treatments-xml/data/03/82/40/03824077FFE1FFCCFF75FAAC4EB752D7.xml

<document id="34A6B5B7AF5AE4361E44DF5A813BA642" ID-CLB-Dataset="2590" ID-DOI="10.11646/zootaxa.5091.3.1" ID-GBIF-Dataset="948cc785-c168-4381-99c5-144ad7941dcf" ID-ISSN="1175-5326" ID-Zenodo-Dep="5848269" ID-ZooBank="747D0DDF-A440-4C7F-91F4-E5B758D28643" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642146842233" checkinUser="plazi" docAuthor="Montoya, Augusto León, Parada-Marín, Henry Mauricio &amp; Ramos-Pastrana, Yardany" docDate="2022" docId="03824077FFE1FFCCFF75FAAC4EB752D7" docLanguage="en" docName="zootaxa.5091.3.1.pdf" docOrigin="Zootaxa 5091 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copestylum vagum" docType="treatment" docVersion="5" lastPageNumber="411" masterDocId="FFBB380FFFE3FFC6FFE2FFBB4B2A5475" masterDocTitle="Description of a new flower fly species of the Copestylum vagum group (Diptera Syrphidae) from pristine Amazonian rainforests of Colombia and Suriname" masterLastPageNumber="415" masterPageNumber="401" pageNumber="403" updateTime="1699302271136" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8F38B4EF573D7BD3285EE8FD5E37A3B7">Description of a new flower fly species of the Copestylum vagum group (Diptera Syrphidae) from pristine Amazonian rainforests of Colombia and Suriname</mods:title>
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<mods:namePart id="26838F5D9069863EE583CE782D43BF31">Montoya, Augusto León</mods:namePart>
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<mods:affiliation id="722FC22A7EEC6F0446C320EB8D08D1C3">Universidad de Antioquia, Instituto de Biología, Grupo de Entomología - GEAU, Calle 67 53 - 108, AA 1226 Medellín, Colombia. aleon. montoya @ udea. edu. co; https: // orcid. org / 0000 - 0003 - 3307 - 034 X</mods:affiliation>
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<mods:namePart id="FD70645897BA0D7B3E07D16AFC40D594">Parada-Marín, Henry Mauricio</mods:namePart>
<mods:nameIdentifier id="25B3374C62A826318FE66C488742B91F" type="ORCID">0000-0002-5387-4495</mods:nameIdentifier>
<mods:affiliation id="1DF27B5B6BB34D296277056D668BDF4D">Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia. &amp; h. parada @ udla. edu. co; https: // orcid. org / 0000 - 0002 - 5387 - 4495</mods:affiliation>
<mods:nameIdentifier id="F0787F2147ED27F6037F6F80B0E272D5" type="email">h.parada@udla.edu.co</mods:nameIdentifier>
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<mods:namePart id="A128B5962346FEF252C31764252AC96B">Ramos-Pastrana, Yardany</mods:namePart>
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<mods:affiliation id="B180A21FD5000A256A19B9CAB4C4B8F1">Universidad de Antioquia, Instituto de Biología, Grupo de Entomología - GEAU, Calle 67 53 - 108, AA 1226 Medellín, Colombia. aleon. montoya @ udea. edu. co; https: // orcid. org / 0000 - 0003 - 3307 - 034 X &amp; Universidad de la Amazonia, Grupo de Investigación en Entomología Universidad de la Amazonia - GIEU-, Laboratório de Entomologia, Av, 11 5 - 69 Juan XXIII, Florencia, Caquetá, Colombia.</mods:affiliation>
<mods:nameIdentifier id="9E387792555655E1DAD67A576DCE30B2" type="email">aleon.montoya@udea.edu.co</mods:nameIdentifier>
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<treatment id="03824077FFE1FFCCFF75FAAC4EB752D7" ID-DOI="http://doi.org/10.5281/zenodo.5848259" ID-GBIF-Taxon="192128720" ID-Zenodo-Dep="5848259" LSID="urn:lsid:plazi:treatment:03824077FFE1FFCCFF75FAAC4EB752D7" httpUri="http://treatment.plazi.org/id/03824077FFE1FFCCFF75FAAC4EB752D7" lastPageId="10" lastPageNumber="411" pageId="2" pageNumber="403">
<subSubSection id="C331A2EAFFE1FFC4FF75FAAC494F5144" box="[151,613,1303,1330]" pageId="2" pageNumber="403" type="nomenclature">
<paragraph id="8B94F161FFE1FFC4FF75FAAC494F5144" blockId="2.[151,613,1303,1330]" box="[151,613,1303,1330]" pageId="2" pageNumber="403">
<heading id="D0DC460DFFE1FFC4FF75FAAC494F5144" bold="true" box="[151,613,1303,1330]" fontSize="11" level="1" pageId="2" pageNumber="403" reason="1">
<emphasis id="B95F2D73FFE1FFC4FF75FAAC494F5144" bold="true" box="[151,613,1303,1330]" pageId="2" pageNumber="403">
The 
<taxonomicName id="4C2B8AE2FFE1FFC4FF2DFAAC4A9C5144" ID-CoL="8BKPY" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[207,438,1303,1330]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE1FFC4FF2DFAAC4A9C5144" bold="true" box="[207,438,1303,1330]" italics="true" pageId="2" pageNumber="403">Copestylum vagum</emphasis>
</taxonomicName>
species group
</emphasis>
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</paragraph>
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<subSubSection id="C331A2EAFFE1FFC4FF75FAE449905200" pageId="2" pageNumber="403" type="diagnosis">
<paragraph id="8B94F161FFE1FFC4FF75FAE449905200" blockId="2.[151,1437,1375,1798]" pageId="2" pageNumber="403">
<emphasis id="B95F2D73FFE1FFC4FF75FAE44A20510C" bold="true" box="[151,266,1375,1401]" pageId="2" pageNumber="403">Diagnosis</emphasis>
(adapted from 
<bibRefCitation id="EFBA8C90FFE1FFC4FE54FAE44947510C" author="Thompson, F. C." box="[438,621,1375,1401]" pageId="2" pageNumber="403" refId="ref13414" refString="Thompson, F. C. (2006) Primer Taller de identificacion Syrphidae del Neotropico. February 21 th to 27 th 2006. Universidad del Valle, Facultad de Ciencias, Cali, Colombia, 860 pp." type="book" year="2006">Thompson 2006</bibRefCitation>
and 
<bibRefCitation id="EFBA8C90FFE1FFC4FD46FAE44853510F" author="Ricarte, A. &amp; Marcos-Garcia, M. A. &amp; Hancock, E. G. &amp; Rotheray, G. E." box="[676,889,1375,1402]" pageId="2" pageNumber="403" pagination="0142441" refId="ref12857" refString="Ricarte, A., Marcos-Garcia, M. A., Hancock, E. G. &amp; Rotheray, G. E. (2015) Neotropical Copestylum Macquart (Diptera: Syrphidae) breeding in fruits and flowers, including 7 new species. PLOS ONE, 10 (11), e 0142441. https: // doi. org / 10.1371 / journal. pone. 0142441" type="journal article" year="2015">
Ricarte 
<emphasis id="B95F2D73FFE1FFC4FD1FFADB4804510C" box="[765,814,1375,1401]" italics="true" pageId="2" pageNumber="403">et al</emphasis>
. 2015
</bibRefCitation>
). The 
<taxonomicName id="4C2B8AE2FFE1FFC4FC5CFADB4F00510C" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[958,1066,1376,1402]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE1FFC4FC5CFADB4F00510C" box="[958,1066,1376,1402]" italics="true" pageId="2" pageNumber="403">C. vagum</emphasis>
</taxonomicName>
group can be distinguished from the other 
<taxonomicName id="4C2B8AE2FFE1FFC4FF1DFA384AF151E8" box="[255,475,1411,1437]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95F2D73FFE1FFC4FF1DFA384AA951E8" box="[255,387,1411,1437]" italics="true" pageId="2" pageNumber="403">Copestylum</emphasis>
species
</taxonomicName>
groups by: 1) face vittate but not tomentose and with background colour shiny yellow, green, or orange, 2) face not particularly extended, 3) scutum with mixed length pile, not forming distinct layers and most orientated posteriorly, 4) no outstanding, long, thick setae along the rear margin of the scutum, although long pile may be present, 5) apex of the scutellum with a linear depression, 6) calypter dark, brown to black or with brown border, 7) tergum 1 and most of tergum 2 with a pale mark contrasting with the colour of the scutum, scutellum and the rest of the abdomen, and 8) male genitalia with aedeagal hood projecting ventrally from the hypandrium, superior lobe with smooth margins.
</paragraph>
</subSubSection>
<subSubSection id="C331A2EAFFE1FFC4FF25F9C44EBA5370" pageId="2" pageNumber="403" type="discussion">
<paragraph id="8B94F161FFE1FFC4FF25F9C44EBA5370" blockId="2.[151,1437,1375,1798]" pageId="2" pageNumber="403">
<emphasis id="B95F2D73FFE1FFC4FF25F9C44A1152EC" bold="true" box="[199,315,1663,1689]" pageId="2" pageNumber="403">Remarks.</emphasis>
The new species here described agrees with the diagnostic characters proposed for the 
<taxonomicName id="4C2B8AE2FFE1FFC4FACCF93B4EB752EC" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[1326,1437,1664,1690]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE1FFC4FACCF93B4EB752EC" box="[1326,1437,1664,1690]" italics="true" pageId="2" pageNumber="403">C. vagum</emphasis>
</taxonomicName>
group, but can easily be recognized within this group by the unique and striking male genitalia, characterized by the epandrium and cercus black; epandrium with a dorsal extension, a novel character among the group, in addition to the surstylus L-shaped, with two pairs of rounded ridges in the dorsal edge, similar to small deer antlers in velvet.
</paragraph>
</subSubSection>
<caption id="DF54A1E9FFE0FFC5FF75FAF14A6351DF" ID-DOI="http://doi.org/10.5281/zenodo.5848271" ID-Zenodo-Dep="5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="3" pageNumber="404" startId="3.[151,250,1354,1378]" targetBox="[189,1396,373,1329]" targetPageId="3">
<paragraph id="8B94F161FFE0FFC5FF75FAF14A6351DF" blockId="3.[151,1436,1354,1450]" pageId="3" pageNumber="404">
<emphasis id="B95F2D73FFE0FFC5FF75FAF14A3F5117" bold="true" box="[151,277,1354,1378]" pageId="3" pageNumber="404">FIGURE 1.</emphasis>
<taxonomicName id="4C2B8AE2FFE0FFC5FEFEFAF14ADA5117" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[284,496,1354,1378]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="404" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE0FFC5FEFEFAF14ADA5117" box="[284,496,1354,1378]" italics="true" pageId="3" pageNumber="404">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFE0FFC5FE1AFAF049605117" bold="true" box="[504,586,1355,1378]" pageId="3" pageNumber="404">
<taxonomicNameLabel id="A26C9008FFE0FFC5FE1AFAF049605117" box="[504,586,1355,1378]" pageId="3" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (LEUA–00000035890): 
<emphasis id="B95F2D73FFE0FFC5FC11FAF14F265117" bold="true" box="[1011,1036,1354,1378]" pageId="3" pageNumber="404">A.</emphasis>
Head, frontal view; 
<emphasis id="B95F2D73FFE0FFC5FB00FAF04FD05117" bold="true" box="[1250,1274,1355,1378]" pageId="3" pageNumber="404">B.</emphasis>
Habitus, dorsal view; 
<emphasis id="B95F2D73FFE0FFC5FF35FAD54BC551F3" bold="true" box="[215,239,1390,1414]" pageId="3" pageNumber="404">C.</emphasis>
Thorax, dorsal view, detail of maculae and pile patterns on scutum; 
<emphasis id="B95F2D73FFE0FFC5FC52FAD448E351F2" bold="true" box="[944,969,1391,1415]" pageId="3" pageNumber="404">D.</emphasis>
Lateral view, 
<emphasis id="B95F2D73FFE0FFC5FBBEFAD44F5E51F2" bold="true" box="[1116,1140,1391,1415]" pageId="3" pageNumber="404">E.</emphasis>
Latero-posterior view, detail of male genitalia.
</paragraph>
</caption>
<caption id="DF54A1E9FFE7FFC2FF75F88648515390" ID-DOI="http://doi.org/10.5281/zenodo.5848273" ID-Zenodo-Dep="5848273" httpUri="https://zenodo.org/record/5848273/files/figure.png" pageId="4" pageNumber="405" startId="4.[151,250,1853,1877]" targetBox="[175,1411,993,1828]" targetPageId="4">
<paragraph id="8B94F161FFE7FFC2FF75F88648515390" blockId="4.[151,1437,1853,2021]" pageId="4" pageNumber="405">
<emphasis id="B95F2D73FFE7FFC2FF75F8864A3F5320" bold="true" box="[151,277,1853,1877]" pageId="4" pageNumber="405">FIGURE 2.</emphasis>
<taxonomicName id="4C2B8AE2FFE7FFC2FEFCF8864AD85320" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[286,498,1853,1877]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE7FFC2FEFCF8864AD85320" box="[286,498,1853,1877]" italics="true" pageId="4" pageNumber="405">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFE7FFC2FE19F88649675320" bold="true" box="[507,589,1853,1877]" pageId="4" pageNumber="405">
<taxonomicNameLabel id="A26C9008FFE7FFC2FE19F88649675320" box="[507,589,1853,1877]" pageId="4" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype (LEUA–00000035890). 
<emphasis id="B95F2D73FFE7FFC2FC1BF8864F385320" bold="true" box="[1017,1042,1853,1877]" pageId="4" pageNumber="405">A.</emphasis>
Abdomen, lateral view; 
<emphasis id="B95F2D73FFE7FFC2FAF6F8864E015320" bold="true" box="[1300,1323,1853,1877]" pageId="4" pageNumber="405">B.</emphasis>
Abdomen, ventral view; 
<emphasis id="B95F2D73FFE7FFC2FEC6F8DA4A17530C" bold="true" box="[292,317,1889,1913]" pageId="4" pageNumber="405">C.</emphasis>
Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); 
<emphasis id="B95F2D73FFE7FFC2FE7DF83E4A9D53E8" bold="true" box="[415,439,1925,1949]" pageId="4" pageNumber="405">D.</emphasis>
Epandrium, dorsal view, detail of the dorsal extension (white arrow); 
<emphasis id="B95F2D73FFE7FFC2FB91F83E4FA053E8" bold="true" box="[1139,1162,1925,1949]" pageId="4" pageNumber="405">E.</emphasis>
Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: 
<emphasis id="B95F2D73FFE7FFC2FCFAF812481253B4" bold="true" box="[792,824,1961,1985]" pageId="4" pageNumber="405">Ah</emphasis>
= Aedeagal hood; 
<emphasis id="B95F2D73FFE7FFC2FC1BF8124F0B53B4" bold="true" box="[1017,1057,1961,1985]" pageId="4" pageNumber="405">Cer</emphasis>
= Cercus; 
<emphasis id="B95F2D73FFE7FFC2FB73F8124F9453B4" bold="true" box="[1169,1214,1961,1985]" pageId="4" pageNumber="405">Epd</emphasis>
= Epandrium; 
<emphasis id="B95F2D73FFE7FFC2FABBF8124EAD53B4" bold="true" box="[1369,1415,1961,1985]" pageId="4" pageNumber="405">Hyp</emphasis>
= Hypandrium; 
<emphasis id="B95F2D73FFE7FFC2FEC7F8764A6A5390" bold="true" box="[293,320,1997,2021]" pageId="4" pageNumber="405">Sa</emphasis>
= Surstylar apodeme; 
<emphasis id="B95F2D73FFE7FFC2FDFDF876491E5390" bold="true" box="[543,564,1997,2021]" pageId="4" pageNumber="405">Sl</emphasis>
= Superior lobe; 
<emphasis id="B95F2D73FFE7FFC2FD3DF876482C5390" bold="true" box="[735,774,1997,2021]" pageId="4" pageNumber="405">Sur</emphasis>
= Surstyle.
</paragraph>
</caption>
<caption id="DF54A1E9FFE6FFC3FF75FA8B48395119" ID-DOI="http://doi.org/10.5281/zenodo.5848275" ID-Zenodo-Dep="5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,250,1328,1352]" targetBox="[169,1428,273,1305]" targetPageId="5">
<paragraph id="8B94F161FFE6FFC3FF75FA8B48395119" blockId="5.[151,1436,1328,1388]" pageId="5" pageNumber="406">
<emphasis id="B95F2D73FFE6FFC3FF75FA8B4A39513D" bold="true" box="[151,275,1328,1352]" pageId="5" pageNumber="406">FIGURE 3.</emphasis>
<taxonomicName id="4C2B8AE2FFE6FFC3FEF8FA8B4AC6513D" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[282,492,1328,1352]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE6FFC3FEF8FA8B4AC6513D" box="[282,492,1328,1352]" italics="true" pageId="5" pageNumber="406">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFE6FFC3FE11FA8B4969513D" bold="true" box="[499,579,1328,1352]" pageId="5" pageNumber="406">
<taxonomicNameLabel id="A26C9008FFE6FFC3FE11FA8B4969513D" box="[499,579,1328,1352]" pageId="5" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, paratype (RMNH-collection). 
<emphasis id="B95F2D73FFE6FFC3FC34FA8B48C5513D" bold="true" box="[982,1007,1328,1352]" pageId="5" pageNumber="406">A.</emphasis>
Head, frontal view; 
<emphasis id="B95F2D73FFE6FFC3FB22FA8B4FF2513D" bold="true" box="[1216,1240,1328,1352]" pageId="5" pageNumber="406">B.</emphasis>
Head, lateral view, 
<emphasis id="B95F2D73FFE6FFC3FF75FAEF4B9A5119" bold="true" box="[151,176,1364,1388]" pageId="5" pageNumber="406">C.</emphasis>
Habitus, lateral view; 
<emphasis id="B95F2D73FFE6FFC3FE74FAEF4A855119" bold="true" box="[406,431,1364,1388]" pageId="5" pageNumber="406">D.</emphasis>
Thorax and abdomen, dorsal view.
</paragraph>
</caption>
<caption id="DF54A1E9FFE5FFC0FF75FE6B4D81559E" ID-Table-UUID="DF54A1E9FFE5FFC0FF75FE6B4D81559E" box="[151,1707,464,491]" httpUri="http://table.plazi.org/id/DF54A1E9FFE5FFC0FF75FE6B4D81559E" pageId="6" pageNumber="407" startId="6.[151,244,464,491]" targetBox="[160,2007,505,1697]" targetIsTable="true" targetPageId="6">
<paragraph id="8B94F161FFE5FFC0FF75FE6B4D81559E" blockId="6.[151,1707,464,491]" box="[151,1707,464,491]" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0FF75FE6B4A3A559E" bold="true" box="[151,272,464,491]" pageId="6" pageNumber="407">TABLE 1.</emphasis>
Morphological differences between males of 
<taxonomicName id="4C2B8AE2FFE5FFC0FCF0FE6B48D0559E" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[786,1018,464,491]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE5FFC0FCF0FE6B48D0559E" box="[786,1018,464,491]" italics="true" pageId="6" pageNumber="407">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFE5FFC0FBE0FE6B4F71559E" bold="true" box="[1026,1115,464,491]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="A26C9008FFE5FFC0FBE0FE6B4F71559E" box="[1026,1115,464,491]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and similar species of the 
<taxonomicName id="4C2B8AE2FFE5FFC0FA64FE6B4D76559E" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[1414,1628,464,491]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE5FFC0FA64FE6B4D76559E" box="[1414,1628,464,491]" italics="true" pageId="6" pageNumber="407">Copestylum vagum</emphasis>
</taxonomicName>
group.
</paragraph>
</caption>
<paragraph id="8B94F161FFE5FFC0FF42FE414CB1522C" pageId="6" pageNumber="407">
<table id="F92B03C1FFE50039FF42FE424CFD52D4" box="[160,2007,505,1697]" colsContinueIn="7.[151,2014,733,1500]" gridcols="6" gridrows="10" pageId="6" pageNumber="407">
<tr id="351BF323FFE50039FF42FE424CFD5666" box="[160,2007,505,531]" gridrow="0" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FE424A675666" box="[160,333,505,531]" gridcol="0" gridrow="0" pageId="6" pageNumber="407">Character</th>
<th id="76CA9A5FFFE50039FE83FE42498E5666" box="[353,676,505,531]" gridcol="1" gridrow="0" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0FE83FE4249BA5667" bold="true" box="[353,656,505,530]" pageId="6" pageNumber="407">
<taxonomicName id="4C2B8AE2FFE5FFC0FE83FE4249135667" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[353,569,505,530]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE5FFC0FE83FE4249135667" bold="true" box="[353,569,505,530]" italics="true" pageId="6" pageNumber="407">Copestylum enriquei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C9008FFE5FFC0FDDDFE4249BA5667" box="[575,656,505,530]" pageId="6" pageNumber="407" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CA9A5FFFE50039FD59FE4248E65666" box="[699,972,505,531]" gridcol="2" gridrow="0" pageId="6" pageNumber="407">
<taxonomicName id="4C2B8AE2FFE5FFC0FD59FE4248A85667" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[699,898,505,530]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE5FFC0FD59FE4248A85667" bold="true" box="[699,898,505,530]" italics="true" pageId="6" pageNumber="407">Copestylum vagum</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE50039FC05FE424E2D5666" box="[999,1287,505,531]" gridcol="3" gridrow="0" pageId="6" pageNumber="407">
<taxonomicName id="4C2B8AE2FFE5FFC0FC05FE424FE25667" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[999,1224,505,530]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="tenorium">
<emphasis id="B95F2D73FFE5FFC0FC05FE424FE25667" bold="true" box="[999,1224,505,530]" italics="true" pageId="6" pageNumber="407">Copestylum tenorium</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE50039FAC5FE424D505666" box="[1319,1658,505,531]" gridcol="4" gridrow="0" pageId="6" pageNumber="407">
<taxonomicName id="4C2B8AE2FFE5FFC0FAC5FE424D085667" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[1319,1570,505,530]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="musicanum">
<emphasis id="B95F2D73FFE5FFC0FAC5FE424D085667" bold="true" box="[1319,1570,505,530]" italics="true" pageId="6" pageNumber="407">Copestylum musicanum</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE50039F973FE424CFD5666" box="[1681,2007,505,531]" gridcol="5" gridrow="0" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0F973FE424CB85667" bold="true" box="[1681,1938,505,530]" italics="true" pageId="6" pageNumber="407">Copestylum chapadensis</emphasis>
</th>
</tr>
<tr id="351BF323FFE50039FF42FD994CFD5616" box="[160,2007,546,611]" gridrow="1" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FD994A675616" box="[160,333,546,611]" gridcol="0" gridrow="1" pageId="6" pageNumber="407">
Body length 
<emphasis id="B95F2D73FFE5FFC0FF42FDF14BF05616" bold="true" box="[160,218,586,611]" pageId="6" pageNumber="407">Head</emphasis>
</th>
<td id="76CA9A5FFFE50039FE83FD99498E5616" box="[353,676,546,611]" gridcol="1" gridrow="1" pageId="6" pageNumber="407">6.7−7.1</td>
<td id="76CA9A5FFFE50039FD59FD9948E65616" box="[699,972,546,611]" gridcol="2" gridrow="1" pageId="6" pageNumber="407">8.5−9.3</td>
<td id="76CA9A5FFFE50039FC05FD994E2D5616" box="[999,1287,546,611]" gridcol="3" gridrow="1" pageId="6" pageNumber="407">8.3−10</td>
<td id="76CA9A5FFFE50039FAC5FD994D505616" box="[1319,1658,546,611]" gridcol="4" gridrow="1" pageId="6" pageNumber="407">8.2−9.1</td>
<td id="76CA9A5FFFE50039F973FD994CFD5616" box="[1681,2007,546,611]" gridcol="5" gridrow="1" pageId="6" pageNumber="407">9.0−9.6</td>
</tr>
<tr id="351BF323FFE50039FF42FDC94CFD56DA" box="[160,2007,626,687]" gridrow="2" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FDC94A6756DA" box="[160,333,626,687]" gridcol="0" gridrow="2" pageId="6" pageNumber="407">Facial lateral vitta</th>
<td id="76CA9A5FFFE50039FE83FDC9498E56DA" box="[353,676,626,687]" gridcol="1" gridrow="2" pageId="6" pageNumber="407">Strong and wide, wider than the gena</td>
<td id="76CA9A5FFFE50039FD59FDC948E656DA" box="[699,972,626,687]" gridcol="2" gridrow="2" pageId="6" pageNumber="407">Narrower than the gena</td>
<td id="76CA9A5FFFE50039FC05FDC94E2D56DA" box="[999,1287,626,687]" gridcol="3" gridrow="2" pageId="6" pageNumber="407">Narrower than the gena</td>
<td id="76CA9A5FFFE50039FAC5FDC94D5056DA" box="[1319,1658,626,687]" gridcol="4" gridrow="2" pageId="6" pageNumber="407">Narrower than the gena</td>
<td id="76CA9A5FFFE50039F973FDC94CFD56DA" box="[1681,2007,626,687]" gridcol="5" gridrow="2" pageId="6" pageNumber="407">Very wide and of uniform width</td>
</tr>
<tr id="351BF323FFE50039FF42FD054CFD5751" box="[160,2007,702,804]" gridrow="3" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FD054A675751" box="[160,333,702,804]" gridcol="0" gridrow="3" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0FF42FD054BDA56A2" bold="true" box="[160,240,702,727]" pageId="6" pageNumber="407">Thorax</emphasis>
Dorsal margin of male eyes
</th>
<td id="76CA9A5FFFE50039FE83FD05498E5751" box="[353,676,702,804]" gridcol="1" gridrow="3" pageId="6" pageNumber="407">Sloping</td>
<td id="76CA9A5FFFE50039FD59FD0548E65751" box="[699,972,702,804]" gridcol="2" gridrow="3" pageId="6" pageNumber="407">Sloping</td>
<td id="76CA9A5FFFE50039FC05FD054E2D5751" box="[999,1287,702,804]" gridcol="3" gridrow="3" pageId="6" pageNumber="407">Flattened</td>
<td id="76CA9A5FFFE50039FAC5FD054D505751" box="[1319,1658,702,804]" gridcol="4" gridrow="3" pageId="6" pageNumber="407">Sloping</td>
<td id="76CA9A5FFFE50039F973FD054CFD5751" box="[1681,2007,702,804]" gridcol="5" gridrow="3" pageId="6" pageNumber="407">Sloping</td>
</tr>
<tr id="351BF323FFE50039FF42FCEC4CFD5075" box="[160,2007,855,1024]" gridrow="4" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FCEC4A675075" box="[160,333,855,1024]" gridcol="0" gridrow="4" pageId="6" pageNumber="407">Scutum pattern</th>
<td id="76CA9A5FFFE50039FE83FCEC498E5075" box="[353,676,855,1024]" gridcol="1" gridrow="4" pageId="6" pageNumber="407">Orange except for the wide medial vitta, which is dark and metallic, ending before the prescutellar region, with the apical margin M-shaped</td>
<td id="76CA9A5FFFE50039FD59FCEC48E65075" box="[699,972,855,1024]" gridcol="2" gridrow="4" pageId="6" pageNumber="407">Orange except for yellow anterior and lateral margins, the middle region from scutum to scutellum dark and metallic</td>
<td id="76CA9A5FFFE50039FC05FCEC4E2D5075" box="[999,1287,855,1024]" gridcol="3" gridrow="4" pageId="6" pageNumber="407">Dark greenish-orange with a central, brown prescutellar macula and similar coloured maculae on the scutum</td>
<td id="76CA9A5FFFE50039FAC5FCEC4D505075" box="[1319,1658,855,1024]" gridcol="4" gridrow="4" pageId="6" pageNumber="407">Orange except for yellow anterior and lateral margins and brown prescutellar macula and two brown, medial vitta ending before the transverse suture</td>
<td id="76CA9A5FFFE50039F973FCEC4CFD5075" box="[1681,2007,855,1024]" gridcol="5" gridrow="4" pageId="6" pageNumber="407">Orange with a brown macula reaching the scutellar region, scutellar depression black</td>
</tr>
<tr id="351BF323FFE50039FF42FB884CFD5005" box="[160,2007,1075,1136]" gridrow="5" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FB884A675005" box="[160,333,1075,1136]" gridcol="0" gridrow="5" pageId="6" pageNumber="407">Setae above wing insertion</th>
<td id="76CA9A5FFFE50039FE83FB88498E5005" box="[353,676,1075,1136]" gridcol="1" gridrow="5" pageId="6" pageNumber="407">2</td>
<td id="76CA9A5FFFE50039FD59FB8848E65005" box="[699,972,1075,1136]" gridcol="2" gridrow="5" pageId="6" pageNumber="407">2</td>
<td id="76CA9A5FFFE50039FC05FB884E2D5005" box="[999,1287,1075,1136]" gridcol="3" gridrow="5" pageId="6" pageNumber="407">4</td>
<td id="76CA9A5FFFE50039FAC5FB884D505005" box="[1319,1658,1075,1136]" gridcol="4" gridrow="5" pageId="6" pageNumber="407">3</td>
<td id="76CA9A5FFFE50039F973FB884CFD5005" box="[1681,2007,1075,1136]" gridcol="5" gridrow="5" pageId="6" pageNumber="407">3</td>
</tr>
<tr id="351BF323FFE50039FF42FB184CFD5095" box="[160,2007,1187,1248]" gridrow="6" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FB184A675095" box="[160,333,1187,1248]" gridcol="0" gridrow="6" pageId="6" pageNumber="407">Pleuron</th>
<td id="76CA9A5FFFE50039FE83FB18498E5095" box="[353,676,1187,1248]" gridcol="1" gridrow="6" pageId="6" pageNumber="407">Extensively yellow</td>
<td id="76CA9A5FFFE50039FD59FB1848E65095" box="[699,972,1187,1248]" gridcol="2" gridrow="6" pageId="6" pageNumber="407">With black maculae</td>
<td id="76CA9A5FFFE50039FC05FB184E2D5095" box="[999,1287,1187,1248]" gridcol="3" gridrow="6" pageId="6" pageNumber="407">Extensively yellow</td>
<td id="76CA9A5FFFE50039FAC5FB184D505095" box="[1319,1658,1187,1248]" gridcol="4" gridrow="6" pageId="6" pageNumber="407">Extensively yellow except slightly darkened on the katatergum</td>
<td id="76CA9A5FFFE50039F973FB184CFD5095" box="[1681,2007,1187,1248]" gridcol="5" gridrow="6" pageId="6" pageNumber="407">Extensively yellow</td>
</tr>
<tr id="351BF323FFE50039FF42FAA84CFD510C" box="[160,2007,1299,1401]" gridrow="7" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FAA84A67510C" box="[160,333,1299,1401]" gridcol="0" gridrow="7" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0FF42FAA84BF95159" bold="true" box="[160,211,1299,1324]" pageId="6" pageNumber="407">Legs</emphasis>
Metafemur
</th>
<td id="76CA9A5FFFE50039FE83FAA8498E510C" box="[353,676,1299,1401]" gridcol="1" gridrow="7" pageId="6" pageNumber="407">Orange</td>
<td id="76CA9A5FFFE50039FD59FAA848E6510C" box="[699,972,1299,1401]" gridcol="2" gridrow="7" pageId="6" pageNumber="407">Orange except for black or blackened apex</td>
<td id="76CA9A5FFFE50039FC05FAA84E2D510C" box="[999,1287,1299,1401]" gridcol="3" gridrow="7" pageId="6" pageNumber="407">Greenish-orange</td>
<td id="76CA9A5FFFE50039FAC5FAA84D50510C" box="[1319,1658,1299,1401]" gridcol="4" gridrow="7" pageId="6" pageNumber="407">Orange on the basal area</td>
<td id="76CA9A5FFFE50039F973FAA84CFD510C" box="[1681,2007,1299,1401]" gridcol="5" gridrow="7" pageId="6" pageNumber="407">Orange</td>
</tr>
<tr id="351BF323FFE50039FF42FA174CFD5278" box="[160,2007,1452,1549]" gridrow="8" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42FA174A675278" box="[160,333,1452,1549]" gridcol="0" gridrow="8" pageId="6" pageNumber="407">Tibia</th>
<td id="76CA9A5FFFE50039FE83FA17498E5278" box="[353,676,1452,1549]" gridcol="1" gridrow="8" pageId="6" pageNumber="407">Dark-brown, except yellow on basal 1/4, black pilose on these areas</td>
<td id="76CA9A5FFFE50039FD59FA1748E65278" box="[699,972,1452,1549]" gridcol="2" gridrow="8" pageId="6" pageNumber="407">Black, black pilose</td>
<td id="76CA9A5FFFE50039FC05FA174E2D5278" box="[999,1287,1452,1549]" gridcol="3" gridrow="8" pageId="6" pageNumber="407">Black, black pilose</td>
<td id="76CA9A5FFFE50039FAC5FA174D505278" box="[1319,1658,1452,1549]" gridcol="4" gridrow="8" pageId="6" pageNumber="407">Darkened, black pilose</td>
<td id="76CA9A5FFFE50039F973FA174CFD5278" box="[1681,2007,1452,1549]" gridcol="5" gridrow="8" pageId="6" pageNumber="407">Black, black pilose</td>
</tr>
<tr id="351BF323FFE50039FF42F9FB4CFD52D4" box="[160,2007,1600,1697]" gridrow="9" pageId="6" pageNumber="407">
<th id="76CA9A5FFFE50039FF42F9FB4A6752D4" box="[160,333,1600,1697]" gridcol="0" gridrow="9" pageId="6" pageNumber="407">Tarsomeres</th>
<td id="76CA9A5FFFE50039FE83F9FB498E52D4" box="[353,676,1600,1697]" gridcol="1" gridrow="9" pageId="6" pageNumber="407">Tarsomeres 1 and 2 orange, tarsomeres 3–5 black, black pilose</td>
<td id="76CA9A5FFFE50039FD59F9FB48E652D4" box="[699,972,1600,1697]" gridcol="2" gridrow="9" pageId="6" pageNumber="407">Basitarsomere orange, tarsomeres 3–5 black, black pilose</td>
<td id="76CA9A5FFFE50039FC05F9FB4E2D52D4" box="[999,1287,1600,1697]" gridcol="3" gridrow="9" pageId="6" pageNumber="407">Black, black pilose</td>
<td id="76CA9A5FFFE50039FAC5F9FB4D5052D4" box="[1319,1658,1600,1697]" gridcol="4" gridrow="9" pageId="6" pageNumber="407">Basotarsomere of meso- and metalegs contrastingly pale, black pilose</td>
<td id="76CA9A5FFFE50039F973F9FB4CFD52D4" box="[1681,2007,1600,1697]" gridcol="5" gridrow="9" pageId="6" pageNumber="407">Tarsi reddish, black pilose</td>
</tr>
</table>
</paragraph>
<paragraph id="8B94F161FFE5FFC0F95AF90A4CC552BF" blockId="6.[1720,2031,1713,1738]" box="[1720,2031,1713,1738]" pageId="6" pageNumber="407">
<emphasis id="B95F2D73FFE5FFC0F95AF90A4CC552BF" box="[1720,2031,1713,1738]" italics="true" pageId="6" pageNumber="407">......continued on the next page</emphasis>
</paragraph>
<caption id="DF54A1E9FFE4FFC1FF75FD0E4AAF56BB" box="[151,389,693,718]" isContinuationCaption="true" pageId="7" pageNumber="408" startId="7.[151,235,693,718]" targetBox="[151,2014,733,1500]" targetIsTable="true" targetPageId="7">
<paragraph id="8B94F161FFE4FFC1FF75FD0E4AAF56BB" blockId="7.[151,389,693,718]" box="[151,389,693,718]" pageId="7" pageNumber="408">
<emphasis id="B95F2D73FFE4FFC1FF75FD0E4A2E56BB" bold="true" box="[151,260,693,718]" pageId="7" pageNumber="408">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B94F161FFE4FFC1FF75FD664CAC5105" pageId="7" pageNumber="408">
<table id="F92B03C1FFE40039FF75FD664CF451A9" box="[151,2014,733,1500]" colsContinueFrom="6.[160,2007,505,1697]" gridcols="6" gridrows="6" pageId="7" pageNumber="408">
<tr id="351BF323FFE40039FF75FD664CF45683" box="[151,2014,733,758]" gridrow="0" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FD664A145683" box="[151,318,733,758]" gridcol="0" gridrow="0" pageId="7" pageNumber="408">Character</th>
<th id="76CA9A5FFFE40039FEB5FD6649B75683" box="[343,669,733,758]" gridcol="1" gridrow="0" pageId="7" pageNumber="408">
<emphasis id="B95F2D73FFE4FFC1FEB5FD6649AC5683" bold="true" box="[343,646,733,758]" pageId="7" pageNumber="408">
<taxonomicName id="4C2B8AE2FFE4FFC1FEB5FD6649055683" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[343,559,733,758]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFE4FFC1FEB5FD6649055683" bold="true" box="[343,559,733,758]" italics="true" pageId="7" pageNumber="408">Copestylum enriquei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C9008FFE4FFC1FDD4FD6649AC5683" box="[566,646,733,758]" pageId="7" pageNumber="408" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CA9A5FFFE40039FD50FD6648ED5683" box="[690,967,733,758]" gridcol="2" gridrow="0" pageId="7" pageNumber="408">
<taxonomicName id="4C2B8AE2FFE4FFC1FD50FD6648525683" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[690,888,733,758]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFE4FFC1FD50FD6648525683" bold="true" box="[690,888,733,758]" italics="true" pageId="7" pageNumber="408">Copestylum vagum</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE40039FC3CFD664E2E5683" box="[990,1284,733,758]" gridcol="3" gridrow="0" pageId="7" pageNumber="408">
<taxonomicName id="4C2B8AE2FFE4FFC1FC3CFD664F955683" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[990,1215,733,758]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="tenorium">
<emphasis id="B95F2D73FFE4FFC1FC3CFD664F955683" bold="true" box="[990,1215,733,758]" italics="true" pageId="7" pageNumber="408">Copestylum tenorium</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE40039FAFCFD664D5E5683" box="[1310,1652,733,758]" gridcol="4" gridrow="0" pageId="7" pageNumber="408">
<taxonomicName id="4C2B8AE2FFE4FFC1FAFCFD664D335683" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[1310,1561,733,758]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="musicanum">
<emphasis id="B95F2D73FFE4FFC1FAFCFD664D335683" bold="true" box="[1310,1561,733,758]" italics="true" pageId="7" pageNumber="408">Copestylum musicanum</emphasis>
</taxonomicName>
</th>
<th id="76CA9A5FFFE40039F965FD664CF45683" box="[1671,2014,733,758]" gridcol="5" gridrow="0" pageId="7" pageNumber="408">
<emphasis id="B95F2D73FFE4FFC1F965FD664CA25683" bold="true" box="[1671,1928,733,758]" italics="true" pageId="7" pageNumber="408">Copestylum chapadensis</emphasis>
</th>
</tr>
<tr id="351BF323FFE40039FF75FCBD4CF457A6" box="[151,2014,774,979]" gridrow="1" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FCBD4A1457A6" box="[151,318,774,979]" gridcol="0" gridrow="1" pageId="7" pageNumber="408">Terga 2−4</th>
<td id="76CA9A5FFFE40039FEB5FCBD49B757A6" box="[343,669,774,979]" gridcol="1" gridrow="1" pageId="7" pageNumber="408">Terga 2–4 yellow, brownish infuscated apically; Terga 3–4 black pilose on apical margin</td>
<td id="76CA9A5FFFE40039FD50FCBD48ED57A6" box="[690,967,774,979]" gridcol="2" gridrow="1" pageId="7" pageNumber="408">Terga 2–4 yellow except for a black apex; Tergum 4 white pilose</td>
<td id="76CA9A5FFFE40039FC3CFCBD4E2E57A6" box="[990,1284,774,979]" gridcol="3" gridrow="1" pageId="7" pageNumber="408">Terga 2–4 greenish-orange except for a black apex; Terga 3–4 black pilose</td>
<td id="76CA9A5FFFE40039FAFCFCBD4D5E57A6" box="[1310,1652,774,979]" gridcol="4" gridrow="1" pageId="7" pageNumber="408">Tergum 2 yellow except for black posterior margin; Terga 3–4 black, black pilose</td>
<td id="76CA9A5FFFE40039F965FCBD4CF457A6" box="[1671,2014,774,979]" gridcol="5" gridrow="1" pageId="7" pageNumber="408">Terga 2–4 yellow except black posteriorly; pile short, black, yellow in front of the first black band, on lateral margins, and the broad base of tergum 4. Tergum 5 wholly black</td>
</tr>
<tr id="351BF323FFE40039FF75FBBD4CF4506A" box="[151,2014,1030,1055]" gridrow="2" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FBBD4CF4506A" box="[151,2014,1030,1055]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="7" pageNumber="408">
<emphasis id="B95F2D73FFE4FFC1FF75FBBD4BD1506A" bold="true" box="[151,251,1030,1055]" pageId="7" pageNumber="408">Genitalia</emphasis>
</th>
</tr>
<tr id="351BF323FFE40039FF75FB944CF450E5" box="[151,2014,1071,1168]" gridrow="3" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FB944A1450E5" box="[151,318,1071,1168]" gridcol="0" gridrow="3" pageId="7" pageNumber="408">Cercus</th>
<td id="76CA9A5FFFE40039FEB5FB9449B750E5" box="[343,669,1071,1168]" gridcol="1" gridrow="3" pageId="7" pageNumber="408">Black, globular and roundtipped</td>
<td id="76CA9A5FFFE40039FD50FB9448ED50E5" box="[690,967,1071,1168]" gridcol="2" gridrow="3" pageId="7" pageNumber="408">Orange, variable in shape, usually slightly longer than basally wide</td>
<td id="76CA9A5FFFE40039FC3CFB944E2E50E5" box="[990,1284,1071,1168]" gridcol="3" gridrow="3" pageId="7" pageNumber="408">Orange, tapered and about as long as basally wide</td>
<td id="76CA9A5FFFE40039FAFCFB944D5E50E5" box="[1310,1652,1071,1168]" gridcol="4" gridrow="3" pageId="7" pageNumber="408">Orange, L-shaped</td>
<td id="76CA9A5FFFE40039F965FB944CF450E5" box="[1671,2014,1071,1168]" gridcol="5" gridrow="3" pageId="7" pageNumber="408">No information available</td>
</tr>
<tr id="351BF323FFE40039FF75FB784CF45151" box="[151,2014,1219,1316]" gridrow="4" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FB784A145151" box="[151,318,1219,1316]" gridcol="0" gridrow="4" pageId="7" pageNumber="408">Surstylus</th>
<td id="76CA9A5FFFE40039FEB5FB7849B75151" box="[343,669,1219,1316]" gridcol="1" gridrow="4" pageId="7" pageNumber="408">Orange, L-shaped, with two pairs of rounded ridges in the dorsal edge</td>
<td id="76CA9A5FFFE40039FD50FB7848ED5151" box="[690,967,1219,1316]" gridcol="2" gridrow="4" pageId="7" pageNumber="408">Apically tapering</td>
<td id="76CA9A5FFFE40039FC3CFB784E2E5151" box="[990,1284,1219,1316]" gridcol="3" gridrow="4" pageId="7" pageNumber="408">Subrectangular abruptly indented below the middle</td>
<td id="76CA9A5FFFE40039FAFCFB784D5E5151" box="[1310,1652,1219,1316]" gridcol="4" gridrow="4" pageId="7" pageNumber="408">L-shaped, basal section longer than the apical section</td>
<td id="76CA9A5FFFE40039F965FB784CF45151" box="[1671,2014,1219,1316]" gridcol="5" gridrow="4" pageId="7" pageNumber="408">No information available</td>
</tr>
<tr id="351BF323FFE40039FF75FAEC4CF451A9" box="[151,2014,1367,1500]" gridrow="5" pageId="7" pageNumber="408">
<th id="76CA9A5FFFE40039FF75FAEC4A1451A9" box="[151,318,1367,1500]" gridcol="0" gridrow="5" pageId="7" pageNumber="408">Superior lobe (Apex of hypandrium)</th>
<td id="76CA9A5FFFE40039FEB5FAEC49B751A9" box="[343,669,1367,1500]" gridcol="1" gridrow="5" pageId="7" pageNumber="408">Hook-shaped, with a kinked apex strongly curved downward</td>
<td id="76CA9A5FFFE40039FD50FAEC48ED51A9" box="[690,967,1367,1500]" gridcol="2" gridrow="5" pageId="7" pageNumber="408">With a kinked apex</td>
<td id="76CA9A5FFFE40039FC3CFAEC4E2E51A9" box="[990,1284,1367,1500]" gridcol="3" gridrow="5" pageId="7" pageNumber="408">Hook-shaped and curved unevenly with a longer section above the base and tapering to a point</td>
<td id="76CA9A5FFFE40039FAFCFAEC4D5E51A9" box="[1310,1652,1367,1500]" gridcol="4" gridrow="5" pageId="7" pageNumber="408">With a kinked apex strongly curved downward</td>
<td id="76CA9A5FFFE40039F965FAEC4CF451A9" box="[1671,2014,1367,1500]" gridcol="5" gridrow="5" pageId="7" pageNumber="408">No information available</td>
</tr>
</table>
</paragraph>
<caption id="DF54A1E9FFEBFFCEFF75FC404FD95029" ID-DOI="http://doi.org/10.5281/zenodo.5848277" ID-Zenodo-Dep="5848277" httpUri="https://zenodo.org/record/5848277/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,250,1019,1043]" targetBox="[153,1436,490,995]" targetPageId="8">
<paragraph id="8B94F161FFEBFFCEFF75FC404FD95029" blockId="8.[151,1437,1019,1116]" pageId="8" pageNumber="409">
<emphasis id="B95F2D73FFEBFFCEFF75FC404A185066" bold="true" box="[151,306,1019,1043]" pageId="8" pageNumber="409">FIGURE 4. A.</emphasis>
Habitat of 
<taxonomicName id="4C2B8AE2FFEBFFCEFE44FC4749535066" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[422,633,1020,1044]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFEBFFCEFE44FC4749535066" box="[422,633,1020,1044]" italics="true" pageId="8" pageNumber="409">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFEBFFCEFD62FC4749FB5061" bold="true" box="[640,721,1020,1044]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A26C9008FFEBFFCEFD62FC4749FB5061" box="[640,721,1020,1044]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in a pristine forest at Florencia, Caquetá in Colombia. The specimens were collected using a canopy Malaise trap (Rafael &amp; Gorayeb 1982); 
<emphasis id="B95F2D73FFEBFFCEFC82FB9B485D504D" bold="true" box="[864,887,1056,1080]" pageId="8" pageNumber="409">B.</emphasis>
Geographical distribution map showing the collection localities of 
<taxonomicName id="4C2B8AE2FFEBFFCEFEF6FBFF4ACC502E" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[276,486,1092,1116]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFEBFFCEFEF6FBFF4ACC502E" box="[276,486,1092,1116]" italics="true" pageId="8" pageNumber="409">Copestylum enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFEBFFCEFE0FFBFF49145029" bold="true" box="[493,574,1092,1116]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="A26C9008FFEBFFCEFE0FFBFF49145029" box="[493,574,1092,1116]" pageId="8" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the Amazon rainforest of Colombia and Suriname (Red triangle).
</paragraph>
</caption>
<subSection id="E2A4E88AFFEBFFCEFF75FB154F38512D" pageId="8" pageNumber="409" type="multiple">
<subSubSection id="C331A2EAFFEBFFCEFF75FB154F38512D" pageId="8" pageNumber="409" type="distribution">
<paragraph id="8B94F161FFEBFFCEFF75FB15484150BD" blockId="8.[151,875,1197,1224]" box="[151,875,1197,1224]" pageId="8" pageNumber="409">
<heading id="D0DC460DFFEBFFCEFF75FB15484150BD" bold="true" box="[151,875,1197,1224]" fontSize="11" level="1" pageId="8" pageNumber="409" reason="1">
<emphasis id="B95F2D73FFEBFFCEFF75FB15484150BD" bold="true" box="[151,875,1197,1224]" pageId="8" pageNumber="409">
Additional records of species of the 
<taxonomicName id="4C2B8AE2FFEBFFCEFDA3FB16483750BD" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[577,797,1197,1224]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEBFFCEFDA3FB16483750BD" bold="true" box="[577,797,1197,1224]" italics="true" pageId="8" pageNumber="409">Copestylum vagum</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
<paragraph id="8B94F161FFEBFFCEFF75FB4D4F38512D" blockId="8.[151,1437,1270,1368]" pageId="8" pageNumber="409">
New distributional records of 
<taxonomicName id="4C2B8AE2FFEBFFCEFE3FFB4D494A5165" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[477,608,1270,1296]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="tenorium">
<emphasis id="B95F2D73FFEBFFCEFE3FFB4D494A5165" box="[477,608,1270,1296]" italics="true" pageId="8" pageNumber="409">C. tenorium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2B8AE2FFEBFFCEFD70FB4D48015165" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[658,811,1270,1296]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="araceorum">
<emphasis id="B95F2D73FFEBFFCEFD70FB4D48015165" box="[658,811,1270,1296]" italics="true" pageId="8" pageNumber="409">C. araceorum</emphasis>
</taxonomicName>
are here provided. 
<taxonomicName id="4C2B8AE2FFEBFFCEFC18FB4D4FC25165" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[1018,1256,1270,1296]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="tenorium">
<emphasis id="B95F2D73FFEBFFCEFC18FB4D4FC25165" box="[1018,1256,1270,1296]" italics="true" pageId="8" pageNumber="409">Copestylum tenorium</emphasis>
</taxonomicName>
was exclusively known from 
<collectingCountry id="F33CB1F1FFEBFFCEFECAFAA14A885141" box="[296,418,1306,1332]" name="Costa Rica" pageId="8" pageNumber="409">Costa Rica</collectingCountry>
and 
<taxonomicName id="4C2B8AE2FFEBFFCEFE3BFAA1495F5141" authorityName="Ricarte &amp; Rotheray" authorityYear="2015" box="[473,629,1306,1332]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="araceorum">
<emphasis id="B95F2D73FFEBFFCEFE3BFAA1495F5141" box="[473,629,1306,1332]" italics="true" pageId="8" pageNumber="409">C. araceorum</emphasis>
</taxonomicName>
was described from material collected in 
<collectingCountry id="F33CB1F1FFEBFFCEFBA9FAA14FEF5141" box="[1099,1221,1306,1332]" name="Costa Rica" pageId="8" pageNumber="409">Costa Rica</collectingCountry>
and 
<collectingCountry id="F33CB1F1FFEBFFCEFB1EFAA14E765141" box="[1276,1372,1306,1332]" name="Ecuador" pageId="8" pageNumber="409">Ecuador</collectingCountry>
. Both species are recorded for the first time in the Amazonian rainforest of 
<collectingCountry id="F33CB1F1FFEBFFCEFC7EFA854F24512D" box="[924,1038,1342,1368]" name="Colombia" pageId="8" pageNumber="409">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
</subSection>
<subSection id="E2A4E88AFFEAFFCCFF75FAAD4EB752D7" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="multiple">
<subSubSection id="C331A2EAFFEAFFCFFF75FAAD4FF75258" pageId="9" pageNumber="410" type="description">
<paragraph id="8B94F161FFEAFFCFFF75FAAD4F095144" blockId="9.[151,1059,1302,1329]" box="[151,1059,1302,1329]" pageId="9" pageNumber="410">
<heading id="D0DC460DFFEAFFCFFF75FAAD4F095144" bold="true" box="[151,1059,1302,1329]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
<emphasis id="B95F2D73FFEAFFCFFF75FAAD4F095144" bold="true" box="[151,1059,1302,1329]" pageId="9" pageNumber="410">
Adjustments to existing identification keys of the 
<taxonomicName id="4C2B8AE2FFEAFFCFFCE2FAAC48595144" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[768,883,1303,1329]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEAFFCFFCE2FAAC48595144" bold="true" box="[768,883,1303,1329]" italics="true" pageId="9" pageNumber="410">C. vagum</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
<paragraph id="8B94F161FFEAFFCFFF75FAE44FF75258" blockId="9.[151,1436,1375,1582]" pageId="9" pageNumber="410">
The last key of 
<bibRefCitation id="EFBA8C90FFEAFFCFFEADFAE44AC0510C" author="Curran, C. H." box="[335,490,1375,1401]" pageId="9" pageNumber="410" pagination="1 - 23" refId="ref11428" refString="Curran, C. H. (1930) New species of Volucellinae from America (Syrphidae, Diptera). American Museum Novitates, 413, 1 - 23." type="journal article" year="1930">Curran (1930</bibRefCitation>
, 
<bibRefCitation id="EFBA8C90FFEAFFCFFE1AFAE4491B510C" author="Curran, C. H." box="[504,561,1375,1401]" pageId="9" pageNumber="410" pagination="1 - 7" refId="ref11485" refString="Curran, C. H. (1939) Synopsis of the American species of Volucella (Syrphidae; Diptera). American Museum Novitates, 127, 1 - 7." type="journal article" year="1939">1939</bibRefCitation>
) covered most of the species included in the 
<taxonomicName id="4C2B8AE2FFEAFFCFFBDDFAE44F87510D" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[1087,1197,1375,1401]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEAFFCFFBDDFAE44F87510D" box="[1087,1197,1375,1401]" italics="true" pageId="9" pageNumber="410">C. vagum</emphasis>
</taxonomicName>
group. 
<bibRefCitation id="EFBA8C90FFEAFFCFFAEAFAE44BC851E8" author="Ricarte, A. &amp; Marcos-Garcia, M. A. &amp; Hancock, E. G. &amp; Rotheray, G. E." pageId="9" pageNumber="410" pagination="0142441" refId="ref12857" refString="Ricarte, A., Marcos-Garcia, M. A., Hancock, E. G. &amp; Rotheray, G. E. (2015) Neotropical Copestylum Macquart (Diptera: Syrphidae) breeding in fruits and flowers, including 7 new species. PLOS ONE, 10 (11), e 0142441. https: // doi. org / 10.1371 / journal. pone. 0142441" type="journal article" year="2015">
Ricarte 
<emphasis id="B95F2D73FFEAFFCFFA80FAE44EBF510C" box="[1378,1429,1375,1401]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
. (2015)
</bibRefCitation>
partially reviewed the 
<taxonomicName id="4C2B8AE2FFEAFFCFFE05FA38497E51E9" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[487,596,1411,1437]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEAFFCFFE05FA38497E51E9" box="[487,596,1411,1437]" italics="true" pageId="9" pageNumber="410">C. vagum</emphasis>
</taxonomicName>
group, providing a key for seven species, of which five were described as new. We provide modifications to the previous keys of the 
<taxonomicName id="4C2B8AE2FFEAFFCFFCCAFA1C48BE51B5" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[808,916,1447,1473]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEAFFCFFCCAFA1C48BE51B5" box="[808,916,1447,1473]" italics="true" pageId="9" pageNumber="410">C. vagum</emphasis>
</taxonomicName>
group to distinguish 
<taxonomicName id="4C2B8AE2FFEAFFCFFB63FA1C4FD451B5" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" box="[1153,1278,1447,1473]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFEAFFCFFB63FA1C4FD451B5" box="[1153,1278,1447,1473]" italics="true" pageId="9" pageNumber="410">C. enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFEAFFCFFAE4FA1C4E7551B4" bold="true" box="[1286,1375,1447,1473]" pageId="9" pageNumber="410">
<taxonomicNameLabel id="A26C9008FFEAFFCFFAE4FA1C4E7551B4" box="[1286,1375,1447,1473]" pageId="9" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from the other eighteen species included in the group. The new key is based on characters provided by 
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, 
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), 
<bibRefCitation id="EFBA8C90FFEAFFCFFF04FA544A9B527F" author="Thompson, F. C." box="[230,433,1519,1546]" pageId="9" pageNumber="410" refId="ref13414" refString="Thompson, F. C. (2006) Primer Taller de identificacion Syrphidae del Neotropico. February 21 th to 27 th 2006. Universidad del Valle, Facultad de Ciencias, Cali, Colombia, 860 pp." type="book" year="2006">Thompson (2006)</bibRefCitation>
, and 
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, in addition to some new provided here. Images and information of 
<typeStatus id="54904FC3FFEAFFCFFF54F9A84BCC5258" box="[182,230,1555,1581]" pageId="9" pageNumber="410">type</typeStatus>
specimens of nine species (see material and methods) are available at the AMNH website.
</paragraph>
</subSubSection>
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<keyLead id="30DA5254FFEAFFCFFF75F9E34EB65218" box="[151,1436,1624,1646]" pageId="9" pageNumber="410">1. Scutellum with a preapical depression..................................................................... 2</keyLead>
</paragraph>
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- Scutellum without a preapical depression.................... other 
<taxonomicName id="4C2B8AE2FFEAFFCFFC7CF9CE4F7052FE" box="[926,1114,1653,1675]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B95F2D73FFEAFFCFFC7CF9CE4F2552FF" box="[926,1039,1653,1674]" italics="true" pageId="9" pageNumber="410">Copestylum</emphasis>
species
</taxonomicName>
(couplet 48, p. 
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)
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<keyLead id="30DA5254FFEAFFCFFF75F9294EB652D2" box="[151,1436,1682,1704]" pageId="9" pageNumber="410">2. Calypter dark, brown to black or with brown border.......................................................... 3</keyLead>
</paragraph>
<paragraph id="8B94F161FFEAFFCFFF75F90B4EB652B3" blockId="9.[151,1437,1624,2027]" box="[151,1436,1712,1734]" pageId="9" pageNumber="410">
<keyLead id="30DA5254FFEAFFCFFF75F90B4EB652B3" box="[151,1436,1712,1734]" pageId="9" pageNumber="410">
- Calypter entirely pale, yellow to orange..................... other 
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<emphasis id="B95F2D73FFEAFFCFFC7CF90B4F2552B0" box="[926,1039,1712,1733]" italics="true" pageId="9" pageNumber="410">Copestylum</emphasis>
species
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(couplet 43, p. 
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3. Face with a pale background colour (
<figureCitation id="1310EDE4FFEAFFCFFDBAF97649F95297" box="[600,723,1741,1763]" captionStart="FIGURE 3" captionStartId="5.[151,250,1328,1352]" captionTargetBox="[169,1428,273,1305]" captionTargetId="figure-61@5.[158,1428,273,1305]" captionTargetPageId="5" captionText="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="9" pageNumber="410">Figs 3A–3D</figureCitation>
in 
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Ricarte 
<emphasis id="B95F2D73FFEAFFCFFCA4F97548595297" box="[838,883,1741,1763]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
. 2015
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); Tergum 1 and anterior part of tergum 2 with a conspicuous pale mark compared to the rest of the abdominal tergites (Fig. 11A in 
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Ricarte 
<emphasis id="B95F2D73FFEAFFCFFBA5F9504F5B528A" box="[1095,1137,1770,1792]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
. 2015
</bibRefCitation>
)................................................................................................. 4 (
<taxonomicName id="4C2B8AE2FFEAFFCFFB4DF8B34E215368" ID-CoL="8BKPY" baseAuthorityName="Wiedemann" baseAuthorityYear="1830" box="[1199,1291,1800,1821]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="vagum">
<emphasis id="B95F2D73FFEAFFCFFB4DF8B34E215368" box="[1199,1291,1800,1821]" italics="true" pageId="9" pageNumber="410">C. vagum</emphasis>
</taxonomicName>
species group)
</keyLead>
</paragraph>
<paragraph id="8B94F161FFEAFFCFFF75F89E4EB75300" blockId="9.[151,1437,1624,2027]" pageId="9" pageNumber="410">
<keyLead id="30DA5254FFEAFFCFFF75F89E4EB75300" pageId="9" pageNumber="410">
- Face with a dark background colour (Figs 24A–24B in 
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Ricarte 
<emphasis id="B95F2D73FFEAFFCFFCB6F89D48AA534F" box="[852,896,1829,1851]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
. 2015
</bibRefCitation>
); Tergum 1 and anterior part of tergum 2 with a similar colour pattern to the rest of the abdominal tergites (Fig. 
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Ricarte 
<emphasis id="B95F2D73FFEAFFCFFC24F8F848DA5322" box="[966,1008,1858,1880]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
. 2015
</bibRefCitation>
).............................................................................(
<taxonomicName id="4C2B8AE2FFEAFFCFFCCFF8DB48935300" ID-CoL="8BKGT" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[813,953,1888,1909]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="cinctiventre">
<emphasis id="B95F2D73FFEAFFCFFCCFF8DB48935300" box="[813,953,1888,1909]" italics="true" pageId="9" pageNumber="410">C. cinctiventre</emphasis>
</taxonomicName>
species group) (to couplet 
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Ricarte 
<emphasis id="B95F2D73FFEAFFCFFAD6F8DB4E755300" box="[1332,1375,1888,1909]" italics="true" pageId="9" pageNumber="410">et al</emphasis>
2015
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)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFEAFFCFFF75F8C64EB653B8" pageId="9" pageNumber="410">
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<keyLead id="30DA5254FFEAFFCFFF75F8C64EB653C5" pageId="9" pageNumber="410">
4. Bristles of the thorax entirely pale; at least metatibiae black to dark-brown, sometimes yellow on basal 1/4 such as in 
<taxonomicName id="4C2B8AE2FFEAFFCFFA64F8C64A1853C5" ID-CoL="9HZSD" authority="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana, 2022" authorityName="Montoya &amp; Parada-Marín &amp; Ramos-Pastrana" authorityYear="2022" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="enriquei" status="sp. nov.">
<emphasis id="B95F2D73FFEAFFCFFA64F8C64A1853C5" italics="true" pageId="9" pageNumber="410">C. enriquei</emphasis>
</taxonomicName>
<emphasis id="B95F2D73FFEAFFCFFEDAF8214AAE53DA" bold="true" box="[312,388,1946,1967]" pageId="9" pageNumber="410">
<taxonomicNameLabel id="A26C9008FFEAFFCFFEDAF8214AAE53DA" box="[312,388,1946,1967]" pageId="9" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
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, 
<figureCitation id="1310EDE4FFEAFFCFFDD9F821495C53C5" box="[571,630,1946,1968]" captionStart="FIGURE 3" captionStartId="5.[151,250,1328,1352]" captionTargetBox="[169,1428,273,1305]" captionTargetId="figure-61@5.[158,1428,273,1305]" captionTargetPageId="5" captionText="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="9" pageNumber="410">3C–D</figureCitation>
)................................................................. 5
</keyLead>
</paragraph>
<paragraph id="8B94F161FFEAFFCFFF75F8034EB653B8" blockId="9.[151,1437,1624,2027]" box="[151,1436,1976,1998]" pageId="9" pageNumber="410">
<keyLead id="30DA5254FFEAFFCFFF75F8034EB653B8" box="[151,1436,1976,1998]" pageId="9" pageNumber="410">- Bristles of the thorax yellow and black; tibiae entirely orange................................................. 21</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8B94F161FFEAFFCFFF75F86E4EB6539E" blockId="9.[151,1437,1624,2027]" box="[151,1436,2005,2027]" pageId="9" pageNumber="410">
<keyLead id="30DA5254FFEAFFCFFF75F86E4EB6539E" box="[151,1436,2005,2027]" pageId="9" pageNumber="410">5. Scutellum with entire wrinkled fascia basally or with subtriangular, roughened flat areas on each side.................. 6</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FF234EB654D8" blockId="10.[151,1437,151,1699]" box="[151,1436,151,173]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FF234EB654D8" box="[151,1436,151,173]" pageId="10" pageNumber="411">- Scutellum without such areas............................................................................ 7</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8B94F161FFE9FFCCFF75FF0E4EB75492" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FF0E4EB75492" pageId="10" pageNumber="411">
6. Scutellum with entire roughened fascia basally; wing extensively microtrichose apically, costal cell microtrichose in apical half, almost all of the cells R4+5 and DM microtrichose............................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFBA0FF694E66549D" ID-CoL="8BKHJ" authority="(Curran, 1939)" baseAuthorityName="Curran" baseAuthorityYear="1939" box="[1090,1356,210,232]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="curiosum">
<emphasis id="B95F2D73FFE9FFCCFBA0FF694F9D549D" box="[1090,1207,210,232]" italics="true" pageId="10" pageNumber="411">C. curiosum</emphasis>
(
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)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFAB5FF6A4EBC5492" box="[1367,1430,209,231]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
]
</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FF4B4EB75557" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FF4B4EB75557" pageId="10" pageNumber="411">
- Scutellum with subtriangular, roughened flat areas on each side; wing extensively bare apically, costal cell completely bare, cells R4+5 and DM bare except apical fourth or less.................................. 
<taxonomicName id="4C2B8AE2FFE9FFCCFB8EFEB64E665556" ID-CoL="8BKGV" authority="(Curran, 1939)" baseAuthorityName="Curran" baseAuthorityYear="1939" box="[1132,1356,269,291]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="circe">
<emphasis id="B95F2D73FFE9FFCCFB8EFEB64F9D5557" box="[1132,1207,269,290]" italics="true" pageId="10" pageNumber="411">C. circe</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFB26FEB64E6F5556" author="Curran, C. H." box="[1220,1349,269,291]" pageId="10" pageNumber="411" pagination="1 - 7" refId="ref11485" refString="Curran, C. H. (1939) Synopsis of the American species of Volucella (Syrphidae; Diptera). American Museum Novitates, 127, 1 - 7." type="journal article" year="1939">Curran, 1939</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFAB5FEB74EBC5557" box="[1367,1430,268,290]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8B94F161FFE9FFCCFF75FE904EB6554A" blockId="10.[151,1437,151,1699]" box="[151,1436,298,320]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FE904EB6554A" box="[151,1436,298,320]" pageId="10" pageNumber="411">7. Terga 3–4 with yellow marks or completely black........................................................... 12</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FEF34EB65528" blockId="10.[151,1437,151,1699]" box="[151,1436,327,349]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FEF34EB65528" box="[151,1436,327,349]" pageId="10" pageNumber="411">- Terga 3–4 green to orange with a narrow, black, apical margin.................................................. 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75FEDE4EB655E2" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75FEDE4EB6550F" blockId="10.[151,1437,151,1699]" box="[151,1436,357,379]" pageId="10" pageNumber="411">
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8. Thoracic setae black (Fig 
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- Thoracic setae orange (Fig 
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); mid femora with at least some pale pile......................... 10
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9. Face with central vitta strongly pigmented, shining black (
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); pleuron with a U-shaped yellow macula (Fig 
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); femora jet black..................................................................................... 
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- Face with central vitta faintly pigmented black (
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); pleuron with a W-shaped black macula (Fig 
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); femora dark yellow... 
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10. Eyes more than half as long as high (
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); face with central vitta present, even if faint (
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)................................. 
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); face without central vitta (
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. 2015
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) .................................................................................................. 11
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11. Tibiae jet black; male cerci triangular-shaped (Fig 19A in 
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Ricarte 
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. 2015
</bibRefCitation>
)....................................................................................... 
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<emphasis id="B95F2D73FFE9FFCCFCD5FD7E488C56AF" box="[823,934,709,730]" italics="true" pageId="10" pageNumber="411">C. tigrinum</emphasis>
Ricarte &amp; Hancock
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<emphasis id="B95F2D73FFE9FFCCFB88FD7E4F5756AF" box="[1130,1149,709,730]" italics="true" pageId="10" pageNumber="411">in</emphasis>
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Ricarte 
<emphasis id="B95F2D73FFE9FFCCFB2CFD7E4FD556AF" box="[1230,1279,709,730]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
2015
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<keyLead id="30DA5254FFE9FFCCFF75FD584EB65760" pageId="10" pageNumber="411">
- Tibiae dark yellow; male genitalia large, cerci elongate, L-shaped (Fig 17A in 
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Ricarte 
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. 2015
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)................................................................................... 
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<emphasis id="B95F2D73FFE9FFCCFC24FCBB4F655760" box="[966,1103,768,789]" italics="true" pageId="10" pageNumber="411">C. musicanum</emphasis>
(
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)
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, 
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<keyLead id="30DA5254FFE9FFCCFF75FCA64EB65747" box="[151,1436,797,819]" pageId="10" pageNumber="411">12. Mesonotum wholly pale pilose on posterior half............................................................ 13</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FC804EB6573A" blockId="10.[151,1437,151,1699]" box="[151,1436,826,848]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FC804EB6573A" box="[151,1436,826,848]" pageId="10" pageNumber="411">- Mesonotum broadly black pilose in front of scutellum....................................................... 18</keyLead>
</paragraph>
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13. Gena and face separated by a very broad brown vitta (
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, 
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); scutum orange except for the wide medial vitta, which is dark and metallic, ending before the prescutellar region, with the apical margin M-shaped (
<figureCitation id="1310EDE4FFE9FFCCFB44FCCE4E3757FF" box="[1190,1309,885,907]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="10" pageNumber="411">Figs 1B–1C</figureCitation>
, 
<figureCitation id="1310EDE4FFE9FFCCFAC5FCCE4E6D57FF" box="[1319,1351,885,906]" captionStart="FIGURE 3" captionStartId="5.[151,250,1328,1352]" captionTargetBox="[169,1428,273,1305]" captionTargetId="figure-61@5.[158,1428,273,1305]" captionTargetPageId="5" captionText="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="10" pageNumber="411">3D</figureCitation>
; 
<bibRefCitation id="EFBA8C90FFE9FFCCFAB0FCCE4A6C57D2" author="Reemer, M." pageId="10" pageNumber="411" pagination="97 - 142" refId="ref12659" refString="Reemer, M. (2016) Syrphidae (Diptera) of Surinam: Eristalinae and synthesis. Tijdschrift voor Entomologie, 159, 97 - 142. https: // doi. org / 10.1163 / 22119434 - 15902002" type="journal article" year="2016">Reemer 2016: 100</bibRefCitation>
, Fig. 15, dorsal habitus); tibiae dark-brown, except yellow on basal 1/4 (
<figureCitation id="1310EDE4FFE9FFCCFBECFC294F9B57D2" box="[1038,1201,914,936]" captionStart="FIGURE 1" captionStartId="3.[151,250,1354,1378]" captionTargetBox="[189,1396,373,1329]" captionTargetId="figure-40@3.[189,1398,373,1329]" captionTargetPageId="3" captionText="FIGURE 1. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890): A. Head, frontal view; B. Habitus, dorsal view; C. Thorax, dorsal view, detail of maculae and pile patterns on scutum; D. Lateral view, E. Latero-posterior view, detail of male genitalia." figureDoi="http://doi.org/10.5281/zenodo.5848271" httpUri="https://zenodo.org/record/5848271/files/figure.png" pageId="10" pageNumber="411">Figs 1A, 1D–1E</figureCitation>
, 
<figureCitation id="1310EDE4FFE9FFCCFB5EFC294FD357DD" box="[1212,1273,914,936]" captionStart="FIGURE 3" captionStartId="5.[151,250,1328,1352]" captionTargetBox="[169,1428,273,1305]" captionTargetId="figure-61@5.[158,1428,273,1305]" captionTargetPageId="5" captionText="FIGURE 3. Copestylum enriquei sp. nov., female, paratype (RMNH-collection). A. Head, frontal view; B. Head, lateral view, C. Habitus, lateral view; D. Thorax and abdomen, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5848275" httpUri="https://zenodo.org/record/5848275/files/figure.png" pageId="10" pageNumber="411">3C–D</figureCitation>
); epandrium and cercus black (
<figureCitation id="1310EDE4FFE9FFCCFE84FC144AF657B3" box="[358,476,943,966]" captionStart="FIGURE 2" captionStartId="4.[151,250,1853,1877]" captionTargetBox="[175,1411,993,1828]" captionTargetId="figure-379@4.[175,1411,993,1828]" captionTargetPageId="4" captionText="FIGURE 2. Copestylum enriquei sp. nov., male, holotype (LEUA–00000035890). A. Abdomen, lateral view; B. Abdomen, ventral view; C. Whole genitalia including epandrium, cercus, and surstylus, lateral view, detail of the dorsal extension of the epandrium (white arrow); D. Epandrium, dorsal view, detail of the dorsal extension (white arrow); E. Hypandrium, ventral view. Abbreviations used in male genitalia structures are as follows: Ah = Aedeagal hood; Cer = Cercus; Epd = Epandrium; Hyp = Hypandrium; Sa= Surstylar apodeme; Sl= Superior lobe; Sur = Surstyle." figureDoi="http://doi.org/10.5281/zenodo.5848273" httpUri="https://zenodo.org/record/5848273/files/figure.png" pageId="10" pageNumber="411">Figs 2A–2E</figureCitation>
)............................................. 
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<emphasis id="B95F2D73FFE9FFCCFBE3FC0B4F4757B0" box="[1025,1133,944,965]" italics="true" pageId="10" pageNumber="411">C. enriquei</emphasis>
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<keyLead id="30DA5254FFE9FFCCFF75FC764EB6578A" pageId="10" pageNumber="411">- Gena and face separated by a narrower brownish vitta; scutum orange, medial vitta with a different pattern; tibiae extensively black; epandrium and cercus orange..................................................................... 14</keyLead>
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<keyLead id="30DA5254FFE9FFCCFF75FBB34EB65068" box="[151,1436,1031,1053]" pageId="10" pageNumber="411">14. Scutellum black pilose on disc.......................................................................... 15</keyLead>
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<keyLead id="30DA5254FFE9FFCCFF75FB9E4EB6504E" box="[151,1436,1061,1083]" pageId="10" pageNumber="411">- Scutellum pale pilose on disc........................................................................... 16</keyLead>
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15. Pleuron black-marked; dorsal margin of eyes sloping (Fig. 
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Ricarte 
<emphasis id="B95F2D73FFE9FFCCFC6DFBF848935022" box="[911,953,1090,1112]" italics="true" pageId="10" pageNumber="411">et al</emphasis>
. 2015
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- Pleuron without black marks, uniformly yellow and orange; dorsal margin of eyes flattened (Fig. 
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Ricarte 
<emphasis id="B95F2D73FFE9FFCCFB1DFBC64E0350E7" box="[1279,1321,1149,1170]" italics="true" pageId="10" pageNumber="411">et al</emphasis>
. 2015
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)......................................... 
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<emphasis id="B95F2D73FFE9FFCCFD4CFB21480B50C5" box="[686,801,1178,1200]" italics="true" pageId="10" pageNumber="411">C. tenorium</emphasis>
Ricarte &amp; Rotheray
</taxonomicName>
<emphasis id="B95F2D73FFE9FFCCFC0BFB2048D650C5" box="[1001,1020,1179,1200]" italics="true" pageId="10" pageNumber="411">in</emphasis>
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Ricarte 
<emphasis id="B95F2D73FFE9FFCCFBAFFB204F5450DA" box="[1101,1150,1178,1200]" italics="true" pageId="10" pageNumber="411">et al.</emphasis>
2015
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, 
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<keyLead id="30DA5254FFE9FFCCFF75FB0C4EB650B8" box="[151,1436,1207,1229]" pageId="10" pageNumber="411">16. Gena and face separated by a very broad brownish vitta...................................................... 17</keyLead>
</paragraph>
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<keyLead id="30DA5254FFE9FFCCFF75FB6E4EB6509F" box="[151,1436,1236,1259]" pageId="10" pageNumber="411">
- No brown vitta separating face and gena............. 
<taxonomicName id="4C2B8AE2FFE9FFCCFCEAFB6E4F08509E" ID-CoL="8BKG9" authority="(Curran, 1926)" baseAuthorityName="Curran" baseAuthorityYear="1926" box="[776,1058,1237,1259]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="brevifacies">
<emphasis id="B95F2D73FFE9FFCCFCEAFB6E48A6509F" box="[776,908,1237,1258]" italics="true" pageId="10" pageNumber="411">C. brevifacies</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFC7BFB6E4F30509E" author="Curran, C. H." box="[921,1050,1237,1259]" pageId="10" pageNumber="411" pagination="50 - 66" refId="ref11380" refString="Curran, C. H. (1926) Partial synopsis of American species of Volucella with notes on Wiedemann's types. Annals of the Entomological Society of America, 18, 50 - 66. https: // doi. org / 10.1093 / aesa / 19.1.50" type="journal article" year="1926">Curran, 1926</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFBCFFB6E4FB8509E" box="[1069,1170,1237,1259]" name="Argentina" pageId="10" pageNumber="411">Argentina</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFB79FB6E4FF2509E" box="[1179,1240,1237,1259]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFB00FB6E4E05509F" box="[1250,1327,1237,1258]" name="Guyana" pageId="10" pageNumber="411">Guyana</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFADAFB6F4EBD509F" box="[1336,1431,1236,1258]" name="Suriname" pageId="10" pageNumber="411">Suriname</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75FB494EB6512A" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75FB494EB65150" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FB494EB65150" pageId="10" pageNumber="411">
17. Mesonotum with the median third black; black vitta separating face and gena very wide and of uniform width.................................................................. 
<taxonomicName id="4C2B8AE2FFE9FFCCFC88FAAB4FB85153" ID-CoL="8FLQ5" authority="(Curran, 1930)" baseAuthorityName="Curran" baseAuthorityYear="1930" box="[874,1170,1295,1318]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="chapadensis">
<emphasis id="B95F2D73FFE9FFCCFC88FAAB48D75151" box="[874,1021,1295,1317]" italics="true" pageId="10" pageNumber="411">C. chapadensis</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFBE8FAAB4FA15153" author="Curran, C. H." box="[1034,1163,1296,1318]" pageId="10" pageNumber="411" pagination="1 - 23" refId="ref11428" refString="Curran, C. H. (1930) New species of Volucellinae from America (Syrphidae, Diptera). American Museum Novitates, 413, 1 - 23." type="journal article" year="1930">Curran, 1930</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFB7FFAB44FF65150" box="[1181,1244,1295,1317]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
, Brazilian Amazon]
</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FA964EB6512A" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FA964EB6512A" pageId="10" pageNumber="411">
- Mesonotum usually all ferruginous on the disc (if with black vitta it is partly linear); vitta separating face and gena usually narrow, sometimes broad above.................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFC2EFAF14FE3512A" ID-CoL="8BKCY" authority="Curran, 1930" authorityName="Curran" authorityYear="1930" box="[972,1225,1354,1375]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="B95F2D73FFE9FFCCFC2EFAF14F6E512A" box="[972,1092,1354,1375]" italics="true" pageId="10" pageNumber="411">C. bequaerti</emphasis>
<bibRefCitation id="EFBA8C90FFE9FFCCFBA8FAF14FE3512A" author="Curran, C. H." box="[1098,1225,1354,1375]" pageId="10" pageNumber="411" pagination="1 - 23" refId="ref11428" refString="Curran, C. H. (1930) New species of Volucellinae from America (Syrphidae, Diptera). American Museum Novitates, 413, 1 - 23." type="journal article" year="1930">Curran, 1930</bibRefCitation>
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFB36FAF14E6B5115" box="[1236,1345,1354,1376]" name="Guatemala" pageId="10" pageNumber="411">Guatemala</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFAA9FAF24EBD512A" box="[1355,1431,1353,1375]" name="Mexico" pageId="10" pageNumber="411">Mexico</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75FAD34EB651EF" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75FAD34EB65108" blockId="10.[151,1437,151,1699]" box="[151,1436,1383,1405]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FAD34EB65108" box="[151,1436,1383,1405]" pageId="10" pageNumber="411">18. Metatibiae black..................................................................................... 19</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FA3E4EB651EF" blockId="10.[151,1437,151,1699]" box="[151,1436,1412,1435]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FA3E4EB651EF" box="[151,1436,1412,1435]" pageId="10" pageNumber="411">
- Metatibiae orange................................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFC19FA3E4FCB51EE" ID-CoL="8BKJB" authority="(Curran, 1936)" baseAuthorityName="Curran" baseAuthorityYear="1936" box="[1019,1249,1413,1435]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="flukei">
<emphasis id="B95F2D73FFE9FFCCFC19FA3E4F6651EF" box="[1019,1100,1413,1434]" italics="true" pageId="10" pageNumber="411">C. flukei</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFBBBFA3E4FF051EE" author="Curran, C. H." box="[1113,1242,1413,1435]" pageId="10" pageNumber="411" pagination="1 - 17" refId="ref11459" refString="Curran, C. H. (1936) New Neotropical Syrphidae (Diptera). American Museum Novitates, 882, 1 - 17." type="journal article" year="1936">Curran, 1936</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFB0FFA3E4E0651EE" box="[1261,1324,1413,1435]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFAD4FA3F4EBD51EF" box="[1334,1431,1412,1434]" name="Colombia" pageId="10" pageNumber="411">Colombia</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75FA194EB65187" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75FA194EB651C2" blockId="10.[151,1437,151,1699]" box="[151,1436,1442,1464]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FA194EB651C2" box="[151,1436,1442,1464]" pageId="10" pageNumber="411">19. Mesonotum with a large, median black spot before the scutellum, ventral scutellar fringe black....................... 20</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75FA7B4EB65187" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FA7B4EB65187" pageId="10" pageNumber="411">
- Mesonotum without median black macula anterior to scutellum; ventral scutellar fringe yellow.......................................................................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFBE2FA664FCB5186" ID-CoL="8BKKK" authority="(Curran, 1936)" baseAuthorityName="Curran" baseAuthorityYear="1936" box="[1024,1249,1501,1523]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="lanei">
<emphasis id="B95F2D73FFE9FFCCFBE2FA664F665187" box="[1024,1100,1501,1522]" italics="true" pageId="10" pageNumber="411">C. lanei</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFBBBFA664FF05186" author="Curran, C. H." box="[1113,1242,1501,1523]" pageId="10" pageNumber="411" pagination="1 - 17" refId="ref11459" refString="Curran, C. H. (1936) New Neotropical Syrphidae (Diptera). American Museum Novitates, 882, 1 - 17." type="journal article" year="1936">Curran, 1936</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFB0FFA664E065186" box="[1261,1324,1501,1523]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFAD4FA674EBD5187" box="[1334,1431,1500,1522]" name="Colombia" pageId="10" pageNumber="411">Colombia</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75FA414EB75258" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75FA414EB6527A" blockId="10.[151,1437,151,1699]" box="[151,1436,1529,1552]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75FA414EB6527A" box="[151,1436,1529,1552]" pageId="10" pageNumber="411">
20. Coxae yellow pilose............. 
<taxonomicName id="4C2B8AE2FFE9FFCCFDA4FA41487E5265" ID-CoL="8BKLC" authority="(Curran, 1936)" baseAuthorityName="Curran" baseAuthorityYear="1936" box="[582,852,1530,1552]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="mocanum">
<emphasis id="B95F2D73FFE9FFCCFDA4FA4149955265" box="[582,703,1530,1552]" italics="true" pageId="10" pageNumber="411">C. mocanum</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFD29FA4148675265" author="Curran, C. H." box="[715,845,1530,1552]" pageId="10" pageNumber="411" pagination="1 - 17" refId="ref11459" refString="Curran, C. H. (1936) New Neotropical Syrphidae (Diptera). American Museum Novitates, 882, 1 - 17." type="journal article" year="1936">Curran, 1936</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFCBDFA4148E95265" box="[863,963,1530,1552]" name="Colombia" pageId="10" pageNumber="411">Colombia</collectingCountry>
, Brazilian Amazon, 
<collectingCountry id="F33CB1F1FFE9FFCCFB64FA414FDA5265" box="[1158,1264,1530,1552]" name="Guatemala" pageId="10" pageNumber="411">Guatemala</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFB18FA404E025265" box="[1274,1320,1531,1552]" name="Peru" pageId="10" pageNumber="411">Peru</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFAD0FA424EBD527A" box="[1330,1431,1529,1551]" name="Venezuela" pageId="10" pageNumber="411">Venezuela</collectingCountry>
]
</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75F9A34EB75258" blockId="10.[151,1437,151,1699]" box="[151,1437,1559,1582]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75F9A34EB75258" box="[151,1437,1559,1582]" pageId="10" pageNumber="411">
- Coxae black pilose........................................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFBBDF9A34E66525B" ID-CoL="8BKNS" authority="(Curran, 1939)" baseAuthorityName="Curran" baseAuthorityYear="1939" box="[1119,1356,1559,1582]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="rurale">
<emphasis id="B95F2D73FFE9FFCCFBBDF9A34F9D5259" box="[1119,1207,1559,1581]" italics="true" pageId="10" pageNumber="411">C. rurale</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFB26F9A34E6F525B" author="Curran, C. H." box="[1220,1349,1560,1582]" pageId="10" pageNumber="411" pagination="1 - 7" refId="ref11485" refString="Curran, C. H. (1939) Synopsis of the American species of Volucella (Syrphidae; Diptera). American Museum Novitates, 127, 1 - 7." type="journal article" year="1939">Curran, 1939</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFAB5F9AC4EBC5258" box="[1367,1430,1559,1581]" name="Brazil" pageId="10" pageNumber="411">Brazil</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30DFE9C4FFE9FFCCFF75F98E4EB752D7" pageId="10" pageNumber="411">
<paragraph id="8B94F161FFE9FFCCFF75F98E4EB65212" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75F98E4EB65212" pageId="10" pageNumber="411">
21. Face green or yellow, without median black vitta; terga 3−5 entirely black pilose................................................................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFCC2F9E94F15521D" ID-CoL="8BKQJ" authority="(Williston, 1891)" baseAuthorityName="Williston" baseAuthorityYear="1891" box="[800,1087,1618,1640]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B95F2D73FFE9FFCCFCC2F9E948BE521D" box="[800,916,1618,1640]" italics="true" pageId="10" pageNumber="411">C. virescens</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFC43F9E94F12521D" author="Williston, S. W." box="[929,1080,1618,1640]" pageId="10" pageNumber="411" pagination="57 - 72" refId="ref13517" refString="Williston, S. W. (1891 - 1892) Fam. Syrphidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali - Americana, Zoologia- Insecta-Deptera, 3, pp. 57 - 72 (1891.12), pp. 73 - 78 (1892.05)." type="journal article" year="1891">Williston, 1891</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFBA8F9E94F84521D" box="[1098,1198,1618,1640]" name="Colombia" pageId="10" pageNumber="411">Colombia</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFB5AF9E94E08521D" box="[1208,1314,1618,1640]" name="Costa Rica" pageId="10" pageNumber="411">Costa Rica</collectingCountry>
, 
<collectingCountry id="F33CB1F1FFE9FFCCFACEF9EA4EBD5212" box="[1324,1431,1617,1639]" name="Guatemala" pageId="10" pageNumber="411">Guatemala</collectingCountry>
]
</keyLead>
</paragraph>
<paragraph id="8B94F161FFE9FFCCFF75F9CB4EB752D7" blockId="10.[151,1437,151,1699]" pageId="10" pageNumber="411">
<keyLead id="30DA5254FFE9FFCCFF75F9CB4EB752D7" pageId="10" pageNumber="411">
- Face with broad median blackish vitta; terga 3−5 extensively white pilose on basal 1/2...................................................................................................... 
<taxonomicName id="4C2B8AE2FFE9FFCCFBA1F9364E7052D6" ID-CoL="8D9G6" authority="(Curran, 1925)" baseAuthorityName="Curran" baseAuthorityYear="1925" box="[1091,1370,1677,1699]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="varichaeta">
<emphasis id="B95F2D73FFE9FFCCFBA1F9364FEC52D7" box="[1091,1222,1677,1698]" italics="true" pageId="10" pageNumber="411">C. varichaeta</emphasis>
(
<bibRefCitation id="EFBA8C90FFE9FFCCFB30F9364E7952D6" author="Curran, C. H." box="[1234,1363,1677,1699]" pageId="10" pageNumber="411" pagination="243 - 253" refId="ref11352" refString="Curran, C. H. (1925) New American Diptera. I. Annals and Magazine of Natural History, 9, 243 - 253." type="journal article" year="1925">Curran, 1925</bibRefCitation>
)
</taxonomicName>
[
<collectingCountry id="F33CB1F1FFE9FFCCFA84F9374EBD52D7" box="[1382,1431,1676,1698]" name="Peru" pageId="10" pageNumber="411">Peru</collectingCountry>
]
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</subSection>
</treatment>
</document>