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<document id="D311C6FA559649C839BD7642E87962F0" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630475279EF4EFECA75D604E5" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Canthidium lebasi Harold 1867" docType="treatment" docVersion="2" lastPageNumber="30" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="30" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="86C186A3BD0684FE214621ECAE409183">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="3D4D56FE425D36CAABB18DCCEC92F4E2">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="CCE8CAB42A971617BEA8AF25663CB48E">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="432421E3F369A7970E6786029AC3B73A">Cupello, Mario</mods:namePart>
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<mods:affiliation id="E795A685BFE7C27DED1C3694BC0F4C9A">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<paragraph id="9546C77030475279EF4EFECA74D600B8" blockId="29.[601,986,365,427]" box="[601,986,365,392]" pageId="29" pageNumber="30">
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<taxonomicName id="52F9BCF330475279EF4EFECA74D600B8" ID-CoL="5WXF3" authority="Harold, 1867" authorityName="Harold" authorityYear="1867" box="[601,986,365,392]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lebasi">
<emphasis id="A78D1B6230475279EF4EFECA743400B7" bold="true" box="[601,824,365,392]" italics="true" pageId="29" pageNumber="30">Canthidium lebasi</emphasis>
Harold, 1867
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<figureCitation id="0DC2DBF530475279EFF1FE3674410094" box="[742,845,401,427]" captionStart="Fig" captionStartId="30.[189,232,1928,1954]" captionTargetBox="[206,1380,313,1848]" captionTargetId="figure-18@30.[193,1394,265,1880]" captionTargetPageId="30" captionText="Fig. 13. Species distribution. A. Bradypodidium venezuelense (Ferreira &amp; Galileo, 1993). B. Dorsal view of B. venezuelense. C. Canthidium lebasi Harold, 1867. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945492" httpUri="https://zenodo.org/record/13945492/files/figure.png" pageId="29" pageNumber="30">Fig. 13C</figureCitation>
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<emphasis id="A78D1B6230475279EDAAFE75769400D3" box="[189,408,466,492]" italics="true" pageId="29" pageNumber="30">Canthidium lebasi</emphasis>
<bibRefCitation id="F168BA8130475279ECB3FE75757300D3" author="Harold E." box="[420,639,466,492]" pageId="29" pageNumber="30" pagination="1 - 61" refId="ref123143" refString="Harold E. 1867 a. Zur Kenntniss der Gattung Canthidium und ihrer nachsten Verwandten. Coleopterologische Hefte 1: 1 - 61. Available from https: // www. biodiversitylibrary. org / page / 16069237 [accessed 16 Feb. 2024]." type="journal article" year="1867">Harold, 1867a: 38</bibRefCitation>
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(original description). Type locality: 
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. Name-bearing type: 
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(MNHN), examined by FZVM.
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<taxonomicName id="52F9BCF330475279EDAAFD907690036E" ID-CoL="5WXF3" authorityName="Harold" authorityYear="1867" box="[189,412,567,593]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lebasi">Canthidium lebasi</taxonomicName>
–
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<treatmentCitation id="1458E16130475279ECDFFD9075B5036E" author="Candeze E." box="[456,697,567,593]" page="330" pageId="29" pageNumber="30" year="1891">
<bibRefCitation id="F168BA8130475279ECDFFD9075B5036E" author="Candeze E." box="[456,697,567,593]" pageId="29" pageNumber="30" refId="ref113476" refString="Candeze E. 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888), 15 e Memoire (1). Coleopteres - Lamellicornes determines par le Dr E. Candeze. Annales de la Societe entomologique de France 60: [329] - 332. Available from https: // www. biodiversitylibrary. org / page / 32547689 [accessed 14 Feb. 2024]." type="journal volume" year="1891">Candèze 1891: 330</bibRefCitation>
</treatmentCitation>
(list from 
<collectingCountry id="EDEE87E030475279EE5CFD9074C5036E" box="[843,969,567,593]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
). 
<emphasis id="A78D1B6230475279EEF4FD9F74F1036E" box="[995,1021,568,593]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279E91BFD9073C9036E" author="Roze J. A." box="[1036,1221,567,594]" page="44" pageId="29" pageNumber="30" year="1955">
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</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030475279EDFDFDFD7664034B" box="[234,360,602,628]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
). 
<emphasis id="A78D1B6230475279EC6AFDFC769B034B" box="[381,407,603,628]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279EC88FDFD75F1034A" author="Vulcano M. A. &amp; Pereira F. S." box="[415,765,602,629]" page="595" pageId="29" pageNumber="30" year="1967">
<bibRefCitation id="F168BA8130475279EC88FDFD75F1034A" author="Vulcano M. A. &amp; Pereira F. S." box="[415,765,602,629]" pageId="29" pageNumber="30" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 595</bibRefCitation>
</treatmentCitation>
(key). 
<emphasis id="A78D1B6230475279EE42FDFC7463034B" box="[853,879,603,628]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279EE61FDFD7355034B" author="Havranek D." box="[886,1113,602,628]" page="61" pageId="29" pageNumber="30" year="1989">
<bibRefCitation id="F168BA8130475279EE61FDFD7355034B" author="Havranek D." box="[886,1113,602,628]" pageId="29" pageNumber="30" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
</treatmentCitation>
(list). 
<emphasis id="A78D1B6230475279E9BCFDFC73C9034B" box="[1195,1221,603,628]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279E9DBFDFD761403A8" author="Escobar F." page="206" pageId="29" pageNumber="30" year="2000">
<bibRefCitation id="F168BA8130475279E9DBFDFD761403A8" author="Escobar F." pageId="29" pageNumber="30" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 206</bibRefCitation>
</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030475279ECABFDDA753903A8" box="[444,565,637,663]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
). 
<emphasis id="A78D1B6230475279EF54FDD9755103A8" box="[579,605,638,663]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279EF73FDDA747903A8" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[612,885,637,663]" page="137" pageId="29" pageNumber="30" year="2001">
<bibRefCitation id="F168BA8130475279EF73FDDA747903A8" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[612,885,637,663]" pageId="29" pageNumber="30" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina 
<emphasis id="A78D1B6230475279EFD4FDD975F003A8" box="[707,764,637,663]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2001: 137
</bibRefCitation>
</treatmentCitation>
(checklist for 
<collectingCountry id="EDEE87E030475279E90DFDDA739F03A8" box="[1050,1171,637,663]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
). — Pulido-Herrera 
<emphasis id="A78D1B6230475279EDFFFD06762B0385" box="[232,295,672,698]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2007: 307 (cited for Andean region of 
<collectingCountry id="EDEE87E030475279EFE9FD07747B0385" box="[766,887,672,698]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
). 
<emphasis id="A78D1B6230475279EE9AFD0674AB0385" box="[909,935,673,698]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279EEB9FD06737B0385" author="Lozano C." box="[942,1143,672,699]" page="86" pageId="29" pageNumber="30" year="2010">
<bibRefCitation id="F168BA8130475279EEB9FD06737B0385" author="Lozano C." box="[942,1143,672,699]" pageId="29" pageNumber="30" pagination="83 - 89" refId="ref128409" refString="Lozano C. 2010. Nuevos registros de escarabajos coprofagos para la Orchila, Sierra de Perija, Venezuela. Boletin del Centro de Investigaciones Biologicas 44 (1): 83 - 89." type="journal article" year="2010">Lozano 2010: 86</bibRefCitation>
</treatmentCitation>
(list). 
<emphasis id="A78D1B6230475279E9DDFD0673E80385" box="[1226,1252,673,698]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279E9FCFD0776AE03E1" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." page="108" pageId="29" pageNumber="30" year="2013">
<bibRefCitation id="F168BA8130475279E9FCFD0776AE03E1" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." pageId="29" pageNumber="30" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris 
<emphasis id="A78D1B6230475279EDFDFD63762903E1" box="[234,293,708,734]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2013: 108
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(list).
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</paragraph>
<paragraph id="9546C77030475279EDAAFD4073A5023E" blockId="29.[189,1399,567,839]" box="[189,1193,743,769]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="B5E9E05E30475279EDAAFD4073A5023E" box="[189,1193,743,769]" pageId="29" pageNumber="30">
<taxonomicName id="52F9BCF330475279EDAAFD407558023E" ID-CoL="5WXF3" authorityName="Harold" authorityYear="1867" box="[189,596,743,769]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lebasi" subGenus="Eucanthidium">
<emphasis id="A78D1B6230475279EDAAFD407644023E" box="[189,328,743,769]" italics="true" pageId="29" pageNumber="30">Canthidium</emphasis>
(
<emphasis id="A78D1B6230475279EC4EFD4076F2023E" box="[345,510,743,769]" italics="true" pageId="29" pageNumber="30">Eucanthidium</emphasis>
) 
<emphasis id="A78D1B6230475279EF19FD407558023E" box="[526,596,743,769]" italics="true" pageId="29" pageNumber="30">lebasi</emphasis>
</taxonomicName>
<emphasis id="A78D1B6230475279EF4BFD4F7567023E" box="[604,619,744,769]" italics="true" pageId="29" pageNumber="30">–</emphasis>
<treatmentCitation id="1458E16130475279EF65FD4074C0023E" author="Martinez A. &amp; Halffter G." box="[626,972,743,769]" page="32" pageId="29" pageNumber="30" year="1986">
<bibRefCitation id="F168BA8130475279EF65FD4074C0023E" author="Martinez A. &amp; Halffter G." box="[626,972,743,769]" pageId="29" pageNumber="30" pagination="19 - 40" refId="ref130097" refString="Martinez A. &amp; Halffter G. 1986. Situacion del genero Canthidium. Acta Zoologica Mexicana 18: 19 - 40. https: // doi. org / 10.21829 / azm. 1986.13182047" type="journal article" year="1986">Martínez &amp; Halffter 1986: 32</bibRefCitation>
</treatmentCitation>
(list, distribution).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030475279EDAAFCAD753F0278" blockId="29.[189,1399,567,839]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="B5E9E05E30475279EDAAFCAD753F0278" pageId="29" pageNumber="30">
<taxonomicName id="52F9BCF330475279EDAAFCAD7531021B" ID-CoL="5WXF3" authorityName="Harold" authorityYear="1867" box="[189,573,778,804]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="lebasi" subGenus="Canthidium">
<emphasis id="A78D1B6230475279EDAAFCAD7644021B" box="[189,328,778,804]" italics="true" pageId="29" pageNumber="30">Canthidium</emphasis>
(
<emphasis id="A78D1B6230475279EC4DFCAD76E9021B" box="[346,485,778,804]" italics="true" pageId="29" pageNumber="30">Canthidium</emphasis>
) 
<emphasis id="A78D1B6230475279ECE0FCAD7531021B" box="[503,573,778,804]" italics="true" pageId="29" pageNumber="30">lebasi</emphasis>
</taxonomicName>
– Cupello 2018: 460 (catalogue). 
<emphasis id="A78D1B6230475279EECFFCAC74FE021B" box="[984,1010,779,804]" italics="true" pageId="29" pageNumber="30">—</emphasis>
<treatmentCitation id="1458E16130475279EEEEFCAD727A021B" ID-DOI="http://doi.org/10.5281/zenodo.3684789" ID-Zenodo-Dep="3684789" author="Hielkema A. J. &amp; Hielkema M. A." box="[1017,1398,778,804]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFFBFFDDFF48FA27346EFDFE" page="47" pageId="29" pageNumber="30" year="2019">
<bibRefCitation id="F168BA8130475279EEEEFCAD727A021B" author="Hielkema A. J. &amp; Hielkema M. A." box="[1017,1398,778,804]" pageId="29" pageNumber="30" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 47</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30475279EDAAFCC9765202E4" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="9546C77030475279EDAAFCC976A602B7" blockId="29.[189,426,878,904]" box="[189,426,878,904]" pageId="29" pageNumber="30">
<heading id="CE0E701C30475279EDAAFCC976A602B7" bold="true" box="[189,426,878,904]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="A78D1B6230475279EDAAFCC976A602B7" bold="true" box="[189,426,878,904]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030475279EDAAFC3A765202E4" blockId="29.[189,1398,924,987]" pageId="29" pageNumber="30">
<materialsCitation id="2591CD2D30475279EDAAFC3A74960288" box="[189,922,924,952]" collectingDate="1992-05" country="Venezuela" location="Tachira" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Tachira">
<collectingCountry id="EDEE87E030475279EDAAFC3A76670288" box="[189,363,925,951]" name="Venezuela" pageId="29" pageNumber="30">VENEZUELA</collectingCountry>
‒ 
<emphasis id="A78D1B6230475279EC93FC3B76EB0288" bold="true" box="[388,487,924,951]" pageId="29" pageNumber="30">
<collectingRegion id="573D099230475279EC93FC3B76EB0288" box="[388,487,924,951]" country="Venezuela" name="Tachira" pageId="29" pageNumber="30">Táchira</collectingRegion>
</emphasis>
• 2 specs; Chorro; 
<date id="E147E1B030475279EFD5FC3974300288" box="[706,828,925,952]" pageId="29" pageNumber="30" value="1992-05">
<collectingDate id="F103185830475279EFD5FC3974300288" box="[706,828,925,952]" pageId="29" pageNumber="30" value="1992-05">May 1992</collectingDate>
</date>
; CEMT
</materialsCitation>
. ‒ 
<materialsCitation id="2591CD2D30475279EEA1FC3A765602E4" collectingDate="1989-11-26" collectorName="La Victoria" country="Venezuela" location="Aragua" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Aragua">
<emphasis id="A78D1B6230475279EEA1FC3A73190288" bold="true" box="[950,1045,925,951]" pageId="29" pageNumber="30">
<collectingRegion id="573D099230475279EEA1FC3A73190288" box="[950,1045,925,951]" country="Venezuela" name="Aragua" pageId="29" pageNumber="30">Aragua</collectingRegion>
</emphasis>
• 
<specimenCount id="83FF0CF930475279E93CFC3A73730287" box="[1067,1151,925,952]" count="1" pageId="29" pageNumber="30" type="generic">1 spec.</specimenCount>
; 
<collectorName id="380CA2A630475279E99DFC39721C0288" box="[1162,1296,925,952]" pageId="29" pageNumber="30">La Victoria</collectorName>
; 
<date id="E147E1B030475279E80CFC3A77F702E4" pageId="29" pageNumber="30" value="1989-11-26">
<collectingDate id="F103185830475279E80CFC3A77F702E4" pageId="29" pageNumber="30" value="1989-11-26">26 Nov. 1989</collectingDate>
</date>
; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30475279EDAAFBA575D604E5" pageId="29" pageNumber="30" type="distribution">
<paragraph id="9546C77030475279EDAAFBA5765A0523" blockId="29.[189,342,1026,1052]" box="[189,342,1026,1052]" pageId="29" pageNumber="30">
<heading id="CE0E701C30475279EDAAFBA5765A0523" bold="true" box="[189,342,1026,1052]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="A78D1B6230475279EDAAFBA5765A0523" bold="true" box="[189,342,1026,1052]" pageId="29" pageNumber="30">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030475279EDAAFB9776E60575" blockId="29.[189,490,1072,1098]" box="[189,490,1072,1098]" pageId="29" pageNumber="30">
<collectingCountry id="EDEE87E030475279EDAAFB97763D0575" box="[189,305,1072,1098]" name="Colombia" pageId="29" pageNumber="30">Colombia</collectingCountry>
and 
<collectingCountry id="EDEE87E030475279EC7EFB9776EA0575" box="[361,486,1072,1098]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030475279EDAAFBD576E105B3" blockId="29.[189,493,1138,1164]" box="[189,493,1138,1164]" pageId="29" pageNumber="30">
<heading id="CE0E701C30475279EDAAFBD576E105B3" bold="true" box="[189,493,1138,1164]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="A78D1B6230475279EDAAFBD576E105B3" bold="true" box="[189,493,1138,1164]" pageId="29" pageNumber="30">
Subregions of 
<collectingCountry id="EDEE87E030475279EC78FBD576E105B3" box="[367,493,1138,1164]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030475279EDAAFB07756705E1" blockId="29.[189,1399,1184,1246]" pageId="29" pageNumber="30">
Maracaibo Depression, System of hills and low sierras Lara-Falcón, Andes mountains, Central 
<collectingRegion id="573D099230475279E823FB06727B0584" box="[1332,1399,1185,1211]" country="New Zealand" name="West Coast" pageId="29" pageNumber="30">Coast</collectingRegion>
Mountain Range and Guiana Shield.
</paragraph>
<paragraph id="9546C77030475279EDAAFAA276A80420" blockId="29.[189,420,1285,1311]" box="[189,420,1285,1311]" pageId="29" pageNumber="30">
<heading id="CE0E701C30475279EDAAFAA276A80420" bold="true" box="[189,420,1285,1311]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="A78D1B6230475279EDAAFAA276A80420" bold="true" box="[189,420,1285,1311]" pageId="29" pageNumber="30">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030475279EDAAFA9375D604E5" blockId="29.[189,1399,1331,1498]" pageId="29" pageNumber="30">
<bibRefCitation id="F168BA8130475279EDAAFA9376640471" author="Roze J. A." box="[189,360,1331,1358]" pageId="29" pageNumber="30" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279EC60FA9376FA0471" box="[375,502,1332,1358]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230475279EF16FA9375B70471" box="[513,699,1332,1358]" country="Venezuela" name="Distrito Capital" pageId="29" pageNumber="30">Distrito Capital</collectingRegion>
: Caracas). — 
<bibRefCitation id="F168BA8130475279EE77FA9373BB0471" author="Vulcano M. A. &amp; Pereira F. S." box="[864,1207,1331,1358]" pageId="29" pageNumber="30" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 595</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279E9D1FA93724A0471" box="[1222,1350,1332,1358]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
). — 
<bibRefCitation id="F168BA8130475279EDAAFAF0751A044E" author="Martinez A. &amp; Halffter G." box="[189,534,1367,1393]" pageId="29" pageNumber="30" pagination="19 - 40" refId="ref130097" refString="Martinez A. &amp; Halffter G. 1986. Situacion del genero Canthidium. Acta Zoologica Mexicana 18: 19 - 40. https: // doi. org / 10.21829 / azm. 1986.13182047" type="journal article" year="1986">Martínez &amp; Halffter 1986: 32</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279EF31FAF075AA044E" box="[550,678,1367,1393]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
). — 
<bibRefCitation id="F168BA8130475279EFCAFAF074B1044E" author="Havranek D." box="[733,957,1367,1393]" pageId="29" pageNumber="30" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279EEDAFAF07340044E" box="[973,1100,1367,1393]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230475279E941FAF073B8044E" box="[1110,1204,1367,1393]" country="Venezuela" name="Tachira" pageId="29" pageNumber="30">Táchira</collectingRegion>
: outskirts of 
<collectingCountry id="EDEE87E030475279E85DFAF0762204AB" name="Saint Kitts and Nevis" pageId="29" pageNumber="30">San Cristóbal</collectingCountry>
). — 
<bibRefCitation id="F168BA8130475279EC72FADC752704AB" author="Lozano C." box="[357,555,1402,1429]" pageId="29" pageNumber="30" pagination="83 - 89" refId="ref128409" refString="Lozano C. 2010. Nuevos registros de escarabajos coprofagos para la Orchila, Sierra de Perija, Venezuela. Boletin del Centro de Investigaciones Biologicas 44 (1): 83 - 89." type="journal article" year="2010">Lozano 2010: 86</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279EF2CFADD75B604AB" box="[571,698,1402,1428]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230475279EFD3FADD740B04AB" box="[708,775,1402,1428]" country="Venezuela" name="Zulia" pageId="29" pageNumber="30">Zulia</collectingRegion>
). — 
<bibRefCitation id="F168BA8130475279EE28FADD738504AB" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[831,1161,1402,1429]" pageId="29" pageNumber="30" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris 
<emphasis id="A78D1B6230475279EEC6FADC730904AB" box="[977,1029,1402,1428]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2013: 108
</bibRefCitation>
(
<collectingCountry id="EDEE87E030475279E98FFADD721B04AB" box="[1176,1303,1402,1428]" name="Venezuela" pageId="29" pageNumber="30">Venezuela</collectingCountry>
: 
<collectingRegion id="573D099230475279E837FADD727A04AB" box="[1312,1398,1402,1428]" country="Venezuela" name="Aragua" pageId="29" pageNumber="30">Aragua</collectingRegion>
[actually, 
<collectingRegion id="573D099230475279EC26FA3A769B0488" box="[305,407,1437,1463]" country="Venezuela" name="Guarico" pageId="29" pageNumber="30">Guárico</collectingRegion>
]: Altagracia de Orituco; 
<collectingRegion id="573D099230475279EFAFFA3A74190488" box="[696,789,1437,1463]" country="Venezuela" name="Bolivar" pageId="29" pageNumber="30">Bolívar</collectingRegion>
: 
<collectingCountry id="EDEE87E030475279EE37FA3A74460488" box="[800,842,1437,1463]" name="Iceland" pageId="29" pageNumber="30">Isla</collectingCountry>
de Anacoco; Miranda: Altos de Pipe; Yaracuy: Hacienda Guáquira; Zulia: Rosário de Perijá).
</paragraph>
</subSubSection>
</treatment>
</document>