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<mods:title id="72D5808CD8E684996D96856313D6FFA1">Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae) from the southern South American Transition Zone and boundaries: taxonomic overview with four new species</mods:title>
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<emphasis id="8FABEAC8FFD79A58FF03EBDC4FA4F946" bold="true" box="[151,405,1698,1725]" italics="true" pageId="28" pageNumber="74">Liogenys xanthocera</emphasis>
Harold, 1869
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<figureCitation id="25E42A5FFFD79A58FF03EBB84EC2F91A" box="[151,243,1735,1761]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="28" pageNumber="74">Figs. 10</figureCitation>
, 
<figureCitation id="25E42A5FFFD79A58FF6BEBB84F1AF91A" box="[255,299,1735,1761]" captionStart="FIGURE 13" captionStartId="32.[151,250,1909,1933]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure-19@32.[214,1373,181,1885]" captionTargetPageId="32" captionText="FIGURE 13. Liogenys maxillaricuspis Cherman, new species: A) head in ventral view, C) labium, G) right maxilla; L. brachyclypeata Cherman, new species: B) head in ventral view, D) labium; L. calcarata Frey, 1970: E) left maxilla; L. xanthocera Harold, 1869: F) right maxilla. Legends: A, B) CLP: clypeus, LBM: labium, LBR: labrum, MXL: maxilla, MXLP: maxillary palp. C, D) LGL: ligula, LPLP: labial palp, MTM: mentum, PMTM: postmentum." figureDoi="http://doi.org/10.5281/zenodo.4359222" httpUri="https://zenodo.org/record/4359222/files/figure.png" pageId="28" pageNumber="74">13F</figureCitation>
, 
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<emphasis id="8FABEAC8FFD79A58FF03EA734F4AF8D8" box="[151,379,1803,1827]" italics="true" pageId="28" pageNumber="74">Liogenys pallidicornis</emphasis>
<bibRefCitation id="D94E4B2BFFD79A58FE15EA744C6EF8D8" author="Blanchard, E." box="[385,607,1803,1827]" pageId="28" pageNumber="74" refId="ref27011" refString="Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp." type="book" year="1851">Blanchard, 1851: 168</bibRefCitation>
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; Lacordaire 1855: 269 (generic redescription); 
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(key).
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<emphasis id="8FABEAC8FFD79A58FF03EA554F44F8B9" box="[151,373,1834,1858]" italics="true" pageId="28" pageNumber="74">Liogenys xanthocerus</emphasis>
<bibRefCitation id="D94E4B2BFFD79A58FEEFEA554C09F8B9" author="Harold, E. F. von" box="[379,568,1834,1858]" pageId="28" pageNumber="74" pagination="122 - 125" refId="ref27868" refString="Harold, E. F. von (1869 a) Abanderungen vergebener Namen. Coleopterologische Hefte, 1, 122 - 125." type="journal article" year="1869">Harold 1869a: 123</bibRefCitation>
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(new replacement name for 
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<emphasis id="8FABEAC8FFD79A58FCCDEA544DC4F8B9" box="[857,1013,1834,1858]" italics="true" pageId="28" pageNumber="74">L. pallidicornis</emphasis>
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); 
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<bibRefCitation id="D94E4B2BFFD79A58FB9FEA554AE7F8B9" author="Harold, E. F. von" box="[1035,1238,1834,1858]" pageId="28" pageNumber="74" refId="ref27892" refString="Harold, E. F. von (1869 b) Scarabaeidae. In: Gemminger, M. &amp; von Harold, E. F. (Eds.), Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus, Vol. 4. E. H. Gummi, Munich, pp. 976 - 1346." type="book" year="1869">Harold 1869b: 1140</bibRefCitation>
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(checklist, synonymy); 
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(checklist); 
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<bibRefCitation id="D94E4B2BFFD79A58FD98EA374C84F89A" author="Bruch, C." box="[524,693,1864,1889]" pageId="28" pageNumber="74" pagination="181 - 225" refId="ref27050" refString="Bruch, C. (1911) Catalogo sistematico de los Coleopteros de la Republica Argentina. Revista del Museo de la Plata, 17, 181 - 225. https: // doi. org / 10.5962 / bhl. title. 8799" type="journal article" year="1911">Bruch 1911: 200</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation id="3C7E10CBFFD79A58FCA0EA374A22F89A" author="Dalla Torre, K. W. von" box="[820,1043,1864,1889]" page="318" pageId="28" pageNumber="74" year="1913">
<bibRefCitation id="D94E4B2BFFD79A58FCA0EA374A22F89A" author="Dalla Torre, K. W. von" box="[820,1043,1864,1889]" pageId="28" pageNumber="74" pagination="291 - 450" refId="ref27433" refString="Dalla Torre, K. W. von (1913) Scarabaeidae: Melolonthinae IV. Pars 50. In: S. Schenkling (Ed.), Coleopterorum catalogus. Vol. XX. Junk, W., Berlin, pp. 291 - 450." type="book chapter" year="1913">Dalla Torre 1913: 318</bibRefCitation>
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(checklist); 
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<bibRefCitation id="D94E4B2BFFD79A58FB05EA364B2EF89A" author="Frey, G." box="[1169,1311,1864,1889]" pageId="28" pageNumber="74" pagination="38 - 64" refId="ref27688" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey 1969: 42</bibRefCitation>
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(key; junior synonym of 
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<emphasis id="8FABEAC8FFD79A58FED2EA174FD2F884" box="[326,483,1895,1919]" italics="true" pageId="28" pageNumber="74">L. pallidicornis</emphasis>
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[sic]); 
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<bibRefCitation id="D94E4B2BFFD79A58FDBFEA184CD5F884" author="Krajcik, M." box="[555,740,1895,1919]" pageId="28" pageNumber="74" refId="ref27990" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. 5 th Edition. Animma. x, Plzen, 278 pp." type="book" year="2012">Krajčík 2012: 145</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation id="3C7E10CBFFD79A58FCF6EA184A7AF884" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[866,1099,1895,1919]" page="4" pageId="28" pageNumber="74" year="2017">
<bibRefCitation id="D94E4B2BFFD79A58FCF6EA184A7AF884" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[866,1099,1895,1919]" pageId="28" pageNumber="74" pagination="1 - 120" refId="ref27290" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman 
<emphasis id="8FABEAC8FFD79A58FC51EA174DC8F884" box="[965,1017,1895,1919]" italics="true" pageId="28" pageNumber="74">et al.</emphasis>
2017: 4
</bibRefCitation>
</treatmentCitation>
(generic history).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BD6036DAFFD79A58FF03EAF94D9BF865" blockId="28.[151,1437,1803,1981]" box="[151,938,1926,1950]" pageId="28" pageNumber="74">
<treatmentCitationGroup id="9DCF11F4FFD79A58FF03EAF94D9BF865" box="[151,938,1926,1950]" pageId="28" pageNumber="74">
<taxonomicName id="7ADF4D59FFD79A58FF03EAF94F5EF865" authority="Frey 1969" box="[151,367,1926,1950]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="74" phylum="Arthropoda" rank="species" species="xanthoceris">
<emphasis id="8FABEAC8FFD79A58FF03EAF94F5EF865" box="[151,367,1926,1950]" italics="true" pageId="28" pageNumber="74">Liogenys xanthoceris</emphasis>
</taxonomicName>
[sic]: 
<treatmentCitation id="3C7E10CBFFD79A58FE24EAF94C0FF865" author="Frey, G." box="[432,574,1926,1950]" page="57" pageId="28" pageNumber="74" year="1969">
<bibRefCitation id="D94E4B2BFFD79A58FE24EAF94C0FF865" author="Frey, G." box="[432,574,1926,1950]" pageId="28" pageNumber="74" pagination="38 - 64" refId="ref27688" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera-Melolonthinae-Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey 1969: 57</bibRefCitation>
</treatmentCitation>
(synonym of 
<taxonomicName id="7ADF4D59FFD79A58FD5FEAF84D59F865" authorityName="Blanchard" authorityYear="1851" box="[715,872,1926,1950]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="74" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="8FABEAC8FFD79A58FD5FEAF84D59F865" box="[715,872,1926,1950]" italics="true" pageId="28" pageNumber="74">L. pallidicornis</emphasis>
</taxonomicName>
[sic]).
</treatmentCitationGroup>
</paragraph>
<paragraph id="BD6036DAFFD79A58FF03EADA4BBDF847" blockId="28.[151,1437,1803,1981]" box="[151,1420,1956,1981]" pageId="28" pageNumber="74">
<treatmentCitationGroup id="9DCF11F4FFD79A58FF03EADA4BBDF847" box="[151,1420,1956,1981]" pageId="28" pageNumber="74">
<treatmentCitation id="3C7E10CBFFD79A58FF03EADA4C54F846" author="Blackwelder, R. E." box="[151,613,1956,1981]" page="227" pageId="28" pageNumber="74" year="1944">
<taxonomicName id="7ADF4D59FFD79A58FF03EADA4C54F846" authority=": Blackwelder 1944: 227" authorityName="Blackwelder" authorityPageNumber="227" authorityYear="1944" box="[151,613,1956,1981]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="74" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFD79A58FF03EADA4F5AF847" box="[151,363,1956,1980]" italics="true" pageId="28" pageNumber="74">Liogenys xanthocera</emphasis>
: 
<bibRefCitation id="D94E4B2BFFD79A58FEECEADB4C54F846" author="Blackwelder, R. E." box="[376,613,1956,1981]" pageId="28" pageNumber="74" pagination="189 - 265" refId="ref26953" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2" type="book chapter" year="1944">Blackwelder 1944: 227</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(checklist); 
<treatmentCitation id="3C7E10CBFFD79A58FD77EADA4DBFF847" author="Evans, A. V." box="[739,910,1956,1981]" page="215" pageId="28" pageNumber="74" year="2003">
<bibRefCitation id="D94E4B2BFFD79A58FD77EADA4DBFF847" author="Evans, A. V." box="[739,910,1956,1981]" pageId="28" pageNumber="74" pagination="1 - 458" refId="ref27511" refString="Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211 (1), 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 215</bibRefCitation>
</treatmentCitation>
(checklist); 
<treatmentCitation id="3C7E10CBFFD79A58FB98EADA4B25F846" author="Evans, A. V. &amp; Smith, A. B. T." box="[1036,1300,1956,1981]" page="183" pageId="28" pageNumber="74" year="2009">
<bibRefCitation id="D94E4B2BFFD79A58FB98EADA4B25F846" author="Evans, A. V. &amp; Smith, A. B. T." box="[1036,1300,1956,1981]" pageId="28" pageNumber="74" pagination="1 - 344" refId="ref27603" refString="Evans, A. V. &amp; Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3. Papers in Entomology, 1, 1 - 344." type="journal article" year="2009">Evans &amp; Smith 2009: 183</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="E9A06652FFD69A59FF03EAE84D36F82F" ID-DOI="http://doi.org/10.5281/zenodo.4359210" ID-Zenodo-Dep="4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="29" pageNumber="75" startId="29.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="29">
<paragraph id="BD6036DAFFD69A59FF03EAE84D36F82F" blockId="29.[151,1436,1943,2004]" pageId="29" pageNumber="75">
<emphasis id="8FABEAC8FFD69A59FF03EAE84F11F854" bold="true" box="[151,288,1943,1967]" pageId="29" pageNumber="75">FIGURE 10.</emphasis>
<taxonomicName id="7ADF4D59FFD69A59FEB3EAE74CB8F84B" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[295,649,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="75" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFD69A59FEB3EAE74FCAF84B" box="[295,507,1944,1968]" italics="true" pageId="29" pageNumber="75">Liogenys xanthocera</emphasis>
Harold, 1869
</taxonomicName>
, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral.
</paragraph>
</caption>
<caption id="E9A06652FFD59A5AFF03EAE84ABCF82F" ID-DOI="http://doi.org/10.5281/zenodo.4359214" ID-Zenodo-Dep="4359214" httpUri="https://zenodo.org/record/4359214/files/figure.png" pageId="30" pageNumber="76" startId="30.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="30">
<paragraph id="BD6036DAFFD59A5AFF03EAE84ABCF82F" blockId="30.[151,1436,1943,2004]" pageId="30" pageNumber="76">
<emphasis id="8FABEAC8FFD59A5AFF03EAE84F2FF854" bold="true" box="[151,286,1943,1967]" pageId="30" pageNumber="76">FIGURE 11.</emphasis>
Dimorphism in clypeus. 
<taxonomicName id="7ADF4D59FFD59A5AFD8AEAE74D68F84B" authority="Moser, 1924" authorityName="Moser" authorityYear="1924" box="[542,857,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="76" phylum="Arthropoda" rank="species" species="flaveola">
<emphasis id="8FABEAC8FFD59A5AFD8AEAE74CE0F84B" box="[542,721,1944,1968]" italics="true" pageId="30" pageNumber="76">Liogenys flaveola</emphasis>
<bibRefCitation id="D94E4B2BFFD59A5AFD43EAE74D68F84B" author="Moser, J." box="[727,857,1944,1968]" pageId="30" pageNumber="76" pagination="114 - 136" refId="ref28322" refString="Moser, J. (1924) Beitrag zur Kenntnis der Melolonthiden (Col.). Stettiner Entomologische Zeitung, 84, 114 - 136." type="journal article" year="1924">Moser, 1924</bibRefCitation>
</taxonomicName>
: A) male, B) female; 
<taxonomicName id="7ADF4D59FFD59A5AFBA6EAE74B2AF84B" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[1074,1307,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="76" phylum="Arthropoda" rank="species" species="flavida">
<emphasis id="8FABEAC8FFD59A5AFBA6EAE74AA3F84B" box="[1074,1170,1944,1968]" italics="true" pageId="30" pageNumber="76">L. flavida</emphasis>
<bibRefCitation id="D94E4B2BFFD59A5AFB0DEAE74B2AF84B" author="Moser, J." box="[1177,1307,1944,1968]" pageId="30" pageNumber="76" pagination="95 - 167" refId="ref28263" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
: C) male, D) female; 
<taxonomicName id="7ADF4D59FFD59A5AFF7DEAC34FD0F82F" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[233,481,1980,2004]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="76" phylum="Arthropoda" rank="species" species="kuntzeni">
<emphasis id="8FABEAC8FFD59A5AFF7DEAC34F69F82F" box="[233,344,1980,2004]" italics="true" pageId="30" pageNumber="76">L. kuntzeni</emphasis>
<bibRefCitation id="D94E4B2BFFD59A5AFECBEAC34FD0F82F" author="Moser, J." box="[351,481,1980,2004]" pageId="30" pageNumber="76" pagination="133 - 182" refId="ref28284" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und S ʾ d-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
</taxonomicName>
: E) male, F) female; 
<taxonomicName id="7ADF4D59FFD59A5AFD23EAC34D86F82F" authority="Moser, 1918" authorityName="Moser" authorityYear="1918" box="[695,951,1980,2004]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="76" phylum="Arthropoda" rank="species" species="rufoflava">
<emphasis id="8FABEAC8FFD59A5AFD23EAC34D1FF82F" box="[695,814,1980,2004]" italics="true" pageId="30" pageNumber="76">L. rufoflava</emphasis>
<bibRefCitation id="D94E4B2BFFD59A5AFCA1EAC34D86F82F" author="Moser, J." box="[821,951,1980,2004]" pageId="30" pageNumber="76" pagination="95 - 167" refId="ref28263" refString="Moser, J. (1918) Neue Amerikanische Melolonthiden. Stettiner Entomologische Zeitung, 79, 95 - 167." type="journal article" year="1918">Moser, 1918</bibRefCitation>
</taxonomicName>
: G) male, H) female.
</paragraph>
</caption>
<caption id="E9A06652FFD49A5BFF03EB874A91F8CF" ID-DOI="http://doi.org/10.5281/zenodo.4359218" ID-Zenodo-Dep="4359218" httpUri="https://zenodo.org/record/4359218/files/figure.png" pageId="31" pageNumber="77" startId="31.[151,250,1784,1808]" targetBox="[151,1435,181,1760]" targetPageId="31">
<paragraph id="BD6036DAFFD49A5BFF03EB874A91F8CF" blockId="31.[151,1436,1784,1844]" pageId="31" pageNumber="77">
<emphasis id="8FABEAC8FFD49A5BFF03EB874F10F8EB" bold="true" box="[151,289,1784,1808]" pageId="31" pageNumber="77">FIGURE 12.</emphasis>
Dimorphism in clypeus. 
<taxonomicName id="7ADF4D59FFD49A5BFDB3EB864D45F8EB" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" box="[551,884,1784,1808]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="77" phylum="Arthropoda" rank="species" species="lucialmeidae" status="sp. nov.">
<emphasis id="8FABEAC8FFD49A5BFDB3EB864D3CF8EB" box="[551,781,1784,1808]" italics="true" pageId="31" pageNumber="77">Liogenys lucialmeidae</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="8FABEAC8FFD49A5BFCEBEB874DCCF8EB" bold="true" box="[895,1021,1784,1808]" pageId="31" pageNumber="77">
<taxonomicNameLabel id="949857B3FFD49A5BFCEBEB874DCCF8EB" box="[895,1021,1784,1808]" pageId="31" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</emphasis>
: A) male, B) female; 
<taxonomicName id="7ADF4D59FFD49A5BFB74EB864EF4F8C8" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="77" phylum="Arthropoda" rank="species" species="martinezi" status="sp. nov.">
<emphasis id="8FABEAC8FFD49A5BFB74EB864B6DF8EB" box="[1248,1372,1785,1808]" italics="true" pageId="31" pageNumber="77">L. martinezi</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="8FABEAC8FFD49A5BFF5BEA634F7DF8CF" bold="true" box="[207,332,1820,1844]" pageId="31" pageNumber="77">
<taxonomicNameLabel id="949857B3FFD49A5BFF5BEA634F7DF8CF" box="[207,332,1820,1844]" pageId="31" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</emphasis>
: C) male, D) female; 
<taxonomicName id="7ADF4D59FFD49A5BFDBEEA624D79F8CF" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" box="[554,840,1820,1844]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="77" phylum="Arthropoda" rank="species" species="maxillaricuspis" status="sp. nov.">
<emphasis id="8FABEAC8FFD49A5BFDBEEA624CD0F8CF" box="[554,737,1820,1844]" italics="true" pageId="31" pageNumber="77">L. maxillaricuspis</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="8FABEAC8FFD49A5BFCC5EA634DFFF8CF" bold="true" box="[849,974,1820,1844]" pageId="31" pageNumber="77">
<taxonomicNameLabel id="949857B3FFD49A5BFCC5EA634DFFF8CF" box="[849,974,1820,1844]" pageId="31" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</emphasis>
: E) male, F) female.
</paragraph>
</caption>
<caption id="E9A06652FFEB9A64FF03EA0A4DE8F803" ID-DOI="http://doi.org/10.5281/zenodo.4359222" ID-Zenodo-Dep="4359222" httpUri="https://zenodo.org/record/4359222/files/figure.png" pageId="32" pageNumber="78" startId="32.[151,250,1909,1933]" targetBox="[214,1373,181,1885]" targetPageId="32">
<paragraph id="BD6036DAFFEB9A64FF03EA0A4DE8F803" blockId="32.[151,1437,1909,2041]" pageId="32" pageNumber="78">
<emphasis id="8FABEAC8FFEB9A64FF03EA0A4F11F876" bold="true" box="[151,288,1909,1933]" pageId="32" pageNumber="78">FIGURE 13.</emphasis>
<taxonomicName id="7ADF4D59FFEB9A64FEB2EA094CBAF876" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" box="[294,651,1909,1933]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="78" phylum="Arthropoda" rank="species" species="maxillaricuspis" status="sp. nov.">
<emphasis id="8FABEAC8FFEB9A64FEB2EA094C14F876" box="[294,549,1909,1933]" italics="true" pageId="32" pageNumber="78">Liogenys maxillaricuspis</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="8FABEAC8FFEB9A64FD01EA0A4D23F876" bold="true" box="[661,786,1909,1933]" pageId="32" pageNumber="78">
<taxonomicNameLabel id="949857B3FFEB9A64FD01EA0A4D23F876" box="[661,786,1909,1933]" pageId="32" pageNumber="78" rank="species">new species</taxonomicNameLabel>
</emphasis>
: A) head in ventral view, C) labium, G) right maxilla; 
<taxonomicName id="7ADF4D59FFEB9A64FAAAEA094F51F84A" authority="Cherman" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="78" phylum="Arthropoda" rank="species" species="brachyclypeata" status="sp. nov.">
<emphasis id="8FABEAC8FFEB9A64FAAAEA094EC9F84A" italics="true" pageId="32" pageNumber="78">L. brachyclypeata</emphasis>
Cherman
</taxonomicName>
, 
<emphasis id="8FABEAC8FFEB9A64FEFEEAE64FD9F84A" bold="true" box="[362,488,1945,1969]" pageId="32" pageNumber="78">
<taxonomicNameLabel id="949857B3FFEB9A64FEFEEAE64FD9F84A" box="[362,488,1945,1969]" pageId="32" pageNumber="78" rank="species">new species</taxonomicNameLabel>
</emphasis>
: B) head in ventral view, D) labium; 
<taxonomicName id="7ADF4D59FFEB9A64FCFFEAE54A51F84A" authority="Frey, 1970" authorityName="Frey" authorityYear="1970" box="[875,1120,1945,1969]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="78" phylum="Arthropoda" rank="species" species="calcarata">
<emphasis id="8FABEAC8FFEB9A64FCFFEAE54DD9F84A" box="[875,1000,1945,1969]" italics="true" pageId="32" pageNumber="78">L. calcarata</emphasis>
<bibRefCitation id="D94E4B2BFFEB9A64FC64EAE64A51F84A" author="Frey, G." box="[1008,1120,1945,1969]" pageId="32" pageNumber="78" pagination="281 - 287" refId="ref27721" refString="Frey, G. (1970) Beitrag zur kenntnis der Sudamerikanischen Melolonthiden (Col.). Entomologische Arbeiten aus dem Museum G. Frey, 21, 281 - 287." type="journal article" year="1970">Frey, 1970</bibRefCitation>
</taxonomicName>
: E) left maxilla; 
<taxonomicName id="7ADF4D59FFEB9A64FA9AEAE54F10F82E" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="78" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFEB9A64FA9AEAE54BADF84A" box="[1294,1436,1945,1969]" italics="true" pageId="32" pageNumber="78">L. xanthocera</emphasis>
Harold, 1869
</taxonomicName>
: F) right maxilla. Legends: A, B) CLP: clypeus, LBM: labium, LBR: labrum, MXL: maxilla, MXLP: maxillary palp. C, D) LGL: ligula, LPLP: labial palp, MTM: mentum, PMTM: postmentum.
</paragraph>
</caption>
<caption id="E9A06652FFEA9A65FF03EAE74FDCF803" ID-DOI="http://doi.org/10.5281/zenodo.4359226" ID-Zenodo-Dep="4359226" httpUri="https://zenodo.org/record/4359226/files/figure.png" pageId="33" pageNumber="79" startId="33.[151,250,1944,1968]" targetBox="[214,1377,192,1909]" targetPageId="33">
<paragraph id="BD6036DAFFEA9A65FF03EAE74FDCF803" blockId="33.[151,1436,1944,2040]" pageId="33" pageNumber="79">
<emphasis id="8FABEAC8FFEA9A65FF03EAE74F11F84B" bold="true" box="[151,288,1944,1968]" pageId="33" pageNumber="79">FIGURE 14.</emphasis>
Geographical distribution of known 
<taxonomicName id="7ADF4D59FFEA9A65FD0EEAE64CC4F84B" box="[666,757,1945,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="8FABEAC8FFEA9A65FD0EEAE64CC4F84B" box="[666,757,1945,1968]" italics="true" pageId="33" pageNumber="79">Liogenys</emphasis>
</taxonomicName>
species occurring in the STZ and its boundaries: A) general, B, C, D) detail. Image legend: shaped points: already known species; colors: provinces of Neotropical, South American Transition Zone (STZ), and Andean regions.
</paragraph>
</caption>
<subSubSection id="F5C56551FFE99A66FF03EDE84CB9FD52" pageId="34" pageNumber="80" type="materials_examined">
<paragraph id="BD6036DAFFE99A66FF03EDE84CB8FE0E" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF03EDE84F76FF4B" bold="true" box="[151,327,150,177]" pageId="34" pageNumber="80">Type material.</emphasis>
Male 
<typeStatus id="62648878FFE99A66FE07EDE84FCBFF4A" box="[403,506,151,177]" pageId="34" pageNumber="80" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="7ADF4D59FFE99A66FDB7EDE84D10FF4A" authorityName="Blanchard" authorityYear="1851" box="[547,801,151,177]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="8FABEAC8FFE99A66FDB7EDE84D10FF4A" box="[547,801,151,177]" italics="true" pageId="34" pageNumber="80">Liogenys pallidicornis</emphasis>
</taxonomicName>
here designated (
<collectionCode id="DBCEAE1FFFE99A66FC78EDE84A78FF4A" box="[1004,1097,151,177]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="80">MNHN</collectionCode>
): [white handwritten] “6051 / 34”, [light green, typeset and handwritten] “Patagonie / d’Orbigny / 
<collectionCode id="DBCEAE1FFFE99A66FC30EDC34A48FF2E" box="[932,1145,187,215]" pageId="34" pageNumber="80">MUSÉUM PARIS</collectionCode>
”, [green handwritten] “ 
<taxonomicName id="7ADF4D59FFE99A66FA17EDC44F16FF02" authorityName="Blanchard" authorityYear="1851" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="pallidicornis">L. pallidicornis</taxonomicName>
. / Cat. Mus. / Baie de San Blas (Patagonie) / M. D’Orbigny”, [white typeset] “Southern Neotropical Scarabs / database # 
<collectingCountry id="C5C8764AFFE99A66FEE2EC7D4FA9FEE6" box="[374,408,258,285]" name="American Samoa" pageId="34" pageNumber="80">AS</collectingCountry>
2615155 / 
<taxonomicName id="7ADF4D59FFE99A66FD9BEC7C4CC9FEE6" authorityName="Harold" authorityYear="1869" box="[527,760,259,285]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFE99A66FD9BEC7C4CC9FEE6" box="[527,760,259,285]" italics="true" pageId="34" pageNumber="80">Liogenys xanthocera</emphasis>
</taxonomicName>
/ Harold 1869 ³ / DET:A.B. 
<collectionCode id="DBCEAE1FFFE99A66FBA1EC7C4A76FEE6" box="[1077,1095,259,285]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="80">T</collectionCode>
. 
<collectionCode id="DBCEAE1FFFE99A66FBDEEC7C4A99FEE6" box="[1098,1192,259,285]" pageId="34" pageNumber="80">SMITH</collectionCode>
2008”, [red, outlined, typeset and handwritten] “ 
<taxonomicName id="7ADF4D59FFE99A66FE57EC584C63FEBA" box="[451,594,295,321]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ PALLIDICORNIS / 
<bibRefCitation id="D94E4B2BFFE99A66FCC3EC584A7CFEBA" author="Blanchard, E." box="[855,1101,294,321]" pageId="34" pageNumber="80" refId="ref27011" refString="Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp." type="book" year="1851">BLANCHARD, 1851</bibRefCitation>
/ 
<typeStatus id="62648878FFE99A66FBFCEC584B39FEBA" box="[1128,1288,295,321]" pageId="34" pageNumber="80" type="lectotype">LECTOTYPE</typeStatus>
/ Des. Cherman M. A.”. Male 
<typeStatus id="62648878FFE99A66FEFDEC344C31FE9E" box="[361,512,331,357]" pageId="34" pageNumber="80" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="DBCEAE1FFFE99A66FD9BEC344C59FE9E" box="[527,616,331,357]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="34" pageNumber="80">MNHN</collectionCode>
) bearing the labels: [white handwritten] “6051 / 34”, [white handwritten] “1052”, [light green typeset and handwritten] “Patagonie / d’Orbigny / 
<collectionCode id="DBCEAE1FFFE99A66FC24EC0F4AB2FE72" box="[944,1155,367,395]" pageId="34" pageNumber="80">MUSÉUM PARIS</collectionCode>
”, [white typeset] “Southern Neotropical Scarabs / database # 
<collectingCountry id="C5C8764AFFE99A66FDABECED4C50FE56" box="[575,609,402,429]" name="American Samoa" pageId="34" pageNumber="80">AS</collectingCountry>
2615156 / 
<taxonomicName id="7ADF4D59FFE99A66FD75ECEC4DFFFE56" authorityName="Harold" authorityYear="1869" box="[737,974,403,429]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFE99A66FD75ECEC4DFFFE56" box="[737,974,403,429]" italics="true" pageId="34" pageNumber="80">Liogenys xanthocera</emphasis>
</taxonomicName>
/ Harold 1869 ³ / DET: A.B. 
<collectionCode id="DBCEAE1FFFE99A66FABDECEC4B0AFE56" box="[1321,1339,403,429]" name="Tavera, Department of Geology and Geophysics" pageId="34" pageNumber="80">T</collectionCode>
. 
<collectionCode id="DBCEAE1FFFE99A66FAAAECEC4BADFE56" box="[1342,1436,403,429]" pageId="34" pageNumber="80">SMITH</collectionCode>
2008”, [red, outlined, typeset and handwritten] “ 
<taxonomicName id="7ADF4D59FFE99A66FD5CECC84D66FE2A" box="[712,855,439,465]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ PALLIDICORNIS / ³ / 
<bibRefCitation id="D94E4B2BFFE99A66FB06ECC84BBBFE2A" author="Blanchard, E." box="[1170,1418,438,465]" pageId="34" pageNumber="80" refId="ref27011" refString="Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp." type="book" year="1851">BLANCHARD, 1851</bibRefCitation>
/ 
<typeStatus id="62648878FFE99A66FF03ECA44F4EFE0E" box="[151,383,475,501]" pageId="34" pageNumber="80" type="paralectotype">PARALECTOTYPE</typeStatus>
/ Des. Cherman M. A.”
</paragraph>
<paragraph id="BD6036DAFFE99A66FF53EC804CB9FD52" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EC804F95FDE3" bold="true" box="[199,420,510,537]" pageId="34" pageNumber="80">Non-type material</emphasis>
(14). 
<materialsCitation id="0DB73C87FFE99A66FE78EC804BA9FDE2" ID-GBIF-Occurrence="3003785305" box="[492,1432,511,537]" collectionCode="ZMHB" country="Argentina" location="Patagonia" pageId="34" pageNumber="80" specimenCount="1" specimenCount-female="1">
<collectingCountry id="C5C8764AFFE99A66FE78EC804CA2FDE2" box="[492,659,511,537]" name="Argentina" pageId="34" pageNumber="80">ARGENTINA</collectingCountry>
. 
<location id="B8006001FFE99A66FD0BEC804D23FDE2" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FD0BEC804D23FDE2" box="[671,786,511,537]" country="Argentina" name="Patagonia" pageId="34" pageNumber="80">Patagonia</location>
: without locality, date, and collector, 
<specimenCount id="ABD9FD53FFE99A66FB50EC804B16FDE2" box="[1220,1319,511,537]" pageId="34" pageNumber="80" type="female">1 female</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FAADEC804BA1FDE2" box="[1337,1424,511,537]" pageId="34" pageNumber="80">ZMHB</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FF03EF5C4DE4FDC6" ID-GBIF-Occurrence="3003785476" box="[151,981,547,573]" collectingDate="1944-12" collectionCode="CMNC" collectorName="Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="34" pageNumber="80" specimenCount="2" specimenCount-male="2" stateProvince="Ciudad Autonoma de Buenos Aires">
<emphasis id="8FABEAC8FFE99A66FF03EF5C4F02FDC6" box="[151,307,547,573]" italics="true" pageId="34" pageNumber="80">
<collectingRegion id="7F1BF838FFE99A66FF03EF5C4F1CFDC6" box="[151,301,547,573]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="34" pageNumber="80">Buenos Aires</collectingRegion>
:
</emphasis>
<collectingMunicipality id="5D04ACA0FFE99A66FEADEF5C4F45FDC6" box="[313,372,547,573]" pageId="34" pageNumber="80">Puán</collectingMunicipality>
, 
<location id="B8006001FFE99A66FEEBEF5C4FCEFDC6" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FEEBEF5C4FCEFDC6" box="[383,511,547,573]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="34" pageNumber="80" stateProvince="Ciudad Autonoma de Buenos Aires">Felipe Solá</location>
, 
<date id="C961101AFFE99A66FD9EEF5C4C42FDC6" box="[522,627,547,573]" pageId="34" pageNumber="80" value="1944-12">
<collectingDate id="D925E9F2FFE99A66FD9EEF5C4C42FDC6" box="[522,627,547,573]" pageId="34" pageNumber="80" value="1944-12">XII.1944</collectingDate>
</date>
, A. 
<collectorName id="102A530CFFE99A66FD0AEF5C4D37FDC6" box="[670,774,547,573]" pageId="34" pageNumber="80">Martínez</collectorName>
, 
<specimenCount id="ABD9FD53FFE99A66FC84EF5C4D57FDC6" box="[784,870,547,573]" pageId="34" pageNumber="80" type="male">2 males</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FCE0EF5C4DFCFDC6" box="[884,973,547,573]" pageId="34" pageNumber="80">CMNC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FC74EF5C4F6CFD9A" ID-GBIF-Occurrence="3003785348" collectingDate="1946-02" collectionCode="II, CMNC" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="34" pageNumber="80" specimenCount="4" specimenCount-male="4">
<collectingMunicipality id="5D04ACA0FFE99A66FC74EF5C4A2AFDC6" box="[992,1051,547,573]" pageId="34" pageNumber="80">Puán</collectingMunicipality>
, 
<location id="B8006001FFE99A66FBB1EF5C4A97FDC6" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FBB1EF5C4A97FDC6" box="[1061,1190,547,573]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="34" pageNumber="80">Felipe Solá</location>
, 
<date id="C961101AFFE99A66FB25EF5C4B34FDC6" box="[1201,1285,547,573]" pageId="34" pageNumber="80" value="1946-02">
<collectingDate id="D925E9F2FFE99A66FB25EF5C4B34FDC6" box="[1201,1285,547,573]" pageId="34" pageNumber="80" value="1946-02">
<collectionCode id="DBCEAE1FFFE99A66FB25EF5C4AF4FDC6" box="[1201,1221,547,573]" pageId="34" pageNumber="80">II</collectionCode>
.1946
</collectingDate>
</date>
, 
<collectorName id="102A530CFFE99A66FA9AEF5C4BA9FDC6" box="[1294,1432,547,573]" pageId="34" pageNumber="80">A. Martínez</collectorName>
, 
<specimenCount id="ABD9FD53FFE99A66FF03EF384EDCFD9A" box="[151,237,583,609]" pageId="34" pageNumber="80" type="male">4 males</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FF68EF384F64FD9A" box="[252,341,583,609]" pageId="34" pageNumber="80">CMNC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FEFCEF384DCCFD9A" ID-GBIF-Occurrence="3003785351" box="[360,1021,582,609]" collectingDate="1951-12" collectionCode="MZSP" collectorName="Felipe Sola, A" country="Argentina" location="Martinez" municipality="Puan" pageId="34" pageNumber="80" specimenCount="3" specimenCount-male="3">
<collectingMunicipality id="5D04ACA0FFE99A66FEFCEF384F92FD9A" box="[360,419,583,609]" pageId="34" pageNumber="80">Puán</collectingMunicipality>
, 
<collectorName id="102A530CFFE99A66FE39EF384C7EFD9A" box="[429,591,583,609]" pageId="34" pageNumber="80">Felipe Solá, A</collectorName>
. 
<location id="B8006001FFE99A66FDCEEF384CF3FD9A" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FDCEEF384CF3FD9A" box="[602,706,583,609]" country="Argentina" municipality="Puan" name="Martinez" pageId="34" pageNumber="80">Martínez</location>
, 
<date id="C961101AFFE99A66FD59EF394D04FD9A" box="[717,821,582,609]" pageId="34" pageNumber="80" value="1951-12">
<collectingDate id="D925E9F2FFE99A66FD59EF394D04FD9A" box="[717,821,582,609]" pageId="34" pageNumber="80" value="1951-12">XII.1951</collectingDate>
</date>
, 
<specimenCount id="ABD9FD53FFE99A66FCD5EF384DA7FD9A" box="[833,918,583,609]" pageId="34" pageNumber="80" type="male">3 males</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FC32EF384DC4FD9A" box="[934,1013,583,609]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="80" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FB9CEF384FBBFD7E" ID-GBIF-Occurrence="3003785453" collectingDate="1959-11" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="34" pageNumber="80" specimenCount="1" specimenCount-male="1">
<collectingMunicipality id="5D04ACA0FFE99A66FB9CEF384A72FD9A" box="[1032,1091,583,609]" pageId="34" pageNumber="80">Puán</collectingMunicipality>
, 
<location id="B8006001FFE99A66FBDAEF384AFFFD9A" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FBDAEF384AFFFD9A" box="[1102,1230,583,609]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="34" pageNumber="80">Felipe Solá</location>
, 
<date id="C961101AFFE99A66FB4EEF394B08FD9A" box="[1242,1337,582,609]" pageId="34" pageNumber="80" value="1959-11">
<collectingDate id="D925E9F2FFE99A66FB4EEF394B08FD9A" box="[1242,1337,582,609]" pageId="34" pageNumber="80" value="1959-11">XI.1959</collectingDate>
</date>
, 
<collectorName id="102A530CFFE99A66FAD7EF384EE1FD7E" pageId="34" pageNumber="80">A. Martínez</collectorName>
, 
<specimenCount id="ABD9FD53FFE99A66FF4FEF144F14FD7E" box="[219,293,619,645]" pageId="34" pageNumber="80" type="male">1 male</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FEA7EF144FB3FD7E" box="[307,386,619,645]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="80" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FE00EF144A12FD7E" ID-GBIF-Occurrence="3003785368" box="[404,1059,619,645]" collectingDate="1960-12" collectionCode="MZSP" collectorName="A. Martinez" country="Argentina" location="Felipe Sola" municipality="Puan" pageId="34" pageNumber="80" specimenCount="2" specimenCount-male="2">
<collectingMunicipality id="5D04ACA0FFE99A66FE00EF144FFEFD7E" box="[404,463,619,645]" pageId="34" pageNumber="80">Puán</collectingMunicipality>
, 
<location id="B8006001FFE99A66FE4DEF144C68FD7E" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FE4DEF144C68FD7E" box="[473,601,619,645]" country="Argentina" municipality="Puan" name="Felipe Sola" pageId="34" pageNumber="80">Felipe Solá</location>
, 
<date id="C961101AFFE99A66FDF0EF144CFCFD7E" box="[612,717,619,645]" pageId="34" pageNumber="80" value="1960-12">
<collectingDate id="D925E9F2FFE99A66FDF0EF144CFCFD7E" box="[612,717,619,645]" pageId="34" pageNumber="80" value="1960-12">XII.1960</collectingDate>
</date>
, 
<collectorName id="102A530CFFE99A66FD41EF144D6EFD7E" box="[725,863,619,645]" pageId="34" pageNumber="80">A. Martínez</collectorName>
, 
<specimenCount id="ABD9FD53FFE99A66FCFDEF144D8FFD7E" box="[873,958,619,645]" pageId="34" pageNumber="80" type="male">2 males</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FC58EF144A2AFD7E" box="[972,1051,619,645]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="34" pageNumber="80" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="0DB73C87FFE99A66FBB9EF144CB5FD52" ID-GBIF-Occurrence="3003785381" collectingDate="1946-11" collectionCode="CMNC" collectorName="A. Martinez" country="Argentina" location="Estancia Barrau" pageId="34" pageNumber="80" specimenCount="1" specimenCount-male="1">
<location id="B8006001FFE99A66FBB9EF144AD0FD7E" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FBB9EF144AD0FD7E" box="[1069,1249,619,645]" country="Argentina" name="Estancia Barrau" pageId="34" pageNumber="80">Estancia Barraú</location>
, 
<locationDeviation id="ACD22FF8FFE99A66FB7FEF144E8EFD52" location="Villa Iris" pageId="34" pageNumber="80">
<quantity id="7A279B3FFFE99A66FB7FEF144B00FD7E" box="[1259,1329,619,645]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="80" unit="km" value="30.0">30 km</quantity>
SW Villa Iris
</locationDeviation>
, 
<date id="C961101AFFE99A66FF5FEFF04F18FD52" box="[203,297,655,681]" pageId="34" pageNumber="80" value="1946-11">
<collectingDate id="D925E9F2FFE99A66FF5FEFF04F18FD52" box="[203,297,655,681]" pageId="34" pageNumber="80" value="1946-11">XI.1946</collectingDate>
</date>
, 
<collectorName id="102A530CFFE99A66FEA0EFF04F8FFD52" box="[308,446,655,681]" pageId="34" pageNumber="80">A. Martínez</collectorName>
, 
<specimenCount id="ABD9FD53FFE99A66FE5DEFF04C25FD52" box="[457,532,655,681]" pageId="34" pageNumber="80" type="male">1 male</specimenCount>
(
<collectionCode id="DBCEAE1FFFE99A66FDB7EFF04C4DFD52" box="[547,636,655,681]" pageId="34" pageNumber="80">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F5C56551FFE99A66FF53EFCD4C4BFC32" pageId="34" pageNumber="80" type="diagnosis">
<paragraph id="BD6036DAFFE99A66FF53EFCD4C4BFC32" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EFCD4F0BFD37" bold="true" box="[199,314,690,716]" pageId="34" pageNumber="80">Diagnosis</emphasis>
. Body elongate; body, head, and pronotum in general dark purple; elytra glabrous, shiny yellow, sides subparallel (
<figureCitation id="25E42A5FFFE99A66FEF1EFA84FFEFD0A" box="[357,463,727,753]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10A</figureCitation>
); clypeal emargination deep, narrow and rounded; clypeal lateral margin convex and produced, obtuse angle between outer side of anterior teeth and clypeal lateral projection (
<figureCitation id="25E42A5FFFE99A66FB04EF844AC6FCEE" box="[1168,1271,763,789]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10C</figureCitation>
); antenna with 10 antennomeres, yellow, club nearly twice as long as the funicle; posterior corners subangulate, obtuse; mesotibia cylindrical in cross section (
<figureCitation id="25E42A5FFFE99A66FE4CEE3C4C73FCA6" box="[472,578,835,861]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10D</figureCitation>
); pygidium slightly convex; wide, subtrapezoidal; pygidial disc glabrous or with unconspicous bristles throughout (
<figureCitation id="25E42A5FFFE99A66FDC5EE184C86FC7A" box="[593,695,871,897]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10E</figureCitation>
); males with inner margin of metatibia not carinate; transverse carina weakly defined; protarsomere II elongate; parameres narrowed at the midline; splitted after the midline; inner margins straight; apex spatulate (
<figureCitation id="25E42A5FFFE99A66FD9DEED04C5CFC32" box="[521,621,943,969]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F5C56551FFE99A66FF53EEAD4B70F8FE" pageId="34" pageNumber="80" type="description">
<paragraph id="BD6036DAFFE99A66FF53EEAD4B70F8FE" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EEAD4F5CFC17" bold="true" box="[199,365,978,1004]" pageId="34" pageNumber="80">Redescription</emphasis>
. Length: 
<quantity id="7A279B3FFFE99A66FE4DEEAC4C52FC16" box="[473,611,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.449999999999999" metricValueMax="7.6" metricValueMin="7.3" pageId="34" pageNumber="80" unit="mm" value="7.449999999999999" valueMax="7.6" valueMin="7.3">7.3–7.6 mm</quantity>
; width: 
<quantity id="7A279B3FFFE99A66FD28EEAC4D77FC16" box="[700,838,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="3.9" metricValueMin="3.8" pageId="34" pageNumber="80" unit="mm" value="3.8499999999999996" valueMax="3.9" valueMin="3.8">3.8–3.9 mm</quantity>
. Dark purple in body, head, and pronotum, elytra dull yellow. 
<emphasis id="8FABEAC8FFE99A66FF7AEE884F1AFBEA" box="[238,299,1015,1041]" italics="true" pageId="34" pageNumber="80">Head</emphasis>
. distance between eyes more than three times the width of one eye; clypeal emargination deep, narrow, and rounded; outer sides of anterior teeth parallel; clypeal lateral margin convex and produced, forming a rounded projection; distance between clypeal lateral projection and anterior tooth longer than the basal width of anterior tooth, distance between clypeal lateral projection and anterior margin of eye longer than one eye; obtuse angle between outer side of anterior teeth and clypeal lateral projection; antenna with 10 antennomeres, club nearly twice as long as and lighter in color than funicle. 
<emphasis id="8FABEAC8FFE99A66FDCEE9D44C9AFB3E" box="[602,683,1195,1221]" italics="true" pageId="34" pageNumber="80">Thorax</emphasis>
: anterior margin of pronotum straight; pronotal disc with punctures coarse, shallow and sparse; posterior corners subangulate, obtuse; hypomere, mesepisternum, sides of metaventrite, and metacoxae with sparse, long bristles; distance between mesocoxae and metacoxae less than twice the metacoxa width; scutellum ogival to rounded, punctate throughout. 
<emphasis id="8FABEAC8FFE99A66FC8AE8684D55FACA" box="[798,868,1303,1329]" italics="true" pageId="34" pageNumber="80">Elytra</emphasis>
: shiny, glabrous; elytral suture and elytron unicolored, elevated; two inner pair of elytral ridges more defined than the other two ridges. 
<emphasis id="8FABEAC8FFE99A66FBF1E8444AABFAAF" box="[1125,1178,1339,1364]" italics="true" pageId="34" pageNumber="80">Legs</emphasis>
: procoxa with bristles; protibia with three teeth, middle and apical teeth equal in size, all three teeth equally spaced; mesotibia cylindrical in cross section, mesotibia dorsally smooth with two transverse carinae, the apical carina complete; inner margin of male metatibia not carinate; surface finely sculptured; metatibia with one transverse carina present posteriorly and weakly defined dorsally; metatibial apical spurs of equal lengths, equal to the diameter of the tibial apex; metatarsomere I shorter than metatarsomere II; in males protarsomeres and mesotarsomeres I–IV enlarged, protarsomere II long; protarsomeres and mesotarsomeres equally enlarged; claw bifid, symmetrical, superior tooth of claw almost three times longer and narrower than the inferior tooth; distance between teeth longer than the length of inferior tooth. 
<emphasis id="8FABEAC8FFE99A66FF75EB244F7AF98E" box="[225,331,1627,1653]" italics="true" pageId="34" pageNumber="80">Abdomen</emphasis>
: ventrites with sparse bristles; propygidium glabrous; pygidium slightly convex; subtrapezoidal; pygidial width not exceeding distance between spiracles of propygidium; pygidial disc with inconspicuous bristles, finely and very sparsely punctate; pygidial apex subquadrate. 
<emphasis id="8FABEAC8FFE99A66FCC5EBDC4DFAF947" box="[849,971,1699,1724]" italics="true" pageId="34" pageNumber="80">Parameres</emphasis>
: basal region wider than both sections of the parameres at its maximum width, parameres narrowed at the midline; parameral split after the midline; inner margins straight; apex spatulate, angulate; parameres straight or slightly convex in lateral view (
<figureCitation id="25E42A5FFFE99A66FB58EB944B05F8FE" box="[1228,1332,1771,1797]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="34" pageNumber="80">Fig. 10G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F5C56551FFE99A66FF53EA704D54F889" pageId="34" pageNumber="80" type="materials_examined">
<paragraph id="BD6036DAFFE99A66FF53EA704F1CF8B6" blockId="34.[151,1437,150,2013]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EA704F52F8D3" bold="true" box="[199,355,1806,1833]" pageId="34" pageNumber="80">Type locality</emphasis>
. 
<materialsCitation id="0DB73C87FFE99A66FEE7EA704F18F8B6" ID-GBIF-Occurrence="3003785311" collectorName="de San-Blas" country="Argentina" latitude="-40.552223" location="Baie de San-Blas" longLatPrecision="19" longitude="-62.234444" pageId="34" pageNumber="80" specimenCount="1" stateProvince="Ciudad Autonoma de Buenos Aires">
<collectingCountry id="C5C8764AFFE99A66FEE7EA704C2BF8D2" box="[371,538,1807,1833]" name="Argentina" pageId="34" pageNumber="80">ARGENTINA</collectingCountry>
. 
<location id="B8006001FFE99A66FDBCEA704CDEF8D2" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFE99A66FDBCEA704CDEF8D2" box="[552,751,1807,1833]" country="Argentina" latitude="-40.552223" longLatPrecision="19" longitude="-62.234444" name="Baie de San-Blas" pageId="34" pageNumber="80" stateProvince="Ciudad Autonoma de Buenos Aires">
Baie 
<collectorName id="102A530CFFE99A66FDF2EA704CDEF8D2" box="[614,751,1807,1833]" pageId="34" pageNumber="80">de San-Blas</collectorName>
</location>
(Patagonie) [
<collectingRegion id="7F1BF838FFE99A66FC18EA704A1BF8D2" box="[908,1066,1807,1833]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="34" pageNumber="80">Buenos Aires</collectingRegion>
, Bahia de San Blas, 
<geoCoordinate id="D8EB501DFFE99A66FAB6EA704BADF8D2" box="[1314,1436,1807,1833]" degrees="40" direction="south" minutes="33" orientation="latitude" pageId="34" pageNumber="80" precision="15" seconds="8" value="-40.552223">40°33’8’’S</geoCoordinate>
<geoCoordinate id="D8EB501DFFE99A66FF03EA4D4F13F8B6" box="[151,290,1842,1869]" degrees="62" direction="west" minutes="14" orientation="longitude" pageId="34" pageNumber="80" precision="15" seconds="4" value="-62.234444">62°14’4’’W</geoCoordinate>
]
</materialsCitation>
.
</paragraph>
<paragraph id="BD6036DAFFE99A66FF53EA294D54F889" blockId="34.[151,1437,150,2013]" box="[199,869,1878,1906]" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EA294FCFF88B" bold="true" box="[199,510,1878,1904]" pageId="34" pageNumber="80">Geographical distribution</emphasis>
. 
<collectingCountry id="C5C8764AFFE99A66FD98EA284C9FF88A" box="[524,686,1879,1905]" name="Argentina" pageId="34" pageNumber="80">ARGENTINA</collectingCountry>
(
<collectingRegion id="7F1BF838FFE99A66FD28EA284D68F889" box="[700,857,1879,1906]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="34" pageNumber="80">Buenos Aires</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="F5C56551FFE99A60FF53EA044F67FDE2" lastPageId="36" lastPageNumber="82" pageId="34" pageNumber="80" type="discussion">
<paragraph id="BD6036DAFFE99A67FF53EA044DE9FEE6" blockId="34.[151,1437,150,2013]" lastBlockId="35.[151,1437,150,285]" lastPageId="35" lastPageNumber="81" pageId="34" pageNumber="80">
<emphasis id="8FABEAC8FFE99A66FF53EA044F05F86E" bold="true" box="[199,308,1915,1941]" pageId="34" pageNumber="80">Remarks</emphasis>
. 
<taxonomicName id="7ADF4D59FFE99A66FED6EA044C1CF86E" authorityName="Harold" authorityYear="1869" box="[322,557,1915,1941]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFE99A66FED6EA044C1CF86E" box="[322,557,1915,1941]" italics="true" pageId="34" pageNumber="80">Liogenys xanthocera</emphasis>
</taxonomicName>
is easily recognized among the other 
<taxonomicName id="7ADF4D59FFE99A66FC43EA044A0DF86F" box="[983,1084,1915,1940]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="genus">
<emphasis id="8FABEAC8FFE99A66FC43EA044A0DF86F" box="[983,1084,1915,1940]" italics="true" pageId="34" pageNumber="80">Liogenys</emphasis>
</taxonomicName>
by its characteristic dark body, pronotum, and scutellum contrasting with the elytra dull yellow; and by the club nearly twice the length of the funicle. The teeth of the galea are reduced (
<figureCitation id="25E42A5FFFE99A66FDEBEABC4CD3F826" box="[639,738,1987,2013]" captionStart="FIGURE 13" captionStartId="32.[151,250,1909,1933]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure-19@32.[214,1373,181,1885]" captionTargetPageId="32" captionText="FIGURE 13. Liogenys maxillaricuspis Cherman, new species: A) head in ventral view, C) labium, G) right maxilla; L. brachyclypeata Cherman, new species: B) head in ventral view, D) labium; L. calcarata Frey, 1970: E) left maxilla; L. xanthocera Harold, 1869: F) right maxilla. Legends: A, B) CLP: clypeus, LBM: labium, LBR: labrum, MXL: maxilla, MXLP: maxillary palp. C, D) LGL: ligula, LPLP: labial palp, MTM: mentum, PMTM: postmentum." figureDoi="http://doi.org/10.5281/zenodo.4359222" httpUri="https://zenodo.org/record/4359222/files/figure.png" pageId="34" pageNumber="80">Fig. 13F</figureCitation>
), as in 
<taxonomicName id="7ADF4D59FFE99A66FCA7EABC4A2FF826" authority="Moser" authorityName="Moser" authorityYear="1918" box="[819,1054,1987,2013]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="80" phylum="Arthropoda" rank="species" species="sinuaticeps">
<emphasis id="8FABEAC8FFE99A66FCA7EABC4DFEF827" box="[819,975,1987,2012]" italics="true" pageId="34" pageNumber="80">L. sinuaticeps</emphasis>
Moser
</taxonomicName>
(
<bibRefCitation id="D94E4B2BFFE99A66FBB8EABC4B27F826" author="Cherman, M. A. &amp; Moron, M. A. &amp; Almeida, L. M." box="[1068,1302,1987,2013]" pageId="34" pageNumber="80" pagination="744 - 770" refId="ref27371" refString="Cherman, M. A., Moron, M. A. &amp; Almeida, L. M. (2016) Phylogenetic relationships within Diplotaxini Kirby (Coleoptera: Melolonthidae: Melolonthinae) with emphasis on Liogenys Guerin-Meneville. Systematic Entomology, 41, 744 - 770. https: // doi. org / 10.1111 / syen. 12188" type="journal article" year="2016">
Cherman 
<emphasis id="8FABEAC8FFE99A66FB08EABC4AE5F826" box="[1180,1236,1987,2013]" italics="true" pageId="34" pageNumber="80">et al.</emphasis>
2016
</bibRefCitation>
, 
<bibRefCitation id="D94E4B2BFFE99A66FAB6EABC4B6EF826" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[1314,1375,1987,2013]" pageId="34" pageNumber="80" pagination="1 - 120" refId="ref27290" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">2017</bibRefCitation>
). The specimen from 
<quantity id="7A279B3FFFE89A67FEDDEDE84FA3FF4A" box="[329,402,151,177]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="81" unit="km" value="30.0">30 km</quantity>
SW Villa Iris, Puán, 
<collectingRegion id="7F1BF838FFE89A67FD1DEDE84D13FF4A" box="[649,802,151,177]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="35" pageNumber="81">Buenos Aires</collectingRegion>
[
<geoCoordinate id="D8EB501DFFE89A67FCA6EDE94DE0FF4A" box="[818,977,150,177]" degrees="38" direction="south" minutes="21" orientation="latitude" pageId="35" pageNumber="81" precision="1" seconds="15.8" value="-38.35439">38°21’15.8’’S</geoCoordinate>
<geoCoordinate id="D8EB501DFFE89A67FC4EEDE94AB4FF4A" box="[986,1157,150,177]" degrees="63" direction="west" minutes="20" orientation="longitude" pageId="35" pageNumber="81" precision="1" seconds="37.4" value="-63.34372">63°20’37.4’’W</geoCoordinate>
] shows variations in the body color, entirely dull yellow, the lateral margin of clypeus more produced, and the parameres strongly convex. As we have only this specimen with these single features, we presume that those differences are only intraspecific variations, needing further material to determine if it is a different species.
</paragraph>
<caption id="E9A06652FFE89A67FF03EAEC4F0FF808" ID-DOI="http://doi.org/10.5281/zenodo.4359228" ID-Zenodo-Dep="4359228" httpUri="https://zenodo.org/record/4359228/files/figure.png" pageId="35" pageNumber="81" startId="35.[151,250,1939,1963]" targetBox="[187,1409,323,1905]" targetPageId="35">
<paragraph id="BD6036DAFFE89A67FF03EAEC4F0FF808" blockId="35.[151,1437,1939,2035]" pageId="35" pageNumber="81">
<emphasis id="8FABEAC8FFE89A67FF03EAEC4F13F850" bold="true" box="[151,290,1939,1963]" pageId="35" pageNumber="81">FIGURE 15.</emphasis>
Geographical distribution of 
<taxonomicName id="7ADF4D59FFE89A67FDC1EAEB4C81F850" authority="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida, 2020" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" box="[597,688,1940,1963]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="81" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="8FABEAC8FFE89A67FDC1EAEB4C81F850" box="[597,688,1940,1963]" italics="true" pageId="35" pageNumber="81">Liogenys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="949857B3FFE89A67FD2CEAEB4D05F850" box="[696,820,1939,1963]" pageId="35" pageNumber="81" rank="species">new species</taxonomicNameLabel>
occurring in the STZ and its boundaries: A) general, B, C) detail. Image legend: shaped points: new species; colors: provinces of Neotropical, South American Transition Zone (STZ), and Andean regions.
</paragraph>
</caption>
<paragraph id="BD6036DAFFEF9A60FF53EDE84CDEFE72" blockId="36.[151,1436,151,537]" pageId="36" pageNumber="82">
<materialsCitation id="0DB73C87FFEF9A60FF53EDE84CDAFE72" ID-GBIF-Occurrence="3003785306" collectingDate="1829-01-14" collectingDateMax="1829-02-18" collectingDateMin="1829-01-14" collectionCode="MNHN" collectorName="A. D. d'Orbigny. The" country="Argentina" county="d'Orbigny. The region" location="The" municipality="Bahia San Blas" pageId="36" pageNumber="82" specimenCount="2" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="lectotype">
<location id="B8006001FFEF9A60FF53EDE84EC2FF4A" LSID="urn:lsid:plazi:treatment:357687CCFFD79A60FF03EBDC4F67FDE2:B8006001FFEF9A60FF53EDE84EC2FF4A" box="[199,243,151,177]" country="Argentina" county="d'Orbigny. The region" municipality="Bahia San Blas" name="The" pageId="36" pageNumber="82" stateProvince="Ciudad Autonoma de Buenos Aires">The</location>
<typeStatus id="62648878FFEF9A60FF68EDE84F52FF4A" box="[252,355,151,177]" pageId="36" pageNumber="82" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="25E42A5FFFEF9A60FEE1EDE84FEAFF4A" box="[373,475,151,177]" captionStart="FIGURE 10" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-20@29.[203,1385,181,1920]" captionTargetPageId="29" captionText="FIGURE 10. Liogenys xanthocera Harold, 1869, male lectotype views: A) dorsal, B) labels, C) clypeus and pronotum, D) lateral, E) pygidium, and parameres views: F) dorsal, G) lateral." figureDoi="http://doi.org/10.5281/zenodo.4359210" httpUri="https://zenodo.org/record/4359210/files/figure.png" pageId="36" pageNumber="82">Fig. 10B</figureCitation>
) was collected 
<collectingMunicipality id="5D04ACA0FFEF9A60FD04EDE84D72FF4A" box="[656,835,151,177]" pageId="36" pageNumber="82">Bahía San Blas</collectingMunicipality>
, 
<collectingRegion id="7F1BF838FFEF9A60FCC5EDE84DDDFF4A" box="[849,1004,151,177]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="36" pageNumber="82">Buenos Aires</collectingRegion>
, 
<collectingCountry id="C5C8764AFFEF9A60FC6CEDE84A56FF4A" box="[1016,1127,151,177]" name="Argentina" pageId="36" pageNumber="82">Argentina</collectingCountry>
during the period from 
<date id="C961101AFFEF9A60FA14EDE84FF3FF2E" pageId="36" pageNumber="82" value="1829-01-14" valueMax="1829-02-18" valueMin="1829-01-14">
<collectingDate id="D925E9F2FFEF9A60FA14EDE84FF3FF2E" pageId="36" pageNumber="82" value="1829-01-14" valueMax="1829-02-18" valueMin="1829-01-14">14 January–18 February 1829</collectingDate>
</date>
by 
<collectorName id="102A530CFFEF9A60FE7CEDC44CE0FF2E" box="[488,721,187,213]" pageId="36" pageNumber="82">
A.D. 
<collectingCounty id="54014E56FFEF9A60FDB1EDC44CE0FF2E" box="[549,721,187,213]" pageId="36" pageNumber="82">d’Orbigny. The</collectingCounty>
</collectorName>
region around the nearby village of “Patagones” (now known as Viedma, 
<collectingRegion id="7F1BF838FFEF9A60FF69EDA04F44FF02" box="[253,373,223,249]" country="Argentina" name="Rio Negro" pageId="36" pageNumber="82">Río Negro</collectingRegion>
) was collectively called Patagonia (Patagonie) at the time (see 
<bibRefCitation id="D94E4B2BFFEF9A60FBD5EDA04AF8FF02" author="Smith, A. B. T." box="[1089,1225,223,249]" pageId="36" pageNumber="82" pagination="81 - 103" refId="ref28536" refString="Smith, A. B. T. (2017) Southern South American Trogidae (Coleoptera): verification of type specimens and notes on geographic distribution. Revista Chilena de Entomologia, 43, 81 - 103." type="journal article" year="2017">Smith 2017</bibRefCitation>
). The “1052” code number on the 
<typeStatus id="62648878FFEF9A60FED5EC7C4FE9FEE6" box="[321,472,259,285]" pageId="36" pageNumber="82" type="paralectotype">paralectotype</typeStatus>
corresponds with an entry in the original field notes of d’Orbigny (available from the 
<collectionCode id="DBCEAE1FFFEF9A60FF03EC584EDCFEBA" box="[151,237,295,321]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="36" pageNumber="82">MNHN</collectionCode>
library). D’Orbigny made these observations (text transcribed and translated): “we found this species in great numbers in January and February. They emerged from the earth every day, in numbers. We never saw them during the day, when they were hiding underground”
</materialsCitation>
.
</paragraph>
<paragraph id="BD6036DAFFEF9A60FF53ECEC4F67FDE2" blockId="36.[151,1436,151,537]" pageId="36" pageNumber="82">
<bibRefCitation id="D94E4B2BFFEF9A60FF53ECEC4FBDFE56" author="Blanchard, E." box="[199,396,402,429]" pageId="36" pageNumber="82" refId="ref27011" refString="Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp." type="book" year="1851">Blanchard (1851)</bibRefCitation>
described two species in 
<taxonomicName id="7ADF4D59FFEF9A60FD3BECEC4D25FE57" authorityName="Cherman &amp; Basílio &amp; Mise &amp; Frisch &amp; Smith &amp; Almeida" authorityYear="2020" box="[687,788,403,428]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="82" phylum="Arthropoda" rank="genus">
<emphasis id="8FABEAC8FFEF9A60FD3BECEC4D25FE57" box="[687,788,403,428]" italics="true" pageId="36" pageNumber="82">Liogenys</emphasis>
</taxonomicName>
with the epithet “ 
<taxonomicName id="7ADF4D59FFEF9A60FC4BECEC4A5FFE56" authorityName="Blanchard" authorityYear="1851" box="[991,1134,403,429]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="82" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="8FABEAC8FFEF9A60FC4BECEC4A5FFE56" box="[991,1134,403,429]" italics="true" pageId="36" pageNumber="82">pallidicornis</emphasis>
</taxonomicName>
”, the first one in page 167 and the other in the next page. Harold (1869) corrected this homonym by replacing the name of the second one described, as 
<taxonomicName id="7ADF4D59FFEF9A60FEBAECA44FF8FE0E" authorityName="Harold" authorityYear="1869" box="[302,457,475,501]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="82" phylum="Arthropoda" rank="species" species="xanthocera">
<emphasis id="8FABEAC8FFEF9A60FEBAECA44FF8FE0E" box="[302,457,475,501]" italics="true" pageId="36" pageNumber="82">L. xanthocera</emphasis>
</taxonomicName>
. The first one described, 
<taxonomicName id="7ADF4D59FFEF9A60FD70ECA44AAEFE0E" authority="Blanchard, 1851" authorityName="Blanchard" authorityYear="1851" box="[740,1183,474,501]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="82" phylum="Arthropoda" rank="species" species="pallidicornis">
<emphasis id="8FABEAC8FFEF9A60FD70ECA44DEEFE0E" box="[740,991,475,501]" italics="true" pageId="36" pageNumber="82">Liogenys pallidicornis</emphasis>
<bibRefCitation id="D94E4B2BFFEF9A60FC72ECA44AAEFE0E" author="Blanchard, E." box="[998,1183,474,501]" pageId="36" pageNumber="82" refId="ref27011" refString="Blanchard, E. (1851) Museum d'Histoire Naturelle de Paris. Catalogue de la Collection Entomologique. Classe des Insectes. Ordre des Coleopteres. Tome I. Gide et Baudry, Paris, 240 pp." type="book" year="1851">Blanchard, 1851</bibRefCitation>
</taxonomicName>
is a valid name (
<bibRefCitation id="D94E4B2BFFEF9A60FACFECA44F7BFDE2" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." pageId="36" pageNumber="82" pagination="1 - 120" refId="ref27290" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman 
<emphasis id="8FABEAC8FFEF9A60FF5AEC804F31FDE2" box="[206,256,511,537]" italics="true" pageId="36" pageNumber="82">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>