329 lines
22 KiB
XML
329 lines
22 KiB
XML
|
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.87779" ID-Pensoft-Pub="1314-2003-222-75" ID-Pensoft-UUID="348E6053A6D55787993694B23682FDE9" ModsDocID="1314-2003-222-75" checkinTime="1679617121532" checkinUser="pensoft" docAuthor="Maas, Paul J. M., Maas-van de Kamer, Hiltje, Andre, Thiago, Skinner, David, Valderrama, Eugenio & Specht, Chelsea D." docDate="2023" docId="21E5187F33005474943314D0CC978B38" docLanguage="en" docName="PhytoKeys 222: 75-127" docOrigin="PhytoKeys 222" docPubDate="2023-03-22" docSource="http://dx.doi.org/10.3897/phytokeys.222.87779" docTitle="Costus obscurus D. Skinner & Maas 2023, sp. nov." docType="treatment" docVersion="1" id="348E6053A6D55787993694B23682FDE9" lastPageNumber="75" masterDocId="348E6053A6D55787993694B23682FDE9" masterDocTitle="Eighteen new species of Neotropical Costaceae (Zingiberales)" masterLastPageNumber="127" masterPageNumber="75" pageNumber="75" updateTime="1679617121532" updateUser="pensoft">
|
|||
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo>
|
|||
|
|
<mods:title>Eighteen new species of Neotropical Costaceae (Zingiberales)</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Maas, Paul J. M.</mods:namePart>
|
|||
|
|
<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Maas-van de Kamer, Hiltje</mods:namePart>
|
|||
|
|
<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Andre, Thiago</mods:namePart>
|
|||
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4148-3662</mods:nameIdentifier>
|
|||
|
|
<mods:affiliation>Universidade de Brasilia, Departamento de Botanica, Campus Universitario Darcy Ribeiro, Asa Norte, Brasilia (DF), Brazil</mods:affiliation>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Skinner, David</mods:namePart>
|
|||
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4585-925X</mods:nameIdentifier>
|
|||
|
|
<mods:affiliation>Le Jardin Ombrage, Tallahassee, (Private botanical garden, Botanic Gardens Conservation International - BGCI - registration ID 50148), Florida, USA</mods:affiliation>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Valderrama, Eugenio</mods:namePart>
|
|||
|
|
<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:name type="personal">
|
|||
|
|
<mods:role>
|
|||
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart>Specht, Chelsea D.</mods:namePart>
|
|||
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7746-512X</mods:nameIdentifier>
|
|||
|
|
<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
|
|||
|
|
<mods:nameIdentifier type="email">cdspecht@cornell.edu</mods:nameIdentifier>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem type="host">
|
|||
|
|
<mods:titleInfo>
|
|||
|
|
<mods:title>PhytoKeys</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part>
|
|||
|
|
<mods:date>2023</mods:date>
|
|||
|
|
<mods:detail type="pubDate">
|
|||
|
|
<mods:number>2023-03-22</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail type="volume">
|
|||
|
|
<mods:number>222</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent unit="page">
|
|||
|
|
<mods:start>75</mods:start>
|
|||
|
|
<mods:end>127</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:location>
|
|||
|
|
<mods:url>http://dx.doi.org/10.3897/phytokeys.222.87779</mods:url>
|
|||
|
|
</mods:location>
|
|||
|
|
<mods:classification>journal article</mods:classification>
|
|||
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.87779</mods:identifier>
|
|||
|
|
<mods:identifier type="Pensoft-Pub">1314-2003-222-75</mods:identifier>
|
|||
|
|
<mods:identifier type="Pensoft-UUID">348E6053A6D55787993694B23682FDE9</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment LSID="urn:lsid:plazi:treatment:21E5187F33005474943314D0CC978B38" httpUri="http://treatment.plazi.org/id/21E5187F33005474943314D0CC978B38" lastPageNumber="75" pageId="0" pageNumber="75">
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="nomenclature">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<taxonomicName LSID="21E5187F-3300-5474-9433-14D0CC978B38" authority="D. Skinner & Maas" authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus" status="sp. nov.">Costus obscurus D.Skinner & Maas</taxonomicName>
|
|||
|
|
<taxonomicNameLabel pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="diagnosis">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus obscurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. (Fig.
|
|||
|
|
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Costus obscurus D. Skinner & Maas A cultivated plant from Plowman's type collection showing upright orientation of leaf lamina B inflorescence C cultivated plant showing colours and upright orientation of leaf lamina D photo showing villose indument on leaf sheaths E Indument on adaxial leaf surface F indument on abaxial leaf surface A, B cultivated specimen from Lyon Arboretum, Hawaii L- 81.0901 grown from seeds wild collected by Tim Plowman from the plant accessioned as Plowman 5054 C-F photos taken in Pantiacolla, Madre de Dios, Peru prior to collecting seeds for D. Skinner R 3322, currently in cultivation. Photos A-F by Dave Skinner." figureDoi="10.3897/phytokeys.222.87779.figure14" httpUri="https://binary.pensoft.net/fig/828353" pageId="0" pageNumber="75">14</figureCitation>
|
|||
|
|
) can be recognized by its large, dark green leaves with dark purple undersides and densely whitish villose sheaths. It could superficially be confused with
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. erythrophyllus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="erythrophyllus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. erythrophyllus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Loes. and
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. acreanus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="acreanus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. acreanus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Loes.) Maas, but these species never have densely villose sheaths.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption doi="10.3897/phytokeys.222.87779.figure14" httpUri="https://binary.pensoft.net/fig/828353" pageId="0" pageNumber="75" start="Figure 14" startId="F14">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Figure 14.</emphasis>
|
|||
|
|
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus obscurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
D.Skinner & Maas
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
|
|||
|
|
cultivated plant from
|
|||
|
|
<normalizedToken originalValue="Plowman’s">Plowman's</normalizedToken>
|
|||
|
|
type collection showing upright orientation of leaf lamina
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">B</emphasis>
|
|||
|
|
inflorescence
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">C</emphasis>
|
|||
|
|
cultivated plant showing colours and upright orientation of leaf lamina
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
|
|||
|
|
photo showing villose indument on leaf sheaths
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
|
|||
|
|
Indument on adaxial leaf surface
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">F</emphasis>
|
|||
|
|
indument on abaxial leaf surface
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">A, B</emphasis>
|
|||
|
|
cultivated specimen from Lyon Arboretum, Hawaii
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">L-81.0901</emphasis>
|
|||
|
|
grown from seeds wild collected by Tim Plowman from the plant accessioned as
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman 5054</emphasis>
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">C-F</emphasis>
|
|||
|
|
photos taken in Pantiacolla, Madre de Dios,
|
|||
|
|
<normalizedToken originalValue="Perú">Peru</normalizedToken>
|
|||
|
|
prior to collecting seeds for
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">D.Skinner R3322</emphasis>
|
|||
|
|
, currently in cultivation. Photos
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">A-F</emphasis>
|
|||
|
|
by Dave Skinner.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="materials_examined">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<typeStatus>Type</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<materialsCitation collectingDate="1974-02-09" collectorName="Plowman, Davis" country="Peru" elevation="600" location="Salvacion" municipality="Shintuya" specimenCount="1" stateProvince="Madre de Dios" typeStatus="holotype">
|
|||
|
|
<collectingCountry name="Peru">Peru</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingRegion country="Peru" name="Madre de Dios">Madre de Dios</collectingRegion>
|
|||
|
|
: Prov. Manu, near
|
|||
|
|
<collectingMunicipality>Shintuya</collectingMunicipality>
|
|||
|
|
, on road to
|
|||
|
|
<normalizedToken originalValue="Salvación">
|
|||
|
|
<location LSID="urn:lsid:plazi:treatment:21E5187F33005474943314D0CC978B38:159633C0596A1646DE2248E752E51879" country="Peru" municipality="Shintuya" name="Salvacion" stateProvince="Madre de Dios">Salvacion</location>
|
|||
|
|
</normalizedToken>
|
|||
|
|
,
|
|||
|
|
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
|
|||
|
|
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<collectingDate value="1974-02-09">9 Feb 1974</collectingDate>
|
|||
|
|
,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">
|
|||
|
|
<collectorName>Plowman</collectorName>
|
|||
|
|
&
|
|||
|
|
<collectorName>Davis</collectorName>
|
|||
|
|
5054
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<typeStatus>holotype</typeStatus>
|
|||
|
|
USM; isotypes GH, L1483069, L1480371, L1480372)
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="description">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Description.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Herb</emphasis>
|
|||
|
|
1-1.5 m tall.
|
|||
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Leaves</emphasis>
|
|||
|
|
sheaths 10-20 mm diam; ligule truncate, 10-15 mm long; petiole 8-12 mm long; sheaths, ligule and petiole densely to rather densely whitish villose; lamina narrowly elliptic, 32-45
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
9-17 cm, adaxially dark green, rather densely to densely whitish villose, abaxially dark purple, densely to sparsely whitish puberulous, base acute to cordate, apex acuminate (acumen 5-10 mm long).
|
|||
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Inflorescence</emphasis>
|
|||
|
|
ovoid, 8-13
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
5-7 cm, terminating a leafy shoot; bracts, appendages of bracts, bracteole, calyx, ovary, and capsule rather densely to sparsely puberulous.
|
|||
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Flowers</emphasis>
|
|||
|
|
abaxially oriented; bracts green in the exposed part, red or green in the covered part, coriaceous, broadly ovate-triangular, 35-45
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
30-35 mm; appendages green, erect, broadly ovate-triangular, 15-20
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
18-20 mm, or almost absent, apex rounded; bracteole boat-shaped, 35-37 mm long; calyx red, 25-27 mm long, deeply split on one side, lobes deltate to broadly triangular, ca. 2 mm long; corolla cream to yellow, ca. 75 mm long, glabrous, lobes narrowly elliptic, 55-60 mm long; labellum pale yellow, distal edge horizontally spreading, broadly obovate, ca. 60-70
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
60-70 mm, lateral lobes strongly striped with pink or red, middle lobe reflexed, with yellow honey mark, margin irregularly dentate to lobulate; stamen white to pink, 35-45
|
|||
|
|
<normalizedToken originalValue="×">x</normalizedToken>
|
|||
|
|
12-15 mm, not exceeding the labellum, apex pinkish red, deeply and irregularly lobed, anther 7-8 mm long.
|
|||
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Capsule</emphasis>
|
|||
|
|
ellipsoid, 12-15 mm long.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="distribution">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
Peru (Cuzco,
|
|||
|
|
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
|||
|
|
, Madre de Dios, San
|
|||
|
|
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
|||
|
|
) (Fig.
|
|||
|
|
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution maps for new species of Costaceae. Costus gibbosus - Costus whiskeycola m A Costus gibbosus D. Skinner & Maas B Costus mollissimus Maas & H. Maas C Costus obscurus D. Skinner & Maas D Costus oreophilus Maas & D. Skinner E Costus pitalito C. D. Specht & H. Maas F Costus prancei Maas & H. Maas G Costus pseudospiralis Maas & H. Maas H Costus rubineus D. Skinner & Maas I Costus whiskeycola Maas & H. Maas." figureDoi="10.3897/phytokeys.222.87779.figure22" httpUri="https://binary.pensoft.net/fig/828361" pageId="0" pageNumber="75">22C</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="habitat">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Habitat and ecology.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">In forests, in wet, shady areas, often in disturbed places, at elevations of 450-1300 m. Flowering in the rainy season.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="etymology">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Etymology.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus obscurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. is known for its leaves having a very dark green adaxial surface and dark purple abaxial surface, and for growing in dark and shady places, hence the specific epithet obscurus (
|
|||
|
|
<normalizedToken originalValue="‘obscurus’">'obscurus'</normalizedToken>
|
|||
|
|
means 'dark, shady,
|
|||
|
|
<normalizedToken originalValue="“indistinct’">"indistinct'</normalizedToken>
|
|||
|
|
in Latin). This species might also be considered
|
|||
|
|
<normalizedToken originalValue="“obscure”">"obscure"</normalizedToken>
|
|||
|
|
(in English: unclear, uncertain, unknown, or in doubt) because it has been confused by the authors in the past as either
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. erythrophyllus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="erythrophyllus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. erythrophyllus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Loes. or as
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. acreanus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="acreanus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. acreanus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Loes.) Maas. Only recently has it been determined to be an undescribed species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="paratypes">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Paratypes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Peru. Cuzco</emphasis>
|
|||
|
|
: Prov. La
|
|||
|
|
<normalizedToken originalValue="Convención">Convencion</normalizedToken>
|
|||
|
|
, Distr. Echarati, Cashiriari-3 well site, 5 km S of Camisea River, 700 m, 2 Sep 1998,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">
|
|||
|
|
<normalizedToken originalValue="Nuñez">Nunez</normalizedToken>
|
|||
|
|
V. et al. 23785
|
|||
|
|
</emphasis>
|
|||
|
|
(F).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">
|
|||
|
|
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
|||
|
|
</emphasis>
|
|||
|
|
: Prov. Leoncio Prado, Puente Pucayaco,
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Pucayaco, road from Tingo Maria to Tocache, 570 m, 30 Mar 1976,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman & Kennedy 5775</emphasis>
|
|||
|
|
(U); cultivated in Tocache, besides Maior Plaza, originally collected by
|
|||
|
|
<normalizedToken originalValue="José">Jose</normalizedToken>
|
|||
|
|
Schunke V. at Quebrada Cachiyaco de Lopuna, near Mina de Sal, S of Tocache, 31 Mar 1976,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman & Kennedy 5791</emphasis>
|
|||
|
|
(GH, U); cultivated in Jardin Botanico de Tingo Maria, from material collected by Plowman in Ramal de Aspusana, 65 km N of Tingo Maria, on the border of
|
|||
|
|
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
|||
|
|
and San
|
|||
|
|
<normalizedToken originalValue="Martín">Martin</normalizedToken>
|
|||
|
|
, 5 Jul 1978,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">
|
|||
|
|
Plowman &
|
|||
|
|
<normalizedToken originalValue="Ramírez">Ramirez</normalizedToken>
|
|||
|
|
R. 7595
|
|||
|
|
</emphasis>
|
|||
|
|
(F, U).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Madre de Dios</emphasis>
|
|||
|
|
: Prov. Manu, Cerro de Pantiacolla,
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Palotoa, 10-15 km NNW of Shintuya, 700-1300 m, 16 Dec 1985,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Foster et al. 10937</emphasis>
|
|||
|
|
(F); Pantiacolla, serrania across
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Alto Madre de Dios from Shintuya, 450-650 m, 28 Oct 1979,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Gentry 27304</emphasis>
|
|||
|
|
(MO, U).
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Cultivated material</emphasis>
|
|||
|
|
: Harvard Forest Greenhouses, Petersham, Massachusetts, USA, cultivated from
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman & Davis 5054</emphasis>
|
|||
|
|
, Peru,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman & Davis 5054 A</emphasis>
|
|||
|
|
(F, L); cultivated from Peru,
|
|||
|
|
<normalizedToken originalValue="Huánuco">Huanuco</normalizedToken>
|
|||
|
|
, 8 km NW of Tingo Maria, along quebrada on road to Shinchi Roca, 900 m, 14 Nov 2016,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Skinner R3383</emphasis>
|
|||
|
|
(BH). Peru, Madre de Dios, Pantiacolla, 25 Jan 2013,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Skinner R3322</emphasis>
|
|||
|
|
(UC).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="notes">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Notes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
Plants of
|
|||
|
|
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus obscurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. have a compact appearance, with leaves closely spaced together along the shoot and generally pointed upward rather than horizontally or drooping. The
|
|||
|
|
<normalizedToken originalValue="leaves’">leaves'</normalizedToken>
|
|||
|
|
dark green adaxial surface and dark purple abaxial surface are striking in appearance.
|
|||
|
|
<taxonomicName authorityName="D. Skinner & Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus obscurus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="obscurus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus obscurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be confused with
|
|||
|
|
<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. erythrophyllus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="erythrophyllus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">C. erythrophyllus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Loes., but can be distinguished from that species by its shorter, truncate ligule instead of a long and deeply 2-lobed ligule as well as by having smooth rather than (mostly) plicate leaves.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
This new species has been widely cultivated from the type collection
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Plowman 5054</emphasis>
|
|||
|
|
.
|
|||
|
|
<normalizedToken originalValue="Plowman’s">Plowman's</normalizedToken>
|
|||
|
|
journal indicates the distribution of live plants to Marie Selby Botanical Garden and Lyon Arboretum, which is accessioned as L-81.0901. It has sometimes been confused with the famous "El Whiskey" plant from Colombia, but it can be easily distinguished by the indumenta on the adaxial and abaxial leaf surfaces.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|