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<mods:title id="3B5A6521C37E0647202BFBC865F23345">On the Lomechusini fauna of the Palaearctic and Oriental regions. XXVI. New species, a new synonymy, and additional records (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="8AAC905E81F19B91A8B887204AE13DEC">Assing, Volker</mods:namePart>
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<taxonomicName id="4C474D30FFEF3B68FC95FED473459940" ID-CoL="8Y7D2" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[854,1045,266,294]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="species" species="taurus" status="sp. nov.">
<emphasis id="B933EAA1FFEF3B68FC95FED473459940" box="[854,1045,266,294]" italics="true" pageId="15" pageNumber="48">Orphnebius taurus</emphasis>
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<paragraph id="8BF836B3FFEF3B68FC95FE8F748A9903" blockId="15.[854,1383,266,357]" box="[854,986,337,357]" pageId="15" pageNumber="48">
(
<figureCitation id="137C2A36FFEF3B68FC99FE8F74CE9903" box="[858,926,337,357]" captionStart="Figs 1–15" captionStartId="5.[174,214,2002,2024]" captionTargetBox="[174,1419,255,1990]" captionTargetId="figure-112@5.[173,1421,254,1994]" captionTargetPageId="5" captionText="Figs 1–15:Myrmoecia urartu (1), Pella feldmanni (2), Orphnebius taurus (3–5), O. fortesetosus (6–7), O. morulus (8–9), O. globifer (10–11), andO. elevatus (12–15): habitus (1–3); head and pronotum (4–5); forebody (6, 8, 10, 12–13); abdomen (7, 9, 11); anterolateral portion of abdomen (14); tergites VII–VIII (15). Scale bars: 1–3: 1.0 mm; 4–15: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742770" httpUri="https://zenodo.org/record/5742770/files/figure.png" pageId="15" pageNumber="48">Figs 3–5</figureCitation>
, 
<figureCitation id="137C2A36FFEF3B68FC66FE8F74869903" box="[933,982,337,357]" captionStart="Figs 79–95" captionStartId="10.[212,252,2002,2024]" captionTargetBox="[215,1459,258,1989]" captionTargetId="figure-121@10.[211,1459,254,1994]" captionTargetPageId="10" captionText="Figs 79–95: Myrmoecia urartu (79), Pella feldmanni (80–83), Orphnebius cachemiricus (84–86), O. fortesetosus (87–90), and O. taurus(91–95): spermatheca (79); median lobe of aedeagus in lateral and in ventral view (80–81, 84–85, 87–88, 92–93); male tergite VIII (82, 91); male sternite VIII (83); paramere (86, 89, 95); apical portion of paramere (90); apical portion of median lobe of aedeagus (94). Scale bars: 79–93, 95: 0.2 mm; 94: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742775" httpUri="https://zenodo.org/record/5742775/files/figure.png" pageId="15" pageNumber="48">91–95</figureCitation>
)
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<subSubSection id="C35D6538FFEF3B68FCF3FE5E7221999E" pageId="15" pageNumber="48" type="materials_examined">
<paragraph id="8BF836B3FFEF3B68FCF3FE5E7221999E" blockId="15.[816,1428,383,504]" pageId="15" pageNumber="48">
<emphasis id="B933EAA1FFEF3B68FCF3FE5E749A99F1" bold="true" box="[816,970,383,408]" pageId="15" pageNumber="48">Type material</emphasis>
: 
<materialsCitation id="3B2F3CEEFFEF3B68FC18FE5E7221999E" collectingDate="2005-05-03" collectingDateMax="2005-05-19" collectingDateMin="2005-05-03" collectionCode="V, MMB" collectorName="E. Jendek &amp; O. Sausa" country="Cambodia" elevation="175" latitude="11.566667" location="Tatai river" longLatPrecision="1295" longitude="103.11667" pageId="15" pageNumber="48" specimenCount="2" stateProvince="Koh Kong" typeStatus="holotype">
<typeStatus id="54FC8811FFEF3B68FC18FE5E731499FE" box="[987,1092,384,408]" pageId="15" pageNumber="48" type="holotype">Holotype</typeStatus>
: “SW 
<collectingCountry id="F3507623FFEF3B68FB65FE5E724899FE" box="[1190,1304,384,408]" name="Cambodia" pageId="15" pageNumber="48">Cambodia</collectingCountry>
, 
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SE 
<collectingRegion id="4983F851FFEF3B68FCF3FE7E74F199DE" box="[816,929,416,440]" country="Cambodia" name="Koh Kong" pageId="15" pageNumber="48">Koh Kong</collectingRegion>
, 
<quantity id="4CBF9B56FFEF3B68FC6EFE7E734999D1" box="[941,1049,416,440]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="15" pageNumber="48" unit="m" value="175.0" valueMax="300.0" valueMin="50.0">
<elevation id="006AD180FFEF3B68FC6EFE7E734999D1" box="[941,1049,416,440]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="15" pageNumber="48" unit="m" value="175.0" valueMax="300.0" valueMin="50.0">50–300 m</elevation>
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, 
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<collectingDate id="EFBDE99BFFEF3B68FBE6FE7E73FB99DE" box="[1061,1195,416,440]" pageId="15" pageNumber="48" value="2005-05-03" valueMax="2005-05-19" valueMin="2005-05-03">3.–19.v.2005</collectingDate>
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, 
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, 
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, 
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, 
<collectorName id="26B25365FFEF3B68FC60FE1E734199BE" box="[931,1041,448,472]" pageId="15" pageNumber="48">E. Jendek</collectorName>
&amp; 
<collectorName id="26B25365FFEF3B68FB80FE1E73CE99BF" box="[1091,1182,444,473]" pageId="15" pageNumber="48">O. Šauša</collectorName>
leg. / 
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<taxonomicName id="4C474D30FFEF3B68FCF3FE3E74AB9991" authority="Assing, 2019" authorityName="Assing" authorityYear="2019" box="[816,1019,480,504]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="48" phylum="Arthropoda" rank="species" species="taurus" status="sp. nov.">
<emphasis id="B933EAA1FFEF3B68FCF3FE3E74AB9991" box="[816,1019,480,504]" italics="true" pageId="15" pageNumber="48">Orphnebius taurus</emphasis>
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, det. 
<collectionCode id="ED56AE76FFEF3B68FBAFFE3E732E9991" box="[1132,1150,480,503]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="15" pageNumber="48" type="Museum">V</collectionCode>
. Assing 2018” (
<collectionCode id="ED56AE76FFEF3B68FAE3FE3E7235999E" box="[1312,1381,480,504]" country="Czech Republic" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="15" pageNumber="48">MMB</collectionCode>
).
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</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFEF3B68FCF3FDFE73039A1E" pageId="15" pageNumber="48" type="etymology">
<paragraph id="8BF836B3FFEF3B68FCF3FDFE73039A1E" blockId="15.[816,1428,544,632]" pageId="15" pageNumber="48">
<emphasis id="B933EAA1FFEF3B68FCF3FDFE74F79A5E" bold="true" box="[816,935,544,568]" pageId="15" pageNumber="48">Etymology</emphasis>
: The specific epithet (Latin: bull) is a noun in apposition and alludes to the two long horn-shaped processes of the mandibles.
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFEF3B77FCF3FD417558999E" lastPageId="16" lastPageNumber="49" pageId="15" pageNumber="48" type="description">
<paragraph id="8BF836B3FFEF3B68FCF3FD4172499BFE" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">
<emphasis id="B933EAA1FFEF3B68FCF3FD4174E29AD1" bold="true" box="[816,946,671,695]" pageId="15" pageNumber="48">Description</emphasis>
: Small species; body length 3.0 mm; length of forebody 
<quantity id="4CBF9B56FFEF3B68FC7DFD1E73429AB1" box="[958,1042,704,728]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="48" unit="mm" value="1.4">1.4 mm</quantity>
. Habitus as in 
<figureCitation id="137C2A36FFEF3B68FB05FD1E72539ABE" box="[1222,1283,704,728]" captionStart="Figs 1–15" captionStartId="5.[174,214,2002,2024]" captionTargetBox="[174,1419,255,1990]" captionTargetId="figure-112@5.[173,1421,254,1994]" captionTargetPageId="5" captionText="Figs 1–15:Myrmoecia urartu (1), Pella feldmanni (2), Orphnebius taurus (3–5), O. fortesetosus (6–7), O. morulus (8–9), O. globifer (10–11), andO. elevatus (12–15): habitus (1–3); head and pronotum (4–5); forebody (6, 8, 10, 12–13); abdomen (7, 9, 11); anterolateral portion of abdomen (14); tergites VII–VIII (15). Scale bars: 1–3: 1.0 mm; 4–15: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742770" httpUri="https://zenodo.org/record/5742770/files/figure.png" pageId="15" pageNumber="48">Fig. 3</figureCitation>
. Coloration: head blackish with the anterior portion brown; pronotum pitchy-black; elytra yellowish-brown with the scutellar portion diffusely darker; abdomen pale yellowish-brown; legs yellowish-brown with dark-yellowish tarsi; antennae dark-brown with antennomeres II–V reddish; maxillary palpi brown with the apical palpomere yellow.
</paragraph>
<paragraph id="8BF836B3FFEF3B68FCF3FC7E723F9D31" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">
Head (
<figureCitation id="137C2A36FFEF3B68FCBEFC7E748A9BDE" box="[893,986,928,952]" captionStart="Figs 1–15" captionStartId="5.[174,214,2002,2024]" captionTargetBox="[174,1419,255,1990]" captionTargetId="figure-112@5.[173,1421,254,1994]" captionTargetPageId="5" captionText="Figs 1–15:Myrmoecia urartu (1), Pella feldmanni (2), Orphnebius taurus (3–5), O. fortesetosus (6–7), O. morulus (8–9), O. globifer (10–11), andO. elevatus (12–15): habitus (1–3); head and pronotum (4–5); forebody (6, 8, 10, 12–13); abdomen (7, 9, 11); anterolateral portion of abdomen (14); tergites VII–VIII (15). Scale bars: 1–3: 1.0 mm; 4–15: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742770" httpUri="https://zenodo.org/record/5742770/files/figure.png" pageId="15" pageNumber="48">Figs 4–5</figureCitation>
) strongly transverse, in the middle of dorsal surface with pronounced tubercle-shaped elevation; both mandibles with a long and erect horn-shaped process originating from their bases (male secondary sexual character?). Eyes very large, longer than distance from posterior margin of eye to posterior constriction of head. Antenna approximately 1.0 mm long, distinctly incrassate apically, and somewhat clavate, with antennomeres VI–X much larger than antennomeres II–V; antennomere I large, long, and distinctly compressed, nearly as long as the combined length of antennomeres II–V; antennomeres VI–X strongly transverse and distinctly asymmetric; antennomere XI approximately as long as the combined length of antennomeres VIII–X.
</paragraph>
<paragraph id="8BF836B3FFEF3B68FCF3FABE74E79E71" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">Pronotum strongly transverse and rather small in relation to head, approximately 1.65 times as broad as long and approximately as broad as head, moderately convex in cross-section; posterior angles obsolete; disc impunctate; lateral margins each with three long and stout darkbrown setae.</paragraph>
<paragraph id="8BF836B3FFEF3B68FCF3F9FE721F9EF1" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">Elytra approximately 0.8 times as long as pronotum, strongly widened posteriad; punctation sparse and fine. Hind wings fully developed. Metatarsomere I approximately as long as the combined length of II and III.</paragraph>
<paragraph id="8BF836B3FFEF3B68FCF3F97E723B9F31" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">Abdomen broad, gradually tapering from base towards apex; segments III–VI with broad and dorsally truncate paratergites; tergites III–VI practically impunctate; tergite VII with dense and long striae except near anterior margin; tergite VIII laterally with a cluster of dense thin setae on either side, posterior margin broadly convex.</paragraph>
<paragraph id="8BF836B3FFEF3B68FC86F8BE74E8907E" blockId="15.[816,1428,671,2104]" pageId="15" pageNumber="48">
: hemi-tergites IX and tergite X with dense and long pubescence; sternite VIII as in 
<figureCitation id="137C2A36FFEF3B68FB4FF85E73859FFE" box="[1164,1237,1920,1944]" captionStart="Figs 79–95" captionStartId="10.[212,252,2002,2024]" captionTargetBox="[215,1459,258,1989]" captionTargetId="figure-121@10.[211,1459,254,1994]" captionTargetPageId="10" captionText="Figs 79–95: Myrmoecia urartu (79), Pella feldmanni (80–83), Orphnebius cachemiricus (84–86), O. fortesetosus (87–90), and O. taurus(91–95): spermatheca (79); median lobe of aedeagus in lateral and in ventral view (80–81, 84–85, 87–88, 92–93); male tergite VIII (82, 91); male sternite VIII (83); paramere (86, 89, 95); apical portion of paramere (90); apical portion of median lobe of aedeagus (94). Scale bars: 79–93, 95: 0.2 mm; 94: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742775" httpUri="https://zenodo.org/record/5742775/files/figure.png" pageId="15" pageNumber="48">Fig. 91</figureCitation>
; median lobe of aedeagus (
<figureCitation id="137C2A36FFEF3B68FC61F87E73499FDE" box="[930,1049,1952,1976]" captionStart="Figs 79–95" captionStartId="10.[212,252,2002,2024]" captionTargetBox="[215,1459,258,1989]" captionTargetId="figure-121@10.[211,1459,254,1994]" captionTargetPageId="10" captionText="Figs 79–95: Myrmoecia urartu (79), Pella feldmanni (80–83), Orphnebius cachemiricus (84–86), O. fortesetosus (87–90), and O. taurus(91–95): spermatheca (79); median lobe of aedeagus in lateral and in ventral view (80–81, 84–85, 87–88, 92–93); male tergite VIII (82, 91); male sternite VIII (83); paramere (86, 89, 95); apical portion of paramere (90); apical portion of median lobe of aedeagus (94). Scale bars: 79–93, 95: 0.2 mm; 94: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742775" httpUri="https://zenodo.org/record/5742775/files/figure.png" pageId="15" pageNumber="48">Figs 92–94</figureCitation>
) 
<quantity id="4CBF9B56FFEF3B68FBE8F87E73D99FD1" box="[1067,1161,1952,1976]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="48" unit="mm" value="0.65">0.65 mm</quantity>
long and of distinctive shape; ventral process apically acute and conspicuously structured; paramere (
<figureCitation id="137C2A36FFEF3B68FBDCF83E733A9F9E" box="[1055,1130,2016,2040]" captionStart="Figs 79–95" captionStartId="10.[212,252,2002,2024]" captionTargetBox="[215,1459,258,1989]" captionTargetId="figure-121@10.[211,1459,254,1994]" captionTargetPageId="10" captionText="Figs 79–95: Myrmoecia urartu (79), Pella feldmanni (80–83), Orphnebius cachemiricus (84–86), O. fortesetosus (87–90), and O. taurus(91–95): spermatheca (79); median lobe of aedeagus in lateral and in ventral view (80–81, 84–85, 87–88, 92–93); male tergite VIII (82, 91); male sternite VIII (83); paramere (86, 89, 95); apical portion of paramere (90); apical portion of median lobe of aedeagus (94). Scale bars: 79–93, 95: 0.2 mm; 94: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742775" httpUri="https://zenodo.org/record/5742775/files/figure.png" pageId="15" pageNumber="48">Fig. 95</figureCitation>
) 
<quantity id="4CBF9B56FFEF3B68FBBBF83E73879F91" box="[1144,1239,2016,2040]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="15" pageNumber="48" unit="mm" value="0.57">0.57 mm</quantity>
long, paramerite very slender.
</paragraph>
<paragraph id="8BF836B3FFEF3B68FC86F7FE74EF905E" blockId="15.[816,1428,671,2104]" box="[837,959,2080,2104]" pageId="15" pageNumber="48">: unknown.</paragraph>
<paragraph id="8BF836B3FFF03B77FF17FF217558999E" blockId="16.[212,823,255,504]" pageId="16" pageNumber="49">
<emphasis id="B933EAA1FFF03B77FF17FF2176F4997E" bold="true" box="[212,420,255,280]" pageId="16" pageNumber="49">Comparative notes</emphasis>
: Based on the large head (in relation to the pronotum) and the slender paramerite, 
<taxonomicName id="4C474D30FFF03B77FD0DFEFE74619951" authorityName="Assing" authorityYear="2019" box="[718,817,288,311]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="49" phylum="Arthropoda" rank="species" species="taurus">
<emphasis id="B933EAA1FFF03B77FD0DFEFE74619951" box="[718,817,288,311]" italics="true" pageId="16" pageNumber="49">O. taurus</emphasis>
</taxonomicName>
is assigned to the 
<taxonomicName id="4C474D30FFF03B77FE50FE9E755D993E" authorityName="Assing" authorityYear="2016" box="[403,525,320,344]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="49" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="B933EAA1FFF03B77FE50FE9E755D993E" box="[403,525,320,344]" italics="true" pageId="16" pageNumber="49">O. dilatatus</emphasis>
</taxonomicName>
subgroup of the 
<taxonomicName id="4C474D30FFF03B77FD07FE9E7460993E" authorityName="Eppelsheim" authorityYear="1895" box="[708,816,320,344]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="49" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="B933EAA1FFF03B77FD07FE9E7460993E" box="[708,816,320,344]" italics="true" pageId="16" pageNumber="49">O. hauseri</emphasis>
</taxonomicName>
group. Among the species of the 
<taxonomicName id="4C474D30FFF03B77FD89FEBE75E5991E" authorityName="Eppelsheim" authorityYear="1895" box="[586,693,352,376]" class="Insecta" family="Staphylinidae" genus="Orphnebius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="49" phylum="Arthropoda" rank="species" species="hauseri">
<emphasis id="B933EAA1FFF03B77FD89FEBE75E5991E" box="[586,693,352,376]" italics="true" pageId="16" pageNumber="49">O. hauseri</emphasis>
</taxonomicName>
group, the new species is characterized particularly by the conspicuous modifications of the (male?) head (median tubercle; two long mandibular processes) and by the shape of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C35D6538FFF03B77FF17FDC176679AFE" pageId="16" pageNumber="49" type="distribution">
<paragraph id="8BF836B3FFF03B77FF17FDC176679AFE" blockId="16.[212,823,543,664]" pageId="16" pageNumber="49">
<emphasis id="B933EAA1FFF03B77FF17FDC175169A51" bold="true" box="[212,582,543,568]" pageId="16" pageNumber="49">Distribution and natural history</emphasis>
: The 
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locality is situated near 
<collectingRegion id="4983F851FFF03B77FEA8FD9E768C9A3E" box="[363,476,576,600]" country="Cambodia" name="Koh Kong" pageId="16" pageNumber="49">Koh Kong</collectingRegion>
in Southwest 
<collectingCountry id="F3507623FFF03B77FDBCFD9E75BF9A3E" box="[639,751,576,600]" name="Cambodia" pageId="16" pageNumber="49">Cambodia</collectingCountry>
at an altitude between 50 and 
<specimenCount id="9D41FD3AFFF03B77FE22FDBE757A9A11" box="[481,554,608,632]" pageId="16" pageNumber="49" type="generic">
<quantity id="4CBF9B56FFF03B77FE22FDBE75769A11" box="[481,550,608,632]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="49" unit="m" value="300.0">300 m</quantity>
.
</specimenCount>
Additional data are not available.
</paragraph>
</subSubSection>
</treatment>
</document>