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<document id="9C68D4980667F3320ECC7A19F490D1FA" ID-CLB-Dataset="54646" ID-DOI="10.11646/zootaxa.5278.1.1" ID-GBIF-Dataset="1528757a-5499-40b9-a728-c5bf0c770b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce &amp; Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFACFFEAFF6D69DACDE2FB2A" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma Archibald &amp; Rasnitsyn 2023, n. gen." docType="treatment" docUuid="531F3657-F3D9-4A4D-86B7-1C146E74D627" docUuidSource="ZooBank" docVersion="9" lastPageNumber="8" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="8" updateTime="1728574925831" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="AC5E976484EEF653141DB430269DA310">Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:namePart id="94EA5F4FAA314A2723D105C962DCB17F">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="8B5C3293D2E6E688AE76FACD3A485ABC">Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 &amp; Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America &amp; Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:namePart id="94343BAF88A8913CB7A8AE113D946812">Rasnitsyn, Alexandr P.</mods:namePart>
<mods:affiliation id="AD954ECC65B800F2939CE9DD3D9B173F">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia &amp; Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<emphasis id="1B95B80DFFACFFEAFF6D69DAC953FD60" bold="true" box="[151,299,655,681]" italics="true" pageId="7" pageNumber="8">Leptostigma</emphasis>
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<figureCitation id="B1DA789AFFACFFEAFF6D6982C917FD38" box="[151,367,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="7" pageNumber="8">Figs. 3A–D, 3G–K</figureCitation>
, 
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.
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<emphasis id="1B95B80DFFACFFEAFF6D684EC95BFCFA" bold="true" box="[151,291,795,819]" pageId="7" pageNumber="8">
<typeStatus id="F65ADABDFFACFFEAFF6D684EC8B4FCFA" box="[151,204,795,819]" pageId="7" pageNumber="8">Type</typeStatus>
species.
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<taxonomicName id="EEE11F9CFFACFFEAFED06849CA62FCFA" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[298,538,795,819]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFED06849CA62FCFA" box="[298,538,795,819]" italics="true" pageId="7" pageNumber="8">Leptostigma brevilatum</emphasis>
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<emphasis id="1B95B80DFFACFFEAFF6D6834C96AFCB2" bold="true" box="[151,274,865,891]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
Distinguished from 
<taxonomicName id="EEE11F9CFFACFFEAFE016834CAFDFCB2" authorityName="Nel" authorityYear="2004" box="[507,645,865,891]" class="Insecta" family="Cimbicidae" genus="Cenocimbex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1B95B80DFFACFFEAFE016834CAFDFCB2" box="[507,645,865,891]" italics="true" pageId="7" pageNumber="8">Cenocimbex</emphasis>
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by lacking dark triangular sclerite on R basal to pterostigma reaching 1-Rs+M near 1m-cu (absent or undetectable being small, pale); wing narrower (length/width 2.8 or more); cell 3rm long (2r-m to 3r-m distinctly longer than length of 3r-m); hind wing with cu-a well before middle of cell mcu; distinguished from 
<taxonomicName id="EEE11F9CFFACFFEAFE8A6898CA6DFC2E" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[368,533,973,999]" class="Insecta" family="Cimbicidae" genus="Allenbycimbex" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1B95B80DFFACFFEAFE8A6898CA6DFC2E" box="[368,533,973,999]" italics="true" pageId="7" pageNumber="8">Allenbycimbex</emphasis>
</taxonomicName>
by pterostigma narrow, maximum height about equal to height to C to R basal to pterostigma.
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<paragraph id="295E641FFFACFFEAFF3D6F40CB14FB9A" blockId="7.[151,1436,865,1252]" pageId="7" pageNumber="8">
<emphasis id="1B95B80DFFACFFEAFF3D6F40C9F4FBE6" bold="true" box="[199,396,1045,1071]" pageId="7" pageNumber="8">Species included</emphasis>
. The 
<typeStatus id="F65ADABDFFACFFEAFE316F43C983FBF9" box="[459,507,1046,1072]" pageId="7" pageNumber="8">type</typeStatus>
species and 
<taxonomicName id="EEE11F9CFFACFFEAFD726F43CB63FBE6" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[648,795,1045,1071]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFD726F43CB63FBE6" box="[648,795,1045,1071]" italics="true" pageId="7" pageNumber="8">L. longiclava</emphasis>
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<emphasis id="1B95B80DFFACFFEAFCD86F40CB18FBE6" bold="true" box="[802,864,1045,1071]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFCD86F40CB18FBE6" box="[802,864,1045,1071]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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, 
<taxonomicName id="EEE11F9CFFACFFEAFC946F43CC50FBE6" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[878,1064,1045,1071]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFC946F43CC50FBE6" box="[878,1064,1045,1071]" italics="true" pageId="7" pageNumber="8">L. longipallidum</emphasis>
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<emphasis id="1B95B80DFFACFFEAFBD56F40CC15FBE6" bold="true" box="[1071,1133,1045,1071]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFBD56F40CC15FBE6" box="[1071,1133,1045,1071]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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, 
<taxonomicName id="EEE11F9CFFACFFEAFB816F43CD28FBE6" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[1147,1360,1045,1071]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFB816F43CD28FBE6" box="[1147,1360,1045,1071]" italics="true" pageId="7" pageNumber="8">L. longitenebricum</emphasis>
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<emphasis id="1B95B80DFFACFFEAFAAD6F40CDEDFBE6" bold="true" box="[1367,1429,1045,1071]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFAAD6F40CDEDFBE6" box="[1367,1429,1045,1071]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="EEE11F9CFFACFFEAFF6D6F6FC95CFB9A" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[151,292,1082,1107]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="proxivena" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFF6D6F6FC95CFB9A" box="[151,292,1082,1107]" italics="true" pageId="7" pageNumber="8">L. proxivena</emphasis>
</taxonomicName>
<emphasis id="1B95B80DFFACFFEAFED16F6CC911FB9A" bold="true" box="[299,361,1081,1107]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFED16F6CC911FB9A" box="[299,361,1081,1107]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="EEE11F9CFFACFFEAFE826F6FCA7AFB9A" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[376,514,1081,1107]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fasciatum" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFE826F6FCA7AFB9A" box="[376,514,1081,1107]" italics="true" pageId="7" pageNumber="8">L. fasciatum</emphasis>
</taxonomicName>
<emphasis id="1B95B80DFFACFFEAFDF36F6CCA30FB9A" bold="true" box="[521,584,1081,1107]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFDF36F6CCA30FB9A" box="[521,584,1081,1107]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="EEE11F9CFFACFFEAFD7C6F6FCB5EFB9A" authority="Archibald &amp; Rasnitsyn, 2023" authorityName="Archibald &amp; Rasnitsyn" authorityYear="2023" box="[646,806,1081,1107]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alaemacula" status="sp. nov.">
<emphasis id="1B95B80DFFACFFEAFD7C6F6FCB5EFB9A" box="[646,806,1081,1107]" italics="true" pageId="7" pageNumber="8">L. alaemacula</emphasis>
</taxonomicName>
<emphasis id="1B95B80DFFACFFEAFCD76F6CCB14FB9A" bold="true" box="[813,876,1081,1107]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="00A60576FFACFFEAFCD76F6CCB14FB9A" box="[813,876,1081,1107]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="61FB3794FFACFFEAFF3D6F08CA17FB52" pageId="7" pageNumber="8" type="etymology">
<paragraph id="295E641FFFACFFEAFF3D6F08CA17FB52" blockId="7.[151,1436,865,1252]" pageId="7" pageNumber="8">
<emphasis id="1B95B80DFFACFFEAFF3D6F08C936FBBE" bold="true" box="[199,334,1117,1143]" pageId="7" pageNumber="8">Etymology.</emphasis>
From the Greek 
<emphasis id="1B95B80DFFACFFEAFDED6F08CA22FBBE" box="[535,602,1117,1143]" italics="true" pageId="7" pageNumber="8">leptos</emphasis>
, meaning thin, and 
<emphasis id="1B95B80DFFACFFEAFCB96F0BCBF6FBBE" box="[835,910,1118,1143]" italics="true" pageId="7" pageNumber="8">stigma</emphasis>
, for the pterostigma. The name is neuter (see Examples below Art. 30.1.2, 
<bibRefCitation id="4D7019EEFFACFFEAFE246FD4CA1AFB52" author="ICZN" box="[478,610,1153,1179]" pageId="7" pageNumber="8" refId="ref17880" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp. [https: // www. iczn. org / the-code / the-code-online /]" type="book" year="1999">ICZN 1999</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61FB3794FFACFFEAFF3D6FF0CDE2FB2A" pageId="7" pageNumber="8" type="distribution">
<paragraph id="295E641FFFACFFEAFF3D6FF0CDE2FB2A" blockId="7.[151,1436,865,1252]" pageId="7" pageNumber="8">
<emphasis id="1B95B80DFFACFFEAFF3D6FF0C9EAFB76" bold="true" box="[199,402,1189,1215]" pageId="7" pageNumber="8">Locality and age.</emphasis>
<materialsCitation id="99896E42FFACFFEAFE636FF0CA3DFB2A" ID-GBIF-Occurrence="4100387318" collectionCode="A" country="United States of America" county="Republic" location="Klondike Mountain Formation" municipality="Tom Thumb Tuff Member" pageId="7" pageNumber="8" specimenCode="A0307" specimenCount="1" stateProvince="British Columbia">
<collectingCounty id="C03F1C93FFACFFEAFE636FF0CA79FB76" box="[409,513,1189,1215]" pageId="7" pageNumber="8">Republic</collectingCounty>
, Washington, 
<collectingCountry id="51F6248FFFACFFEAFD676FF0CBC0FB76" box="[669,952,1189,1215]" name="United States of America" pageId="7" pageNumber="8">United States of America</collectingCountry>
, 
<collectingMunicipality id="C93AFE65FFACFFEAFC386FF3CC90FB76" box="[962,1256,1189,1216]" pageId="7" pageNumber="8">Tom Thumb Tuff Member</collectingMunicipality>
of the 
<location id="2C3E32C4FFACFFEAFACE6FF0C9F8FB2A" LSID="urn:lsid:plazi:treatment:A148D509FFACFFEAFF6D69DACDE2FB2A:2C3E32C4FFACFFEAFACE6FF0C9F8FB2A" country="United States of America" county="Republic" municipality="Tom Thumb Tuff Member" name="Mountain Formation" pageId="7" pageNumber="8" stateProvince="British Columbia">Klondike Mountain Formation</location>
exposure 
<specimenCode id="7947CC64FFACFFEAFE0B6F9CCA39FB2A" box="[497,577,1225,1251]" collectionCode="A" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="7" pageNumber="8" type="Herbarium">A0307</specimenCode>
;
</materialsCitation>
<materialsCitation id="99896E42FFACFFEAFDB66F9CCDEEFB2A" ID-GBIF-Occurrence="4100387308" box="[588,1430,1225,1252]" collectionCode="A" country="United States of America" county="Republic" location="Klondike Mountain Formation" municipality="Tom Thumb Tuff Member" pageId="7" pageNumber="8" specimenCode="A0307" specimenCount="1" stateProvince="British Columbia">
and McAbee, 
<collectingRegion id="EB25AAFDFFACFFEAFD106F9CCBD6FB2A" box="[746,942,1225,1251]" country="Canada" name="British Columbia" pageId="7" pageNumber="8">British Columbia</collectingRegion>
, 
<collectingCountry id="51F6248FFFACFFEAFC406F9CCC69FB2A" box="[954,1041,1225,1251]" name="Canada" pageId="7" pageNumber="8">Canada</collectingCountry>
, Hoodoo Face beds; mid-Ypresian
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>