212 lines
28 KiB
XML
212 lines
28 KiB
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<document id="89EC9646A73950A7CF0D71900FEF0880" ID-CLB-Dataset="6423" ID-DOI="10.11646/zootaxa.4968.1.1" ID-GBIF-Dataset="8f948bf9-6e6b-4167-aa50-bf648409d0e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4745566" ID-ZooBank="09F4CC3C-879C-4FCD-94D5-9ADE4A81EFAC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1620638208101" checkinUser="plazi" docAuthor="Sinclair, Bradley J. & Shamshev, Igor V." docDate="2021" docId="0E6887D4FF9FFFB04EBCFDE9FAF1B150" docLanguage="en" docName="zootaxa.4968.1.1.pdf" docOrigin="Zootaxa 4968 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Iteaphila grandis Sinclair & Shamshev 2021, sp. nov." docType="treatment" docUuid="0935B4C3-958D-4D09-89EF-17C3E1A01849" docUuidSource="ZooBank" docVersion="7" lastPageNumber="48" masterDocId="F251FFACFFB1FF9F4E2BFFCBFF9CB20D" masterDocTitle="World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)" masterLastPageNumber="89" masterPageNumber="1" pageNumber="47" updateTime="1698938463478" updateUser="ExternalLinkService">
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<mods:mods id="75050B99B9B9D9142B04D417D35DE49F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4FB54934581C68FB4CD2398A7B29FB73">
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<mods:title id="6D635080E05C87D6600384A3A7A07331">World revision of Iteaphila with unbranched radial vein (Diptera: Empidoidea: Iteaphilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="DB08E4040801E8C28EB340CF99373256" type="personal">
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<mods:role id="A0CE5E4FD0C48C56707E8ED34F2D6D8D">
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<mods:roleTerm id="E7EAB34DA3F703906EFF60E140E13AD5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DE089C9C97EB1D1D03A44CBC50DD7639">Sinclair, Bradley J.</mods:namePart>
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<mods:nameIdentifier id="4EC1BFA334115EA18625ACD3BDD3A0F7" type="ORCID">0000-0001-6413-1606</mods:nameIdentifier>
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<mods:nameIdentifier id="B300F259418E8BC93DE9437B62FE07DC" type="email">bradley.sinclair@canada.ca</mods:nameIdentifier>
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</mods:name>
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<mods:name id="E53FA21C5776252D8C1715948EA56A2D" type="personal">
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<mods:roleTerm id="B301B46B1D3D1AFD149C0022F8F9AEE1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A6FBA9EC19548D9D46B9284D70556B72">Shamshev, Igor V.</mods:namePart>
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<mods:nameIdentifier id="CEE7CDA91726C64F4B9B0EA320AAFB90" type="ORCID">0000-0001-6413-1606</mods:nameIdentifier>
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<mods:nameIdentifier id="772905CC1FE5296070D0AEE60C64AE1F" type="email">bradley.sinclair@canada.ca</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="7A19FCC2C31F09ED96599D5F8F19625E">text</mods:typeOfResource>
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<mods:relatedItem id="ECC04E45241CCEC0C4D48999D45E88A8" type="host">
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<mods:titleInfo id="4DC6F17CA8A7C89A3EAAB4A0577F6638">
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<mods:title id="D6EE9BAAEBA0B63C270E0C719A30CB78">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="90910530C6F66AB78D167073455F2874">
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<mods:date id="6E01B0E961CD4AED84387D2354A1EF76">2021</mods:date>
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<mods:detail id="08B9141C310A5F2FF6F21F50C20681F5" type="pubDate">
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<mods:number id="87C9C38CC18286770B71198A65387849">2021-05-07</mods:number>
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</mods:detail>
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<mods:detail id="DFED985FF47AF0A8456E98A85E25F0D8" type="volume">
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<mods:number id="E02695EA104765BD0BA5EE8270C0B7AA">4968</mods:number>
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</mods:detail>
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<mods:detail id="6DDCB796A82907F7DA958C4E2C74839E" type="issue">
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<mods:number id="EAD2B472E7C22F53396A602A975D3AAC">1</mods:number>
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</mods:detail>
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<mods:start id="E35AE724CA8284BD624B8487192F6A16">1</mods:start>
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<mods:end id="E8DC72F9010295FC3516CEADB4421A86">89</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="104A2827811C776AD28EB191B49226B9">journal article</mods:classification>
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<mods:identifier id="8C0B075FB802791E6F36F26A007A2379" type="CLB-Dataset">6423</mods:identifier>
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<mods:identifier id="6E94B0843CA21A7BF26DDF9A4AED6AC7" type="DOI">10.11646/zootaxa.4968.1.1</mods:identifier>
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<mods:identifier id="063974E08711CCB70540379961302EEB" type="GBIF-Dataset">8f948bf9-6e6b-4167-aa50-bf648409d0e5</mods:identifier>
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<mods:identifier id="3A21EC087B664A45DFBFC12799143939" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BBE31940EF93BBAC16FA7CB827024411" type="Zenodo-Dep">4745566</mods:identifier>
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<mods:identifier id="089F6547ED17A29D0915B11F1042642F" type="ZooBank">09F4CC3C-879C-4FCD-94D5-9ADE4A81EFAC</mods:identifier>
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</mods:mods>
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<treatment id="0E6887D4FF9FFFB04EBCFDE9FAF1B150" ID-DOI="http://doi.org/10.5281/zenodo.4814474" ID-GBIF-Taxon="182399081" ID-Zenodo-Dep="4814474" LSID="urn:lsid:zoobank.org:act:0935B4C3-958D-4D09-89EF-17C3E1A01849" httpUri="http://treatment.plazi.org/id/0E6887D4FF9FFFB04EBCFDE9FAF1B150" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<subSubSection id="CEDB6549FF9FFFB14EBCFDE9FE50B030" box="[151,460,546,573]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="867E36C2FF9FFFB14EBCFDE9FE50B030" blockId="46.[151,964,546,645]" box="[151,460,546,573]" pageId="46" pageNumber="47">
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<heading id="DD3681AEFF9FFFB14EBCFDE9FE50B030" bold="true" box="[151,460,546,573]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
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<emphasis id="B4B5EAD0FF9FFFB14EBCFDE9FE50B030" bold="true" box="[151,460,546,573]" pageId="46" pageNumber="47">
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<taxonomicName id="41C14D41FF9FFFB14EBCFDE9FEFBB030" ID-CoL="9HV7S" authority="Sinclair & Shamshev, 2021" authorityName="Sinclair & Shamshev" authorityYear="2021" box="[151,359,546,573]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
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<emphasis id="B4B5EAD0FF9FFFB14EBCFDE9FEFBB030" bold="true" box="[151,359,546,573]" italics="true" pageId="46" pageNumber="47">Iteaphila grandis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="AF8657ABFF9FFFB14F45FDE8FE50B030" box="[366,460,547,573]" pageId="46" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="CEDB6549FF9FFFB14EBCFD8CFC58B088" pageId="46" pageNumber="47" type="description">
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<paragraph id="867E36C2FF9FFFB14EBCFD8CFECDB06C" blockId="46.[151,964,546,645]" box="[151,337,583,609]" pageId="46" pageNumber="47">
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(
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<figureCitation id="1EFA2A47FF9FFFB14EB4FD8CFF6AB06C" box="[159,246,583,609]" captionStart="FIGURES 15–18" captionStartId="14.[151,264,1979,2003]" captionTargetBox="[160,1425,489,1956]" captionTargetId="figure-144@14.[160,1427,489,1956]" captionTargetPageId="14" captionText="FIGURES 15–18. Species of Iteaphila with unbranched R, lateral view. 15, I. flavipilosa, female, lateral view; 16, I. 4+5 flavipilosa, male, lateral view; 17, I. glabricula sp. nov., male; 18, I. grandis sp. nov., male. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4745580" httpUri="https://zenodo.org/record/4745580/files/figure.png" pageId="46" pageNumber="47">Figs 18</figureCitation>
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,
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<figureCitation id="1EFA2A47FF9FFFB14F29FD8CFEBDB06C" box="[258,289,583,609]" captionStart="FIGURES 45–48" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 45–48. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="46" pageNumber="47">46</figureCitation>
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,
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<figureCitation id="1EFA2A47FF9FFFB14F07FD8CFED5B06C" box="[300,329,583,609]" captionStart="FIGURES 49–52" captionStartId="45.[151,264,1843,1867]" captionTargetBox="[151,1436,381,1820]" captionTargetId="figure-106@45.[151,1436,381,1820]" captionTargetPageId="45" captionText="FIGURES 49–52. Distribution of Iteaphila with unbranched R. 49, I. arnaudi sp. nov., I. bayarea sp. nov., I. grandis sp. 4+5 nov., I. sierrensis sp. nov.; 50, I. brooksi sp. nov., I. lolo sp. nov.; 51, I. flavipilosa (Coquillett); 52, I. glabricula sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4745604" httpUri="https://zenodo.org/record/4745604/files/figure.png" pageId="46" pageNumber="47">49</figureCitation>
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)
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</paragraph>
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<paragraph id="867E36C2FF9FFFB14EBCFDA0FC58B088" blockId="46.[151,964,546,645]" box="[151,964,618,645]" pageId="46" pageNumber="47">
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<uri id="F2503AC0FF9FFFB14EBCFDA0FC58B088" box="[151,964,618,645]" pageId="46" pageNumber="47">
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urn:lsid:zoobank.org:act:
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<uuid id="F2670C17FF9FFFB14F9DFDA1FC58B088" box="[438,964,618,645]" pageId="46" pageNumber="47">0935B4C3-958D-4D09-89EF-17C3E1A01849</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="CEDB6549FF9FFFB14EBCFD78FEEEB1AB" pageId="46" pageNumber="47" type="materials_examined">
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<paragraph id="867E36C2FF9FFFB14EBCFD78FE41B134" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">
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<emphasis id="B4B5EAD0FF9FFFB14EBCFD78FED6B0C1" bold="true" box="[151,330,690,717]" pageId="46" pageNumber="47">Type material.</emphasis>
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<materialsCitation id="36A93C9FFF9FFFB14F7EFD79FC42B118" ID-GBIF-Occurrence="3118988509" collectionCode="CALIF, CAS" collectorName="B. P. Bliven" county="Humboldt Co." latitude="40.965557" location="Fieldbrook" longLatPrecision="19" longitude="-124.03555" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B4B5EAD0FF9FFFB14F7EFD79FE68B0C1" bold="true" box="[341,500,690,716]" pageId="46" pageNumber="47">
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<typeStatus id="597A8860FF9FFFB14F7EFD79FE68B0C1" box="[341,500,690,716]" pageId="46" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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<specimenCount id="90C7FD4BFF9FFFB14FD4FD78FD8AB0C0" box="[511,534,691,717]" pageId="46" pageNumber="47" type="male">♂</specimenCount>
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, labelled: “
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<location id="831E6019FF9FFFB14CB7FD78FC86B0C0" LSID="urn:lsid:plazi:treatment:0E6887D4FF9FFFB04EBCFDE9FAF1B150:831E6019FF9FFFB14CB7FD78FC86B0C0" box="[668,794,691,717]" county="Humboldt Co." latitude="40.965557" longLatPrecision="19" longitude="-124.03555" name="Fieldbrook" pageId="46" pageNumber="47">Fieldbrook</location>
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[
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<geoCoordinate id="E3F55005FF9FFFB14D06FD7FFC29B0C2" box="[813,949,692,719]" degrees="40" direction="north" minutes="57" orientation="latitude" pageId="46" pageNumber="47" precision="15" seconds="56" value="40.965557">40°57′56″N</geoCoordinate>
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<geoCoordinate id="E3F55005FF9FFFB14DEBFD7FFBC0B0C2" box="[960,1116,692,719]" degrees="124" direction="west" minutes="02" orientation="longitude" pageId="46" pageNumber="47" precision="15" seconds="08" value="-124.03555">124°02′08″W</geoCoordinate>
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]/
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<collectingCounty id="6F1F4E4EFF9FFFB14A5DFD78FA86B0C0" box="[1142,1306,691,717]" pageId="46" pageNumber="47">Humboldt Co.</collectingCounty>
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/ Cal. Mar. 24, ’35/
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<collectorName id="2B345314FF9FFFB14ED4FD1CFE1BB0FC" box="[255,391,727,753]" pageId="46" pageNumber="47">B.P. Bliven</collectorName>
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”; “BRUNSON P. BLIVEN/ COLLECTION/ 1981 Accession/
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<collectionCode id="E0D0AE07FF9FFFB14A4FFD1CFB2AB0FC" box="[1124,1206,727,753]" pageId="46" pageNumber="47">CALIF</collectionCode>
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. ACAD. OF SCI.”; “
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<typeStatus id="597A8860FF9FFFB14E8BFD30FEAAB118" box="[160,310,763,789]" pageId="46" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="41C14D41FF9FFFB14F6DFD30FE3BB118" box="[326,423,763,789]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Iteaphila</taxonomicName>
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/ grandis/ Sinclair & Shamshev [red label]” (
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<collectionCode id="E0D0AE07FF9FFFB14DB1FD30FC4AB118" box="[922,982,763,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="46" pageNumber="47" type="Museum">CAS</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="36A93C9FFF9FFFB14DC2FD31FE44B134" ID-GBIF-Occurrence="3118988681" collectionCode="CAS" collectorName="B. P. Bliven" country="United States of America" county="Humboldt Co." latitude="40.965557" location="Same" longLatPrecision="19" longitude="-124.03555" pageId="46" pageNumber="47" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="paratype">
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<emphasis id="B4B5EAD0FF9FFFB14DC2FD31FACBB119" bold="true" box="[1001,1367,762,788]" pageId="46" pageNumber="47">
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<typeStatus id="597A8860FF9FFFB14DC2FD31FB1EB119" box="[1001,1154,762,788]" pageId="46" pageNumber="47" type="paratype">PARATYPE</typeStatus>
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:
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<collectingCountry id="FED67652FF9FFFB14AA6FD31FB5BB119" box="[1165,1223,762,788]" name="United States of America" pageId="46" pageNumber="47">USA</collectingCountry>
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.
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<collectingRegion id="4405F820FF9FFFB14AF8FD31FACEB119" box="[1235,1362,762,788]" country="United States of America" name="California" pageId="46" pageNumber="47">California</collectingRegion>
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:
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</emphasis>
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<location id="831E6019FF9FFFB14B75FD30FA00B118" LSID="urn:lsid:plazi:treatment:0E6887D4FF9FFFB04EBCFDE9FAF1B150:831E6019FF9FFFB14B75FD30FA00B118" box="[1374,1436,763,789]" country="United States of America" name="Same" pageId="46" pageNumber="47" stateProvince="California">Same</location>
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data as holotype (
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<specimenCount id="90C7FD4BFF9FFFB14F74FCD4FE15B134" box="[351,393,799,825]" pageId="46" pageNumber="47" type="male">1 ♂</specimenCount>
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,
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<collectionCode id="E0D0AE07FF9FFFB14FBEFCD4FE4CB134" box="[405,464,799,825]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="46" pageNumber="47" type="Museum">CAS</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="867E36C2FF9FFFB14EECFC89FEEEB1AB" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">
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<emphasis id="B4B5EAD0FF9FFFB14EECFC89FEC1B151" bold="true" box="[199,349,834,860]" pageId="46" pageNumber="47">Recognition.</emphasis>
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This species is distinguished by its very large size (wing length
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<quantity id="41399B27FF9FFFB14A04FC88FB1AB150" box="[1071,1158,835,861]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="4.3">4.3 mm</quantity>
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) and the male terminalia is similar to
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<taxonomicName id="41C14D41FF9FFFB14F0FFCACFE26B18C" authorityName="Sinclair & Shamshev" authorityYear="2021" baseAuthorityName="Melander" box="[292,442,871,897]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="longipalpis">
|
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<emphasis id="B4B5EAD0FF9FFFB14F0FFCACFE26B18C" box="[292,442,871,897]" italics="true" pageId="46" pageNumber="47">I. longipalpis</emphasis>
|
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</taxonomicName>
|
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, but the surstylus is broader, the apex of the postgonite is expanded and the proboscis is very long (
|
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|
<figureCitation id="1EFA2A47FF9FFFB14F3AFC40FEF9B1AB" box="[273,357,907,934]" captionStart="FIGURES 45–48" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 45–48. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="46" pageNumber="47">Fig. 46</figureCitation>
|
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).
|
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</paragraph>
|
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</subSubSection>
|
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<subSubSection id="CEDB6549FF9FFFB04EECFC65FE3EB088" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
|
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|
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<paragraph id="867E36C2FF9FFFB14EECFC65FCA3B6E4" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">
|
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<emphasis id="B4B5EAD0FF9FFFB14EECFC65FEC5B1C5" bold="true" box="[199,345,942,968]" pageId="46" pageNumber="47">Description.</emphasis>
|
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Wing length
|
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<quantity id="41399B27FF9FFFB14FD2FC64FDC9B1C4" box="[505,597,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="46" pageNumber="47" unit="mm" value="4.3">4.3 mm</quantity>
|
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.
|
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<emphasis id="B4B5EAD0FF9FFFB14C49FC64FD03B1C4" bold="true" box="[610,671,943,969]" pageId="46" pageNumber="47">Male</emphasis>
|
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(
|
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<figureCitation id="1EFA2A47FF9FFFB14C9AFC64FC94B1C4" box="[689,776,943,969]" captionStart="FIGURES 15–18" captionStartId="14.[151,264,1979,2003]" captionTargetBox="[160,1425,489,1956]" captionTargetId="figure-144@14.[160,1427,489,1956]" captionTargetPageId="14" captionText="FIGURES 15–18. Species of Iteaphila with unbranched R, lateral view. 15, I. flavipilosa, female, lateral view; 16, I. 4+5 flavipilosa, male, lateral view; 17, I. glabricula sp. nov., male; 18, I. grandis sp. nov., male. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4745580" httpUri="https://zenodo.org/record/4745580/files/figure.png" pageId="46" pageNumber="47">Fig. 18</figureCitation>
|
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). Head dark brown in ground-colour, with pale brown to black setation, occiput thinly to densely greyish pollinose. Eyes holoptic, with upper ommatidia enlarged. Frons represented by small triangular space just above antennae, bare, greyish pollinose. Ocellar triangle prominent, with 2 pairs of long hair-like setae. Postvertical setae subequal to postocular setae, thin; occiput covered with numerous similar setae in lower part, pale brown posterior to mouthparts. Antenna dark brown; scape short, slightly shorter than globular pedicel, both with short setae; postpedicel rather narrow, nearly 4 times longer than basal width, base somewhat expanded and smoothly tapered; stylus
|
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<emphasis id="B4B5EAD0FF9FFFB14D20FB4CFCB9B6AD" box="[779,805,1159,1184]" italics="true" pageId="46" pageNumber="47">ca</emphasis>
|
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|
|
3–4 times longer than wide, sensillum-tipped [broken]; segment 9 tubular, 2–3 times longer than wide. Proboscis very long, projected obliquely; labium nearly twice head height; palpus [broken] slender, projected parallel to labrum.
|
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</paragraph>
|
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<paragraph id="867E36C2FF9FFFB14EECFB38FBC9B420" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">Thorax dark brown in ground-colour with posterior part of postpronotal lobe and postalar tubercle often paler, with brown to black setation; scutum viewed dorsally entirely velvety brown, slightly matt, with pair of faintly shiny vittae, between acr and dc setae; viewed anteriorly densely pruinose with pair of pruinescent vittae; mesopleuron uniformly greyish brown pollinose. Proepisternum with a few hair-like setae. Postpronotal lobe with 1 long thin and numerous shorter setae. Mesonotal setae distinct, thin; acr short, 4-serial, multi-serial anteriorly, arranged in closely spaced irregular rows, lacking on prescutellar depression, distance between paired rows shorter than length of acr; dc uniserial (with some additional setulae in anterior part), offset from row anteriorly, slightly longer than acr, 2–3 distinct prescutellar dc; 1 ph, 1 presut spal (sometimes with additional setulae), 3–5 npl (with a few additional shorter setae), several psut spal setulae, 1 pal and several short setulae, 8 pairs of sctl.</paragraph>
|
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<paragraph id="867E36C2FF9FFFB14EECF9FCFC7BB524" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">Legs, including coxae, almost entirely brownish; knees of fore and mid legs brownish yellow, tarsi somewhat darker. Coxae and trochanters with unmodified hair-like setae. Fore femur with row of short anteroventral hair-like setae, especially distinct apically; and somewhat longer setae on posteroventral, dorsal and posterior faces. Mid femur with moderately long posteroventral bristly setae along entire length, bearing some moderately long setae on dorsal and posterior faces. Hind femur with long anteroventral and dorsal setae along whole length. Tibiae with row of slightly prominent thin posterodorsal setae. Tarsomere 1 of hindleg with pair of rows of stiff ventral setae; tarsomere 5 not flattened on all legs; pulvilli broad, shorter than tarsal claw.</paragraph>
|
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|
<paragraph id="867E36C2FF9FFFB14EECF8F8FBBCB5D0" blockId="46.[151,1437,690,2013]" pageId="46" pageNumber="47">
|
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|
|
Wing distinctly uniformly infuscate; basal costal seta absent; pterostigma distinct brownish, elliptical, overlapping apex of R
|
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|
|
<subScript id="1A453487FF9FFFB14FB4F8A2FE3BB57A" attach="left" box="[415,423,1897,1911]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
|
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|
|
; anal lobe very prominent, acute. Sc complete; R
|
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|
|
<subScript id="1A453487FF9FFFB14DDCF8A2FB8CB57A" attach="left" box="[1015,1040,1897,1911]" fontSize="6" pageId="46" pageNumber="47">2+3</subScript>
|
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|
|
often slightly bowed posterior to pterostigma; R
|
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|
|
<subScript id="1A453487FF9FFFB14F16F846FECBB596" attach="left" box="[317,343,1933,1947]" fontSize="6" pageId="46" pageNumber="47">4+5</subScript>
|
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|
|
unbranched; cell dm broad, longer than basal cells, truncate apically; base of M
|
|||
|
|
<subScript id="1A453487FF9FFFB14AF4F846FB7BB596" attach="left" box="[1247,1255,1933,1947]" fontSize="6" pageId="46" pageNumber="47">2</subScript>
|
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|
|
(crossvein) less than one-third length of dm-m, M branches widely separated; dm-m crossvein slightly concave. Apex of cell cua recurved, CuA+CuP long, pale, ending short of wing margin. Halter dark brown.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="867E36C2FF9EFFB04EECFF5CFE82B06C" blockId="47.[151,1437,151,861]" pageId="47" pageNumber="48">
|
|||
|
|
Abdomen brown, subshiny, finely brownish grey pollinose, covered with long pale brownish setae. Terminalia (
|
|||
|
|
<figureCitation id="1EFA2A47FF9EFFB04E8BFF70FF68B2DB" box="[160,244,187,214]" captionStart="FIGURES 45–48" captionStartId="44.[151,264,1686,1710]" captionTargetBox="[160,1433,194,1631]" captionTargetId="figure-19@44.[151,1436,181,1663]" captionTargetPageId="44" captionText="FIGURES 45–48. Male terminalia of Iteaphila with unbranched R, lateral view. 45, I. gracilis sp. nov.; 46, I. grandis sp. 4+5 nov.; 47, I. lolo sp. nov.; 48, I. longipalpis (Melander). Scale bar = 0.1 mm. Abbreviations: cerc—cercus; epand brg—epandrial bridge; hyprct proc—hypoproct process; ph—phallus; ph gd—phallic guide; sur—surstylus." figureDoi="http://doi.org/10.5281/zenodo.4745600" httpUri="https://zenodo.org/record/4745600/files/figure.png" pageId="47" pageNumber="48">Fig. 46</figureCitation>
|
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|
|
) concolorous with abdomen, large. Hypandrium with numerous short setae, rounded basally, narrowed apically with narrow, folded apex, with roughened posterior apex; gonocoxal apodeme long and narrow; inner apodeme expanded and flattened, shorter than length of gonocoxal apodeme; postgonite long, flattened plate, arched, truncate apically, fused with opposite postgonite with lateral projection. Phallic guide extended well beyond epandrium, parallel with phallus and bent nearly at right angles subapically; bent apical section paired with jagged or toothed surface apically; apex expanded. Epandrium not inflated laterally; dorsal bridge narrow; produced distally into broad dorsally projecting, slightly curved anteriorly surstylus; inner margin of surstylus with short tooth-like projection medially. Phallus slender, arched well beyond epandrium, bent more than right angles subapically; apical section sinuous and slender; ejaculatory apodeme plate-like, rounded, greatly expanded anteriorly, slightly longer than length of gonocoxal apodeme. Cercus short, triangular, one-third length of epandrium; narrowed apically with apex pointed, extended free from epandrium; hypoproct produced into pair of very long, slender processes, longer than cercus.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="867E36C2FF9EFFB04EECFDA0FE3EB088" blockId="47.[151,1437,151,861]" box="[199,418,619,645]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B4B5EAD0FF9EFFB04EECFDA0FEB9B088" bold="true" box="[199,293,619,645]" pageId="47" pageNumber="48">Female.</emphasis>
|
|||
|
|
Unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CEDB6549FF9EFFB04EECFD45FEE5B0C0" pageId="47" pageNumber="48" type="distribution">
|
|||
|
|
<paragraph id="867E36C2FF9EFFB04EECFD45FEE5B0C0" blockId="47.[151,1437,151,861]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B4B5EAD0FF9EFFB04EECFD45FEFDB0A5" bold="true" box="[199,353,654,680]" pageId="47" pageNumber="48">Distribution.</emphasis>
|
|||
|
|
Nearctic:
|
|||
|
|
<collectingCountry id="FED67652FF9EFFB04FFAFD44FD9BB0A4" box="[465,519,655,681]" name="United States of America" pageId="47" pageNumber="48">USA</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<collectingRegion id="4405F820FF9EFFB04C3EFD44FD11B0A7" box="[533,653,655,682]" country="United States of America" name="California" pageId="47" pageNumber="48">California</collectingRegion>
|
|||
|
|
). This species is known only from its
|
|||
|
|
<typeStatus id="597A8860FF9EFFB04A0EFD44FBC9B0A4" box="[1061,1109,655,681]" pageId="47" pageNumber="48">type</typeStatus>
|
|||
|
|
locality in Humboldt County,
|
|||
|
|
<collectingRegion id="4405F820FF9EFFB04EBCFD78FE95B0C0" box="[151,265,691,717]" country="United States of America" name="California" pageId="47" pageNumber="48">California</collectingRegion>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="1EFA2A47FF9EFFB04F33FD78FEF0B0C0" box="[280,364,691,717]" captionStart="FIGURES 49–52" captionStartId="45.[151,264,1843,1867]" captionTargetBox="[151,1436,381,1820]" captionTargetId="figure-106@45.[151,1436,381,1820]" captionTargetPageId="45" captionText="FIGURES 49–52. Distribution of Iteaphila with unbranched R. 49, I. arnaudi sp. nov., I. bayarea sp. nov., I. grandis sp. 4+5 nov., I. sierrensis sp. nov.; 50, I. brooksi sp. nov., I. lolo sp. nov.; 51, I. flavipilosa (Coquillett); 52, I. glabricula sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4745604" httpUri="https://zenodo.org/record/4745604/files/figure.png" pageId="47" pageNumber="48">Fig. 49</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CEDB6549FF9EFFB04EECFD1CFE03B118" pageId="47" pageNumber="48" type="etymology">
|
|||
|
|
<paragraph id="867E36C2FF9EFFB04EECFD1CFE03B118" blockId="47.[151,1437,151,861]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B4B5EAD0FF9EFFB04EECFD1CFED2B0FC" bold="true" box="[199,334,727,753]" pageId="47" pageNumber="48">Etymology.</emphasis>
|
|||
|
|
The specific name is the Latin
|
|||
|
|
<taxonomicName id="41C14D41FF9EFFB04CE4FD1CFCB9B0FC" authorityName="Sinclair & Shamshev" authorityYear="2021" box="[719,805,727,753]" class="Insecta" family="Iteaphilidae" genus="Iteaphila" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="grandis">
|
|||
|
|
<emphasis id="B4B5EAD0FF9EFFB04CE4FD1CFCB9B0FC" box="[719,805,727,753]" italics="true" pageId="47" pageNumber="48">grandis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(large, great), in reference to its large body size and lengthened mouthparts.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="CEDB6549FF9EFFB04EECFCD4FAF1B150" pageId="47" pageNumber="48" type="discussion">
|
|||
|
|
<paragraph id="867E36C2FF9EFFB04EECFCD4FAF1B150" blockId="47.[151,1437,151,861]" pageId="47" pageNumber="48">
|
|||
|
|
<emphasis id="B4B5EAD0FF9EFFB04EECFCD4FEA7B134" bold="true" box="[199,315,799,825]" pageId="47" pageNumber="48">Remarks.</emphasis>
|
|||
|
|
This species was collected in late March, when there are few collection records of the genus at that time of year in
|
|||
|
|
<collectingRegion id="4405F820FF9EFFB04F69FC88FE2BB150" box="[322,439,835,861]" country="United States of America" name="California" pageId="47" pageNumber="48">California</collectingRegion>
|
|||
|
|
. This apparent early flight period may explain the rarity of the species in collections.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|