<documentid="4C2412D7E22F96E5CBE89E1979465510"ID-DOI="10.5852/ejt.2023.865.2097"ID-GBIF-Dataset="3a63a183-743e-4185-8fdc-1080072ada0a"ID-ISSN="2118-9773"ID-Zenodo-Dep="7867448"ID-ZooBank="845BAE1A-0E6E-484B-BE6D-17F866777B51"IM.bibliography_approvedBy="diego"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="diego"checkinTime="1682512407979"checkinUser="felipe"docAuthor="Rivera-Quiroz, F. Andrés & Álvarez-Padilla, Fernando"docDate="2023"docId="03C8E927FF88FFAEFDF7FEF0FC146500"docLanguage="en"docName="ejt-865_1-94.pdf"docOrigin="European Journal of Taxonomy 865"docSource="http://dx.doi.org/10.5852/ejt.2023.865.2097"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Anyphaena noctua Rivera-Quiroz & Álvarez-Padilla 2023, sp. nov."docType="treatment"docVersion="10"lastPageNumber="63"masterDocId="FFF1915FFFB4FF90FF81FFF8FF856160"masterDocTitle="Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico"masterLastPageNumber="94"masterPageNumber="1"pageNumber="61"updateTime="1732489834501"updateUser="ExternalLinkService"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="A704B269E3BD2AC85D71BC6D2CD3238B">Integration or minimalism: twenty-one new species of ghost spiders (Anyphaenidae: Anyphaena) from Mexico</mods:title>
<mods:affiliationid="54C3827AD581494B11246271F978C1ED">Understanding Evolution Research Group, Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.</mods:affiliation>
<mods:affiliationid="500A43F4766F50AB68A04A3F1C83FA3C">Laboratorio de Aracnología, Departamento de Biología Comparada, Facultad de Ciencias, Universidad Nacional Autónoma de México, Distrito Federal, México Ciudad Universitaria, México D. F. Del. Coyoacán, C. P. 04510, Mexico.</mods:affiliation>
<taxonomicNameid="4C6123B2FF88FFACFDF7FEF0FCDE6043"ID-CoL="C66LL"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[630,859,264,291]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="60"pageNumber="61"phylum="Arthropoda"rank="species"species="noctua"status="sp. nov.">
<figureCitationid="135A44B4FF88FFACFD27FEB5FC856008"box="[678,768,333,360]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="60"pageNumber="61">Figs 37</figureCitation>
,
<figureCitationid="135A44B4FF88FFACFC8DFEB6FCE16008"box="[780,868,334,360]"captionStart="Fig"captionStartId="61.[189,232,1927,1953]"captionTargetBox="[217,1392,731,1883]"captionTargetId="figure-161@61.[189,1401,716,1898]"captionTargetPageId="61"captionText="Fig. 38. Anyphaena spp. A–B. A. sofiae sp. nov., paratype, ♀ (CNAN-T01572). A. Epigynum, ventral view. B. Epigynum, dorsal view. –C–D. A. noctua sp. nov., holotype, ♀ (CNAN-T01533). C. Epigynum, ventral view. D. Epigynum, dorsal view. Scale bars: A– B=0.2 mm: C–D =0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.7867572"httpUri="https://zenodo.org/record/7867572/files/figure.png"pageId="60"pageNumber="61">38C–D</figureCitation>
,
<figureCitationid="135A44B4FF88FFACFCF1FEB5FC086008"box="[880,909,333,360]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="60"pageNumber="61">52</figureCitation>
<taxonomicNameid="4C6123B2FF88FFACFEC9FE44FE3C60B5"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[328,441,444,470]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="60"pageNumber="61"phylum="Arthropoda"rank="species"species="noctua"status="sp. nov.">
groups by the following features: atrium small, lateral borders sinuous, hood triangular, posteriorly curved at the center, genital openings at the posterior margin of the atrium, copulatory ducts short, comma-shaped (
<figureCitationid="135A44B4FF88FFACFEFFFDDAFD8C635D"box="[382,521,546,573]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="60"pageNumber="61">Figs 37E–F</figureCitation>
,
<figureCitationid="135A44B4FF88FFACFD95FDDBFDE8635D"box="[532,621,547,573]"captionStart="Fig"captionStartId="61.[189,232,1927,1953]"captionTargetBox="[217,1392,731,1883]"captionTargetId="figure-161@61.[189,1401,716,1898]"captionTargetPageId="61"captionText="Fig. 38. Anyphaena spp. A–B. A. sofiae sp. nov., paratype, ♀ (CNAN-T01572). A. Epigynum, ventral view. B. Epigynum, dorsal view. –C–D. A. noctua sp. nov., holotype, ♀ (CNAN-T01533). C. Epigynum, ventral view. D. Epigynum, dorsal view. Scale bars: A– B=0.2 mm: C–D =0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.7867572"httpUri="https://zenodo.org/record/7867572/files/figure.png"pageId="60"pageNumber="61">38C–D</figureCitation>
<taxonomicNameid="4C6123B2FF88FFACFD57FD69FCA363CA"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[726,806,657,682]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="60"pageNumber="61"phylum="Arthropoda"rank="species"species="noctua">
<taxonomicNameid="4C6123B2FF88FFACFE9CF87FFE7266C1"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[285,503,1927,1953]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="60"pageNumber="61"phylum="Arthropoda"rank="species"species="noctua"status="sp. nov.">
<collectingRegionid="49A596D3FF89FFADFEF0FEA2FDA86014"box="[369,557,346,372]"country="Mexico"name="San Luis Potosi"pageId="61"pageNumber="62">San Luis Potosi</collectingRegion>
Total length 6.1. Carapace light yellow, with two faint darker bands delineating cephalic area and around fovea, ocular quadrangle and clypeus dark brown (
<figureCitationid="135A44B4FF89FFADFCAAFDCCFC41632E"box="[811,964,564,590]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="61"pageNumber="62">Fig. 37A, D</figureCitation>
). Sternum surface white, intercoxal triangles present on all legs. Labium yellow, white at tip, longer than wide. Endites yellow, rectangular, broader at tip (
<figureCitationid="135A44B4FF89FFADFEECFD82FE5F63F4"box="[365,474,634,660]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="61"pageNumber="62">Fig. 37C</figureCitation>
). Chelicerae brown without dorsal pattern (
<figureCitationid="135A44B4FF89FFADFC5EFD82FBF563F5"box="[991,1136,634,661]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="61"pageNumber="62">Fig. 37B, D</figureCitation>
), promargin with four teeth, retromargin with seven to eight denticles. Femora base of legs white, orange-yellow distally, darker from patella to tarsi. Abdomen dorsal surface white and covered by diffuse darker pattern, pattern of lateral and ventral surfaces even fainter, tracheal spiracle closer to epigastric furrow (
<figureCitationid="135A44B4FF8AFFAEFB56FEB4FAEC6006"box="[1239,1385,332,358]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="62"pageNumber="63">Fig. 37A–C</figureCitation>
). Lateral side of espigynal plate flanked by two faint pits. Seminal receptacles close to genital openings. Fertilization ducts short, cylindrical, and entering posterior surface of spermathecae (
<figureCitationid="135A44B4FF8AFFAEFB60FE68FAF760CA"box="[1249,1394,400,426]"captionStart="Fig"captionStartId="60.[189,232,1927,1953]"captionTargetBox="[189,1396,750,1892]"captionTargetId="figure-167@60.[189,1398,750,1896]"captionTargetPageId="60"captionText="Fig. 37. Anyphaena noctua sp. nov., holotype, ♀ (CNAN-T01533). A. Dorsal habitus.B. Lateral habitus. C. Ventral habitus. D. Prosoma, anterior view. E. Epigynum, ventral view. F. Epigynum, dorsal view. Scale bars: A–C=1.0 mm; D =0.5 mm; E–F =0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.7867570"httpUri="https://zenodo.org/record/7867570/files/figure.png"pageId="62"pageNumber="63">Figs 37E–F</figureCitation>
,
<figureCitationid="135A44B4FF8AFFAEFF3CFE4AFE9360AC"box="[189,278,434,460]"captionStart="Fig"captionStartId="61.[189,232,1927,1953]"captionTargetBox="[217,1392,731,1883]"captionTargetId="figure-161@61.[189,1401,716,1898]"captionTargetPageId="61"captionText="Fig. 38. Anyphaena spp. A–B. A. sofiae sp. nov., paratype, ♀ (CNAN-T01572). A. Epigynum, ventral view. B. Epigynum, dorsal view. –C–D. A. noctua sp. nov., holotype, ♀ (CNAN-T01533). C. Epigynum, ventral view. D. Epigynum, dorsal view. Scale bars: A– B=0.2 mm: C–D =0.1 mm."figureDoi="http://doi.org/10.5281/zenodo.7867572"httpUri="https://zenodo.org/record/7867572/files/figure.png"pageId="62"pageNumber="63">38C–D</figureCitation>
). Cephalothorax length 2.71, thoracic width 2.1, cephalic width 1.1. Clypeus height 0.05. Eye diameters: AME 0.07, ALE 0.11, PME 0.1, PLE 0.11. Eye interdistances: AME–AME 0.04, AME–ALE 0.02, ALE–PLE 0.05, PME–PME 0.17, PME–PLE 0.07. Femur lengths: I 2.86, II 2.62, III 2.19, IV 2.95. Leg spination: femur I d1-1-1, p0-0-2, r0-0-1. Tibia I v2-2-0, p1-1-1, r1-1-1. Metatarsus I v2-2-0, p1-1-1, r1-1-1. Femur II d1-1-1, p0-1-1, r0-0-1. Tibia II v2-2-0, p1-1-1, r1-1-1. Metatarsus II v2-2-0, p1-1-1, r1-1-1. Femur III d1-1-1, p0-0-1, r0-0-1. Tibia III v1-1-0, p1-1-1, r1-1-1. Metatarsus III v2-2-2, p1-1-2, r1-1-2. Femur IV d1-1-1, p0-0-1, r0-0-1. Tibia IV v2-2-2, p1-1-1, r1-1-1. Metatarsus IV v2-2-2, p1-1-2, r1-1-2.
<taxonomicNameid="4C6123B2FF89FFADFDAFF870FD1166C1"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[558,660,1927,1954]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="61"pageNumber="62"phylum="Arthropoda"rank="species"species="sofiae"status="sp. nov.">
<taxonomicNameid="4C6123B2FF89FFADFD12F853FC8766A4"authority="Rivera-Quiroz & Álvarez-Padilla, 2023"authorityName="Rivera-Quiroz & Álvarez-Padilla"authorityYear="2023"box="[659,770,1963,1989]"class="Arachnida"family="Anyphaenidae"genus="Anyphaena"kingdom="Animalia"order="Araneae"pageId="61"pageNumber="62"phylum="Arthropoda"rank="species"species="noctua"status="sp. nov.">
<paragraphid="8BDE5831FF8AFFAEFF3CFC94FD8262E6"blockId="62.[189,519,876,902]"box="[189,519,876,902]"pageId="62"pageNumber="63">Only type specimen known.</paragraph>
<figureCitationid="135A44B4FF8AFFAEFDE5FC21FD3E6294"box="[612,699,985,1012]"captionStart="Fig"captionStartId="82.[189,232,1018,1044]"captionTargetBox="[193,1392,269,964]"captionTargetId="figure-209@82.[189,1398,265,976]"captionTargetPageId="82"captionText="Fig. 52. Localities for the species of Anyphaena. Black square=Atotonilco Plots I and II, separated by 400 m, nine species registered: A. bifurcata sp. nov., A. bromelicola Platnick, 1977 (ON619634- 38 Supp. file 1), A. catalina Platnick, 1974 (ON619673-75), A. dulceae sp. nov., A. epicardia sp. nov., A. fernandae sp. nov., A. porta sp. nov., A. salgueiroi sp. nov. and A. sofiae sp. nov. White square=Xamaticpac Plots I and II, plots contiguous, eleven species registered: A. adnani sp. nov., A. alachua Platnick, 1974 (ON619665), A. fraterna (Banks, 1896) (ON619671), A. megamedia sp. nov., A. miniducta sp. nov., A. natachae sp. nov., A. porta sp. nov., A. quadrata sp. nov., A. rebecae sp. nov., A. stigma sp. nov. and A. tonoi sp. nov. Black circle =Xilitla, four species registered: A. franciscoi sp. nov., A. ibarrai sp. nov., A. noctua sp. nov. and A. triangularis sp. nov. White circle =Los Tuxtlas, two species registered: A. jimenezi sp. nov. and A. urieli sp. nov."figureDoi="http://doi.org/10.5281/zenodo.7867604"httpUri="https://zenodo.org/record/7867604/files/figure.png"pageId="62"pageNumber="63">Fig. 52</figureCitation>
<paragraphid="8BDE5831FF8AFFAEFF3CFBBEFC146500"blockId="62.[189,913,1094,1120]"box="[189,913,1094,1120]"pageId="62"pageNumber="63">Collected by beating direct searching over ground vegetation.</paragraph>