex with white scales (
|
|||
|
|
<figureCitation id="13272A7DE751676C081AC0AF2E95FBB0" box="[1139,1246,1015,1041]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38A</figureCitation>
|
|||
|
|
). Pronotum with lateral lobe also wholly covered with dark-grey scales (
|
|||
|
|
<figureCitation id="13272A7DE751676C0E9CC7432911FB94" box="[757,858,1051,1077]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38B</figureCitation>
|
|||
|
|
). TI with internal tympanum small and oval; without external tympanum. FIII 1.2 times as long as TIII; TIII 3.3 times as long as MTaIII (
|
|||
|
|
<figureCitation id="13272A7DE751676C0821C7672EE5FBF8" box="[1096,1198,1087,1113]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38C</figureCitation>
|
|||
|
|
). Fs and Ts generally pale brown, covered with marmoration of dark-grey scales except in basal parts. Tarsi cream-coloured, with apical parts of segments brown. Abdominal tergites and sternites covered with dark-grey scales. Cercus covered with grey scales.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC79728E0F98C" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC7972B40FB48" bold="true" box="[199,267,1231,1257]" pageId="43" pageNumber="344">Male.</emphasis>
|
|||
|
|
FW mostly infumated yellow brown, dorsal field darker at posterior margin (
|
|||
|
|
<figureCitation id="13272A7DE751676C0800C7972E9AFB48" box="[1129,1233,1231,1257]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38A</figureCitation>
|
|||
|
|
); lateral field with dorsal half infumated yellow brown and ventral half infumated black and ventral margin white (
|
|||
|
|
<figureCitation id="13272A7DE751676C08BFC7AB2F75FAAC" box="[1238,1342,1267,1293]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38B</figureCitation>
|
|||
|
|
). Supra-anal plate with last abdominal tergite and epiproct distinctly separated by a transverse suture. Last abdominal tergite transverse. Supra-anal plate less transverse than last abdominal tergite, stout tongue-shaped with apex truncated and straight (
|
|||
|
|
<figureCitation id="13272A7DE751676C0C94C6072B2CFAD8" box="[253,359,1375,1401]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Fig. 38D</figureCitation>
|
|||
|
|
). Subgenital plate black, 1.3 times as wide as long, anterior margin concave, posterior margin obtuse with a few long setae. Paraproct process yellow brown, elongate, cylindrical, curved dorsad and tapering into a subacute apex (
|
|||
|
|
<figureCitation id="13272A7DE751676C0DE0C6FF287BFA60" box="[393,560,1447,1473]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Figs 38D, 38E</figureCitation>
|
|||
|
|
). Genitalia as in
|
|||
|
|
<figureCitation id="13272A7DE751676C0E85C6FF293FFA60" box="[748,884,1447,1473]" captionStart="FIGURE 38" captionStartId="44.[151,250,1943,1967]" captionTargetBox="[183,1407,183,1918]" captionTargetId="figure-17@44.[179,1407,181,1919]" captionTargetPageId="44" captionText="FIGURE 38. Morphology and ♁ genitalia of Ectatoderus dubius sp. nov.: habitus (without posterior legs) in dorsal view (A); head, pronotum and FWs in lateral view (B); posterior leg in external view (C); abdominal apex in dorsal (D) and dorso-lateral (E) views; phallus complex in dorsal (F), ventral (G) and lateral (H) views. Scale bars: 2 mm (A, C), 1 mm (B, D–H)." figureDoi="http://doi.org/10.5281/zenodo.8209511" httpUri="https://zenodo.org/record/8209511/files/figure.png" pageId="43" pageNumber="344">Figs 38F–H</figureCitation>
|
|||
|
|
. Epiphallus (ep) membranous. Lateral valves (lv) forming separate lobes; lobe weakly sclerotized, apex rounded and more sclerotized. Medial valve (mv) strongly sclerotized, curved at base forming a semi-circle; apex pointed, not reaching apex of lateral valve. Ejaculatory duct (ed) surpassing medial valve and lateral valves.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC56F2BE9F9F0" blockId="43.[151,1437,151,1977]" box="[199,418,1591,1617]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC56F2B6EF9F0" bold="true" box="[199,293,1591,1617]" pageId="43" pageNumber="344">Female.</emphasis>
|
|||
|
|
Unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC5032B8EF938" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC5032BA8F9D5" bold="true" box="[199,483,1626,1653]" pageId="43" pageNumber="344">Measurements (in mm).</emphasis>
|
|||
|
|
BL = 8.2; HL = 1.0; PronL = 5.0; PronW = 3.0; FWL = 3.4; FWW = 3.0; FIIIL = 5.0; TIIIL = 4.2; MtaIIIL = 1.3.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC5FB295FF91C" box="[199,788,1699,1725]" pageId="43" pageNumber="344" type="biology_ecology">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC5FB295FF91C" blockId="43.[151,1437,151,1977]" box="[199,788,1699,1725]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC5FB2B67F91C" bold="true" box="[199,300,1699,1725]" pageId="43" pageNumber="344">Ecology.</emphasis>
|
|||
|
|
The species was found in the mossy forest.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC59F2F7BF940" box="[199,1328,1734,1761]" pageId="43" pageNumber="344" type="materials_examined">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC59F2F7BF940" blockId="43.[151,1437,151,1977]" box="[199,1328,1734,1761]" pageId="43" pageNumber="344">
|
|||
|
|
<materialsCitation id="3B743CA5E751676C0CAEC59F2F7BF940" box="[199,1328,1734,1761]" country="Philippines" county="Mindanao Island" location="Barangay Lunotan" municipality="Gingoog City" pageId="43" pageNumber="344" specimenCount="1" stateProvince="Misamis Oriental" typeStatus="holotype">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC59F2B2DF941" bold="true" box="[199,358,1734,1761]" pageId="43" pageNumber="344">
|
|||
|
|
<typeStatus id="54A7885AE751676C0CAEC59F2B49F940" box="[199,258,1735,1761]" pageId="43" pageNumber="344">Type</typeStatus>
|
|||
|
|
locality.
|
|||
|
|
</emphasis>
|
|||
|
|
<collectingCountry id="F30B7668E751676C0D05C59F285FF940" box="[364,532,1735,1761]" name="Philippines" pageId="43" pageNumber="344">PHILIPPINES</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCounty id="62C24E74E751676C0E76C59F2894F940" box="[543,735,1735,1761]" pageId="43" pageNumber="344">Mindanao Island</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingRegion id="49D8F81AE751676C0E83C59F29FAF940" box="[746,945,1735,1761]" country="Philippines" name="Misamis Oriental" pageId="43" pageNumber="344">Misamis Oriental</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="6BC7AC82E751676C0FD2C59F2E1EF940" box="[955,1109,1735,1761]" pageId="43" pageNumber="344">Gingoog City</collectingMunicipality>
|
|||
|
|
,
|
|||
|
|
<location id="8EC36023E751676C0808C59F2F7BF940" LSID="urn:lsid:plazi:treatment:03B587EEE750676C0CFEC52F2903F818:8EC36023E751676C0808C59F2F7BF940" box="[1121,1328,1735,1761]" country="Philippines" county="Mindanao Island" municipality="Gingoog City" name="Barangay Lunotan" pageId="43" pageNumber="344" stateProvince="Misamis Oriental">Barangay Lunotan</location>
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3066573E751676C0CAEC5B22903F818" pageId="43" pageNumber="344" type="distribution">
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC5B22E39F8A4" blockId="43.[151,1437,151,1977]" box="[199,1138,1770,1797]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC5B22B2AF8A5" bold="true" box="[199,353,1770,1796]" pageId="43" pageNumber="344">Distribution.</emphasis>
|
|||
|
|
<collectingCountry id="F30B7668E751676C0D0EC5B32847F8A4" box="[359,524,1771,1797]" name="Philippines" pageId="43" pageNumber="344">PHILIPPINES</collectingCountry>
|
|||
|
|
(Mindanao Island: Gingoog City in
|
|||
|
|
<collectingRegion id="49D8F81AE751676C0FCCC5B32E20F8A4" box="[933,1131,1771,1797]" country="Philippines" name="Misamis Oriental" pageId="43" pageNumber="344">Misamis Oriental</collectingRegion>
|
|||
|
|
)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA336F8E751676C0CAEC4562903F818" blockId="43.[151,1437,151,1977]" pageId="43" pageNumber="344">
|
|||
|
|
<emphasis id="B968EAEAE751676C0CAEC4562822F889" bold="true" box="[199,617,1806,1833]" pageId="43" pageNumber="344">
|
|||
|
|
Calling song (1J, 23.8°C) (
|
|||
|
|
<figureCitation id="13272A7DE751676C0E6FC4562817F889" box="[518,604,1806,1832]" captionStart="FIGURE 39" captionStartId="45.[151,250,1344,1368]" captionTargetBox="[159,1426,191,1304]" captionTargetId="figure-17@45.[151,1436,181,1320]" captionTargetPageId="45" captionText="FIGURE 39. Ectatoderus dubius sp. nov. ♁ calling song: oscillograms showing one echeme (A); spectrogram of the same echeme (B); power spectra of a syllable (C) where the grey line represents the power [dB] (axis name and scale in grey) and the black solid peak represents the relative amplitude (axis name in black)." figureDoi="http://doi.org/10.5281/zenodo.8209513" httpUri="https://zenodo.org/record/8209513/files/figure.png" pageId="43" pageNumber="344">Fig. 39</figureCitation>
|
|||
|
|
).
|
|||
|
|
</emphasis>
|
|||
|
|
The calling song is made up of an echeme consisting of 33 syllables and has a duration of 7.55 s. The first few syllables tend to be of lower amplitudes; amplitude of syllable gradually increases to a maximum and stabilises. The syllable duration is 36.1 ms and average interval between consecutive syllables is 204.8 ms. The frequency spectrum is pure-tonal and forms a harmonic, with the energy peaking at the fundamental frequency. The dominant frequency is 5.72 kHz.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|