treatments-xml/data/03/8F/87/038F8788FF9DFFF2D973FDBDFD6424EF.xml

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<mods:title id="6BFDCD7B41B33D4B300F0C136A3A8FCB">A review of Chelonian type specimens (order Testudines)</mods:title>
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<mods:namePart id="78B42B4D8BF6AA38DEFB373BD5B4EE1D">Iverson, John B.</mods:namePart>
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<mods:date id="38F33BB0EABAD5BA5BC9E83A375CF009">2022</mods:date>
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<treatment id="038F8788FF9DFFF2D973FDBDFD6424EF" ID-DOI="http://doi.org/10.5281/zenodo.6204903" ID-GBIF-Taxon="193856575" ID-Zenodo-Dep="6204903" LSID="urn:lsid:plazi:treatment:038F8788FF9DFFF2D973FDBDFD6424EF" httpUri="http://treatment.plazi.org/id/038F8788FF9DFFF2D973FDBDFD6424EF" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" scope_class="Testudines" scope_family="Pelomedusidae" scope_subOrder="Pleurodira">
<subSubSection id="C33C6515FF9DFFF5D973FDBDFA82206E" box="[810,1303,1770,1797]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B99369EFF9DFFF5D973FDBDFA82206E" blockId="11.[810,1452,1770,1894]" box="[810,1303,1770,1797]" pageId="11" pageNumber="12">
<heading id="D0D181F2FF9DFFF5D973FDBDFA82206E" box="[810,1303,1770,1797]" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
<treatmentCitationGroup id="AB3611B0FF9DFFF5D973FDBDFA82206E" box="[810,1303,1770,1797]" pageId="11" pageNumber="12">
<treatmentCitation id="0A87108FFF9DFFF5D973FDBDFA82206E" author="Schweigger, A. F." box="[810,1303,1770,1797]" page="314" pageId="11" pageNumber="12" year="1812">
<taxonomicName id="4C264D1DFF9DFFF5D973FDBDFA82206E" ID-CoL="76FM7" authority="(Schweigger 1812: 314)" baseAuthorityName="Schweigger" baseAuthorityPageNumber="314" baseAuthorityYear="1812" box="[810,1303,1770,1797]" class="Reptilia" family="Pelomedusidae" genus="Pelusios" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="castaneus">
<emphasis id="B952EA8CFF9DFFF5D973FDBDFA82206E" bold="true" box="[810,1303,1770,1797]" pageId="11" pageNumber="12">
<emphasis id="B952EA8CFF9DFFF5D973FDBDFB96206F" bold="true" box="[810,1027,1770,1797]" italics="true" pageId="11" pageNumber="12">Pelusios castaneus</emphasis>
(
<bibRefCitation id="EFB74B6FFF9DFFF5DE48FDBDFA85206E" author="Schweigger, A. F." box="[1041,1296,1770,1797]" pageId="11" pageNumber="12" pagination="406 - 462" refId="ref81144" refString="Schweigger, A. F. (1812) Prodromus monographiae Cheloniorum. Konigsberger Archiv fur Naturwissenschaft und Mathematik, 1, 271 - 368, 406 - 462." type="journal article" year="1812">Schweigger 1812:314</bibRefCitation>
)
</emphasis>
</taxonomicName>
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</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33C6515FF9DFFF2D973FC5CFD6424EF" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8B99369EFF9DFFF5D973FC5CFA9F200C" blockId="11.[810,1452,1770,1894]" pageId="11" pageNumber="12">
<emphasis id="B952EA8CFF9DFFF5D973FC5CFC35204F" bold="true" box="[810,928,1803,1829]" pageId="11" pageNumber="12">
<typeStatus id="549D883CFF9DFFF5D973FC5CFC0E204F" box="[810,923,1803,1829]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
:
</emphasis>
MNHN 8366, now lost (
<bibRefCitation id="EFB74B6FFF9DFFF5DEB3FC5CFA0A204C" author="Bour, R." box="[1258,1439,1803,1830]" pageId="11" pageNumber="12" pagination="36 - 40" refId="ref67340" refString="Bour, R. (2008 e) Neotype of Emys castanea Schweigger, 1812 (Pelomedusidae). Emys, 15 (4), 36 - 40." type="journal article" year="2008">Bour 2008e:36</bibRefCitation>
); MNHN 2008.0303 designated 
<typeStatus id="549D883CFF9DFFF5DEB2FC7CFAD1202F" box="[1259,1348,1835,1861]" pageId="11" pageNumber="12" type="neotype">neotype</typeStatus>
by 
<bibRefCitation id="EFB74B6FFF9DFFF5DF2AFC7CFC6A200F" author="Bour, R." pageId="11" pageNumber="12" pagination="36 - 40" refId="ref67340" refString="Bour, R. (2008 e) Neotype of Emys castanea Schweigger, 1812 (Pelomedusidae). Emys, 15 (4), 36 - 40." type="journal article" year="2008">Bour (2008e:37</bibRefCitation>
; photographed therein).
</paragraph>
<paragraph id="8B99369EFF9DFFF5D914FCDCFA9620CC" blockId="11.[845,1451,1931,2085]" box="[845,1283,1931,1958]" pageId="11" pageNumber="12">
<taxonomicName id="4C264D1DFF9DFFF5D914FCDCFA9620CC" authority="Bell 1825: 306" authorityName="Bell" authorityPageNumber="306" authorityYear="1825" box="[845,1283,1931,1958]" class="Reptilia" family="Kinosternidae" genus="Sternotherus" kingdom="Animalia" order="Testudines" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="leachianus">
<emphasis id="B952EA8CFF9DFFF5D914FCDCFBC920CF" box="[845,1116,1931,1957]" italics="true" pageId="11" pageNumber="12">Sternotherus leachianus</emphasis>
Bell 1825:306
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFF9DFFF5D914FCFCFA0A2F4F" blockId="11.[845,1451,1931,2085]" pageId="11" pageNumber="12">
<emphasis id="B952EA8CFF9DFFF5D914FCFCFC2F20AF" bold="true" box="[845,954,1963,1989]" pageId="11" pageNumber="12">
<typeStatus id="549D883CFF9DFFF5D914FCFCFC2F20AF" box="[845,954,1963,1989]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectionCode id="ED37AE5BFF9DFFF5D996FCFCFB8520AF" box="[975,1040,1963,1989]" country="United Kingdom" httpUri="http://grbio.org/cool/g6yf-wft0" name="Oxford University Museum of Natural History" pageId="11" pageNumber="12">OUM</collectionCode>
8618 (illustrated in the original description, Pl. XV; reproduced in 
<bibRefCitation id="EFB74B6FFF9DFFF5DF2AFC9CFC632F6C" author="Bour, R." pageId="11" pageNumber="12" pagination="36 - 40" refId="ref67340" refString="Bour, R. (2008 e) Neotype of Emys castanea Schweigger, 1812 (Pelomedusidae). Emys, 15 (4), 36 - 40." type="journal article" year="2008">Bour 2008e:36</bibRefCitation>
), but see Nowak-Kemp &amp; 
<bibRefCitation id="EFB74B6FFF9DFFF5DF2FFCBCFC6D2F4F" author="Fritz, U." pageId="11" pageNumber="12" pagination="1 - 19" refId="ref78148" refString="Nowak-kemp, M. &amp; Fritz, U. (2010) Chelonian type specimens at the Oxford University Museum. Zootaxa, 2604, 1 - 19. https: // doi. org / 10.11646 / zootaxa. 2604.1.1" type="journal article" year="2010">Fritz (2010:17)</bibRefCitation>
concerning another possible 
<typeStatus id="549D883CFF9DFFF5DF19F35CFA0E2F4F" box="[1344,1435,2059,2085]" pageId="11" pageNumber="12" type="syntype">syntype</typeStatus>
.
</paragraph>
<paragraph id="8B99369EFF9AFFF2DAF2FBFCFDC627AF" blockId="12.[171,777,171,294]" box="[171,595,171,197]" pageId="12" pageNumber="13">
<taxonomicName id="4C264D1DFF9AFFF2DAF2FBFCFDC627AF" authority="Gray 1844: 37" authorityName="Gray" authorityPageNumber="37" authorityYear="1844" box="[171,595,171,197]" class="Reptilia" family="Kinosternidae" genus="Sternotherus" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="derbianus">
<emphasis id="B952EA8CFF9AFFF2DAF2FBFCFE2427AF" box="[171,433,171,197]" italics="true" pageId="12" pageNumber="13">Sternotherus derbianus</emphasis>
Gray 1844:37
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFF9AFFF2DAF2FB9CFDD7264C" blockId="12.[171,777,171,294]" pageId="12" pageNumber="13">
<materialsCitation id="3B4E3CC3FF9AFFF2DAF2FB9CFDD7264C" ID-GBIF-Occurrence="3695119755" collectionCode="BMNH" pageId="12" pageNumber="13" specimenCode="BMNH 1947.3" specimenCount="1" typeStatus="holotype">
<emphasis id="B952EA8CFF9AFFF2DAF2FB9CFE8D278F" bold="true" box="[171,280,203,229]" pageId="12" pageNumber="13">
<typeStatus id="549D883CFF9AFFF2DAF2FB9CFE8D278F" box="[171,280,203,229]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<specimenCode id="DB809EE5FF9AFFF2DB7EFB9CFE44278C" box="[295,465,203,230]" collectionCode="BMNH" pageId="12" pageNumber="13">BMNH 1947.3</specimenCode>
.5.77 (formerly 1841.12. 25.1); illustrated in Gray (1855:Pl. XXII; reproduced in 
<bibRefCitation id="EFB74B6FFF9AFFF2DBD3FA5CFDA0264C" author="Bour, R." box="[394,565,267,294]" pageId="12" pageNumber="13" pagination="36 - 40" refId="ref67340" refString="Bour, R. (2008 e) Neotype of Emys castanea Schweigger, 1812 (Pelomedusidae). Emys, 15 (4), 36 - 40." type="journal article" year="2008">Bour 2008e:36</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8B99369EFF9AFFF2DAF2FA1CFED126EF" blockId="12.[171,778,331,901]" pageId="12" pageNumber="13">
<taxonomicName id="4C264D1DFF9AFFF2DAF2FA1CFED126EF" authority="Siebenrock 1906: 38" authorityName="Siebenrock" authorityPageNumber="38" authorityYear="1906" class="Reptilia" family="Kinosternidae" genus="Sternotherus" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="subSpecies" species="nigricans" subSpecies="seychellensis">
<emphasis id="B952EA8CFF9AFFF2DAF2FA1CFDD1260F" box="[171,580,331,357]" italics="true" pageId="12" pageNumber="13">Sternotherus nigricans seychellensis</emphasis>
Siebenrock 1906:38
</taxonomicName>
</paragraph>
<paragraph id="8B99369EFF9AFFF2DAF2FADDFD6424EF" blockId="12.[171,778,331,901]" pageId="12" pageNumber="13">
<emphasis id="B952EA8CFF9AFFF2DAF2FADDFE8326CE" bold="true" box="[171,278,394,420]" pageId="12" pageNumber="13">
<typeStatus id="549D883CFF9AFFF2DAF2FADDFE8326CE" box="[171,278,394,420]" pageId="12" pageNumber="13" type="syntype">Syntypes</typeStatus>
</emphasis>
(at least 2; Hallerman 1998:211): 
<collectionCode id="ED37AE5BFF9AFFF2D8CEFADCFD4B26CF" box="[663,734,395,421]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
13247 (
<bibRefCitation id="EFB74B6FFF9AFFF2DB67FAFCFD1526AF" author="Tiedemann, F. &amp; Haupl, M." box="[318,640,427,453]" pageId="12" pageNumber="13" pagination="1 - 79" refId="ref83047" refString="Tiedemann, F. &amp; Haupl, M. (1980) Typenkatalog der herpetologischen Sammlung: Reptilia. Kataloge der Wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, 4, 1 - 79." type="journal article" year="1980">Tiedemann &amp; Häupl 1980:8</bibRefCitation>
; 
<bibRefCitation id="EFB74B6FFF9AFFF2D8D4FAFCFE18268F" author="Tiedemann, F. &amp; Haupl, M. &amp; Grillitsch, H." pageId="12" pageNumber="13" pagination="1 - 102" refId="ref83082" refString="Tiedemann, F., Haupl, M. &amp; Grillitsch, H. (1994) Katalog der Typen der herpetologischen Sammlung nach dem stand vom 1. Janner 1994. Teil II. Reptilia. Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien, 10 (Vertebrata 4), 1 - 102." type="journal article" year="1994">
Tiedemann 
<emphasis id="B952EA8CFF9AFFF2DAB1FA9BFEB1268F" box="[232,292,459,485]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
1994:12
</bibRefCitation>
; photographed in 
<bibRefCitation id="EFB74B6FFF9AFFF2D838FA9CFC9F268F" author="Vetter, H." box="[609,778,459,485]" pageId="12" pageNumber="13" refId="ref83873" refString="Vetter, H. (2002) Turtles of the World, Vol. 1: Africa, Europe and Western Asia. Edition Chimaira, Frankfurt, 96 pp." type="book" year="2002">Vetter 2002:93</bibRefCitation>
and 
<bibRefCitation id="EFB74B6FFF9AFFF2DB4EFABCFE15256C" author="Vetter, H." box="[279,384,491,518]" pageId="12" pageNumber="13" refId="ref83933" refString="Vetter, H. (2011) Turtles of the World, Vol. 1: Africa, Europe and Western Asia. Second Edition. Edition Chimaira, Frankfurt, 152 pp." type="book" year="2011">2011:145</bibRefCitation>
and 
<bibRefCitation id="EFB74B6FFF9AFFF2DBECFABCFD34256F" author="Stuckas, H. &amp; Gemel, R. &amp; Fritz, U." box="[437,673,491,517]" pageId="12" pageNumber="13" pagination="57116" refId="ref82593" refString="Stuckas, H., Gemel, R. &amp; Fritz, U. (2013) One extinct turtle species less: Pelusios seychellensis is not extinct, it never existed. PLoS ONE, 8 (4), e 57116. https: // doi. org / 10.1371 / journal. pone. 0057116" type="journal article" year="2013">
Stuckas 
<emphasis id="B952EA8CFF9AFFF2D84AFABBFDDF256F" box="[531,586,491,517]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013:3
</bibRefCitation>
; sampled genetically by 
<bibRefCitation id="EFB74B6FFF9AFFF2DBDCF95CFDDB254F" author="Stuckas, H. &amp; Gemel, R. &amp; Fritz, U." box="[389,590,523,549]" pageId="12" pageNumber="13" pagination="57116" refId="ref82593" refString="Stuckas, H., Gemel, R. &amp; Fritz, U. (2013) One extinct turtle species less: Pelusios seychellensis is not extinct, it never existed. PLoS ONE, 8 (4), e 57116. https: // doi. org / 10.1371 / journal. pone. 0057116" type="journal article" year="2013">
Stuckas 
<emphasis id="B952EA8CFF9AFFF2DB86F95BFD81254F" box="[479,532,523,549]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2013
</bibRefCitation>
and 
<bibRefCitation id="EFB74B6FFF9AFFF2D824F95CFEB7252C" author="Kindler, C. &amp; Moosig, M. &amp; Branch, W. R. &amp; Harvey, J. &amp; Kehlmaier, C. &amp; Nagys, Z. T. &amp; Prokop, H. &amp; Siorky, P. &amp; Fritz, U." pageId="12" pageNumber="13" pagination="305 - 321" refId="ref75636" refString="Kindler, C., Moosig, M., Branch, W. R., Harvey, J., Kehlmaier, C., Nagys, Z. T., Prokop, H., Siorky, P. &amp; Fritz, U. (2016) Comparative phylogeographies of six species of hinged terrapins (Pelusios spp.) reveal discordant patterns and unexpected differentiation in the P. castaneus / P. chapini complex and P. rhodesianus. Biological Journal of the Linnaean Society, 117, 305 - 321. https: // doi. org / 10.1111 / bij. 12647" type="journal article" year="2016">
Kindler 
<emphasis id="B952EA8CFF9AFFF2D88FF95BFC9F254F" box="[726,778,523,549]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2016
</bibRefCitation>
) and 
<collectionCode id="ED37AE5BFF9AFFF2DB06F97CFE08252F" box="[351,413,555,581]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="12" pageNumber="13" type="Museum">ZMH</collectionCode>
992–993 (photographed in 
<bibRefCitation id="EFB74B6FFF9AFFF2D88BF97CFEC5250F" author="Bour, R." pageId="12" pageNumber="13" pagination="27 - 41" refId="ref67566" refString="Bour, R. (2013 a) Actualites chez les tortues des Seychelles. Cheloniens, 29, 27 - 41." type="journal article" year="2013">Bour 2013a:37</bibRefCitation>
and 
<bibRefCitation id="EFB74B6FFF9AFFF2DBDDF91CFD1C250F" author="Bour, R. &amp; Gerlach, J." box="[388,649,587,613]" pageId="12" pageNumber="13" pagination="5" refId="ref67789" refString="Bour, R. &amp; Gerlach, J. (2008) Pelusios seychellensis (Siebenrock 1906) - Seychelles Mud Turtle. In: Rhodin, A. G. J., Pritchard, P. C. H., van Dijk, P. P., Saumure, R. A., Buhlmann, K. A. &amp; Iverson, J. B. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, 5, 18.1 - 18.3. https: // doi. org / 10.3854 / crm. 5.018. seychellensis. v 1.2008" type="book chapter" year="2008">Bour &amp; Gerlach 2008:1</bibRefCitation>
[993 only]; listed in Bour 1983:353 as 982–983; repeated by 
<bibRefCitation id="EFB74B6FFF9AFFF2DB57F9DCFE5925CC" author="Iverson, J. B." box="[270,460,651,678]" pageId="12" pageNumber="13" refId="ref74928" refString="Iverson, J. B. (1992) A Revised Checklist with Distribution Maps of the Turtles of the World. Privately printed, Richmond, Indiana, 363 pp." type="book" year="1992">Iverson 1992:67</bibRefCitation>
); 
<collectionCode id="ED37AE5BFF9AFFF2DBBDF9DCFDBE25CF" box="[484,555,651,677]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="12" pageNumber="13">NMW</collectionCode>
13247 erroneously listed as “type” by 
<bibRefCitation id="EFB74B6FFF9AFFF2DB8FF9FCFD4125AC" author="Broadley, D. G." box="[470,724,683,710]" pageId="12" pageNumber="13" pagination="633 - 686" refId="ref68429" refString="Broadley, D. G. (1981 b) A review of the genus Pelusios Wagler in southern Africa (Pleurodira: Pelomedusidae). Occasional Papers of the National Museums and Monuments of Rhodesia, B. Natural Sciences, 6 (9), 633 - 686." type="journal article" year="1981">Broadley (1981b:655)</bibRefCitation>
and as 
<typeStatus id="549D883CFF9AFFF2DB54F99CFEE1258F" box="[269,372,715,741]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
by Bour (1983:353) and 
<bibRefCitation id="EFB74B6FFF9AFFF2D8F6F99CFE25246F" author="Bour, R. &amp; Gerlach, J." pageId="12" pageNumber="13" pagination="5" refId="ref67789" refString="Bour, R. &amp; Gerlach, J. (2008) Pelusios seychellensis (Siebenrock 1906) - Seychelles Mud Turtle. In: Rhodin, A. G. J., Pritchard, P. C. H., van Dijk, P. P., Saumure, R. A., Buhlmann, K. A. &amp; Iverson, J. B. (Eds.). Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project of the IUCN / SSC Tortoise and Freshwater Turtle Specialist Group. Chelonian Research Monographs, 5, 18.1 - 18.3. https: // doi. org / 10.3854 / crm. 5.018. seychellensis. v 1.2008" type="book chapter" year="2008">Bour &amp; Gerlach (2008:1)</bibRefCitation>
, who credited Broadley with the 
<typeStatus id="549D883CFF9AFFF2DB49F85CFEE2244F" box="[272,375,779,805]" pageId="12" pageNumber="13" type="lectotype">lectotype</typeStatus>
designation. Wallerman (1998:211) recorded only a single 
<collectionCode id="ED37AE5BFF9AFFF2D85BF87CFDD5242F" box="[514,576,811,837]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="12" pageNumber="13" type="Museum">ZMH</collectionCode>
specimen (993); whereas 
<bibRefCitation id="EFB74B6FFF9AFFF2DB09F81CFDD4240F" author="Grunwaldt, P. H." box="[336,577,843,869]" pageId="12" pageNumber="13" pagination="1 - 19" refId="ref73584" refString="Grunwaldt, P. H. (1980) Liste der Schildkroten der herpetologischen Sammlung des Zoologischen Instituts und Zoologischen Museum der Universitat Hamburg. (Testudines, Reptilia). Bestandslisten der Herpetologischen Sammlung des Zoologischen Instituts und Zoologischen Museums des Universitat Hamburg, (1), 1 - 19." type="journal article" year="1980">Grünwaldt (1980:17)</bibRefCitation>
listed 
<collectionCode id="ED37AE5BFF9AFFF2D8CFF81CFD41240F" box="[662,724,843,869]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="12" pageNumber="13" type="Museum">ZMH</collectionCode>
982 as 
<emphasis id="B952EA8CFF9AFFF2DB5FF83BFE1324EF" box="[262,390,876,901]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="4C264D1DFF9AFFF2DB5FF83BFE1724EF" baseAuthorityName="Smith" baseAuthorityYear="1838" box="[262,386,876,901]" class="Reptilia" family="Pelomedusidae" genus="Pelusios" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="sinuatus">P. sinuatus</taxonomicName>
,
</emphasis>
and 983 and 993 as 
<taxonomicName id="4C264D1DFF9AFFF2D835F83BFD7F24EF" baseAuthorityName="Bonnaterre" baseAuthorityYear="1789" box="[620,746,875,901]" class="Reptilia" family="Pelomedusidae" genus="Pelusios" kingdom="Animalia" order="Testudines" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="subniger">
<emphasis id="B952EA8CFF9AFFF2D835F83BFD7F24EF" box="[620,746,875,901]" italics="true" pageId="12" pageNumber="13">P. subniger</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>