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<document id="DBE53050969B32FF3F2FB6D71A28A86F" ID-CLB-Dataset="153629" ID-DOI="10.18590/euscorpius.2016.vol2016.iss220.1" ID-GBIF-Dataset="03196918-10d5-44d2-ab42-d38f88314c7b" ID-ISSN="1536-9307" ID-Zenodo-Dep="7124466" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638249082081" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David &amp; Š, V. A." docDate="2016" docId="B07187DFA876FFF1FC07FF329BF2FC60" docLanguage="en" docName="Euscorpius.220.1-133.pdf" docOrigin="Euscorpius 220" docSource="https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Heterometrus Ehrenberg 1828" docType="treatment" docVersion="10" lastPageNumber="96" masterDocId="4C48FFA7A817FF90FFC0FFD79E62FFBD" masterDocTitle="Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n" masterLastPageNumber="133" masterPageNumber="1" pageNumber="96" updateTime="1699471154095" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="989297989256E272BE88EE8DEB4BAF2E">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
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<mods:namePart id="788E0B25C5DD38A7CB3147434E8B2E03">Ranawana, Kithsiri B.</mods:namePart>
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<mods:namePart id="5DF162F243F6044C35C4FC65B12C133D">Hoferek, David</mods:namePart>
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<taxonomicName id="FFD84D4AA876FFF1FC07FF329B75FF40" ID-CoL="4WJC" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" box="[967,1303,229,253]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="97" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA876FFF1FC07FF329A3CFF41" bold="true" box="[967,1118,229,252]" italics="true" pageId="97" pageNumber="96">Heterometrus</emphasis>
Ehrenberg, 1828
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<subSubSection id="70C26542A876FFF1FC11FED39B6FFEA6" box="[977,1293,260,283]" pageId="97" pageNumber="96" type="description">
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(
<figureCitation id="A0E32A4CA876FFF1FC19FED39A50FEA6" box="[985,1074,260,283]" captionStart="Figures 12–15" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="97" pageNumber="96">Figs. 15</figureCitation>
,
<figureCitation id="A0E32A4CA876FFF1FBFDFED39AFFFEA6" box="[1085,1181,260,283]" captionStart-0="Figures 458–461" captionStart-1="Figures 462–469" captionStart-10="Figures 502–503" captionStart-11="Figures 504–511" captionStart-12="Figures 512–515" captionStart-13="Figures 516–519" captionStart-14="Figures 520–525" captionStart-15="Figures 526–529" captionStart-16="Figures 530–534" captionStart-17="Figures 535–536" captionStart-18="Figures 537–538" captionStart-19="Figures 539–543" captionStart-2="Figures 470–475" captionStart-20="Figures 544–546" captionStart-3="Figures 476–479" captionStart-4="Figures 480–481" captionStart-5="Figures 482–485" captionStart-6="Figures 486–489" captionStart-7="Figures 490–493" captionStart-8="Figures 494–499" captionStart-9="Figures 500–501" captionStartId-0="92.[192,278,1853,1877]" captionStartId-1="93.[192,278,1733,1757]" captionStartId-10="103.[192,278,1860,1884]" captionStartId-11="104.[192,278,1369,1393]" captionStartId-12="105.[192,278,1618,1642]" captionStartId-13="106.[192,278,1829,1853]" captionStartId-14="107.[192,278,1460,1484]" captionStartId-15="108.[192,278,1026,1050]" captionStartId-16="109.[192,278,1865,1889]" captionStartId-17="110.[192,278,1349,1373]" captionStartId-18="111.[192,278,1350,1374]" captionStartId-19="112.[192,278,1298,1322]" captionStartId-2="94.[192,278,1782,1806]" captionStartId-20="113.[192,278,1853,1877]" captionStartId-3="95.[192,278,1805,1829]" captionStartId-4="96.[192,278,1874,1898]" captionStartId-5="98.[192,278,1826,1850]" captionStartId-6="99.[192,278,1844,1868]" captionStartId-7="100.[192,278,1805,1829]" captionStartId-8="101.[192,278,1039,1063]" captionStartId-9="102.[192,278,1827,1851]" captionTargetBox-0="[250,1400,172,1819]" captionTargetBox-1="[191,1440,200,1705]" captionTargetBox-10="[191,1440,191,1831]" captionTargetBox-11="[195,1433,177,1322]" captionTargetBox-12="[204,1432,173,1576]" captionTargetBox-13="[213,1417,180,1801]" captionTargetBox-14="[197,1430,202,1414]" captionTargetBox-15="[199,1425,181,994]" captionTargetBox-16="[191,1440,191,1836]" captionTargetBox-17="[213,1421,158,1315]" captionTargetBox-18="[191,1440,191,1322]" captionTargetBox-19="[192,1439,153,1265]" captionTargetBox-2="[213,1432,197,1738]" captionTargetBox-20="[213,1417,167,1825]" captionTargetBox-3="[205,1422,219,1766]" captionTargetBox-4="[191,1440,167,1846]" captionTargetBox-5="[202,1418,200,1789]" captionTargetBox-6="[213,1417,167,1815]" captionTargetBox-7="[208,1424,198,1768]" captionTargetBox-8="[201,1435,171,999]" captionTargetBox-9="[191,1440,191,1798]" captionTargetId-0="figure-11@92.[233,1404,162,1824]" captionTargetId-1="figure-6@93.[192,1439,201,1704]" captionTargetId-10="figure-22@103.[192,1439,192,1830]" captionTargetId-11="figure-11@104.[192,1439,168,1340]" captionTargetId-12="figure-6@105.[192,1439,168,1589]" captionTargetId-13="figure-47@106.[214,1416,181,1800]" captionTargetId-14="figure-6@107.[192,1439,192,1431]" captionTargetId-15="figure-11@108.[192,1439,168,997]" captionTargetId-16="figure-12@109.[192,1439,192,1836]" captionTargetId-17="figure-11@110.[192,1439,142,1320]" captionTargetId-18="figure-393@111.[192,1439,192,1321]" captionTargetId-19="figure-11@112.[192,1439,140,1272]" captionTargetId-2="figure-11@94.[192,1439,192,1754]" captionTargetId-20="figure-30@113.[214,1416,168,1824]" captionTargetId-3="figure-21@95.[192,1439,212,1776]" captionTargetId-4="figure-37@96.[192,1439,168,1845]" captionTargetId-5="figure-11@98.[192,1439,192,1797]" captionTargetId-6="figure-33@99.[214,1416,168,1814]" captionTargetId-7="figure-11@100.[192,1439,189,1776]" captionTargetId-8="figure-6@101.[192,1439,168,1010]" captionTargetId-9="figure-35@102.[192,1439,192,1797]" captionTargetPageId-0="92" captionTargetPageId-1="93" captionTargetPageId-10="103" captionTargetPageId-11="104" captionTargetPageId-12="105" captionTargetPageId-13="106" captionTargetPageId-14="107" captionTargetPageId-15="108" captionTargetPageId-16="109" captionTargetPageId-17="110" captionTargetPageId-18="111" captionTargetPageId-19="112" captionTargetP
,
<figureCitation id="A0E32A4CA876FFF1FB67FED39B64FEA6" box="[1191,1286,260,283]" captionStart="Figures 571–574" captionStartId="116.[192,278,786,810]" captionTargetBox="[195,1423,184,735]" captionTargetId="figure-11@116.[192,1439,168,757]" captionTargetPageId="116" captionText="Figures 571–574: Key characters for distinguishing Sri Lankan Heterometrus. Figures 571–572. Pedipalp chela dorsal view of male, Heterometrus serratus from locality 15CP (571) and Heterometrus gravimanus from locality 15CE (572). Figures 573– 574. Metasomal segments IV-II lateral view of female, Heterometrus indus from locality 15CA (573) and Heterometrus serratus from locality 15CP (574)." figureDoi="http://doi.org/10.5281/zenodo.7124769" httpUri="https://zenodo.org/record/7124769/files/figure.png" pageId="97" pageNumber="96">571–574</figureCitation>
)
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<taxonomicName id="FFD84D4AA876FFF1FCFEFE959AABFEE4" authority="Ehrenberg" authorityName="Ehrenberg" box="[830,1225,322,345]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="97" pageNumber="96" phylum="Arthropoda" rank="subGenus" subGenus="Heterometrus">
<emphasis id="0AACEADBA876FFF1FCFEFE959DEAFEE4" box="[830,904,322,345]" italics="true" pageId="97" pageNumber="96">Buthus</emphasis>
(
<emphasis id="0AACEADBA876FFF1FC68FE959A59FEE4" box="[936,1083,322,345]" italics="true" pageId="97" pageNumber="96">Heterometrus</emphasis>
) Ehrenberg
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in
<bibRefCitation id="5C494B38A876FFF1FACAFE959A45FECA" author="Hemprich &amp; Ehrenberg" bookContentInfo="plate I Buthus; plate II: Androctonus" firstAuthor="Hemprich" journalOrPublisher="Berolini ex Officina Academica, Venditur a Mittlero" pageId="97" pageNumber="96" refId="ref41180" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1828. Zoologica II. Arachnoidea. In: Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg. Decas Prima. Berolini ex Officina Academica, Venditur a Mittlero: plate I Buthus; plate II: Androctonus." title="Zoologica II. Arachnoidea" type="book chapter" volumeTitle="Symbolae Physicae seu Icones et Descriptiones Animalium Evertebratorum Sepositis Insectis quae ex Itinere per Africam Borealem et Asiam Occidentalem. Friderici Guilelmi Hemprich et Christiani Godofredi Ehrenberg, Medicinae et Chirurgiae Doctorum, Studio Novae aut Illustratae Redierunt. Percensuit et Regis Iussu et Impensis Edidit Dr. C. G. Ehrenberg. Decas Prima" year="1828">Hemprich &amp; Ehrenberg, 1828</bibRefCitation>
: pl. 1, figs 1–2 (part, only fig. 2);
<treatmentCitation id="B97910D8A876FFF1FCAEFEA89A84FE2B" author="HEMPRICH, F. W. &amp; C. G. EHRENBERG" box="[878,1254,383,406]" page="351" pageId="97" pageNumber="96" year="1829">
<bibRefCitation id="5C494B38A876FFF1FCAEFEA89A84FE2B" author="Hemprich &amp; Ehrenberg" box="[878,1254,383,406]" firstAuthor="Hemprich" journalOrPublisher="Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin" page="351" pageId="97" pageNumber="96" pagination="348 - 362" part="1" refId="ref41277" refString="HEMPRICH, F. W. &amp; C. G. EHRENBERG. 1829. Vorlaufige Uebersicht der in Nord-Afrika und West- Asien einheimischen Skorpione und deren geographischen Verbreitung. Verhandungen der Gesellschaft der Naturforschender Freunde in Berlin, 1 (6): 348 - 362." title="Vorlaufige Uebersicht der in Nord-Afrika und West- Asien einheimischen Skorpione und deren geographischen Verbreitung" type="journal article" year="1829">Hemprich &amp; Ehrenberg, 1829: 351</bibRefCitation>
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.
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<treatmentCitationGroup id="18C811E7A876FFF1FCFEFE499B21FE4F" pageId="97" pageNumber="96">
<treatmentCitation id="B97910D8A876FFF1FCFEFE499AB3FE08" author="KOVARIK" box="[830,1233,414,437]" page="60" pageId="97" pageNumber="96" year="2004">
<taxonomicName id="FFD84D4AA876FFF1FCFEFE499AB3FE08" ID-CoL="4WJC" authority=": Kovarik, 2004: 1 - 60" authorityName=": Kovarik" authorityPageNumber="1 - 60" authorityYear="2004" box="[830,1233,414,437]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="97" pageNumber="96" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA876FFF1FCFEFE499DB3FE08" box="[830,977,414,437]" italics="true" pageId="97" pageNumber="96">Heterometrus</emphasis>
:
<bibRefCitation id="5C494B38A876FFF1FC25FE499AB3FE08" author="Kovarik" box="[997,1233,414,437]" firstAuthor="Kovarik" page="1 - 60" pageId="97" pageNumber="96" pagination="1 - 60" refId="ref41757" refString="KOVARIK, F. 2004. A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae). Euscorpius, 15: 1 - 60." type="journal article" year="2004">Kovařík, 2004: 1–60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1–32 (complete reference and synonymy list until 2004);
<treatmentCitation id="B97910D8A876FFF1FCAEFE0C9A3EFE4F" author="KOVARIK, F." box="[878,1116,475,498]" page="34" pageId="97" pageNumber="96" year="2009">
<bibRefCitation id="5C494B38A876FFF1FCAEFE0C9A3EFE4F" author="Kovarik" bookContentInfo="170 pp." box="[878,1116,475,498]" firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="34 - 49" pageId="97" pageNumber="96" refId="ref41793" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." title="Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species" type="book" year="2009">Kovařík, 2009: 34–49</bibRefCitation>
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, 73–113, figs. 1–283.
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<typeStatus id="E763886BA876FFF1FCFEFDCF9D15FD92" box="[830,887,536,559]" pageId="97" pageNumber="96">TYPE</typeStatus>
SPECIES.
<taxonomicName id="FFD84D4AA876FFF1FC2CFDCF9DDBFDF3" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="97" pageNumber="96" phylum="Arthropoda" rank="species" species="spinifer" subGenus="Heterometrus">
<emphasis id="0AACEADBA876FFF1FC2CFDCF9A54FD92" box="[1004,1078,536,559]" italics="true" pageId="97" pageNumber="96">Buthus</emphasis>
(
<emphasis id="0AACEADBA876FFF1FB8EFDCF9A83FD92" box="[1102,1249,536,559]" italics="true" pageId="97" pageNumber="96">Heterometrus</emphasis>
)
<emphasis id="0AACEADBA876FFF1FB38FDCF9B2BFD92" box="[1272,1353,536,559]" italics="true" pageId="97" pageNumber="96">spinifer</emphasis>
Ehrenberg, 1828
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, by subsequent designation (
<bibRefCitation id="5C494B38A876FFF1FAC3FDE09D3FFDD0" author="Karsch" firstAuthor="Karsch" page="20" pageId="97" pageNumber="96" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879: 20</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="70C26542A876FFF1FCFEFD449B26FC22" pageId="97" pageNumber="96" type="diagnosis">
<paragraph id="386736C9A876FFF1FCFEFD449B26FC22" blockId="97.[830,1440,659,927]" pageId="97" pageNumber="96">
DIAGNOSIS: Total length
<quantity id="FF209B2CA876FFF1FB94FD449AB9FD17" box="[1108,1243,659,682]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" metricValueMax="1.76" metricValueMin="0.6" pageId="97" pageNumber="96" unit="mm" value="118.0" valueMax="176.0" valueMin="60.0">60–176 mm</quantity>
. Orthobothriotaxy
<typeStatus id="E763886BA876FFF1FCFEFD659D0EFD74" box="[830,876,690,713]" pageId="97" pageNumber="96">type</typeStatus>
<collectorName id="952D531FA876FFF1FCB9FD659D9EFD74" box="[889,1020,690,713]" pageId="97" pageNumber="96">C. Pedipalp</collectorName>
femur with three trichobothria, only one of which resides on internal surface.
<collectorName id="952D531FA876FFF1FB35FD079B30FD5A" box="[1269,1362,720,743]" pageId="97" pageNumber="96">Pedipalp</collectorName>
patella with 19 trichobothria, 3 ventral, and 13 external.
<collectorName id="952D531FA876FFF1FAA2FD389BC2FCBB" box="[1378,1440,751,774]" pageId="97" pageNumber="96">Chela</collectorName>
of pedipalp with 26 trichobothria.
<collectorName id="952D531FA876FFF1FB19FCD99B35FC98" box="[1241,1367,782,805]" pageId="97" pageNumber="96">Retrolateral</collectorName>
pedal spurs absent.
<collectorName id="952D531FA876FFF1FC10FCFB9A37FCFE" box="[976,1109,812,835]" pageId="97" pageNumber="96">Lateroapical</collectorName>
margins of tarsi produced into rounded lobes.
<collectorName id="952D531FA876FFF1FC2FFC9C9A07FCDF" box="[1007,1125,843,866]" pageId="97" pageNumber="96">Metasomal</collectorName>
segments I–IV with paired ventral submedian carinae. Stridulatory organ located on opposing surfaces of pedipalp coxa and first leg.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A876FFF1FCFEFC119BF2FC60" box="[830,1424,966,989]" pageId="97" pageNumber="96" type="distribution">
<paragraph id="386736C9A876FFF1FCFEFC119BF2FC60" blockId="97.[830,1424,966,989]" box="[830,1424,966,989]" pageId="97" pageNumber="96">
DISTRIBUTION. Oriental region from
<collectingCountry id="40CF7659A876FFF1FB08FC119A9DFC60" box="[1224,1279,966,989]" name="India" pageId="97" pageNumber="96">India</collectingCountry>
to
<collectingCountry id="40CF7659A876FFF1FAE1FC119BEEFC60" box="[1313,1420,966,989]" name="Indonesia" pageId="97" pageNumber="96">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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