699 lines
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699 lines
74 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1121.85693" ID-Pensoft-Pub="1313-2970-1121-97" ID-Pensoft-UUID="A978BBB62CF35971A2A2F65B6CD35868" ID-ZooBank="016D0A914E4A4EF5A41B7DAD0A10B408" ModsDocID="1313-2970-1121-97" checkinTime="1663323279549" checkinUser="pensoft" docAuthor="Conradie, Werner, Keates, Chad, Baptista, Ninda L. & Lobon-Rovira, Javier" docDate="2022" docId="47DE685535F0518C8E1AC4092C68D514" docLanguage="en" docName="ZooKeys 1121: 97-143" docOrigin="ZooKeys 1121" docPubDate="2022-09-15" docSource="http://dx.doi.org/10.3897/zookeys.1121.85693" docTitle="Prosymna angolensis Boulenger 1915" docType="treatment" docVersion="2" id="A978BBB62CF35971A2A2F65B6CD35868" lastPageNumber="97" masterDocId="A978BBB62CF35971A2A2F65B6CD35868" masterDocTitle="Taxonomical review of Prosymna angolensis Boulenger, 1915 (Elapoidea, Prosymnidae) with the description of two new species" masterLastPageNumber="143" masterPageNumber="97" pageNumber="97" updateTime="1663323582101" updateUser="ExternalLinkService">
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<mods:title>Taxonomical review of Prosymna angolensis Boulenger, 1915 (Elapoidea, Prosymnidae) with the description of two new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Conradie, Werner</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0805-9683</mods:nameIdentifier>
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<mods:affiliation>Port Elizabeth Museum (Bayworld), P. O. Box 13147, Humewood 6013, Gqeberha, South Africa & Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa & National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">werner@bayworld.co.za</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Keates, Chad</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3125-9536</mods:nameIdentifier>
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<mods:affiliation>Department of Nature Conservation Management, Natural Resource Science and Management Cluster, Faculty of Science, George Campus, Nelson Mandela University, George, South Africa & National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa & South African Institute for Aquatic Biodiversity (SAIAB), Makhanda, South Africa & Department of Zoology and Entomology, Rhodes University, Makhanda, South Africa</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Baptista, Ninda L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2859-6606</mods:nameIdentifier>
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<mods:affiliation>National Geographic Okavango Wilderness Project, Wild Bird Trust, Sandton, South Africa & Instituto Superior de Ciencias da Educacao da Huila (ISCED-Huila), Rua Sarmento Rodrigues, Lubango, Angola & CIBIO-InBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, Laboratorio Associado, Universidade do Porto, Campus Agrario de Vairao, Rua Padre Armando Quintas, 4485 - 661 Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lobon-Rovira, Javier</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4380-9427</mods:nameIdentifier>
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<mods:affiliation>CIBIO-InBIO, Centro de Investigacao em Biodiversidade e Recursos Geneticos, Laboratorio Associado, Universidade do Porto, Campus Agrario de Vairao, Rua Padre Armando Quintas, 4485 - 661 Vairao, Portugal & Departamento de Biologia, Faculdade de Ciencias, Universidade do Porto, 4099 - 002 Porto, Portugal & BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairao, 4485 - 661 Vairao, Portugal</mods:affiliation>
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<mods:date>2022</mods:date>
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<mods:number>2022-09-15</mods:number>
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<mods:detail type="volume">
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<mods:number>1121</mods:number>
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<mods:start>97</mods:start>
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<mods:end>143</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1121.85693</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1121.85693</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1121-97</mods:identifier>
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<mods:identifier type="ZooBank">016D0A914E4A4EF5A41B7DAD0A10B408</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A978BBB62CF35971A2A2F65B6CD35868</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514" httpUri="http://treatment.plazi.org/id/47DE685535F0518C8E1AC4092C68D514" lastPageNumber="97" pageId="0" pageNumber="97">
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<subSubSection pageId="0" pageNumber="97" type="nomenclature">
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<paragraph pageId="0" pageNumber="97">
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<taxonomicName LSID="47DE6855-35F0-518C-8E1A-C4092C68D514" authority="Boulenger, 1915" authorityName="Boulenger" authorityYear="1915" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna angolensis" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="angolensis">Prosymna angolensis Boulenger, 1915</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="97" type="description">
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<paragraph pageId="0" pageNumber="97">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in dorsal colouration of nominotypical Prosymna angolensis A Tundavala, Huila Province, Angola (Photo: Justin R. Nicolau) B Grootfontein, Namibia (Photo: Francois Theart) C Bicuar National Park (CHL 0521), Huila Province, Angola (Photo: Ninda L. Baptista)." figureDoi="10.3897/zookeys.1121.85693.figure4" httpUri="https://binary.pensoft.net/fig/744035" pageId="0" pageNumber="97">Figs 4</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Lateral B ventral C medial D dorsal views of skulls of Prosymna angolensis subgroup. Red arrows depict variation characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure7" httpUri="https://binary.pensoft.net/fig/744038" pageId="0" pageNumber="97">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A Frontal and B posterior views of skulls, and C lateral, dorsal, medial and ventral view of jaw of Prosymna angolensis subgroup. Red arrows depict variation in characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure8" httpUri="https://binary.pensoft.net/fig/744039" pageId="0" pageNumber="97">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. A Dorsal, frontal and ventral view of premaxillae B medial, posterior and dorsal views of palatine; and C lateral, dorsal, and ventral view of maxilla in P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis and P. janii (from left to right)." figureDoi="10.3897/zookeys.1121.85693.figure9" httpUri="https://binary.pensoft.net/fig/744040" pageId="0" pageNumber="97">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neotype of Prosymna angolensis (NMW 19275: 2) from Caconda, Huila Province, Angola (Photos: Alice Schumacher, Natural History Museum Vienna)." figureDoi="10.3897/zookeys.1121.85693.figure10" httpUri="https://binary.pensoft.net/fig/744041" pageId="0" pageNumber="97">
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, 10 Common names: Angolan Shovel-snout snake (English);
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<normalizedToken originalValue="Cobra-de-focinho-de-pá-de-Angola">Cobra-de-focinho-de-pa-de-Angola</normalizedToken>
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(Portuguese).
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="97" type="chresonymy">
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<paragraph pageId="0" pageNumber="97">Chresonymy.1</paragraph>
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<paragraph pageId="0" pageNumber="97">
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<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1867" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna frontalis" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="frontalis">
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<emphasis italics="true" pageId="0" pageNumber="97">Prosymna frontalis</emphasis>
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</taxonomicName>
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:
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<bibRefCitation author="Bocage, JVB" journalOrPublisher="Jornal de Sciencias Mathematicas, Physicas e Naturaes" pageId="0" pageNumber="97" pagination="209 - 227" refId="B5" refString="Bocage, JVB, 1873. Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale. Jornal de Sciencias Mathematicas, Physicas e Naturaes 4: 209 - 227" title="Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale." volume="4" year="1873">Bocage 1873</bibRefCitation>
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: 218,
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<bibRefCitation author="Bocage, JVB" journalOrPublisher="Jornal de Sciencias Mathematicas, Physicas e Naturaes" pageId="0" pageNumber="97" refId="B6" refString="Bocage, JVB, 1882. Noticia acerca de alguns reptis d'Angoche que existem no Museu Nacional de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes. Academia Real das Sciencias de Lisboa VIII(32): 286-290." title="Noticia acerca de alguns reptis d'Angoche que existem no Museu Nacional de Lisboa. Jornal de Sciencias Mathematicas, Physicas e Naturaes. Academia Real das Sciencias de Lisboa VIII (32): 286 - 290." year="1882">1882</bibRefCitation>
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: 288,
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<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">1895</bibRefCitation>
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: 98;
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<bibRefCitation author="Boulenger, GA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="97" refId="B10" refString="Boulenger, GA, 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II. Containing the conclusion of the Colubriae Aglyphae. British Museum (Natural History), London, [xi,] 382 pp." title="Catalogue of the snakes in the British Museum (Natural History). Volume II. Containing the conclusion of the Colubriae Aglyphae. British Museum (Natural History), London, [xi,] 382 pp." year="1894">Boulenger 1894</bibRefCitation>
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: 248,
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<bibRefCitation author="Boulenger, GA" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="97" refId="B11" refString="Boulenger, GA, 1896. Catalogue of the snakes in the British Museum (Natural History). Volume III. Containing the Families Colubridae (Opisthoglyphae and Proteroglyphae), Amblycephalidae, and Viperidae. British Museum (Natural History), London, [xiv,] 727 pp." title="Catalogue of the snakes in the British Museum (Natural History). Volume III. Containing the Families Colubridae (Opisthoglyphae and Proteroglyphae), Amblycephalidae, and Viperidae. British Museum (Natural History), London, [xiv,] 727 pp." year="1896">1896</bibRefCitation>
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: 641.
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</paragraph>
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<paragraph pageId="0" pageNumber="97">
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<taxonomicName authorityName="Bocage" authorityYear="1873" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna ambigua" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="ambigua">
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<emphasis italics="true" pageId="0" pageNumber="97">Prosymna ambigua</emphasis>
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</taxonomicName>
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:
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<bibRefCitation author="Monard, A" journalOrPublisher="Bulletin de la Societe Neuchateloise des sciences Naturelles" pageId="0" pageNumber="97" pagination="89 - 111" refId="B78" refString="Monard, A, 1931. Mission scientifique Suisse dans l'Angola. Resultats scientifiques. Reptiles. Bulletin de la Societe Neuchateloise des sciences Naturelles 33: 89 - 111" title="Mission scientifique Suisse dans l'Angola. Resultats scientifiques. Reptiles." volume="33" year="1931">Monard 1931</bibRefCitation>
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: 104,
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<bibRefCitation author="Monard, A" journalOrPublisher="Arquivos do Museu Bocage" pageId="0" pageNumber="97" pagination="19 - 154" refId="B79" refString="Monard, A, 1937. Contribution a l'Herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154" title="Contribution a l'Herpetologie d'Angola." volume="8" year="1937">1937</bibRefCitation>
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: 123;
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<bibRefCitation author="Mertens, R" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="97" pagination="425 - 443" refId="B72" refString="Mertens, R, 1937. Reptilien und Amphibien aus dem suedlichen Inner-Afrika. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 20: 425 - 443" title="Reptilien und Amphibien aus dem suedlichen Inner-Afrika." volume="20" year="1937">Mertens 1937</bibRefCitation>
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: 13.
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</paragraph>
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<paragraph pageId="0" pageNumber="97">
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<taxonomicName authority="ambigua" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna ambigua subsp. ambigua" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="subSpecies" species="ambigua" subSpecies="ambigua">
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<emphasis italics="true" pageId="0" pageNumber="97">Prosymna ambigua ambigua</emphasis>
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</taxonomicName>
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:
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<bibRefCitation author="Mertens, R" journalOrPublisher="Senckenbergiana" pageId="0" pageNumber="97" pagination="425 - 443" refId="B73" refString="Mertens, R, 1938. Amphibien und Reptilien aus Angola gesmmelt von W. Shack. Senckenbergiana 20: 425 - 443" title="Amphibien und Reptilien aus Angola gesmmelt von W. Shack." volume="20" year="1938">Mertens 1938</bibRefCitation>
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: 439;
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<bibRefCitation author="Loveridge, A" journalOrPublisher="Bulletin of Comparative Zoology" pageId="0" pageNumber="97" pagination="1 - 198" refId="B64" refString="Loveridge, A, 1958. Revision of five African snake genera. Bulletin of Comparative Zoology 1191: 1 - 198" title="Revision of five African snake genera." volume="1191" year="1958">Loveridge 1958</bibRefCitation>
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: 151.
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</paragraph>
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<paragraph pageId="0" pageNumber="97">
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<taxonomicName authorityName="Boulenger" authorityYear="1915" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna angolensis" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="angolensis">
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<emphasis italics="true" pageId="0" pageNumber="97">Prosymna angolensis</emphasis>
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</taxonomicName>
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:
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<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="97" pagination="193 - 223" refId="B12" refString="Boulenger, GA, 1915. A list of the snakes of the Belgian and Portuguese Congo, northern Rhodesia, and Angola. Proceedings of the Zoological Society of London 1915 (2): 193 - 223, DOI: https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" title="A list of the snakes of the Belgian and Portuguese Congo, northern Rhodesia, and Angola." url="https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" volume="1915" year="1915">Boulenger 1915</bibRefCitation>
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: 208;
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<bibRefCitation author="Chabanaud, P" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle" pageId="0" pageNumber="97" pagination="433 - 440" refId="B31" refString="Chabanaud, P, 1916. Revision du genre Prosymna Gray. Bulletin du Museum National d'Histoire Naturelle 1916: 433 - 440" title="Revision du genre Prosymna Gray." volume="1916" year="1916">Chabanaud 1916</bibRefCitation>
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: 439;
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<bibRefCitation author="Monard, A" journalOrPublisher="Arquivos do Museu Bocage" pageId="0" pageNumber="97" pagination="19 - 154" refId="B79" refString="Monard, A, 1937. Contribution a l'Herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154" title="Contribution a l'Herpetologie d'Angola." volume="8" year="1937">Monard 1937</bibRefCitation>
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: 114, 122;
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<bibRefCitation author="Bogert, CM" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="97" pagination="1 - 107" refId="B8" refString="Bogert, CM, 1940. Herpetological Results of the Vernay Angola expedition. Bulletin of the American Museum of Natural History 77: 1 - 107" title="Herpetological Results of the Vernay Angola expedition." volume="77" year="1940">Bogert 1940</bibRefCitation>
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: 59;
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<bibRefCitation author="Mertens, R" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="97" pagination="1 - 172" refId="B74" refString="Mertens, R, 1955. Die Amphibien und Reptilien Suedwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgefuehrten Reise. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 490: 1 - 172" title="Die Amphibien und Reptilien Suedwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgefuehrten Reise." volume="490" year="1955">Mertens 1955</bibRefCitation>
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: 94,
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<bibRefCitation author="Mertens, R" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="97" pagination="1 - 110" refId="B75" refString="Mertens, R, 1971. Die Herpetofauna Suedwest-Afrikas. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 529: 1 - 110" title="Die Herpetofauna Suedwest-Afrikas." volume="529" year="1971">1971</bibRefCitation>
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: 86;
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<bibRefCitation author="Hellmich, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="97" pagination="39 - 80" refId="B53" refString="Hellmich, W, 1957. Die reptilienausbeute der Hamburgischen Angola Expedition. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 55: 39 - 80" title="Die reptilienausbeute der Hamburgischen Angola Expedition." volume="55" year="1957">Hellmich 1957</bibRefCitation>
|
|||
|
|
: 66;
|
|||
|
|
<bibRefCitation author="Loveridge, A" journalOrPublisher="Bulletin of Comparative Zoology" pageId="0" pageNumber="97" pagination="1 - 198" refId="B64" refString="Loveridge, A, 1958. Revision of five African snake genera. Bulletin of Comparative Zoology 1191: 1 - 198" title="Revision of five African snake genera." volume="1191" year="1958">Loveridge 1958</bibRefCitation>
|
|||
|
|
: 149;
|
|||
|
|
<bibRefCitation author="FitzSimons, VFM" journalOrPublisher="Purnell and Sons, Cape Town, South Africa" pageId="0" pageNumber="97" refId="B45" refString="FitzSimons, VFM, 1962. Snakes of southern Africa. Purnell and Sons, Cape Town, South Africa" title="Snakes of southern Africa." year="1962">FitzSimons 1962</bibRefCitation>
|
|||
|
|
: 161,
|
|||
|
|
<bibRefCitation author="FitzSimons, VFM" journalOrPublisher="Annals of the Transvaal Museum" pageId="0" pageNumber="97" pagination="25 - 79" refId="B46" refString="FitzSimons, VFM, 1966. A check-list, with synoptic keys, to the snakes of southern Africa. Annals of the Transvaal Museum 25: 25 - 79" title="A check-list, with synoptic keys, to the snakes of southern Africa." volume="25" year="1966">1966</bibRefCitation>
|
|||
|
|
: 53,
|
|||
|
|
<bibRefCitation author="FitzSimons, VFM" journalOrPublisher="Collins, London" pageId="0" pageNumber="97" refId="B47" refString="FitzSimons, VFM, 1970. A field guide to the snakes of southern Africa. Collins, London" title="A field guide to the snakes of southern Africa." year="1970">1970</bibRefCitation>
|
|||
|
|
: 104;
|
|||
|
|
<bibRefCitation author="Isemonger, RM" journalOrPublisher="Books of Africa, Cape Town" pageId="0" pageNumber="97" refId="B56" refString="Isemonger, RM, 1968. Snakes of Africa. 1st Edn. Books of Africa, Cape Town" title="Snakes of Africa. 1 st Edn." year="1968">Isemonger 1968</bibRefCitation>
|
|||
|
|
: 129;
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
: 512,
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Donker Publishers, Parklands" pageId="0" pageNumber="97" refId="B20" refString="Broadley, DG, 1990. FitzSimons' Snakes of southern Africa. Revised Edition. Jonathan Ball and Ad. Donker Publishers, Parklands" title="FitzSimons' Snakes of southern Africa. Revised Edition. Jonathan Ball and Ad." year="1990">1990</bibRefCitation>
|
|||
|
|
: 227,
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Donker Publishers, Parklands" pageId="0" pageNumber="97" refId="B21" refString="Broadley, DG, 1995. Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." title="Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." year="1995">1995</bibRefCitation>
|
|||
|
|
: 48;
|
|||
|
|
<bibRefCitation author="Auerbach, RD" journalOrPublisher="Mokwepa Consultants, Gaborone, Botswana" pageId="0" pageNumber="97" refId="B1" refString="Auerbach, RD, 1987. The Amphibians and Reptiles of Botswana. Mokwepa Consultants, Gaborone, Botswana" title="The Amphibians and Reptiles of Botswana." year="1987">Auerbach 1987</bibRefCitation>
|
|||
|
|
: 178;
|
|||
|
|
<bibRefCitation author="Branch, WR" journalOrPublisher="Struik Publishers, Cape Town" pageId="0" pageNumber="97" refId="B13" refString="Branch, WR, 1998. Field Guide to the Snakes and other Reptiles of Southern Africa. Revised Edition. Struik Publishers, Cape Town" title="Field Guide to the Snakes and other Reptiles of Southern Africa. Revised Edition." year="1998">Branch 1998</bibRefCitation>
|
|||
|
|
: 84,
|
|||
|
|
<bibRefCitation author="Branch, WR" journalOrPublisher="Struik Publishers, Cape Town" pageId="0" pageNumber="97" refId="B14" refString="Branch, WR, 2018. Snakes of Angola: An annotated checklist. Amphibian & Reptile Conservation 12(2)[General Section]: 41-82[e159]." title="Snakes of Angola: An annotated checklist. Amphibian & Reptile Conservation 12 (2) [General Section]: 41 - 82 [e 159]." year="2018">2018</bibRefCitation>
|
|||
|
|
: 64;
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Edition Chimaira, Frankfurt" pageId="0" pageNumber="97" refId="B23" refString="Broadley, DG, Doria, CT, Wigge, J, 2003. Snakes of Zambia. An Atlas and Field Guide. Edition Chimaira, Frankfurt" title="Snakes of Zambia. An Atlas and Field Guide." year="2003">Broadley et al. 2003</bibRefCitation>
|
|||
|
|
: 187 (in part);
|
|||
|
|
<bibRefCitation author="Marais, J" journalOrPublisher="Struik Publishers, Cape Town" pageId="0" pageNumber="97" refId="B65" refString="Marais, J, 2004. A Complete Guide to the Snakes of Southern Africa, 2nd edn. Struik Publishers, Cape Town" title="A Complete Guide to the Snakes of Southern Africa, 2 nd edn." year="2004">Marais 2004</bibRefCitation>
|
|||
|
|
: 236;
|
|||
|
|
<bibRefCitation author="Broadley, D" journalOrPublisher="Chimaira, Frankfurt" pageId="0" pageNumber="97" refId="B22" refString="Broadley, D, Blaylock, R, 2013. The Snakes of Zimbabwe and Botswana. Chimaira, Frankfurt" title="The Snakes of Zimbabwe and Botswana." year="2013">Broadley and Blaylock 2013</bibRefCitation>
|
|||
|
|
: 219;
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.1016/j.jaridenv.2012.05.003" author="Herrmann, H-W" journalOrPublisher="Journal of Arid Environments" pageId="0" pageNumber="97" pagination="94 - 115" refId="B54" refString="Herrmann, H-W, Branch, WR, 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94 - 115, DOI: https://doi.org/10.1016/j.jaridenv.2012.05.003" title="Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist." url="https://doi.org/10.1016/j.jaridenv.2012.05.003" volume="93" year="2013">Herrmann and Branch 2013</bibRefCitation>
|
|||
|
|
: 8;
|
|||
|
|
<bibRefCitation author="Wallach, V" journalOrPublisher="[type catalogue] CRC Press, Florida" pageId="0" pageNumber="97" refId="B104" refString="Wallach, V, Williams, KL, Boundy, J, 2014. Snakes of the World: A catalogue of living and extinct species. [type catalogue] CRC Press, Florida" title="Snakes of the World: A catalogue of living and extinct species." year="2014">Wallach et al. 2014</bibRefCitation>
|
|||
|
|
: 568;
|
|||
|
|
<bibRefCitation author="Baptista, NL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="97" refId="B3" refString="Baptista, NL, Antonio, T, Branch, WR, 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2)[Special Section]: 96-130[e203]." title="The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13 (2) [Special Section]: 96 - 130 [e 203]." year="2019">Baptista et al. 2019</bibRefCitation>
|
|||
|
|
: 118;
|
|||
|
|
<bibRefCitation author="Chippaux, J-P" journalOrPublisher="Johns Hopkins University Press, Baltimore" pageId="0" pageNumber="97" refId="B33" refString="Chippaux, J-P, Jackson, K, 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, Baltimore" title="Snakes of Central and Western Africa." year="2019">Chippaux and Jackson 2019</bibRefCitation>
|
|||
|
|
: 283,
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1052.64607" author="Ceriaco, LMP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="97" pagination="85 - 156" refId="B29" refString="Ceriaco, LMP, Parrinha, D, Marques, MP, 2021a. Saving collections: Taxonomic revision of the herpetological collection of the Instituto de Investigacao Cientifica Tropical, Lisbon (Portugal) with a protocol to rescue abandoned collections. ZooKeys 1052: 85 - 156, DOI: https://doi.org/10.3897/zookeys.1052.64607" title="Saving collections: Taxonomic revision of the herpetological collection of the Instituto de Investigacao Cientifica Tropical, Lisbon (Portugal) with a protocol to rescue abandoned collections." url="https://doi.org/10.3897/zookeys.1052.64607" volume="1052" year="2021 a">
|
|||
|
|
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
|||
|
|
et al. 2021a
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 16 (in part).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
When
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" author="Boulenger, GA" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="97" pagination="193 - 223" refId="B12" refString="Boulenger, GA, 1915. A list of the snakes of the Belgian and Portuguese Congo, northern Rhodesia, and Angola. Proceedings of the Zoological Society of London 1915 (2): 193 - 223, DOI: https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" title="A list of the snakes of the Belgian and Portuguese Congo, northern Rhodesia, and Angola." url="https://doi.org/10.1111/j.1469-7998.1915.tb07411.x" volume="1915" year="1915">Boulenger (1915)</bibRefCitation>
|
|||
|
|
described
|
|||
|
|
<taxonomicName lsidName="P. angolensis" pageId="0" pageNumber="97" rank="species" species="angolensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">P. angolensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, he did not designate a precise type locality nor a type specimen for that matter. Later,
|
|||
|
|
<bibRefCitation author="Loveridge, A" journalOrPublisher="Bulletin of Comparative Zoology" pageId="0" pageNumber="97" pagination="1 - 198" refId="B64" refString="Loveridge, A, 1958. Revision of five African snake genera. Bulletin of Comparative Zoology 1191: 1 - 198" title="Revision of five African snake genera." volume="1191" year="1958">Loveridge (1958)</bibRefCitation>
|
|||
|
|
proposed designating
|
|||
|
|
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
|||
|
|
as the type locality, but it was
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley (1980)</bibRefCitation>
|
|||
|
|
that finally restricted the type locality to Caconda by designating a lectotype of the material he examined on his visit to Museu Bocage, Lisbon (MBL), Portugal in 1968. The reasons for this change in the proposed type locality, were that the
|
|||
|
|
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
|||
|
|
specimen was unaccounted for, as well as the Caconda material was in the best overall condition to represent the species. He initially designated MBL 1606b as the lectotype, but with the destruction of the MBL collection, he designated one of the remaining Caconda specimens in Naturhistorischen Museums in Wien (NMW 19275b) as the replacement neotype (see
|
|||
|
|
<bibRefCitation author="Gemel, R" journalOrPublisher="Annalen des Naturhistorischen Museums in Wien, Serie B" pageId="0" pageNumber="97" pagination="33 - 248" refId="B48" refString="Gemel, R, Gassner, G, Schweiger, S, 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien-2018. Annalen des Naturhistorischen Museums in Wien, Serie B 121: 33 - 248" title="Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien- 2018." volume="121" year="2019">Gemel et al. 2019</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="97" type="materials_examined">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">Material examined.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" elevation="1662" latitude="-13.73537" location="Huila Province" longLatPrecision="1" longitude="15.0672" specimenCount="1" stateProvince="Huila Province" typeStatus="Neotype">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">
|
|||
|
|
<typeStatus>Neotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
(Fig.
|
|||
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Neotype of Prosymna angolensis (NMW 19275: 2) from Caconda, Huila Province, Angola (Photos: Alice Schumacher, Natural History Museum Vienna)." figureDoi="10.3897/zookeys.1121.85693.figure10" httpUri="https://binary.pensoft.net/fig/744041" pageId="0" pageNumber="97">10</figureCitation>
|
|||
|
|
).
|
|||
|
|
</emphasis>
|
|||
|
|
NHMW 19275:2, collected from Caconda (approx.
|
|||
|
|
<geoCoordinate degrees="13.73537" direction="south" orientation="latitude" precision="1" value="-13.73537">-13.73537</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="15.06720" direction="east" orientation="longitude" precision="1" value="15.0672">15.06720</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.6619999999999997" unit="m" value="1662.0">
|
|||
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6619999999999997" unit="m" value="1662.0">1662 m</quantity>
|
|||
|
|
a.s.l.
|
|||
|
|
</elevation>
|
|||
|
|
),
|
|||
|
|
<collectingRegion country="Angola" name="Huila">
|
|||
|
|
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
|||
|
|
Province
|
|||
|
|
</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<typeStatus>Neotype</typeStatus>
|
|||
|
|
designated by Donald
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley (1980)</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Angola" specimenCount="1" typeStatus="Additional material">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">
|
|||
|
|
<typeStatus isOtherTypeStatus="true">Additional material</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
MBL 1609,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
(no precise locality)
|
|||
|
|
</materialsCitation>
|
|||
|
|
,
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" isTailEnumeration="true" location="Angola" specimenCount="1">
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Bibala" specimenCount="1">
|
|||
|
|
MBL 1605a,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:295CD804B6DE1E2D1211712520B55703" country="Angola" name="Bibala">Bibala</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Bibala" specimenCount="1">
|
|||
|
|
MBL 1605b,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:58D9B6519E196B8010BB83D5CE856DB0" country="Angola" name="Bibala">Bibala</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Bicuar" country="Angola" location="Angola" specimenCount="1">
|
|||
|
|
CHL 0521,
|
|||
|
|
<collectorName>Bicuar</collectorName>
|
|||
|
|
NP,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation accessionNumber="NMW 19275" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Caconda" specimenCount="1">
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NMW19275">NMW 19275</accessionNumber>
|
|||
|
|
:1,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:895C3CA6211DAB987DFC19772C230A3D" country="Angola" name="Caconda">Caconda</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation accessionNumber="NMW 19275" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Caconda" specimenCount="1">
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NMW19275">NMW 19275</accessionNumber>
|
|||
|
|
:2,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:465B911F72A57B105C447828BF75D08F" country="Angola" name="Caconda">Caconda</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Caconda" specimenCount="1">
|
|||
|
|
MBL 1606a,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:8363120212D670DC0F6F9C41325CFB75" country="Angola" name="Caconda">Caconda</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Caconda" specimenCount="1" typeStatus="lectotype">
|
|||
|
|
MBL 1606b (original
|
|||
|
|
<typeStatus>lectotype</typeStatus>
|
|||
|
|
),
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:FE4C03EC484068BFAFF3BC5DD5696693" country="Angola" name="Caconda">Caconda</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Caconda" specimenCount="1">
|
|||
|
|
MBL 1606c,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:C393CC65814D94A5EB2E47337D13C890" country="Angola" name="Caconda">Caconda</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
; MBL 1608, Caconda,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Luanda" specimenCount="1" stateProvince="Luanda">
|
|||
|
|
USBN 20035,
|
|||
|
|
<collectingRegion country="Angola" name="Luanda">Luanda</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation accessionNumber="CAS 84181" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Luanda" specimenCount="1" stateProvince="Luanda">
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/CAS84181">CAS 84181</accessionNumber>
|
|||
|
|
,
|
|||
|
|
<collectingRegion country="Angola" name="Luanda">Luanda</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.828032" unit="mi" value="3.0">3 mile</quantity>
|
|||
|
|
S of airport,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Maconjo" country="Angola" location="Maconge" specimenCount="1">
|
|||
|
|
MBL 1607,
|
|||
|
|
<collectorName>Maconjo</collectorName>
|
|||
|
|
=
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:760D99648E383E210AE5B849F5F04CDF" country="Angola" name="Maconge">Maconge</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation accessionNumber="MCZ 32468" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Missao do Dondi Bela Vista" specimenCount="1">
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MCZ32468">MCZ 32468</accessionNumber>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:B28B5EF501E528A50F9A8F33D3761CBC" country="Angola" name="Missao do Dondi Bela Vista">
|
|||
|
|
<normalizedToken originalValue="Missão">Missao</normalizedToken>
|
|||
|
|
do Dondi Bela Vista
|
|||
|
|
</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Angola" location="Mossamedes" specimenCount="1">
|
|||
|
|
MBL 1604, interior of
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:FD9F55AEEBEFF6624836C3DFDC727B7F" country="Angola" name="Mossamedes">Mossamedes</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
; UM 20178, Goeverega,
|
|||
|
|
<collectingCountry name="Botswana">Botswana</collectingCountry>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Namibia" location="Katima Mulilo" specimenCount="1">
|
|||
|
|
UM 21271, 15 km WSW of
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:DD470192FCEE1C89644627975371DE26" country="Namibia" name="Katima Mulilo">Katima Mulilo</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation accessionNumber="SMF 46614" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Karakuwisa" country="Namibia" location="Kavango" specimenCount="1" stateProvince="Kavango">
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF46614">SMF 46614</accessionNumber>
|
|||
|
|
,
|
|||
|
|
<collectorName>Karakuwisa</collectorName>
|
|||
|
|
,
|
|||
|
|
<collectingRegion country="Namibia" name="Kavango West">Kavango</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Namibia" location="Katima Mulilo" specimenCount="1">
|
|||
|
|
TM 55043,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:FFBEC0E68C3F632731F2606605AC6A11" country="Namibia" name="Katima Mulilo">Katima Mulilo</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Namibia" location="Katima Mulilo" specimenCount="1">
|
|||
|
|
UM 24204,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:18DE4EF4C79E4F63C51BF061CB339E5F" country="Namibia" name="Katima Mulilo">Katima Mulilo</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Namibia" location="Namutoni" specimenCount="1">
|
|||
|
|
SAM ZR16574,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:9A475FEBF3D2CE31D89AE770985916B0" country="Namibia" name="Namutoni">Namutoni</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Namibia" location="Waterberg" specimenCount="1">
|
|||
|
|
NMZB 9532, NE of
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:525947544D0960937CCA8CACA6943F12" country="Namibia" name="Waterberg">Waterberg</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Namibia">Namibia</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Inyokene" country="Zimbabwe" location="Nyamandhlovo" specimenCount="1">
|
|||
|
|
NMZB 13953,
|
|||
|
|
<collectorName>Inyokene</collectorName>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:47ED4427D77927AF3BD0F9132B243F54" country="Zimbabwe" name="Nyamandhlovo">Nyamandhlovo</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Zimbabwe">Zimbabwe</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Malinbdi Siding" country="Zimbabwe" location="Hwange" specimenCount="1">
|
|||
|
|
NMZB 13787,
|
|||
|
|
<collectorName>Malinbdi Siding</collectorName>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:70C7B945DDFD28F50F37F32CFE4F5779" country="Zimbabwe" name="Hwange">Hwange</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Zimbabwe">Zimbabwe</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
;
|
|||
|
|
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="Malinbdi Siding" country="Zimbabwe" location="Hwange" specimenCount="1">
|
|||
|
|
NMZB 13788,
|
|||
|
|
<collectorName>Malinbdi Siding</collectorName>
|
|||
|
|
,
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:47DE685535F0518C8E1AC4092C68D514:D4ED8CC7203AC565D8949383232114A1" country="Zimbabwe" name="Hwange">Hwange</location>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Zimbabwe">Zimbabwe</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption doi="10.3897/zookeys.1121.85693.figure10" httpUri="https://binary.pensoft.net/fig/744041" pageId="0" pageNumber="97" start="Figure 10" startId="F10">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">Figure 10.</emphasis>
|
|||
|
|
<typeStatus>Neotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName authorityName="Boulenger" authorityYear="1915" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna angolensis" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="angolensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Prosymna angolensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/NMW19275">NMW 19275</accessionNumber>
|
|||
|
|
:2) from Caconda,
|
|||
|
|
<collectingRegion country="Angola" name="Huila">
|
|||
|
|
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
|||
|
|
Province
|
|||
|
|
</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry name="Angola">Angola</collectingCountry>
|
|||
|
|
(Photos: Alice Schumacher, Natural History Museum Vienna).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="97" type="description">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">General description.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
See Table
|
|||
|
|
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Summary of morphological features and measurements for Prosymna angolensis group. For abbreviations see Materials and methods. Notes: * including data on the Ebanga (Monard 1937) and Capelongo (Bogert 1940) specimens." httpUri="http://table.plazi.org/id/CE661A2298586FEF7FBADB3D50769868" pageId="0" pageNumber="97" tableUuid="CE661A2298586FEF7FBADB3D50769868">2</tableCitation>
|
|||
|
|
for summarised meristic data. Dorsal scales smooth, arranged in 17-15-15 (rarely 19-15-15) rows at midbody, scale row reduction takes place between ventral scales 16-20 (males) and 14-49 (females); one (sometimes two or three on supracaudal scales) apical pits; 126-163 (126-155 males, 134-163 females) smooth ventral scales; 16-28 (22-28 males, 16-25 females) paired subcaudal scales; rostral is acutely angular horizontally; internasal is single and bandlike; 1 preocular; 1 (rarely 0 or 2) postocular; temporals mostly 1+2 (rarely 2+2, 2+3); mostly 6 supralabials, with 3rd and 4th entering the orbit (rarely 5 (2, 3) or 7 (3, 4)); 7 infralabials, with first 3 in contact with the chin shield (rarely 8 (3)), cloacal scale entire.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Skull osteology and teeth</emphasis>
|
|||
|
|
(Figs
|
|||
|
|
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Lateral B ventral C medial D dorsal views of skulls of Prosymna angolensis subgroup. Red arrows depict variation characters between the P. lisima sp. nov., P. angolensis, P. confusa sp. nov., P. cf. frontalis, and P. janii." figureDoi="10.3897/zookeys.1121.85693.figure7" httpUri="https://binary.pensoft.net/fig/744038" pageId="0" pageNumber="97">7</figureCitation>
|
|||
|
|
</emphasis>
|
|||
|
|
-
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">9).</emphasis>
|
|||
|
|
Based on the examination of a single female specimen (SAM ZR16574, Namutoni, Namibia),
|
|||
|
|
<taxonomicName lsidName="P. angolensis" pageId="0" pageNumber="97" rank="species" species="angolensis">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">P. angolensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
presents a compact and rigid skull, which is common among
|
|||
|
|
<taxonomicName authorityName="Gray" authorityYear="1849" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Prosymna</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species. It has an unfused braincase and nasal bones. Parietals are fused and the fronto-lateral portion presents a sharp edge that forms the edge of the orbital rim. Postorbital bone is absent. Premaxilla has a reduced ascending nasal process with a small groove between the ascending process and frontal portion of the bone. Premaxilla lies between the ventral laminae of nasals with a high profile of the anterior portion which curves shapely to finish on a convex profile. Maxilla and premaxilla are in contact. Nasal bones are reduced and display a wing-shape with a narrower anterior portion. Septomaxilla is a well-developed bone, in broad contact with the premaxilla, frontal, vomer, prefrontal and frontal bones. Vomer well developed with perforated dorsolateral portion of the bone. Maxilla reduced anteriorly with an elongated pick-shaped palatine process, with seven or eight laterally reduced curved tooth loci, followed by four enlarged and lancet-shaped tooth loci. Palatine with three reduced teeth and an enlarged dorsal and curved vomerine process that reinforces the internal portion of the orbit. Pterygoid is a thin elongated bone. Supratemporal is an enlarged bone in broad contact with the quadrate and participates in the lateral movement of the lower jaw. The lower jaw consists of compound, splenial, coronoid, and dentary bones. Coronoid and splenial bones are reduced, almost vestigial. Dentary with eight or nine small sharp tooth loci, with first third clear of any teeth.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Colouration in life</emphasis>
|
|||
|
|
(Fig.
|
|||
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Variation in dorsal colouration of nominotypical Prosymna angolensis A Tundavala, Huila Province, Angola (Photo: Justin R. Nicolau) B Grootfontein, Namibia (Photo: Francois Theart) C Bicuar National Park (CHL 0521), Huila Province, Angola (Photo: Ninda L. Baptista)." figureDoi="10.3897/zookeys.1121.85693.figure4" httpUri="https://binary.pensoft.net/fig/744035" pageId="0" pageNumber="97">4</figureCitation>
|
|||
|
|
).
|
|||
|
|
</emphasis>
|
|||
|
|
The head is yellowish-brown with variable darker black markings that can be absent. Most commonly there is an anterior black band across the frontal, followed by a pair of black blotches around the orbits, supraoculars and parietals. A distinct black nuchal spot or collar is often present. The dorsal colouration varies from having mostly small paired black longitudinal vertebral spots (similar to
|
|||
|
|
<taxonomicName lsidName="P. sundevalli" pageId="0" pageNumber="97" rank="species" species="sundevalli">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">P. sundevalli</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) to a speckled pattern (similar to
|
|||
|
|
<taxonomicName lsidName="P. lineata" pageId="0" pageNumber="97" rank="species" species="lineata">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">P. lineata</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) on a pale yellowish-brown to grey ground colour. Ventrum and outermost two or three dorsal scale rows yellowish white.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<taxonomicName genus="Hemipenis" lsidName="Hemipenis" pageId="0" pageNumber="97" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Hemipenis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Short hemipenis with a length that reaches the 9-10th ventral scales (
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Size</emphasis>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
Males vary from 160-248 (208.7
|
|||
|
|
<normalizedToken originalValue="±">+/-</normalizedToken>
|
|||
|
|
29.8) mm SVL and 22-30 (26.5
|
|||
|
|
<normalizedToken originalValue="±">+/-</normalizedToken>
|
|||
|
|
2.7) mm TL, with the largest male measuring 248+28 = 276 mm (NMZB 9532, NE of Waterberg, Namibia). Females vary from 127-305 (224.7
|
|||
|
|
<normalizedToken originalValue="±">+/-</normalizedToken>
|
|||
|
|
51.1) mm SVL and 12-27 (19.9
|
|||
|
|
<normalizedToken originalValue="±">+/-</normalizedToken>
|
|||
|
|
3.7) mm TL, with the largest female measuring 305+22 = 327 mm (SMF 32541, Cubal, Angola).
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage (1895)</bibRefCitation>
|
|||
|
|
mentioned an unsexed individual (probably a female) that measured 331+29 = 360 mm, but this specimen was unaccounted for in MBL.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="97" type="natural history">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">Natural history.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
This is a semi-fossorial species that feeds exclusively on reptile eggs, using its blade-like rear maxillary teeth to puncture the eggs, similar to other
|
|||
|
|
<taxonomicName authorityName="Gray" authorityYear="1849" class="Reptilia" family="Prosymnidae" genus="Prosymna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prosymna" order="Squamata" pageId="0" pageNumber="97" phylum="Chordata" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="97">Prosymna</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="97" type="distribution">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">Distribution and habitat.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
Currently the species is known to occur in three main geographic clusters: west-central Angola, north-central Namibia and isolated records from the Zambezi Region in north-eastern Namibia, northern Botswana and north-western Zimbabwe (Fig.
|
|||
|
|
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. a Maximum likelihood (IQTREE) concatenated phylogeny with BI BEAST and BI MrBayes support overlain. Each of the species groups are demarcated by coloured vertical bars at the right margin b maximum likelihood (IQTREE) phylogenies (16 S and cyt-b) with gene-specific single-locus species delimitation analyses results overlain. The bars to the right of the phylogeny represent the putative taxa assignments for each analysis and the values beneath the bars denote the total number of putative taxa for each analysis." figureDoi="10.3897/zookeys.1121.85693.figure2" httpUri="https://binary.pensoft.net/fig/744033" pageId="0" pageNumber="97">2</figureCitation>
|
|||
|
|
). However, it is possible that this distribution might be more continuous, given this is a rarely observed species that mostly emerges to the surface only after good rains (
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.1093/biolinnean/blaa076" author="Heinicke, MP" journalOrPublisher="Linnean Society of London" pageId="0" pageNumber="97" pagination="136 - 153" refId="B52" refString="Heinicke, MP, Titus-McQuillan, JE, Daza, JD, Kull, EM, Stanley, EL, Bauer, AM, 2020. Phylogeny and evolution of unique skull morphologies in dietary specialist African shovel-snouted snakes (Lamprophiidae: Prosymna). Biological Journal of the Linnean Society. Linnean Society of London 131 (1): 136 - 153, DOI: https://doi.org/10.1093/biolinnean/blaa076" title="Phylogeny and evolution of unique skull morphologies in dietary specialist African shovel-snouted snakes (Lamprophiidae: Prosymna). Biological Journal of the Linnean Society." url="https://doi.org/10.1093/biolinnean/blaa076" volume="131" year="2020">Heinicke et al. 2020</bibRefCitation>
|
|||
|
|
). The records from Luanda (USBN 20035 and CAS 84181) and northern Angola (IICT/R 14-1957) require verification. This species is associated with savanna with an annual rainfall of 500-1200 mm (
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
). In southwestern Angola it has been found in miombo woodland in sandy soils (
|
|||
|
|
<bibRefCitation author="Baptista, NL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="97" refId="B3" refString="Baptista, NL, Antonio, T, Branch, WR, 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2)[Special Section]: 96-130[e203]." title="The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13 (2) [Special Section]: 96 - 130 [e 203]." year="2019">Baptista et al. 2019</bibRefCitation>
|
|||
|
|
). In the eastern Zambezi Region and northern Botswana, the species is associated with drier savanna in deep Kalahari sands (
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="97" type="localities">
|
|||
|
|
<paragraph pageId="0" pageNumber="97">Localities.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="97">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">Angola</emphasis>
|
|||
|
|
: Bela-Vista (
|
|||
|
|
<normalizedToken originalValue="Missão">Missao</normalizedToken>
|
|||
|
|
do Dondi),
|
|||
|
|
<geoCoordinate degrees="12.36667" direction="south" orientation="latitude" precision="1" value="-12.36667">-12.36667</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="16.20000" direction="east" orientation="longitude" precision="1" value="16.2">16.20000</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Hellmich, W" journalOrPublisher="Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut" pageId="0" pageNumber="97" pagination="39 - 80" refId="B53" refString="Hellmich, W, 1957. Die reptilienausbeute der Hamburgischen Angola Expedition. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 55: 39 - 80" title="Die reptilienausbeute der Hamburgischen Angola Expedition." volume="55" year="1957">Hellmich 1957</bibRefCitation>
|
|||
|
|
: 66); interior of Benguela (
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); Bibala,
|
|||
|
|
<geoCoordinate degrees="14.76667" direction="south" orientation="latitude" precision="1" value="-14.76667">-14.76667</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="13.36667" direction="east" orientation="longitude" precision="1" value="13.36667">13.36667</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Jornal de Sciencias Mathematicas, Physicas e Naturaes" pageId="0" pageNumber="97" pagination="209 - 227" refId="B5" refString="Bocage, JVB, 1873. Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale. Jornal de Sciencias Mathematicas, Physicas e Naturaes 4: 209 - 227" title="Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale." volume="4" year="1873">Bocage 1873</bibRefCitation>
|
|||
|
|
: 218); Bicuar National Park, Woodland trapline 1,
|
|||
|
|
<geoCoordinate degrees="15.09441" direction="south" orientation="latitude" precision="1" value="-15.09441">-15.09441</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="14.83831" direction="east" orientation="longitude" precision="1" value="14.83831">14.83831</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Baptista, NL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="97" refId="B3" refString="Baptista, NL, Antonio, T, Branch, WR, 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2)[Special Section]: 96-130[e203]." title="The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13 (2) [Special Section]: 96 - 130 [e 203]." year="2019">Baptista et al. 2019</bibRefCitation>
|
|||
|
|
: 118); Caconda,
|
|||
|
|
<geoCoordinate degrees="13.73537" direction="south" orientation="latitude" precision="1" value="-13.73537">-13.73537</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="15.06720" direction="east" orientation="longitude" precision="1" value="15.0672">15.06720</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 151); Cubal,
|
|||
|
|
<geoCoordinate degrees="13.03333" direction="south" orientation="latitude" precision="1" value="-13.03333">-13.03333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="14.73333" direction="east" orientation="longitude" precision="1" value="14.73333">14.73333</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Mertens, R" journalOrPublisher="Senckenbergiana" pageId="0" pageNumber="97" pagination="425 - 443" refId="B73" refString="Mertens, R, 1938. Amphibien und Reptilien aus Angola gesmmelt von W. Shack. Senckenbergiana 20: 425 - 443" title="Amphibien und Reptilien aus Angola gesmmelt von W. Shack." volume="20" year="1938">Mertens 1938</bibRefCitation>
|
|||
|
|
: 439); Ebanga, -12.73333, 14.73333 (
|
|||
|
|
<bibRefCitation author="Monard, A" journalOrPublisher="Arquivos do Museu Bocage" pageId="0" pageNumber="97" pagination="19 - 154" refId="B79" refString="Monard, A, 1937. Contribution a l'Herpetologie d'Angola. Arquivos do Museu Bocage 8: 19 - 154" title="Contribution a l'Herpetologie d'Angola." volume="8" year="1937">Monard 1937</bibRefCitation>
|
|||
|
|
: 123);
|
|||
|
|
<normalizedToken originalValue="Huíla">Huila</normalizedToken>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="15.08333" direction="south" orientation="latitude" precision="1" value="-15.08333">-15.08333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="13.55000" direction="east" orientation="longitude" precision="1" value="13.55">13.55000</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); Luanda and 'Luanda, 3 mi S of
|
|||
|
|
<normalizedToken originalValue="airport’">airport'</normalizedToken>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="8.83333" direction="south" orientation="latitude" precision="1" value="-8.83333">-8.83333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="13.26667" direction="east" orientation="longitude" precision="1" value="13.26667">13.26667</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
: 515); Maconjo,
|
|||
|
|
<geoCoordinate degrees="15.016667" direction="south" orientation="latitude" precision="1" value="-15.016667">-15.016667</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="13.2000" direction="east" orientation="longitude" precision="5" value="13.2">13.2000</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); interior of Mossamedes (
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Jornal de Sciencias Mathematicas, Physicas e Naturaes" pageId="0" pageNumber="97" pagination="209 - 227" refId="B5" refString="Bocage, JVB, 1873. Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale. Jornal de Sciencias Mathematicas, Physicas e Naturaes 4: 209 - 227" title="Melanges erpetologiques. II. Sur quelques Reptiles et Batraciens nouveaux, rares ou peu connus d'frique occidentale." volume="4" year="1873">Bocage 1873</bibRefCitation>
|
|||
|
|
: 218); Quibula,
|
|||
|
|
<geoCoordinate degrees="12.28333" direction="south" orientation="latitude" precision="1" value="-12.28333">-12.28333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="14.68333" direction="east" orientation="longitude" precision="1" value="14.68333">14.68333</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); Posto do Milando (
|
|||
|
|
<geoCoordinate degrees="8.81667" direction="south" orientation="latitude" precision="1" value="-8.81667">-8.81667</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="17.56667" direction="east" orientation="longitude" precision="1" value="17.56667">17.56667</geoCoordinate>
|
|||
|
|
) (
|
|||
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1052.64607" author="Ceriaco, LMP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="97" pagination="85 - 156" refId="B29" refString="Ceriaco, LMP, Parrinha, D, Marques, MP, 2021a. Saving collections: Taxonomic revision of the herpetological collection of the Instituto de Investigacao Cientifica Tropical, Lisbon (Portugal) with a protocol to rescue abandoned collections. ZooKeys 1052: 85 - 156, DOI: https://doi.org/10.3897/zookeys.1052.64607" title="Saving collections: Taxonomic revision of the herpetological collection of the Instituto de Investigacao Cientifica Tropical, Lisbon (Portugal) with a protocol to rescue abandoned collections." url="https://doi.org/10.3897/zookeys.1052.64607" volume="1052" year="2021 a">
|
|||
|
|
<normalizedToken originalValue="Ceríaco">Ceriaco</normalizedToken>
|
|||
|
|
et al. 2021a
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 16). Quindumbo,
|
|||
|
|
<geoCoordinate degrees="12.46667" direction="south" orientation="latitude" precision="1" value="-12.46667">-12.46667</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="14.93333" direction="east" orientation="longitude" precision="1" value="14.93333">14.93333</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); Quissange,
|
|||
|
|
<geoCoordinate degrees="12.43333" direction="south" orientation="latitude" precision="1" value="-12.43333">-12.43333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="14.05000" direction="east" orientation="longitude" precision="1" value="14.05">14.05000</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Bocage, JVB" journalOrPublisher="Ministerio da Marinha e das Colonias, Lisbonne, Portugal" pageId="0" pageNumber="97" refId="B7" refString="Bocage, JVB, 1895. Herpetologie d'Angola et du Congo. Ministerio da Marinha e das Colonias, Lisbonne, Portugal" title="Herpetologie d'Angola et du Congo." year="1895">Bocage 1895</bibRefCitation>
|
|||
|
|
: 98); Tundavala,
|
|||
|
|
<geoCoordinate degrees="14.82018" direction="south" orientation="latitude" precision="1" value="-14.82018">-14.82018</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="13.404217" direction="east" orientation="longitude" precision="1" value="13.404217">13.404217</geoCoordinate>
|
|||
|
|
(Justin Nicolau photo).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">Botswana</emphasis>
|
|||
|
|
: Joverega (Geoverega), - 19.13333, 24.25 (
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
: 515).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="97">Namibia</emphasis>
|
|||
|
|
: Grootfontein,
|
|||
|
|
<geoCoordinate degrees="19.55012" direction="south" orientation="latitude" precision="1" value="-19.55012">-19.55012</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="18.10965" direction="east" orientation="longitude" precision="1" value="18.10965">18.10965</geoCoordinate>
|
|||
|
|
(Francois Theart photo); Karakuwisa,
|
|||
|
|
<geoCoordinate degrees="18.933333" direction="south" orientation="latitude" precision="1" value="-18.933332">-18.933333</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="19.733333" direction="east" orientation="longitude" precision="1" value="19.733334">19.733333</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="Mertens, R" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="97" pagination="1 - 172" refId="B74" refString="Mertens, R, 1955. Die Amphibien und Reptilien Suedwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgefuehrten Reise. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 490: 1 - 172" title="Die Amphibien und Reptilien Suedwestafrikas. Aus den Ergebnissen einer im Jahre 1952 ausgefuehrten Reise." volume="490" year="1955">Mertens 1955</bibRefCitation>
|
|||
|
|
: 94); Katima Mulilo, -17.5, 24.266667 (
|
|||
|
|
<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
|
|||
|
|
: 515); Namutoni,
|
|||
|
|
<geoCoordinate degrees="18.807624" direction="south" orientation="latitude" precision="1" value="-18.807625">-18.807624</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate degrees="16.940288" direction="east" orientation="longitude" precision="1" value="16.940289">16.940288</geoCoordinate>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation author="FitzSimons, VFM" journalOrPublisher="Purnell and Sons, Cape Town, South Africa" pageId="0" pageNumber="97" refId="B45" refString="FitzSimons, VFM, 1962. Snakes of southern Africa. Purnell and Sons, Cape Town, South Africa" title="Snakes of southern Africa." year="1962">FitzSimons 1962</bibRefCitation>
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: 161); 15 km WSW of Katima Mulilo,
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<geoCoordinate degrees="17.61448" direction="south" orientation="latitude" precision="1" value="-17.61448">-17.61448</geoCoordinate>
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,
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<geoCoordinate degrees="24.20593" direction="east" orientation="longitude" precision="1" value="24.20593">24.20593</geoCoordinate>
|
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|
2 (
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<bibRefCitation author="Broadley, DG" journalOrPublisher="Nature and Science" pageId="0" pageNumber="97" pagination="481 - 556" refId="B19" refString="Broadley, DG, 1980. A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B. Nature and Science 6: 481 - 556" title="A revision of the African snake genus Prosymna Gray (Colubridae). Occasional papers of the National Museums of Rhodesia, Series B." volume="6" year="1980">Broadley 1980</bibRefCitation>
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: 515); Otjozondjupa Region,
|
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<geoCoordinate degrees="19.08800" direction="south" orientation="latitude" precision="1" value="-19.088">-19.08800</geoCoordinate>
|
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|
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,
|
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<geoCoordinate degrees="18.83300" direction="east" orientation="longitude" precision="1" value="18.833">18.83300</geoCoordinate>
|
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(http://www.the-eis.com/atlas/?q=details/snake-record&occurrence_id=654386).
|
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<emphasis bold="true" pageId="0" pageNumber="97">Zimbabwe</emphasis>
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: Malindi Siding, Hwange,
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<geoCoordinate degrees="18.74885" direction="south" orientation="latitude" precision="1" value="-18.74885">-18.74885</geoCoordinate>
|
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|
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,
|
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<geoCoordinate degrees="27.01852" direction="east" orientation="longitude" precision="1" value="27.01852">27.01852</geoCoordinate>
|
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|
|
(
|
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|
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<bibRefCitation author="Broadley, DG" journalOrPublisher="Donker Publishers, Parklands" pageId="0" pageNumber="97" refId="B21" refString="Broadley, DG, 1995. Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." title="Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." year="1995">Broadley 1995</bibRefCitation>
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: 48); Inyokene, Nyamandlovu, -19.93333, 28.06667 (
|
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|
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<bibRefCitation author="Broadley, DG" journalOrPublisher="Donker Publishers, Parklands" pageId="0" pageNumber="97" refId="B21" refString="Broadley, DG, 1995. Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." title="Geographical Distribution - Prosymna angolensis. African Herp News 23: 48." year="1995">Broadley 1995</bibRefCitation>
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: 48).
|
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</paragraph>
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</subSubSection>
|
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</treatment>
|
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</document>
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