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<document id="E4BF1E9754CEBD57DA688C36E9D71B65" ID-CLB-Dataset="7229" ID-DOI="10.11646/zootaxa.4958.1.18" ID-GBIF-Dataset="5e4f90e6-bed3-4335-85ba-f7dcdbee7f9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4691975" ID-ZooBank="45AA47B6-7C5E-4F04-9B4D-F03B7D235700" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618462074235" checkinUser="plazi" docAuthor="Schuh, Randall T. &amp; Salas, Ruth" docDate="2021" docId="037387810D0AFFA2618C5D47D35EF7CE" docLanguage="en" docName="zootaxa.4958.1.18.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graziaphylus Schuh &amp; Salas 2021, new genus" docType="treatment" docVersion="8" lastPageNumber="321" masterDocId="FF4AFFF90D0BFFA0611B5E05D265F423" masterDocTitle="Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia" masterLastPageNumber="326" masterPageNumber="319" pageNumber="320" updateTime="1698936862551" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2EC391BFAFE462B1A4FEFB29D92A9ACD">Graziaphylus, a new genus and two new species of Eremophila-feeding (Scrophulariceae) Phylinae (Insecta: Heteroptera: Miridae) from Australia</mods:title>
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<mods:name id="1F2C32993B9F954C5C33D6DE61FDAFED" type="personal">
<mods:role id="A30E7A2B30798BA3AE8B6BC27EBC1D65">
<mods:roleTerm id="4D633C7EB01BD9CB0DF0D052443B50EF">Author</mods:roleTerm>
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<mods:namePart id="2FD262F4676A6C5ACB30B2C0D63DCC07">Schuh, Randall T.</mods:namePart>
<mods:nameIdentifier id="14046795151121BEFB3A471DFEAFCF82" type="ORCID">0000-0002-0713-1367</mods:nameIdentifier>
<mods:affiliation id="46F442660F50E6CED9C36C34EF4E2723">Division of Invertebrate Zoology, American Museum of Natural History, New York 10024 USA &amp; schuh @ amnh. org; https: // orcid. org / 0000 - 0002 - 0713 - 1367</mods:affiliation>
<mods:nameIdentifier id="05CB629D8CA7FB2DE7733AF250F730DC" type="email">schuh@amnh.org</mods:nameIdentifier>
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<mods:namePart id="15E6BBA233303B459F0605B0D5A7FDD0">Salas, Ruth</mods:namePart>
<mods:nameIdentifier id="E42EB3E46EA4ED76848B42DEC40FF7D1" type="ORCID">0000-0003-4493-7842</mods:nameIdentifier>
<mods:affiliation id="66A13B4AD83688039B88962955D18BF3">Division of Invertebrate Zoology, American Museum of Natural History, New York 10024 USA &amp; rsalas @ amnh. org; https: // orcid. org / 0000 - 0003 - 4493 - 7842</mods:affiliation>
<mods:nameIdentifier id="1AFB9EFB633EFC0AF95834025BDACC42" type="email">rsalas@amnh.org</mods:nameIdentifier>
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<mods:title id="B89130D7BB905F671D108923ACE944E0">Zootaxa</mods:title>
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<mods:part id="CA3112264039199E534D149480A779CE">
<mods:date id="414F5246024DF4F2D8F9C1E0422B8334">2021</mods:date>
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<mods:number id="3B15682200847A413E263DE817003153">2021-04-14</mods:number>
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<mods:detail id="C140845DA5BF20C7875844F8D16244AC" type="volume">
<mods:number id="D8DA1D6CAB1613090B22D3FB30727027">4958</mods:number>
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<mods:number id="887144D797FB98C683CAB2FE84AEBBBE">1</mods:number>
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<mods:classification id="563353630405A99605AECE5F072048B5">journal article</mods:classification>
<mods:identifier id="D75C5786E3D21C1D80824A4B0F711897" type="CLB-Dataset">7229</mods:identifier>
<mods:identifier id="F0B2E258D496C933152C308D97CE154A" type="DOI">10.11646/zootaxa.4958.1.18</mods:identifier>
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<mods:identifier id="C3160867407D7453342C1137335944F6" type="ISSN">1175-5326</mods:identifier>
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<treatment id="037387810D0AFFA2618C5D47D35EF7CE" ID-DOI="http://doi.org/10.5281/zenodo.4710542" ID-GBIF-Taxon="181530623" ID-Zenodo-Dep="4710542" LSID="urn:lsid:plazi:treatment:037387810D0AFFA2618C5D47D35EF7CE" httpUri="http://treatment.plazi.org/id/037387810D0AFFA2618C5D47D35EF7CE" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320">
<subSubSection id="C3C0651C0D0AFFA1618C5D47D3ACF77E" box="[151,457,834,861]" pageId="1" pageNumber="320" type="nomenclature">
<paragraph id="8B6536970D0AFFA1618C5D47D3ACF77E" blockId="1.[151,457,834,861]" box="[151,457,834,861]" pageId="1" pageNumber="320">
<heading id="D02D81FB0D0AFFA1618C5D47D3ACF77E" bold="true" box="[151,457,834,861]" fontSize="11" level="1" pageId="1" pageNumber="320" reason="1">
<emphasis id="B9AEEA850D0AFFA1618C5D47D3ACF77E" bold="true" box="[151,457,834,861]" pageId="1" pageNumber="320">
<taxonomicName id="4CDA4D140D0AFFA1618C5D47D35DF77E" authority="Schuh &amp; Salas, 2021" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[151,312,834,861]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B9AEEA850D0AFFA1618C5D47D35DF77E" bold="true" box="[151,312,834,861]" italics="true" pageId="1" pageNumber="320">Graziaphylus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A29D57FE0D0AFFA1605C5D46D3ACF77E" box="[327,457,835,861]" pageId="1" pageNumber="320" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3C0651C0D0AFFA1618C5D82D0D2F7BC" box="[151,695,903,927]" pageId="1" pageNumber="320" type="type_taxon">
<paragraph id="8B6536970D0AFFA1618C5D82D0D2F7BC" blockId="1.[151,695,903,927]" box="[151,695,903,927]" pageId="1" pageNumber="320">
<emphasis id="B9AEEA850D0AFFA1618C5D82D343F7BC" bold="true" box="[151,294,903,927]" pageId="1" pageNumber="320">
<typeStatus id="546188350D0AFFA1618C5D82D2A9F7BC" box="[151,204,903,927]" pageId="1" pageNumber="320">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4CDA4D140D0AFFA160375D82D0D2F7BC" authority="Schuh and Salas" authorityName="Schuh and Salas" box="[300,695,903,927]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="species" species="joceliae">
<emphasis id="B9AEEA850D0AFFA160375D82D06EF7BC" box="[300,523,903,927]" italics="true" pageId="1" pageNumber="320">Graziaphylus joceliae</emphasis>
Schuh and Salas
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3C0651C0D0AFFA1618C5DC8D672F7C4" box="[151,1047,973,1000]" pageId="1" pageNumber="320" type="etymology">
<paragraph id="8B6536970D0AFFA1618C5DC8D672F7C4" blockId="1.[151,1437,973,2008]" box="[151,1047,973,1000]" pageId="1" pageNumber="320">
<emphasis id="B9AEEA850D0AFFA1618C5DC8D347F7C4" bold="true" box="[151,290,973,999]" pageId="1" pageNumber="320">Etymology:</emphasis>
From the surname Grazia and the generic name
<taxonomicName id="4CDA4D140D0AFFA162585DC8D1EAF7C4" authorityName="Hahn" authorityYear="1831" box="[835,911,973,999]" class="Insecta" family="Miridae" genus="Phylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D0AFFA162585DC8D1EAF7C4" box="[835,911,973,999]" italics="true" pageId="1" pageNumber="320">Phylus</emphasis>
</taxonomicName>
; masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3C0651C0D0AFFA161DC5DF4D3E4F253" pageId="1" pageNumber="320" type="diagnosis">
<paragraph id="8B6536970D0AFFA161DC5DF4D747F198" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="320">
<emphasis id="B9AEEA850D0AFFA161DC5DF4D321F028" bold="true" box="[199,324,1009,1035]" pageId="1" pageNumber="320">Diagnosis:</emphasis>
<emphasis id="B9AEEA850D0AFFA160515DF4D3E1F028" box="[330,388,1009,1035]" italics="true" pageId="1" pageNumber="320">Male</emphasis>
: Small, total length 2.56–3.18, weakly elongate, broadly ovate, weakly to distinctly flattened, width across pronotum 1.05–1.11, membrane weakly exceeding apex of abdomen; coloration castaneous to nearly black, posterior margin of vertex pale, antennae pale (
<figureCitation id="13E12A120D0AFFA163E75A3CD13BF070" box="[764,862,1081,1107]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Fig. 1, 3</figureCitation>
), legs, except tibiae and tarsi ranging from pale to black/castaneous, tibiae pale, always with black spines with black bases, tarsi pale; head short, broad, eyes somewhat bulging as viewed from above, posterior margin of vertex rounded, nearly straight across (
<figureCitation id="13E12A120D0AFFA165B45A84D77AF0B8" box="[1199,1311,1153,1179]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
); antennae short, slender, not sexually dimorphic, pale (
<figureCitation id="13E12A120D0AFFA163915AA0D091F09C" box="[650,756,1189,1215]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
); claws only moderately elongate, nearly straight over much of length; pulvilli elongate, adnate to claw over most of length of the latter (
<figureCitation id="13E12A120D0AFFA165065ACCD6EBF0C0" box="[1053,1166,1225,1252]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="1" pageNumber="320">Figs. 7, 8</figureCitation>
); parempodia setiform. Dimensions and proportions as in
<tableCitation id="C658032C0D0AFFA163005AE8D015F124" box="[539,624,1261,1287]" captionStart="TABLE 1" captionStartId="5.[151,244,570,597]" captionTargetPageId="5" captionText="TABLE 1. Measurements of Graziaphylus." httpUri="http://table.plazi.org/id/DFA5661F0D0EFFA5618C5C3FD01BF676" pageId="1" pageNumber="320" tableUuid="DFA5661F0D0EFFA5618C5C3FD01BF676">Table 1</tableCitation>
; interocular distance relative to width of head apparently distinctive for two known species. Vestiture composed of reclining simple setae intermixed with flattened lanceolate setae, the latter either dark or silvery in appearance (
<figureCitation id="13E12A120D0AFFA1636B5B30D0BFF16C" box="[624,730,1333,1359]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
,
<figureCitation id="13E12A120D0AFFA163FC5B30D144F16C" box="[743,801,1333,1359]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="1" pageNumber="320">9, 10</figureCitation>
). Legs short, femora flat (
<figureCitation id="13E12A120D0AFFA165595B30D6C8F16C" box="[1090,1197,1333,1359]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
). GENITALIA (
<figureCitation id="13E12A120D0AFFA1647F5B30D36FF150" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1850,1874]" captionTargetBox="[204,1384,181,1826]" captionTargetId="figure-26@6.[204,1384,181,1826]" captionTargetPageId="6" captionText="FIGURES 12–18. Male and female genitalia of Graziaphylus spp (AMNH_PBI 00415251, AMNH_PBI 00415254, AMNH_ PBI 00415323). 12. Phallotheca, G. joceliae. 13. Lateral view of endosoma, G. joceliae. 14. Lateral view of right paramere, G. joceliae. 15. Lateral view of left paramere, G. joceliae. 16. Dorsal view of female bursa copulatrix and gonapophyses 8, G. joceliae. 17. Lateral view of female posterior wall and gonapophyses 9 and conocoxae 9, G. joceliae. 18. Lateral view of endosoma, G. nigripes." figureDoi="http://doi.org/10.5281/zenodo.4691981" httpUri="https://zenodo.org/record/4691981/files/figure.png" pageId="1" pageNumber="320">Figs. 12–15, 18</figureCitation>
): Endosoma with a single strap, apical attenuation single, about the length of the gonopore and with one or two barbs; gonopore relatively large; phallotheca with apical portion at right angles relative to proximal portion; left paramere conventionally phyline, without distinctive structural variation; right paramere lanceolate.
</paragraph>
<paragraph id="8B6536970D0AFFA161DC5BC0D7E1F1FC" blockId="1.[151,1437,973,2008]" box="[199,1412,1477,1504]" pageId="1" pageNumber="320">
Female (
<figureCitation id="13E12A120D0AFFA160325BC0D3F6F1FC" box="[297,403,1477,1503]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 2, 4</figureCitation>
) similar to male in coloration and vestiture, but somewhat larger and more broadly ovoid.
</paragraph>
<paragraph id="8B6536970D0AFFA161DC5BEFD3E4F253" blockId="1.[151,1437,973,2008]" pageId="1" pageNumber="320">
Among Australian
<taxonomicName id="4CDA4D140D0AFFA160B35BECD06FF220" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[424,522,1513,1539]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
<taxonomicName id="4CDA4D140D0AFFA1630C5BECD0CBF220" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[535,686,1513,1539]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D0AFFA1630C5BECD0CBF220" box="[535,686,1513,1539]" italics="true" pageId="1" pageNumber="320">Graziaphylus</emphasis>
</taxonomicName>
is most similar in coloration and shape to
<taxonomicName id="4CDA4D140D0AFFA165A45BECD3BDF204" authority="Weirauch and Schuh (2011)" authorityName="Weirauch and Schuh" authorityYear="2011" class="Insecta" family="Miridae" genus="Xiphoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="320" phylum="Arthropoda" rank="species" species="gilvus">
<emphasis id="B9AEEA850D0AFFA165A45BECD7F9F220" box="[1215,1436,1513,1539]" italics="true" pageId="1" pageNumber="320">Xiphoidellus gilvus</emphasis>
<bibRefCitation id="EF4B4B660D0AFFA1618C5808D3BDF204" author="Weirauch, C. &amp; Schuh, R. T." box="[151,472,1549,1575]" pageId="1" pageNumber="320" pagination="473 - 508" refId="ref4478" refString="Weirauch, C. &amp; Schuh, R. T. (2011) Southern hemisphere distributional patterns in plant bugs (Hemiptera: Miridae: Phylinae): Xiphoidellus, gen. nov. from Australia and Ampimpacoris, gen. nov. from Argentina, show transantarctic relationships. Invertebrate Systematics, 24, 473 - 508. https: // doi. org / 10.1071 / IS 10002" type="journal article" year="2011">Weirauch and Schuh (2011)</bibRefCitation>
</taxonomicName>
, but the latter taxon lacks scalelike setae, has small flaplike pulvilli placed near the midpoint of the claw, and the endosoma forms a complete coil and with a fingerlike process at the apex of the secondary gonopore.
</paragraph>
</subSubSection>
<subSubSection id="C3C0651C0D0AFFA261DC587CD35EF7CE" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320" type="description">
<paragraph id="8B6536970D0AFFA261DC587CD614F5AA" blockId="1.[151,1437,973,2008]" lastBlockId="2.[151,1437,151,1005]" lastPageId="2" lastPageNumber="321" pageId="1" pageNumber="320">
<emphasis id="B9AEEA850D0AFFA161DC587CD33EF2B0" bold="true" box="[199,347,1657,1683]" pageId="1" pageNumber="320">Description:</emphasis>
<emphasis id="B9AEEA850D0AFFA16079587CD3F9F2B0" box="[354,412,1657,1683]" italics="true" pageId="1" pageNumber="320">Male</emphasis>
: Moderately small, broad-bodied, somewhat flattened, total length 2.56–3.18, width across pronotum 1.05–1.11. COLORATION (
<figureCitation id="13E12A120D0AFFA163485898D0A0F294" box="[595,709,1693,1719]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
): Body castaneous to nearly black; antennae pale; legs pale or castaneous/black; tibiae pale, spines with dark bases; membrane of wing, including veins, fumose. SURFACE AND VESTITURE (
<figureCitation id="13E12A120D0AFFA1609358E0D39DF2DC" box="[392,504,1765,1791]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
,
<figureCitation id="13E12A120D0AFFA1631358E0D003F2DC" box="[520,614,1765,1792]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="1" pageNumber="320">5, 9, 10</figureCitation>
): Dorsum smooth, shining, covered with recumbent, dark, simple setae and some dark or silvery scalelike setae. STRUCTURE: Curvature of posterior margin of vertex and eyes more or less conforming to anterior margin of pronotum (
<figureCitation id="13E12A120D0AFFA163E05928D108F364" box="[763,877,1837,1863]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
); posterior margin of vertex and eyes smoothly and rather broadly rounded (
<figureCitation id="13E12A120D0AFFA160C75954D02CF348" box="[476,585,1873,1899]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
); vertex and frons moderately rounded; clypeus barely visible from above; dorsal margin of antennal fossa at level of ventral margin of eye; head short in lateral view, eyes occupying much of height of head; labium reaching to middle trochanters; costal margin of hemelytra broadly convex (
<figureCitation id="13E12A120D0AFFA1643B599CD7E8F390" box="[1312,1421,1945,1971]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
), cuneal fracture pronounced, hemelytra weakly deflexed at fracture (
<figureCitation id="13E12A120D0AFFA1628859B8D19BF3F4" box="[915,1022,1981,2007]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="1" pageNumber="320">Figs. 1, 3</figureCitation>
); pronotum flattened, nearly straight on lateral and posterior margins; mesoscutum moderately exposed, scutellum flattened, weakly elevated anteriorly; claws of moderate length, nearly straight along dorsal margin, sharply bent near apex, pulvilli long, adnate to ventral surface of claw over most of its length (
<figureCitation id="13E12A120D09FFA263795EDAD0B5F4DA" box="[610,720,223,249]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="2" pageNumber="321">Figs. 7, 8</figureCitation>
), parempodia setiform (
<figureCitation id="13E12A120D09FFA262F85EDAD634F4DA" box="[995,1105,223,249]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="2" pageNumber="321">Figs. 7, 8</figureCitation>
). GENITALIA (
<figureCitation id="13E12A120D09FFA264175EDAD2D3F53E" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1850,1874]" captionTargetBox="[204,1384,181,1826]" captionTargetId="figure-26@6.[204,1384,181,1826]" captionTargetPageId="6" captionText="FIGURES 12–18. Male and female genitalia of Graziaphylus spp (AMNH_PBI 00415251, AMNH_PBI 00415254, AMNH_ PBI 00415323). 12. Phallotheca, G. joceliae. 13. Lateral view of endosoma, G. joceliae. 14. Lateral view of right paramere, G. joceliae. 15. Lateral view of left paramere, G. joceliae. 16. Dorsal view of female bursa copulatrix and gonapophyses 8, G. joceliae. 17. Lateral view of female posterior wall and gonapophyses 9 and conocoxae 9, G. joceliae. 18. Lateral view of endosoma, G. nigripes." figureDoi="http://doi.org/10.5281/zenodo.4691981" httpUri="https://zenodo.org/record/4691981/files/figure.png" pageId="2" pageNumber="321">Figs. 12, 15, 18</figureCitation>
): Endosoma formed of a single strap, more-or-less J-shaped, with a slender, nearly straight, apical attenuation about length of secondary gonopore with one or two barbs; secondary gonopore well sclerotized, subtending apical endosomal spine, gonopore sclerite lacking; phallotheca with apical portion conical, at right angle to long, broad basal flap; left paramere of characteristic phyline
<typeStatus id="546188350D09FFA263D85F6AD092F5AA" box="[707,759,367,393]" pageId="2" pageNumber="321">type</typeStatus>
; right paramere small, lanceolate.
</paragraph>
<paragraph id="8B6536970D09FFA261DC5F96D69AF5D6" blockId="2.[151,1437,151,1005]" pageId="2" pageNumber="321">
<emphasis id="B9AEEA850D09FFA261DC5F96D37EF58E" box="[199,283,403,429]" italics="true" pageId="2" pageNumber="321">Female</emphasis>
: Similar to male, but more broadly ovoid; total length 2.98–3.24; width pronotum 1.11–1.26. COLORATION (
<figureCitation id="13E12A120D09FFA260405FB2D3AFF5F2" box="[347,458,439,465]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="2" pageNumber="321">Figs. 2, 4</figureCitation>
): As in male. SURFACE AND VESTITURE (
<figureCitation id="13E12A120D09FFA262C65FB2D63CF5F2" box="[989,1113,439,465]" captionStart="FIGURES 5–11" captionStartId="3.[151,264,1941,1965]" captionTargetBox="[170,1417,669,1917]" captionTargetId="figure-209@3.[170,1417,669,1917]" captionTargetPageId="3" captionText="FIGURES 5–11. Scanning micrographs of morphology of Graziaphylus joceliae female (AMNH_PBI 00415299). 5. Lateral view of head and thorax. 6. Metathoracic scent-efferent system. 7. Frontal view of pretarsus. 8. Detail of pretarsus. 9. Lateral view of head and pronotum. 10. Detail of setae on anterolateral margin of prothorax. 11. Setae on abdominal venter." figureDoi="http://doi.org/10.5281/zenodo.4691979" httpUri="https://zenodo.org/record/4691979/files/figure.png" pageId="2" pageNumber="321">Figs. 9–11</figureCitation>
): As in male. STRUCTURE (
<figureCitation id="13E12A120D09FFA261BB5FDED364F5D6" box="[160,257,475,501]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1637,1661]" captionTargetBox="[161,1432,1039,1602]" captionTargetId="figure-392@2.[151,1436,1029,1614]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus images of Graziaphylus spp. 1. G. joceliae, male holotype (AMNH_PBI 00415222). 2. G. joceliae, female (AMNH_PBI 00415318). 3. G. nigripes, male holotype (AMNH_PBI 00087344). 4. G. nigripes, female (AMNH_PBI 00415325)." figureDoi="http://doi.org/10.5281/zenodo.4691977" httpUri="https://zenodo.org/record/4691977/files/figure.png" pageId="2" pageNumber="321">Fig. 2, 4</figureCitation>
): Macropterous; eyes apparently lacking sexual dimorphism. GENITALIA:
<figureCitation id="13E12A120D09FFA2654C5FDED69EF5D6" box="[1111,1275,475,501]" captionStart="FIGURES 12–18" captionStartId="6.[151,264,1850,1874]" captionTargetBox="[204,1384,181,1826]" captionTargetId="figure-26@6.[204,1384,181,1826]" captionTargetPageId="6" captionText="FIGURES 12–18. Male and female genitalia of Graziaphylus spp (AMNH_PBI 00415251, AMNH_PBI 00415254, AMNH_ PBI 00415323). 12. Phallotheca, G. joceliae. 13. Lateral view of endosoma, G. joceliae. 14. Lateral view of right paramere, G. joceliae. 15. Lateral view of left paramere, G. joceliae. 16. Dorsal view of female bursa copulatrix and gonapophyses 8, G. joceliae. 17. Lateral view of female posterior wall and gonapophyses 9 and conocoxae 9, G. joceliae. 18. Lateral view of endosoma, G. nigripes." figureDoi="http://doi.org/10.5281/zenodo.4691981" httpUri="https://zenodo.org/record/4691981/files/figure.png" pageId="2" pageNumber="321">Figures 16, 17</figureCitation>
.
</paragraph>
<paragraph id="8B6536970D09FFA261DC5FFAD39FF77E" blockId="2.[151,1437,151,1005]" pageId="2" pageNumber="321">
Discussion:Among Australian taxa,
<taxonomicName id="4CDA4D140D09FFA263475FFAD096F63A" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[604,755,511,537]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA263475FFAD096F63A" box="[604,755,511,537]" italics="true" pageId="2" pageNumber="321">Graziaphylus</emphasis>
</taxonomicName>
will be most easily confused with the uniformly dark-colored species of
<taxonomicName id="4CDA4D140D09FFA260145C26D0F4F61E" authority="Weirauch and Schuh" authorityName="Weirauch and Schuh" box="[271,657,547,573]" class="Insecta" family="Miridae" genus="Xiphoidellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA260145C26D3F8F61E" box="[271,413,547,573]" italics="true" pageId="2" pageNumber="321">Xiphoidellus</emphasis>
Weirauch and Schuh
</taxonomicName>
; nonetheless, the male genitalia in the two groups are distinctive and the latter lacks the scalelike setae seen in the former. On a world basis,
<taxonomicName id="4CDA4D140D09FFA262CE5C42D609F642" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[981,1132,583,609]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA262CE5C42D609F642" box="[981,1132,583,609]" italics="true" pageId="2" pageNumber="321">Graziaphylus</emphasis>
</taxonomicName>
species are similar in size and appearance to the multiple dark-colored species of
<taxonomicName id="4CDA4D140D09FFA2620A5C6ED193F6A6" authority="Fieber" authorityName="Fieber" box="[785,1014,619,645]" class="Insecta" family="Miridae" genus="Chlamydatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA2620A5C6ED1C0F6A6" box="[785,933,619,645]" italics="true" pageId="2" pageNumber="321">Chlamydatus</emphasis>
Fieber
</taxonomicName>
in the Northern Hemisphere, but the latter have minute, flaplike pulvilli situated at the midpoint of the claw (
<bibRefCitation id="EF4B4B660D09FFA262CB5C8AD691F68A" author="Schuh, R. T. &amp; Schwartz, M. D." box="[976,1268,654,681]" pageId="2" pageNumber="321" pagination="1 - 55" refId="ref4174" refString="Schuh, R. T. &amp; Schwartz, M. D. (2005) Review of North American Chlamydatus Curtis species, with new synonymy and the description of two new species (Heteroptera: Miridae: Phylinae). American Museum Novitates, 3471, 1 - 55. https: // doi. org / 10.1206 / 0003 - 0082 (2005) 471 % 3 C 0001: RONACC % 3 E 2.0. CO; 2" type="journal article" year="2005">Schuh &amp; Schwartz, 2005</bibRefCitation>
). Among other
<taxonomicName id="4CDA4D140D09FFA2618C5CB6D299F6EE" authorityName="Douglas &amp; Scott" authorityYear="1865" box="[151,252,691,717]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="subFamily" subFamily="Phylinae">Phylinae</taxonomicName>
, genera such as
<taxonomicName id="4CDA4D140D09FFA260AD5CB6D1A9F6EE" authority="Schuh and Schwartz (2004)" authorityName="Schuh and Schwartz" authorityYear="2004" box="[438,972,691,717]" class="Insecta" family="Miridae" genus="Arctostaphylocoris" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA260AD5CB6D0EFF6EE" box="[438,650,691,717]" italics="true" pageId="2" pageNumber="321">Arctostaphylocoris</emphasis>
<bibRefCitation id="EF4B4B660D09FFA263895CB6D1A9F6EE" author="Schuh, R. T. &amp; Schwartz, M. D." box="[658,972,691,717]" pageId="2" pageNumber="321" pagination="1 - 36" refId="ref4094" refString="Schuh, R. T. &amp; Schwartz, M. D. (2004) New genera, new species, new synonyms, and new combinations in North America and Caribbean Phylinae (Heteroptera: Miridae). American Museum Novitates, 3436, 1 - 36. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 436 % 3 C 0001: NGNSNS % 3 E 2.0. CO; 2" type="journal article" year="2004">Schuh and Schwartz (2004)</bibRefCitation>
</taxonomicName>
from the western Nearctic, the Holarctic
<taxonomicName id="4CDA4D140D09FFA2618C5CD2D3E9F6D2" authority="Reuter" authorityName="Reuter" authorityYear="1875" box="[151,396,727,753]" class="Insecta" family="Miridae" genus="Phoenicocoris" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA2618C5CD2D35CF6D2" box="[151,313,727,753]" italics="true" pageId="2" pageNumber="321">Phoenicocoris</emphasis>
Reuter
</taxonomicName>
(
<bibRefCitation id="EF4B4B660D09FFA260805CD2D080F6D2" author="Schwartz, M. D. &amp; Stonedahl, G. M." box="[411,741,727,753]" pageId="2" pageNumber="321" refId="ref4335" refString="Schwartz, M. D. &amp; Stonedahl, G. M. (2004) Revision of Phoenicocoris Reuter with descriptions of three new species from North America and a new genus from Japan (Heteroptera: Miridae: Phylinae). American Museum Novitates, 3464, 55 pp. https: // doi. org / 10.1206 / 0003 - 0082 (2004) 464 % 3 C 0001: ROPRWD % 3 E 2.0. CO; 2" type="book" year="2004">Schwartz &amp; Stonedahl, 2004</bibRefCitation>
), members of the widely distributed
<taxonomicName id="4CDA4D140D09FFA2659F5CD2D73BF6D2" authority="Reuter" authorityName="Reuter" box="[1156,1374,727,753]" class="Insecta" family="Miridae" genus="Sthenaridea" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA2659F5CD2D76EF6D2" box="[1156,1291,727,753]" italics="true" pageId="2" pageNumber="321">Sthenaridea</emphasis>
Reuter
</taxonomicName>
from the Southern Hemisphere, and several of the New World
<taxonomicName id="4CDA4D140D09FFA262025CFED1BBF736" authority="Reuter" authorityName="Reuter" box="[793,990,763,789]" class="Insecta" family="Miridae" genus="Rhinacloa" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA262025CFED1E9F736" box="[793,908,763,789]" italics="true" pageId="2" pageNumber="321">Rhinacloa</emphasis>
Reuter
</taxonomicName>
(
<bibRefCitation id="EF4B4B660D09FFA262F75CFED76DF736" author="Schuh, R. T. &amp; Schwartz, M. D." box="[1004,1288,762,789]" pageId="2" pageNumber="321" pagination="379 - 470" refId="ref4033" refString="Schuh, R. T. &amp; Schwartz, M. D. (1985) Revision of the plant bug genus Rhinacloa Reuter with a phylogenetic analysis (Hemiptera: Miridae). Bulletin of the American Museum of Natural History, 179, 379 - 470. http: // hdl. handle. net / 2246 / 945" type="journal article" year="1985">Schuh &amp; Schwartz, 1985</bibRefCitation>
) species also share a resemblance, including their size and the fact that they all have some scalelike setae on the dorsum and usually on the thoracic pleuron.
</paragraph>
<paragraph id="8B6536970D09FFA261DC5D62D35EF7CE" blockId="2.[151,1437,151,1005]" pageId="2" pageNumber="321">
We tentatively place
<taxonomicName id="4CDA4D140D09FFA260A65D62D031F7A2" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[445,596,871,897]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA260A65D62D031F7A2" box="[445,596,871,897]" italics="true" pageId="2" pageNumber="321">Graziaphylus</emphasis>
</taxonomicName>
in the tribe
<taxonomicName id="4CDA4D140D09FFA263F25D62D101F7A2" box="[745,868,871,897]" pageId="2" pageNumber="321" rank="tribe" tribe="Nasocorini">Nasocorini</taxonomicName>
(see
<bibRefCitation id="EF4B4B660D09FFA262BD5D62D6DFF7A2" author="Schuh, R. T. &amp; Menard, K. L." box="[934,1210,871,897]" pageId="2" pageNumber="321" pagination="1 - 72" refId="ref3976" refString="Schuh, R. T. &amp; Menard, K. L. (2013) A revised classification of the Phylinae (Insecta: Heteroptera: Miridae): Arguments for the placement of genera. American Museum Novitates, 3785, 1 - 72. https: // doi. org / 10.1206 / 3785.2" type="journal article" year="2013">Schuh &amp; Menard, 2013</bibRefCitation>
) based on the size, structure of the male genitalia, and form of the claws and pulvilli. This is the first Australian taxon attributed to the tribe, whose monophyly and accommodation of
<taxonomicName id="4CDA4D140D09FFA263AF5DAAD12EF7EA" authorityName="Schuh &amp; Salas" authorityYear="2021" box="[692,843,943,969]" class="Insecta" family="Miridae" genus="Graziaphylus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="B9AEEA850D09FFA263AF5DAAD12EF7EA" box="[692,843,943,969]" italics="true" pageId="2" pageNumber="321">Graziaphylus</emphasis>
</taxonomicName>
will strongly benefit from further testing using DNA sequence data.
</paragraph>
</subSubSection>
</treatment>
</document>
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