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<document id="A6BA091D60537CD73BC12355FCA6B3F9" ID-CLB-Dataset="21287" ID-DOI="10.1080/00222933.2015.1084057" ID-GBIF-Dataset="fb13a5e8-08b2-4add-9455-79c2fc1b42bc" ID-ISSN="1464-5262" ID-Zenodo-Dep="3990049" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597771790833" checkinUser="carolina" docAuthor="Rossi, Ilana, Amaral, Silvana Vargas, Ribeiro, Giovana Gamino, Cauduro, Guilherme Pinto, Fick, Israel, Valiati, Victor Hugo &amp; Leal-Zanchet, Ana Maria" docDate="2015" docId="03F7B26EFFD2FFB5B1FB1E4FAEB9FAED" docLanguage="en" docName="JNATHIST.50.13-14.787-815.pdf" docOrigin="Journal of Natural History 50" docSource="http://dx.doi.org/10.1080/00222933.2015.1084057" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Geoplaninae Stimpson 1857" docType="treatment" docVersion="8" lastPageNumber="802" masterDocId="FFCECA16FFD7FFBAB06C1C69AD4EFF89" masterDocTitle="Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach" masterLastPageNumber="815" masterPageNumber="787" pageNumber="792" updateTime="1698844695551" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1CCE0CF5BCBDCEBD9F76655319D188F7">Two new Geoplaninae species (Platyhelminthes: Continenticola) from Southern Brazil based on an integrative taxonomic approach</mods:title>
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<mods:namePart id="31D191A5EF9189A10E4CBAB22322F538">Amaral, Silvana Vargas</mods:namePart>
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<mods:namePart id="8F2CACDC0CEBD206F3EBF56D05EABBB3">Fick, Israel</mods:namePart>
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<treatment id="03F7B26EFFD2FFB5B1FB1E4FAEB9FAED" ID-DOI="http://doi.org/10.5281/zenodo.4324237" ID-GBIF-Taxon="166254593" ID-Zenodo-Dep="4324237" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED" httpUri="http://treatment.plazi.org/id/03F7B26EFFD2FFB5B1FB1E4FAEB9FAED" lastPageId="15" lastPageNumber="802" pageId="5" pageNumber="792">
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Subfamily
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<emphasis id="B92ADF6AFFD2FFBFB2791E4FAF82FDB6" bold="true" box="[533,716,550,575]" pageId="5" pageNumber="792">GEOPLANINAE</emphasis>
<bibRefCitation id="EFCF7E89FFD2FFBFB2B91E4FAEC2FDB6" author="Stimpson W" box="[725,908,550,575]" pageId="5" pageNumber="792" pagination="19 - 31" refId="ref13601" refString="Stimpson W. 1857. Prodromus descriptionis animalium evertebratorum, quae in Expeditione ad Oceanum Pacificum Septentrionalem a Republica Federata missa, Johanne Rodgers Duce, observavit et descripsit. Pars I, Turbellaria Dendrocoela. Proc Acad Sci Philadelphia. 9: 19 - 31." type="journal article" year="1857">Stimpson, 1857</bibRefCitation>
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Genus
<taxonomicName id="4C5E78FBFFD2FFBFB27F1E21AE2DFDEB" ID-CoL="7NTR3" authority="Carbayo et al., 2013" authorityName="Carbayo et al. The" authorityYear="2013" box="[531,867,584,610]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFD2FFBFB27F1E21AF21FDEB" bold="true" box="[531,623,584,610]" italics="true" pageId="5" pageNumber="792">Cratera</emphasis>
<bibRefCitation id="EFCF7E89FFD2FFBFB2141E20AE2DFDEB" author="Carbayo F &amp; Alvarez-Presas M &amp; Olivares CT &amp; Marques FPL &amp; Froehlich EM &amp; Riutort M" box="[632,867,585,610]" pageId="5" pageNumber="792" pagination="508 - 528" refId="ref12400" refString="Carbayo F, Alvarez-Presas M, Olivares CT, Marques FPL, Froehlich EM, Riutort M. 2013. Molecular phylogeny of Geoplaninae (Platyhelminthes) challenges current classification: proposal of taxonomic actions. Zool Scr. 42: 508 - 528." type="journal article" year="2013">
Carbayo
<emphasis id="B92ADF6AFFD2FFBFB28E1E20AE58FDEB" box="[738,790,585,610]" italics="true" pageId="5" pageNumber="792">et al</emphasis>
., 2013
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<taxonomicName id="4C5E78FBFFD2FFBFB2661E02AFFAFD0C" ID-CoL="Z8R3" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[522,692,619,645]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD2FFBFB2661E02AFFAFD0C" bold="true" box="[522,692,619,645]" inLineHeading="true" italics="true" pageId="5" pageNumber="792" reason="3">Cratera ochra</emphasis>
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(
<figureCitation id="13651FFDFFD2FFBFB2211EE7AF98FD2E" box="[589,726,654,679]" captionStart-0="Figure 2" captionStart-1="Figure 3" captionStart-2="Figure 4" captionStart-3="Figure 5" captionStart-4="Figure 6" captionStart-5="Figure 7" captionStartId-0="5.[157,222,1748,1771]" captionStartId-1="6.[160,225,1613,1636]" captionStartId-2="7.[160,225,1516,1539]" captionStartId-3="8.[160,225,1407,1430]" captionStartId-4="9.[160,225,1375,1398]" captionStartId-5="10.[160,225,614,637]" captionTargetBox-0="[161,1151,1201,1717]" captionTargetBox-1="[268,1047,142,1582]" captionTargetBox-2="[163,1153,145,1485]" captionTargetBox-3="[163,1153,142,1376]" captionTargetBox-4="[163,1153,152,1344]" captionTargetBox-5="[162,1152,142,583]" captionTargetId-0="figure@5.[161,1151,1201,1717]" captionTargetId-1="figure@6.[268,1048,142,1582]" captionTargetId-2="figure@7.[163,1153,142,1485]" captionTargetId-3="figure@8.[163,1153,142,1376]" captionTargetId-4="figure@9.[163,1153,142,1344]" captionTargetId-5="figure@10.[162,1153,142,583]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="Figure 2. Cratera ochra sp. nov., holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen." captionText-1="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." captionText-2="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." captionText-3="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." captionText-4="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." captionText-5="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi-0="http://doi.org/10.5281/zenodo.3990053" figureDoi-1="http://doi.org/10.5281/zenodo.3990055" figureDoi-2="http://doi.org/10.5281/zenodo.3990057" figureDoi-3="http://doi.org/10.5281/zenodo.3990059" figureDoi-4="http://doi.org/10.5281/zenodo.3990061" figureDoi-5="http://doi.org/10.5281/zenodo.3990063" httpUri-0="https://zenodo.org/record/3990053/files/figure.png" httpUri-1="https://zenodo.org/record/3990055/files/figure.png" httpUri-2="https://zenodo.org/record/3990057/files/figure.png" httpUri-3="https://zenodo.org/record/3990059/files/figure.png" httpUri-4="https://zenodo.org/record/3990061/files/figure.png" httpUri-5="https://zenodo.org/record/3990063/files/figure.png" pageId="5" pageNumber="792">
Figures 2
<emphasis id="B92ADF6AFFD2FFBFB2D61EE7AF86FD2E" box="[698,712,654,679]" italics="true" pageId="5" pageNumber="792">–</emphasis>
7
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)
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<taxonomicName id="4C5E78FBFFD2FFBFB0CC1EAAAC60FD52" authorityName="Stimpson" authorityYear="1857" box="[160,302,707,731]" class="Rhabditophora" family="Geoplanidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="family">Geoplanidae</taxonomicName>
4:
<bibRefCitation id="EFCF7E89FFD2FFBFB13F1EAAAFE9FD52" author="Leal-Zanchet AM &amp; Carbayo F" box="[339,679,707,731]" pageId="5" pageNumber="792" pagination="19 - 25" refId="ref13067" refString="Leal-Zanchet AM, Carbayo F. 2000. Fauna de planarias terrestres (Platyhelminthes, Tricladida, Terricola) da Floresta Nacional de Sao Francisco de Paula, RS, Brasil: Uma analise preliminar. Acta Biol Leopold. 22: 19 - 25." type="journal article" year="2000">Leal-Zanchet &amp; Carbayo, 2000</bibRefCitation>
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<taxonomicName id="4C5E78FBFFD2FFBFB0CC1E8FAC60FD77" authorityName="Stimpson" authorityYear="1857" box="[160,302,742,766]" class="Rhabditophora" family="Geoplanidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="family">Geoplanidae</taxonomicName>
2:
<bibRefCitation id="EFCF7E89FFD2FFBFB13F1E8FAF7BFD77" author="Carbayo F &amp; Leal-Zanchet AM &amp; Vieira EM" box="[339,565,742,766]" pageId="5" pageNumber="792" pagination="1091 - 1104" refId="ref12473" refString="Carbayo F, Leal-Zanchet AM, Vieira EM. 2002. Terrestrial flatworm (Platyhelminthes: Tricladida: Terricola) diversity versus man-induced disturbance in ombrophilous forest in Southern Brazil. Biodivers Conserv. 11: 1091 - 1104." type="journal article" year="2002">Carbayo et al., 2002</bibRefCitation>
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<taxonomicName id="4C5E78FBFFD2FFBFB0CC1F60AC47FCA8" authorityName="Stimpson" authorityYear="1857" box="[160,265,777,801]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFD2FFBFB0CC1F60AC47FCA8" box="[160,265,777,801]" italics="true" pageId="5" pageNumber="792">Geoplana</emphasis>
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sp. 4:
<bibRefCitation id="EFCF7E89FFD2FFBFB13A1F60AFE9FCA8" author="Leal-Zanchet AM &amp; Baptista VA" box="[342,679,777,801]" pageId="5" pageNumber="792" pagination="199 - 207" refId="ref12984" refString="Leal-Zanchet AM, Baptista VA. 2009. Floresta com Araucaria: Ecologia, Conservacao e Desenvolvimento Sustentavel. Ribeirao Preto: Holos. Chapter 19, Planarias terrestres (Platyhelminthes, Tricladida) em remanescentes de Floresta com Araucaria; p. 199 - 207." type="book chapter" year="2009">Leal-Zanchet &amp; Baptista, 2009</bibRefCitation>
</paragraph>
<paragraph id="8BE10378FFD2FFBFB0CC1F42AF25FCCA" blockId="5.[160,679,707,835]" box="[160,619,811,835]" pageId="5" pageNumber="792">
<taxonomicName id="4C5E78FBFFD2FFBFB0CC1F42AC47FCCA" authorityName="Stimpson" authorityYear="1857" box="[160,265,811,835]" class="Rhabditophora" family="Geoplanidae" genus="Geoplana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFD2FFBFB0CC1F42AC47FCCA" box="[160,265,811,835]" italics="true" pageId="5" pageNumber="792">Geoplana</emphasis>
</taxonomicName>
sp. 4:
<bibRefCitation id="EFCF7E89FFD2FFBFB13A1F42AF25FCCA" author="Leal-Zanchet AM &amp; Baptista VA &amp; Campos LM &amp; Raffo JF" box="[342,619,811,835]" pageId="5" pageNumber="792" pagination="25 - 33" refId="ref13031" refString="Leal-Zanchet AM, Baptista VA, Campos LM, Raffo JF. 2011. Spatial and temporal patterns of land flatworm assemblages in Brazilian Araucaria forests. Inv Biol. 130: 25 - 33." type="journal article" year="2011">
Leal-Zanchet
<emphasis id="B92ADF6AFFD2FFBFB1831F42AF6EFCCA" box="[495,544,811,835]" italics="true" pageId="5" pageNumber="792">et al</emphasis>
., 2011
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34450F3FFD2FFB3B0CC1F01AEA9F914" lastPageId="9" lastPageNumber="796" pageId="5" pageNumber="792" type="etymology">
<paragraph id="8BE10378FFD2FFBFB0CC1F01AC53FC08" blockId="5.[160,1021,872,932]" box="[160,285,872,897]" pageId="5" pageNumber="792">
<heading id="D0A9B414FFD2FFBFB0CC1F01AC53FC08" box="[160,285,872,897]" fontSize="11" level="3" pageId="5" pageNumber="792" reason="5">
<emphasis id="B92ADF6AFFD2FFBFB0CC1F01AC53FC08" bold="true" box="[160,285,872,897]" inLineHeading="true" italics="true" pageId="5" pageNumber="792" reason="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFD2FFBFB0CC1FE5AEB3FC2D" blockId="5.[160,1021,872,932]" box="[160,1021,907,932]" pageId="5" pageNumber="792">The specific name refers to the dorsal pigmentation, which is yellow-ochre.</paragraph>
<paragraph id="8BE10378FFD2FFBFB0CC1FA2AC0AFC6D" blockId="5.[160,1155,971,1169]" box="[160,324,971,996]" pageId="5" pageNumber="792">
<heading id="D0A9B414FFD2FFBFB0CC1FA2AC0AFC6D" box="[160,324,971,996]" fontSize="11" level="3" pageId="5" pageNumber="792" reason="5">
<emphasis id="B92ADF6AFFD2FFBFB0CC1FA2AC0AFC6D" bold="true" box="[160,324,971,996]" inLineHeading="true" italics="true" pageId="5" pageNumber="792" reason="3">Type-material</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFD2FFBFB0CC1F87AE77FB18" blockId="5.[160,1155,971,1169]" pageId="5" pageNumber="792">
<materialsCitation id="3B360925FFD2FFBFB0CC1F87AE77FB18" ID-GBIF-Occurrence="2831860315" collectingDate="2013-08-01" collectionCode="MZUSP, PL" collectorName="P. K. Boll" country="Brazil" location="Sao Francisco de Paula" municipality="National Forest of Sao Francisco de Paula" pageId="5" pageNumber="792" specimenCount="1" typeStatus="holotype">
<emphasis id="B92ADF6AFFD2FFBFB0CC1F87AC5FFB8F" bold="true" box="[160,273,1006,1031]" inLineHeading="true" italics="true" pageId="5" pageNumber="792" reason="3">
<typeStatus id="54E5BDDAFFD2FFBFB0CC1F87AC45FB8E" box="[160,267,1006,1031]" pageId="5" pageNumber="792" type="holotype">Holotype</typeStatus>
.
</emphasis>
<emphasis id="B92ADF6AFFD2FFBFB14D1F87ACBEFB8F" bold="true" box="[289,496,1006,1030]" pageId="5" pageNumber="792">
<collectionCode id="ED4F9BBDFFD2FFBFB14D1F87AC39FB8F" box="[289,375,1006,1030]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="5" pageNumber="792">MZUSP</collectionCode>
<collectionCode id="ED4F9BBDFFD2FFBFB1E81F87ACEAFB8F" box="[388,420,1006,1030]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="5" pageNumber="792" type="Museum">PL</collectionCode>
. 1564
</emphasis>
:
<emphasis id="B92ADF6AFFD2FFBFB26E1F87AF6CFB8F" box="[514,546,1006,1030]" italics="true" pageId="5" pageNumber="792">leg</emphasis>
.
<collectorName id="26AB66AEFFD2FFBFB2591F87AFECFB8F" box="[565,674,1006,1030]" pageId="5" pageNumber="792">P. K. Boll</collectorName>
,
<date id="FFE025B8FFD2FFBFB2DE1F87AE2CFB8F" box="[690,866,1006,1030]" pageId="5" pageNumber="792" value="2013-08-01">
<collectingDate id="EFA4DC50FFD2FFBFB2DE1F87AE2CFB8F" box="[690,866,1006,1030]" pageId="5" pageNumber="792" value="2013-08-01">1 August 2013</collectingDate>
</date>
,
<location id="8E8155A3FFD2FFBFB31E1F87A9CDFB8F" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFD2FFBFB31E1F87A9CDFB8F" box="[882,1155,1006,1030]" country="Brazil" municipality="National Forest of Sao Francisco de Paula" name="Sao Francisco de Paula" pageId="5" pageNumber="792">São Francisco de Paula</location>
(
<collectingMunicipality id="6B859902FFD2FFBFB0CA1878AFC9FBA0" box="[166,647,1041,1065]" pageId="5" pageNumber="792">National Forest of São Francisco de Paula</collectingMunicipality>
), RS,
<collectingCountry id="F34943E8FFD2FFBFB2AF1878AFB1FBA0" box="[707,767,1041,1065]" name="Brazil" pageId="5" pageNumber="792">Brazil</collectingCountry>
<emphasis id="B92ADF6AFFD2FFBFB3641878AE5BFBA0" box="[776,789,1041,1065]" italics="true" pageId="5" pageNumber="792">–</emphasis>
anterior tip: transverse sections on 21 slides; anterior region at the level of the ovaries: sagittal sections on 52 slides; prepharyngeal region: transverse sections on nine slides; pharynx: sagittal sections on 34 slides; copulatory apparatus: sagittal sections on 26 slides.
</materialsCitation>
</paragraph>
<caption id="DF2153F0FFD2FFBFB0F11ABDAFBEF881" ID-DOI="http://doi.org/10.5281/zenodo.3990053" ID-Zenodo-Dep="3990053" httpUri="https://zenodo.org/record/3990053/files/figure.png" pageId="5" pageNumber="792" startId="5.[157,222,1748,1771]" targetBox="[161,1151,1201,1717]" targetPageId="5">
<paragraph id="8BE10378FFD2FFBFB0F11ABDAFBEF881" blockId="5.[157,1155,1747,1800]" pageId="5" pageNumber="792">
<emphasis id="B92ADF6AFFD2FFBFB0F11ABDADB7F962" bold="true" box="[157,249,1748,1771]" pageId="5" pageNumber="792">Figure 2.</emphasis>
<taxonomicName id="4C5E78FBFFD2FFBFB16E1ABAACC9F963" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[258,391,1747,1770]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="5" pageNumber="792" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD2FFBFB16E1ABAACC9F963" box="[258,391,1747,1770]" italics="true" pageId="5" pageNumber="792">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD2FFBFB1E31ABDACAEF962" box="[399,480,1748,1771]" pageId="5" pageNumber="792" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen.
</paragraph>
</caption>
<caption id="DF2153F0FFD1FFBCB0CC1A24A96FF916" ID-DOI="http://doi.org/10.5281/zenodo.3990055" ID-Zenodo-Dep="3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="6" pageNumber="793" startId="6.[160,225,1613,1636]" targetBox="[268,1047,142,1582]" targetPageId="6">
<paragraph id="8BE10378FFD1FFBCB0CC1A24A96FF916" blockId="6.[160,1157,1613,1695]" pageId="6" pageNumber="793">
<emphasis id="B92ADF6AFFD1FFBCB0CC1A24ADB5F9ED" bold="true" box="[160,251,1613,1636]" pageId="6" pageNumber="793">Figure 3.</emphasis>
<taxonomicName id="4C5E78FBFFD1FFBCB1681A24ACC6F9ED" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[260,392,1613,1636]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="6" pageNumber="793" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD1FFBCB1681A24ACC6F9ED" box="[260,392,1613,1636]" italics="true" pageId="6" pageNumber="793">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD1FFBCB1FC1A24AC91F9ED" box="[400,479,1613,1636]" pageId="6" pageNumber="793" rank="species">sp. nov.</taxonomicNameLabel>
in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187).
</paragraph>
</caption>
<caption id="DF2153F0FFD0FFBDB0CC1985A97AF9D2" ID-DOI="http://doi.org/10.5281/zenodo.3990057" ID-Zenodo-Dep="3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="7" pageNumber="794" startId="7.[160,225,1516,1539]" targetBox="[163,1153,145,1485]" targetPageId="7">
<paragraph id="8BE10378FFD0FFBDB0CC1985A97AF9D2" blockId="7.[160,1156,1516,1627]" pageId="7" pageNumber="794">
<emphasis id="B92ADF6AFFD0FFBDB0CC1985ADB5F98A" bold="true" box="[160,251,1516,1539]" pageId="7" pageNumber="794">Figure 4.</emphasis>
<taxonomicName id="4C5E78FBFFD0FFBDB16F1985ACC8F98A" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[259,390,1516,1539]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="7" pageNumber="794" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD0FFBDB16F1985ACC8F98A" box="[259,390,1516,1539]" italics="true" pageId="7" pageNumber="794">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD0FFBDB1E21985AC93F98A" box="[398,477,1516,1539]" pageId="7" pageNumber="794" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary.
</paragraph>
</caption>
<caption id="DF2153F0FFDFFFB2B0CC1916ACD8FA58" ID-DOI="http://doi.org/10.5281/zenodo.3990059" ID-Zenodo-Dep="3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="8" pageNumber="795" startId="8.[160,225,1407,1430]" targetBox="[163,1153,142,1376]" targetPageId="8">
<paragraph id="8BE10378FFDFFFB2B0CC1916ACD8FA58" blockId="8.[160,1156,1407,1489]" pageId="8" pageNumber="795">
<emphasis id="B92ADF6AFFDFFFB2B0CC1916ADB4FA1F" bold="true" box="[160,250,1407,1430]" pageId="8" pageNumber="795">Figure 5.</emphasis>
<taxonomicName id="4C5E78FBFFDFFFB2B16F1916ACC8FA1F" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[259,390,1407,1430]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="8" pageNumber="795" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFDFFFB2B16F1916ACC8FA1F" box="[259,390,1407,1430]" italics="true" pageId="8" pageNumber="795">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFDFFFB2B1E11916AC94FA1F" box="[397,474,1407,1430]" pageId="8" pageNumber="795" rank="species">sp. nov.</taxonomicNameLabel>
(A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189.
</paragraph>
</caption>
<paragraph id="8BE10378FFDFFFB3B0CC1992AEA9F914" blockId="8.[160,1156,1531,1695]" lastBlockId="9.[160,1155,1495,1693]" lastPageId="9" lastPageNumber="796" pageId="8" pageNumber="795">
<materialsCitation id="3B360925FFDFFFB2B0CC1992AE6DF99D" ID-GBIF-Occurrence="2831860306" box="[160,803,1531,1556]" collectionCode="PL" country="Brazil" location="Sao Francisco de Paula" pageId="8" pageNumber="795" specimenCount="1" typeStatus="paratype">
<emphasis id="B92ADF6AFFDFFFB2B0CC1992AC55F99D" bold="true" box="[160,283,1531,1556]" inLineHeading="true" italics="true" pageId="8" pageNumber="795" reason="3">
<typeStatus id="54E5BDDAFFDFFFB2B0CC1992AC5BF99D" box="[160,277,1531,1556]" pageId="8" pageNumber="795" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<location id="8E8155A3FFDFFFB2B1401995AF7DF99D" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFDFFFB2B1401995AF7DF99D" box="[300,563,1532,1556]" country="Brazil" name="Sao Francisco de Paula" pageId="8" pageNumber="795">São Francisco de Paula</location>
, RS,
<collectingCountry id="F34943E8FFDFFFB2B2041995AFEAF99D" box="[616,676,1532,1556]" name="Brazil" pageId="8" pageNumber="795">Brazil</collectingCountry>
<emphasis id="B92ADF6AFFDFFFB2B2C11995AFF4F99D" box="[685,698,1532,1556]" italics="true" pageId="8" pageNumber="795">–</emphasis>
<emphasis id="B92ADF6AFFDFFFB2B2AE1995AE6DF99D" bold="true" box="[706,803,1532,1556]" pageId="8" pageNumber="795">
MZU
<collectionCode id="ED4F9BBDFFDFFFB2B36F1995AE6DF99D" box="[771,803,1532,1556]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="795" type="Museum">PL</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B92ADF6AFFDFFFB2B34F1995AE3FF99D" bold="true" box="[803,881,1532,1556]" pageId="8" pageNumber="795">.00187</emphasis>
:
<emphasis id="B92ADF6AFFDFFFB2B3131995AED1F99D" box="[895,927,1532,1556]" italics="true" pageId="8" pageNumber="795">leg</emphasis>
. R.
<materialsCitation id="3B360925FFDFFFB2B3A61995ACBFF9D0" ID-GBIF-Occurrence="2831860313" collectingDate="1998-05-04" collectionCode="PL" country="Brazil" location="Murowaniecki" municipality="National Forest of Sao Francisco de Paula" pageId="8" pageNumber="795" specimenCount="1" typeStatus="paratype">
<location id="8E8155A3FFDFFFB2B3A61995A924F99D" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFDFFFB2B3A61995A924F99D" box="[970,1130,1532,1556]" country="Brazil" municipality="National Forest of Sao Francisco de Paula" name="Murowaniecki" pageId="8" pageNumber="795">Murowaniecki</location>
,
<date id="FFE025B8FFDFFFB2B4191995AC5DF9BE" pageId="8" pageNumber="795" value="1998-05-04">
<collectingDate id="EFA4DC50FFDFFFB2B4191995AC5DF9BE" pageId="8" pageNumber="795" value="1998-05-04">4 May 1998</collectingDate>
</date>
,
<collectingMunicipality id="6B859902FFDFFFB2B14E1A76AE44F9BE" box="[290,778,1567,1591]" pageId="8" pageNumber="795">National Forest of São Francisco de Paula</collectingMunicipality>
<emphasis id="B92ADF6AFFDFFFB2B3791A76AE6CF9BE" box="[789,802,1567,1591]" italics="true" pageId="8" pageNumber="795">–</emphasis>
copulatory apparatus: sagittal sections on 26 slides;
<emphasis id="B92ADF6AFFDFFFB2B1FE1A28ACBFF9D0" bold="true" box="[402,497,1601,1625]" pageId="8" pageNumber="795">
MZU
<collectionCode id="ED4F9BBDFFDFFFB2B1BD1A28ACBFF9D0" box="[465,497,1601,1625]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="8" pageNumber="795" type="Museum">PL</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B92ADF6AFFDFFFB2B19D1A28AF71F9D0" bold="true" box="[497,575,1601,1625]" pageId="8" pageNumber="795">.00188</emphasis>
:
<emphasis id="B92ADF6AFFDFFFB2B2201A28AF22F9D0" box="[588,620,1601,1625]" italics="true" pageId="8" pageNumber="795">leg</emphasis>
<materialsCitation id="3B360925FFDFFFB2B2001A28AFC9F9D0" ID-GBIF-Occurrence="2831860321" box="[620,647,1601,1625]" collectingDate="1998-05-04" collectionCode="PL" country="Brazil" isTailEnumeration="true" location="Murowaniecki" municipality="National Forest of Sao Francisco de Paula" pageId="8" pageNumber="795" specimenCount="1" typeStatus="paratype">. F</materialsCitation>
<materialsCitation id="3B360925FFDFFFB3B2EB1A28AC83F99B" ID-GBIF-Occurrence="2831860307" collectingDate="1998-06-08" collectionCode="PL" country="Brazil" lastPageId="9" lastPageNumber="796" location="Carbayo" municipality="National Forest of Sao Francisco de Paula" pageId="8" pageNumber="795" specimenCount="1" typeStatus="paratype">
.
<location id="8E8155A3FFDFFFB2B2FD1A28AFBEF9D0" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFDFFFB2B2FD1A28AFBEF9D0" box="[657,752,1601,1625]" country="Brazil" municipality="National Forest of Sao Francisco de Paula" name="Carbayo" pageId="8" pageNumber="795">Carbayo</location>
,
<date id="FFE025B8FFDFFFB2B2961A28AECDF9D0" box="[762,899,1601,1625]" pageId="8" pageNumber="795" value="1998-06-08">
<collectingDate id="EFA4DC50FFDFFFB2B2961A28AECDF9D0" box="[762,899,1601,1625]" pageId="8" pageNumber="795" value="1998-06-08">8 June 1998</collectingDate>
</date>
,
<collectingMunicipality id="6B859902FFDFFFB2B3E11A28AC3DF9F5" pageId="8" pageNumber="795">National Forest of São Francisco de Paula</collectingMunicipality>
<emphasis id="B92ADF6AFFDFFFB2B1101A0DACC7F9F5" box="[380,393,1636,1660]" italics="true" pageId="8" pageNumber="795">–</emphasis>
anterior tip: transverse sections on 22 slides; region at the level of the ovaries: sagittal sections on 67 slides; pre-pharyngeal region: transverse sections on eight slides; pharynx: sagittal sections on 24 slides; copulatory apparatus: sagittal sections on 22 slides;
<emphasis id="B92ADF6AFFDEFFB3B1021993AC83F99B" bold="true" box="[366,461,1530,1554]" pageId="9" pageNumber="796">
MZU
<collectionCode id="ED4F9BBDFFDEFFB3B1C11993AC83F99B" box="[429,461,1530,1554]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="9" pageNumber="796" type="Museum">PL</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B92ADF6AFFDEFFB3B1A11993AF55F99B" bold="true" box="[461,539,1530,1554]" pageId="9" pageNumber="796">.00189</emphasis>
:
<emphasis id="B92ADF6AFFDEFFB3B24B1993AF09F99B" box="[551,583,1530,1554]" italics="true" pageId="9" pageNumber="796">leg</emphasis>
<materialsCitation id="3B360925FFDEFFB3B22B1993AF2DF99B" ID-GBIF-Occurrence="2831860316" box="[583,611,1530,1554]" collectingDate="1998-06-08" collectionCode="PL" country="Brazil" isTailEnumeration="true" location="Carbayo" municipality="National Forest of Sao Francisco de Paula" pageId="9" pageNumber="796" specimenCount="1" typeStatus="paratype">. F</materialsCitation>
<materialsCitation id="3B360925FFDEFFB3B20F1993AD8EF9DE" ID-GBIF-Occurrence="2831860305" collectingDate="1998-09-25" collectionCode="PL" country="Brazil" location="Carbayo" municipality="National Forest of Sao Francisco de Paula" pageId="9" pageNumber="796" specimenCount="1" typeStatus="paratype">
.
<location id="8E8155A3FFDEFFB3B2011993AF82F99B" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFDEFFB3B2011993AF82F99B" box="[621,716,1530,1554]" country="Brazil" municipality="National Forest of Sao Francisco de Paula" name="Carbayo" pageId="9" pageNumber="796">Carbayo</location>
,
<date id="FFE025B8FFDEFFB3B2B91993AEFCF99B" box="[725,946,1530,1554]" pageId="9" pageNumber="796" value="1998-09-25">
<collectingDate id="EFA4DC50FFDEFFB3B2B91993AEFCF99B" box="[725,946,1530,1554]" pageId="9" pageNumber="796" value="1998-09-25">25 September 1998</collectingDate>
</date>
,
<collectingMunicipality id="6B859902FFDEFFB3B3D01993ACE4F9BC" pageId="9" pageNumber="796">National Forest of São Francisco de Paula</collectingMunicipality>
<emphasis id="B92ADF6AFFDEFFB3B1D81A75AC8FF9BD" box="[436,449,1564,1588]" italics="true" pageId="9" pageNumber="796">–</emphasis>
copulatory apparatus: horizontal sections on 22 slides;
<emphasis id="B92ADF6AFFDEFFB3B4271A74AD8EF9DE" bold="true" pageId="9" pageNumber="796">
MZU
<collectionCode id="ED4F9BBDFFDEFFB3B0CC1A56AD8EF9DE" box="[160,192,1599,1623]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="9" pageNumber="796" type="Museum">PL</collectionCode>
</emphasis>
</materialsCitation>
<emphasis id="B92ADF6AFFDEFFB3B0AC1A56AC40F9DE" bold="true" box="[192,270,1599,1623]" pageId="9" pageNumber="796">.00190</emphasis>
:
<emphasis id="B92ADF6AFFDEFFB3B14C1A56AC0EF9DE" box="[288,320,1599,1623]" italics="true" pageId="9" pageNumber="796">leg</emphasis>
<materialsCitation id="3B360925FFDEFFB3B12D1A56AC2BF9DE" ID-GBIF-Occurrence="2831860314" box="[321,357,1599,1623]" collectingDate="1998-09-25" collectionCode="PL" country="Brazil" isTailEnumeration="true" location="Carbayo" municipality="National Forest of Sao Francisco de Paula" pageId="9" pageNumber="796" specimenCount="1" typeStatus="paratype">. C</materialsCitation>
<materialsCitation id="3B360925FFDEFFB3B1091A56AEADF914" ID-GBIF-Occurrence="2831860308" collectingDate="2004-10-17" country="Brazil" location="Mata" pageId="9" pageNumber="796" specimenCount="1" typeStatus="paratype">
. Palácios
<date id="FFE025B8FFDEFFB3B1B71A56AFEBF9DE" box="[475,677,1599,1623]" pageId="9" pageNumber="796" value="2004-10-17">
<collectingDate id="EFA4DC50FFDEFFB3B1B71A56AFEBF9DE" box="[475,677,1599,1623]" pageId="9" pageNumber="796" value="2004-10-17">17 October 2004</collectingDate>
</date>
, Research and Conservation Center Pró-
<location id="8E8155A3FFDEFFB3B0CC1A0BAD96F9F3" LSID="urn:lsid:plazi:treatment:03F7B26EFFD2FFB5B1FB1E4FAEB9FAED:8E8155A3FFDEFFB3B0CC1A0BAD96F9F3" box="[160,216,1634,1658]" country="Brazil" name="Mata" pageId="9" pageNumber="796">Mata</location>
<emphasis id="B92ADF6AFFDEFFB3B0891A0BADBCF9F3" box="[229,242,1634,1658]" italics="true" pageId="9" pageNumber="796">–</emphasis>
pre-pharyngeal region: transverse sections on seven slides; pharynx: sagittal sections on 15 slides; copulatory apparatus: sagittal sections on 12 slides
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2153F0FFDEFFB3B0CC1936AE2AFA1D" ID-DOI="http://doi.org/10.5281/zenodo.3990061" ID-Zenodo-Dep="3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="9" pageNumber="796" startId="9.[160,225,1375,1398]" targetBox="[163,1153,152,1344]" targetPageId="9">
<paragraph id="8BE10378FFDEFFB3B0CC1936AE2AFA1D" blockId="9.[160,1155,1375,1428]" pageId="9" pageNumber="796">
<emphasis id="B92ADF6AFFDEFFB3B0CC1936ADB0FAFF" bold="true" box="[160,254,1375,1398]" pageId="9" pageNumber="796">Figure 6.</emphasis>
<taxonomicName id="4C5E78FBFFDEFFB3B16A1936ACC3FAFF" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[262,397,1375,1398]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="9" pageNumber="796" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFDEFFB3B16A1936ACC3FAFF" box="[262,397,1375,1398]" italics="true" pageId="9" pageNumber="796">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFDEFFB3B1FB1936ACA6FAFF" box="[407,488,1375,1398]" pageId="9" pageNumber="796" rank="species">sp. nov.</taxonomicNameLabel>
(A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section.
</paragraph>
</caption>
<caption id="DF2153F0FFDDFFB0B0CC1E0FAEF0FD13" ID-DOI="http://doi.org/10.5281/zenodo.3990063" ID-Zenodo-Dep="3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="10" pageNumber="797" startId="10.[160,225,614,637]" targetBox="[162,1152,142,583]" targetPageId="10">
<paragraph id="8BE10378FFDDFFB0B0CC1E0FAEF0FD13" blockId="10.[160,1155,613,666]" pageId="10" pageNumber="797">
<emphasis id="B92ADF6AFFDDFFB0B0CC1E0FADB4FDF4" bold="true" box="[160,250,614,637]" pageId="10" pageNumber="797">Figure 7.</emphasis>
<taxonomicName id="4C5E78FBFFDDFFB0B16E1E0CACCBFDF5" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[258,389,613,636]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="10" pageNumber="797" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFDDFFB0B16E1E0CACCBFDF5" box="[258,389,613,636]" italics="true" pageId="10" pageNumber="797">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFDDFFB0B1E71E0FAC96FDF4" box="[395,472,614,637]" pageId="10" pageNumber="797" rank="species">sp. nov.</taxonomicNameLabel>
(A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section.
</paragraph>
</caption>
<subSubSection id="C34450F3FFDDFFB0B0CC1EAEAC23FB59" pageId="10" pageNumber="797" type="diagnosis">
<paragraph id="8BE10378FFDDFFB0B0CC1EAEAC5AFD69" blockId="10.[160,1155,711,944]" box="[160,276,711,736]" pageId="10" pageNumber="797">
<heading id="D0A9B414FFDDFFB0B0CC1EAEAC5AFD69" box="[160,276,711,736]" fontSize="11" level="3" pageId="10" pageNumber="797" reason="5">
<emphasis id="B92ADF6AFFDDFFB0B0CC1EAEAC5AFD69" bold="true" box="[160,276,711,736]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC1E82AF3DFC39" blockId="10.[160,1155,711,944]" pageId="10" pageNumber="797">
Specimens of
<taxonomicName id="4C5E78FBFFDDFFB0B12E1E82ACDCFC8A" authorityName="Carbayo et al. The" authorityYear="2013" box="[322,402,747,771]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="10" pageNumber="797" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFDDFFB0B12E1E82ACDCFC8A" box="[322,402,747,771]" italics="true" pageId="10" pageNumber="797">Cratera</emphasis>
</taxonomicName>
with yellow-ochre dorsal colour mainly with dispersed greyish or greyish-brown pigmentation constituting two broad dorsal bands; eyes dorsal; glandular margin with at least four
<typeStatus id="54E5BDDAFFDDFFB0B1A81F59AF4EFCC1" box="[452,512,816,840]" pageId="10" pageNumber="797">types</typeStatus>
of glands; mc: h, 11
<emphasis id="B92ADF6AFFDDFFB0B29E1F59AFB1FCC1" box="[754,767,816,840]" italics="true" pageId="10" pageNumber="797">–</emphasis>
12%; pharynx cylindrical; prostatic vesicle extrabulbar with proximal portion ample and laterally expanded and distal portion tubular; ovovitelline ducts emerging dorsally from the median third of ovaries and ascending anterior to the gonopore.
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC1F8BACF0FBC9" blockId="10.[160,1155,994,1088]" pageId="10" pageNumber="797">
<emphasis id="B92ADF6AFFDDFFB0B0CC1F8BACDBFC73" bold="true" box="[160,405,994,1019]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">Molecular diagnosis.</emphasis>
This species includes all populations that cluster with specimens
<accessionNumber id="940D9E9BFFDDFFB0B0CC186CAC43FB94" box="[160,269,1029,1053]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT250622" pageId="10" pageNumber="797">
KT
<date id="FFE025B8FFDDFFB0B0D1186CAC43FB94" box="[189,269,1029,1053]" pageId="10" pageNumber="797" value="22-06-25">250622</date>
</accessionNumber>
to
<accessionNumber id="940D9E9BFFDDFFB0B15A186CACEAFB94" box="[310,420,1029,1053]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KT250624" pageId="10" pageNumber="797">
KT
<date id="FFE025B8FFDDFFB0B138186CACEAFB94" box="[340,420,1029,1053]" pageId="10" pageNumber="797" value="24-06-25">250624</date>
</accessionNumber>
(see Supplementary material,
<tableCitation id="C6DC36C3FFDDFFB0B36C186CAE2CFB94" box="[768,866,1029,1053]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="10" pageNumber="797">Table S1</tableCitation>
), with significant support in phylogenetic analyses.
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC1818AC79FB03" blockId="10.[160,1155,1137,1232]" box="[160,311,1137,1162]" pageId="10" pageNumber="797">
<heading id="D0A9B414FFDDFFB0B0CC1818AC79FB03" box="[160,311,1137,1162]" fontSize="11" level="3" pageId="10" pageNumber="797" reason="5">
<emphasis id="B92ADF6AFFDDFFB0B0CC1818AC79FB03" bold="true" box="[160,311,1137,1162]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">Type-locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC18FCAC23FB59" blockId="10.[160,1155,1137,1232]" pageId="10" pageNumber="797">
São Francisco de Paula (National Forest of São Francisco de Paula), state of
<collectingRegion id="499ACD9AFFDDFFB0B46918FCADA7FB59" country="Brazil" name="Rio Grande do Sul" pageId="10" pageNumber="797">Rio Grande do Sul</collectingRegion>
(RS),
<collectingCountry id="F34943E8FFDDFFB0B14618D1AC27FB59" box="[298,361,1208,1232]" name="Brazil" pageId="10" pageNumber="797">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34450F3FFDDFFB0B0CC1968AF93FAB4" box="[160,733,1281,1341]" pageId="10" pageNumber="797" type="distribution">
<paragraph id="8BE10378FFDDFFB0B0CC1968AC62FA93" blockId="10.[160,733,1281,1341]" box="[160,300,1281,1306]" pageId="10" pageNumber="797">
<heading id="D0A9B414FFDDFFB0B0CC1968AC62FA93" box="[160,300,1281,1306]" fontSize="11" level="3" pageId="10" pageNumber="797" reason="5">
<emphasis id="B92ADF6AFFDDFFB0B0CC1968AC62FA93" bold="true" box="[160,300,1281,1306]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC194CAF93FAB4" blockId="10.[160,733,1281,1341]" box="[160,733,1317,1341]" pageId="10" pageNumber="797">
<collectingRegion id="499ACD9AFFDDFFB0B0CC194CAC21FAB4" box="[160,367,1317,1341]" country="Brazil" name="Rio Grande do Sul" pageId="10" pageNumber="797">Rio Grande do Sul</collectingRegion>
(São Francisco de Paula),
<collectingCountry id="F34943E8FFDDFFB0B2F7194CAF94FAB4" box="[667,730,1317,1341]" name="Brazil" pageId="10" pageNumber="797">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34450F3FFDDFFB5B0CC1907AEB9FAED" lastPageId="15" lastPageNumber="802" pageId="10" pageNumber="797" type="description">
<paragraph id="8BE10378FFDDFFB0B0CC1907AC68FA0E" blockId="10.[160,1156,1390,1727]" box="[160,294,1390,1415]" pageId="10" pageNumber="797">
<heading id="D0A9B414FFDDFFB0B0CC1907AC68FA0E" box="[160,294,1390,1415]" fontSize="11" level="3" pageId="10" pageNumber="797" reason="5">
<emphasis id="B92ADF6AFFDDFFB0B0CC1907AC68FA0E" bold="true" box="[160,294,1390,1415]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFDDFFB0B0CC19F8AE2DF99B" blockId="10.[160,1156,1390,1727]" pageId="10" pageNumber="797">
<emphasis id="B92ADF6AFFDDFFB0B0CC19F8AC21FA23" bold="true" box="[160,367,1425,1450]" inLineHeading="true" italics="true" pageId="10" pageNumber="797" reason="3">External features.</emphasis>
Body broad with parallel margins and dorsal surface convex; anterior tip rounded and posterior tip pointed (
<figureCitation id="13651FFDFFDDFFB0B21F19DCAFADFA44" box="[627,739,1461,1485]" captionStart="Figure 2" captionStartId="5.[157,222,1748,1771]" captionTargetBox="[161,1151,1201,1717]" captionTargetId="figure@5.[161,1151,1201,1717]" captionTargetPageId="5" captionText="Figure 2. Cratera ochra sp. nov., holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.3990053" httpUri="https://zenodo.org/record/3990053/files/figure.png" pageId="10" pageNumber="797">Figure 2A</figureCitation>
). When creeping, maximum length reaches
<quantity id="4CA6AE9DFFDDFFB0B16F19BEAC15FA66" box="[259,347,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="797" unit="mm" value="85.0">85 mm</quantity>
. After fixation, maximum length was
<quantity id="4CA6AE9DFFDDFFB0B34819BEAE37FA66" box="[804,889,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="10" pageNumber="797" unit="mm" value="52.0">52 mm</quantity>
(
<tableCitation id="C6DC36C3FFDDFFB0B3E219BEAEA8FA66" box="[910,998,1495,1519]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="10" pageNumber="797">Table 1</tableCitation>
). Mouth and gonopore located at the posterior third of the body (
<tableCitation id="C6DC36C3FFDDFFB0B36F1993AE18F99B" box="[771,854,1530,1554]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="10" pageNumber="797">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BE10378FFDDFFB1B0D21A74AE5EFBFB" blockId="10.[160,1156,1390,1727]" lastBlockId="11.[160,1156,837,1346]" lastPageId="11" lastPageNumber="798" pageId="10" pageNumber="797">
In living animals, dorsal surface with dispersed greyish or greyish-brown pigmentation constituting two broad dorsal bands; the dorsal yellow-ochre ground colour is mainly visible between the dorsal bands and on the body margins. The ground colour is darker on the body margins than on the rest of the dorsal surface. The dorsal pigmentation concentrates at the anterior extremity (
<figureCitation id="13651FFDFFDDFFB0B3331ACEAE92F936" box="[863,988,1703,1727]" captionStart="Figure 2" captionStartId="5.[157,222,1748,1771]" captionTargetBox="[161,1151,1201,1717]" captionTargetId="figure@5.[161,1151,1201,1717]" captionTargetPageId="5" captionText="Figure 2. Cratera ochra sp. nov., holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.3990053" httpUri="https://zenodo.org/record/3990053/files/figure.png" pageId="10" pageNumber="797">Figure 2A</figureCitation>
). Under the stereomicroscope, a greyish pigmentation forms two densely pigmented dorsal bands, as well as contours the anterior tip (
<figureCitation id="13651FFDFFDCFFB1B2201F0EAF8AFCF6" box="[588,708,871,895]" captionStart="Figure 2" captionStartId="5.[157,222,1748,1771]" captionTargetBox="[161,1151,1201,1717]" captionTargetId="figure@5.[161,1151,1201,1717]" captionTargetPageId="5" captionText="Figure 2. Cratera ochra sp. nov., holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.3990053" httpUri="https://zenodo.org/record/3990053/files/figure.png" pageId="11" pageNumber="798">Figures 2B</figureCitation>
,
<figureCitation id="13651FFDFFDCFFB1B2B81F0EAFBCFCF6" box="[724,754,871,895]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">3A</figureCitation>
). Posterior to the anterior tip, this pigmentation becomes more irregular and loosely arranged in flecks over the dorsum, making the yellow-ochre ground colour visible (
<figureCitation id="13651FFDFFDCFFB1B2A31FC4AE09FC4C" box="[719,839,941,965]" captionStart="Figure 2" captionStartId="5.[157,222,1748,1771]" captionTargetBox="[161,1151,1201,1717]" captionTargetId="figure@5.[161,1151,1201,1717]" captionTargetPageId="5" captionText="Figure 2. Cratera ochra sp. nov., holotype, in dorsal view. (A) Photograph of the live specimen. (B) Dorsal pattern of pigmentation of the preserved specimen." figureDoi="http://doi.org/10.5281/zenodo.3990053" httpUri="https://zenodo.org/record/3990053/files/figure.png" pageId="11" pageNumber="798">Figures 2B</figureCitation>
,
<figureCitation id="13651FFDFFDCFFB1B33B1FC4AE3DFC4C" box="[855,883,941,965]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">3B</figureCitation>
). These flecks form the dorsal bands as well as paramarginal stripes. The dorsal bands become inconspicuous towards the posterior tip, where the pigmentation is more irregularly distributed (
<figureCitation id="13651FFDFFDCFFB1B0CB187CAC5CFBA4" box="[167,274,1045,1069]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">Figure 3B</figureCitation>
). Ventral surface pale yellow. After fixation, the dorsal pigmentation becomes greyish and mainly greyish-brown next to the margins; dorsal ground colour becomes pale yellow, as well as the ventral surface of the body.
</paragraph>
<caption id="DF2153F0FFDCFFB1B0CC1CE7AE25FF2C" box="[160,875,142,165]" pageId="11" pageNumber="798">
<paragraph id="8BE10378FFDCFFB1B0CC1CE7AE25FF2C" blockId="11.[160,875,142,165]" box="[160,875,142,165]" pageId="11" pageNumber="798">
<emphasis id="B92ADF6AFFDCFFB1B0CC1CE7ADBDFF2C" bold="true" box="[160,243,142,165]" pageId="11" pageNumber="798">Table 1.</emphasis>
Measurements, in mm, of specimens of
<taxonomicName id="4C5E78FBFFDCFFB1B2FC1CE7AE5AFF2C" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[656,788,142,165]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="11" pageNumber="798" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFDCFFB1B2FC1CE7AE5AFF2C" box="[656,788,142,165]" italics="true" pageId="11" pageNumber="798">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFDCFFB1B3701CE7AE25FF2C" box="[796,875,142,165]" pageId="11" pageNumber="798" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BE10378FFDCFFB1B11C1CDBA90EFD24" pageId="11" pageNumber="798">
<table id="F95EF1D8FFDC0045B0C71CDBA920FD24" box="[171,1134,178,685]" gridcols="6" gridrows="21" pageId="11" pageNumber="798">
<tr id="356E013AFFDC0045B0C71CDBA920FF4C" box="[171,1134,178,197]" gridrow="0" pageId="11" pageNumber="798" rowspan-0="1">
<th id="76BF6846FFDC0045B11C1CDBACB0FF4C" box="[368,510,178,197]" gridcol="1" gridrow="0" pageId="11" pageNumber="798">Holotype MZUSP</th>
<th id="76BF6846FFDC0045B2721CDBAFDBFF4C" box="[542,661,178,197]" gridcol="2" gridrow="0" pageId="11" pageNumber="798">Paratype MZU</th>
<th id="76BF6846FFDC0045B2D11CDBAE7DFF4C" box="[701,819,178,197]" gridcol="3" gridrow="0" pageId="11" pageNumber="798">Paratype MZU</th>
<th id="76BF6846FFDC0045B3371CDBAE9CFF4C" box="[859,978,178,197]" gridcol="4" gridrow="0" pageId="11" pageNumber="798">Paratype MZU</th>
<th id="76BF6846FFDC0045B39B1CDBA920FF4C" box="[1015,1134,178,197]" gridcol="5" gridrow="0" pageId="11" pageNumber="798">Paratype MZU</th>
</tr>
<tr id="356E013AFFDC0045B0C71CA3A920FF54" box="[171,1134,202,221]" gridrow="1" pageId="11" pageNumber="798" rowspan-0="1">
<td id="76BF6846FFDC0045B11C1CA3ACB0FF54" box="[368,510,202,221]" gridcol="1" gridrow="1" pageId="11" pageNumber="798">PL.1564</td>
<td id="76BF6846FFDC0045B2721CA3AFDBFF54" box="[542,661,202,221]" gridcol="2" gridrow="1" pageId="11" pageNumber="798">PL.00187</td>
<td id="76BF6846FFDC0045B2D11CA3AE7DFF54" box="[701,819,202,221]" gridcol="3" gridrow="1" pageId="11" pageNumber="798">PL.00188</td>
<td id="76BF6846FFDC0045B3371CA3AE9CFF54" box="[859,978,202,221]" gridcol="4" gridrow="1" pageId="11" pageNumber="798">PL.00189</td>
<td id="76BF6846FFDC0045B39B1CA3A920FF54" box="[1015,1134,202,221]" gridcol="5" gridrow="1" pageId="11" pageNumber="798">PL.00190</td>
</tr>
<tr id="356E013AFFDC0045B0C71C83A920FF74" box="[171,1134,234,253]" gridrow="2" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71C83AC1BFF74" box="[171,341,234,253]" gridcol="0" gridrow="2" pageId="11" pageNumber="798">Maximum length in</th>
<td id="76BF6846FFDC0045B11C1C83ACB0FF74" box="[368,510,234,253]" gridcol="1" gridrow="2" pageId="11" pageNumber="798">54</td>
<td id="76BF6846FFDC0045B2721C83AFDBFF74" box="[542,661,234,253]" gridcol="2" gridrow="2" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11C83AE7DFF74" box="[701,819,234,253]" gridcol="3" gridrow="2" pageId="11" pageNumber="798">70</td>
<td id="76BF6846FFDC0045B3371C83AE9CFF74" box="[859,978,234,253]" gridcol="4" gridrow="2" pageId="11" pageNumber="798">72</td>
<td id="76BF6846FFDC0045B39B1C83A920FF74" box="[1015,1134,234,253]" gridcol="5" gridrow="2" pageId="11" pageNumber="798">40</td>
</tr>
<tr id="356E013AFFDC0045B0C71D6BA920FE9C" box="[171,1134,258,277]" gridrow="3" pageId="11" pageNumber="798" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76BF6846FFDC0045B0C71D6BAC1BFE9C" box="[171,341,258,277]" gridcol="0" gridrow="3" pageId="11" pageNumber="798">extension</th>
</tr>
<tr id="356E013AFFDC0045B0C71D73A920FEA4" box="[171,1134,282,301]" gridrow="4" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71D73AC1BFEA4" box="[171,341,282,301]" gridcol="0" gridrow="4" pageId="11" pageNumber="798">Maximum width in</th>
<td id="76BF6846FFDC0045B11C1D73ACB0FEA4" box="[368,510,282,301]" gridcol="1" gridrow="4" pageId="11" pageNumber="798">4.5</td>
<td id="76BF6846FFDC0045B2721D73AFDBFEA4" box="[542,661,282,301]" gridcol="2" gridrow="4" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11D73AE7DFEA4" box="[701,819,282,301]" gridcol="3" gridrow="4" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B3371D73AE9CFEA4" box="[859,978,282,301]" gridcol="4" gridrow="4" pageId="11" pageNumber="798">4</td>
<td id="76BF6846FFDC0045B39B1D73A920FEA4" box="[1015,1134,282,301]" gridcol="5" gridrow="4" pageId="11" pageNumber="798">3</td>
</tr>
<tr id="356E013AFFDC0045B0C71D5BA920FECC" box="[171,1134,306,325]" gridrow="5" pageId="11" pageNumber="798" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76BF6846FFDC0045B0C71D5BAC1BFECC" box="[171,341,306,325]" gridcol="0" gridrow="5" pageId="11" pageNumber="798">extension</th>
</tr>
<tr id="356E013AFFDC0045B0C71D23A920FED4" box="[171,1134,330,349]" gridrow="6" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71D23AC1BFED4" box="[171,341,330,349]" gridcol="0" gridrow="6" pageId="11" pageNumber="798">Length at rest</th>
<td id="76BF6846FFDC0045B11C1D23ACB0FED4" box="[368,510,330,349]" gridcol="1" gridrow="6" pageId="11" pageNumber="798">19</td>
<td id="76BF6846FFDC0045B2721D23AFDBFED4" box="[542,661,330,349]" gridcol="2" gridrow="6" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11D23AE7DFED4" box="[701,819,330,349]" gridcol="3" gridrow="6" pageId="11" pageNumber="798">34</td>
<td id="76BF6846FFDC0045B3371D23AE9CFED4" box="[859,978,330,349]" gridcol="4" gridrow="6" pageId="11" pageNumber="798">45</td>
<td id="76BF6846FFDC0045B39B1D23A920FED4" box="[1015,1134,330,349]" gridcol="5" gridrow="6" pageId="11" pageNumber="798">30</td>
</tr>
<tr id="356E013AFFDC0045B0C71D0BA920FEFC" box="[171,1134,354,373]" gridrow="7" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71D0BAC1BFEFC" box="[171,341,354,373]" gridcol="0" gridrow="7" pageId="11" pageNumber="798">Width at rest</th>
<td id="76BF6846FFDC0045B11C1D0BACB0FEFC" box="[368,510,354,373]" gridcol="1" gridrow="7" pageId="11" pageNumber="798">8</td>
<td id="76BF6846FFDC0045B2721D0BAFDBFEFC" box="[542,661,354,373]" gridcol="2" gridrow="7" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11D0BAE7DFEFC" box="[701,819,354,373]" gridcol="3" gridrow="7" pageId="11" pageNumber="798">7</td>
<td id="76BF6846FFDC0045B3371D0BAE9CFEFC" box="[859,978,354,373]" gridcol="4" gridrow="7" pageId="11" pageNumber="798">6</td>
<td id="76BF6846FFDC0045B39B1D0BA920FEFC" box="[1015,1134,354,373]" gridcol="5" gridrow="7" pageId="11" pageNumber="798">3</td>
</tr>
<tr id="356E013AFFDC0045B0C71D13A920FE04" box="[171,1134,378,397]" gridrow="8" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71D13AC1BFE04" box="[171,341,378,397]" gridcol="0" gridrow="8" pageId="11" pageNumber="798">Length*</th>
<td id="76BF6846FFDC0045B11C1D13ACB0FE04" box="[368,510,378,397]" gridcol="1" gridrow="8" pageId="11" pageNumber="798">52</td>
<td id="76BF6846FFDC0045B2721D13AFDBFE04" box="[542,661,378,397]" gridcol="2" gridrow="8" pageId="11" pageNumber="798">44</td>
<td id="76BF6846FFDC0045B2D11D13AE7DFE04" box="[701,819,378,397]" gridcol="3" gridrow="8" pageId="11" pageNumber="798">54</td>
<td id="76BF6846FFDC0045B3371D13AE9CFE04" box="[859,978,378,397]" gridcol="4" gridrow="8" pageId="11" pageNumber="798">53</td>
<td id="76BF6846FFDC0045B39B1D13A920FE04" box="[1015,1134,378,397]" gridcol="5" gridrow="8" pageId="11" pageNumber="798">36</td>
</tr>
<tr id="356E013AFFDC0045B0C71DFBA920FE2C" box="[171,1134,402,421]" gridrow="9" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71DFBAC1BFE2C" box="[171,341,402,421]" gridcol="0" gridrow="9" pageId="11" pageNumber="798">Width*</th>
<td id="76BF6846FFDC0045B11C1DFBACB0FE2C" box="[368,510,402,421]" gridcol="1" gridrow="9" pageId="11" pageNumber="798">5</td>
<td id="76BF6846FFDC0045B2721DFBAFDBFE2C" box="[542,661,402,421]" gridcol="2" gridrow="9" pageId="11" pageNumber="798">5</td>
<td id="76BF6846FFDC0045B2D11DFBAE7DFE2C" box="[701,819,402,421]" gridcol="3" gridrow="9" pageId="11" pageNumber="798">6.5</td>
<td id="76BF6846FFDC0045B3371DFBAE9CFE2C" box="[859,978,402,421]" gridcol="4" gridrow="9" pageId="11" pageNumber="798">5.5</td>
<td id="76BF6846FFDC0045B39B1DFBA920FE2C" box="[1015,1134,402,421]" gridcol="5" gridrow="9" pageId="11" pageNumber="798">4</td>
</tr>
<tr id="356E013AFFDC0045B0C71DC3A920FE34" box="[171,1134,426,445]" gridrow="10" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71DC3AC1BFE34" box="[171,341,426,445]" gridcol="0" gridrow="10" pageId="11" pageNumber="798">DM*</th>
<td id="76BF6846FFDC0045B11C1DC3ACB0FE34" box="[368,510,426,445]" gridcol="1" gridrow="10" pageId="11" pageNumber="798">38 (73)</td>
<td id="76BF6846FFDC0045B2721DC3AFDBFE34" box="[542,661,426,445]" gridcol="2" gridrow="10" pageId="11" pageNumber="798">33 (75)</td>
<td id="76BF6846FFDC0045B2D11DC3AE7DFE34" box="[701,819,426,445]" gridcol="3" gridrow="10" pageId="11" pageNumber="798">42 (78)</td>
<td id="76BF6846FFDC0045B3371DC3AE9CFE34" box="[859,978,426,445]" gridcol="4" gridrow="10" pageId="11" pageNumber="798">39 (73.5)</td>
<td id="76BF6846FFDC0045B39B1DC3A920FE34" box="[1015,1134,426,445]" gridcol="5" gridrow="10" pageId="11" pageNumber="798">26 (72)</td>
</tr>
<tr id="356E013AFFDC0045B0C71DABA920FE5C" box="[171,1134,450,469]" gridrow="11" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71DABAC1BFE5C" box="[171,341,450,469]" gridcol="0" gridrow="11" pageId="11" pageNumber="798">DG*</th>
<td id="76BF6846FFDC0045B11C1DABACB0FE5C" box="[368,510,450,469]" gridcol="1" gridrow="11" pageId="11" pageNumber="798">46 (88.5)</td>
<td id="76BF6846FFDC0045B2721DABAFDBFE5C" box="[542,661,450,469]" gridcol="2" gridrow="11" pageId="11" pageNumber="798">39 (88.5)</td>
<td id="76BF6846FFDC0045B2D11DABAE7DFE5C" box="[701,819,450,469]" gridcol="3" gridrow="11" pageId="11" pageNumber="798">47 (87)</td>
<td id="76BF6846FFDC0045B3371DABAE9CFE5C" box="[859,978,450,469]" gridcol="4" gridrow="11" pageId="11" pageNumber="798">47 (88.5)</td>
<td id="76BF6846FFDC0045B39B1DABA920FE5C" box="[1015,1134,450,469]" gridcol="5" gridrow="11" pageId="11" pageNumber="798">33 (91.5)</td>
</tr>
<tr id="356E013AFFDC0045B0C71DB3A920FE64" box="[171,1134,474,493]" gridrow="12" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71DB3AC1BFE64" box="[171,341,474,493]" gridcol="0" gridrow="12" pageId="11" pageNumber="798">DMG*</th>
<td id="76BF6846FFDC0045B11C1DB3ACB0FE64" box="[368,510,474,493]" gridcol="1" gridrow="12" pageId="11" pageNumber="798">8</td>
<td id="76BF6846FFDC0045B2721DB3AFDBFE64" box="[542,661,474,493]" gridcol="2" gridrow="12" pageId="11" pageNumber="798">6</td>
<td id="76BF6846FFDC0045B2D11DB3AE7DFE64" box="[701,819,474,493]" gridcol="3" gridrow="12" pageId="11" pageNumber="798">5</td>
<td id="76BF6846FFDC0045B3371DB3AE9CFE64" box="[859,978,474,493]" gridcol="4" gridrow="12" pageId="11" pageNumber="798">8</td>
<td id="76BF6846FFDC0045B39B1DB3A920FE64" box="[1015,1134,474,493]" gridcol="5" gridrow="12" pageId="11" pageNumber="798">7</td>
</tr>
<tr id="356E013AFFDC0045B0C71D9BA920FD8C" box="[171,1134,498,517]" gridrow="13" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71D9BAC1BFD8C" box="[171,341,498,517]" gridcol="0" gridrow="13" pageId="11" pageNumber="798">DPVP</th>
<td id="76BF6846FFDC0045B11C1D9BACB0FD8C" box="[368,510,498,517]" gridcol="1" gridrow="13" pageId="11" pageNumber="798">4.2</td>
<td id="76BF6846FFDC0045B2721D9BAFDBFD8C" box="[542,661,498,517]" gridcol="2" gridrow="13" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11D9BAE7DFD8C" box="[701,819,498,517]" gridcol="3" gridrow="13" pageId="11" pageNumber="798">5.5</td>
<td id="76BF6846FFDC0045B3371D9BAE9CFD8C" box="[859,978,498,517]" gridcol="4" gridrow="13" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B39B1D9BA920FD8C" box="[1015,1134,498,517]" gridcol="5" gridrow="13" pageId="11" pageNumber="798">1.2</td>
</tr>
<tr id="356E013AFFDC0045B0C71E63A920FD94" box="[171,1134,522,541]" gridrow="14" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71E63AC1BFD94" box="[171,341,522,541]" gridcol="0" gridrow="14" pageId="11" pageNumber="798">Ovaries</th>
<td id="76BF6846FFDC0045B11C1E63ACB0FD94" box="[368,510,522,541]" gridcol="1" gridrow="14" pageId="11" pageNumber="798">13.5 (26)</td>
<td id="76BF6846FFDC0045B2721E63AFDBFD94" box="[542,661,522,541]" gridcol="2" gridrow="14" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11E63AE7DFD94" box="[701,819,522,541]" gridcol="3" gridrow="14" pageId="11" pageNumber="798">12 (22)</td>
<td id="76BF6846FFDC0045B3371E63AE9CFD94" box="[859,978,522,541]" gridcol="4" gridrow="14" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B39B1E63A920FD94" box="[1015,1134,522,541]" gridcol="5" gridrow="14" pageId="11" pageNumber="798">–</td>
</tr>
<tr id="356E013AFFDC0045B0C71E4BA920FDBC" box="[171,1134,546,565]" gridrow="15" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71E4BAC1BFDBC" box="[171,341,546,565]" gridcol="0" gridrow="15" pageId="11" pageNumber="798">Anteriormost testes</th>
<td id="76BF6846FFDC0045B11C1E4BACB0FDBC" box="[368,510,546,565]" gridcol="1" gridrow="15" pageId="11" pageNumber="798">15 (29)</td>
<td id="76BF6846FFDC0045B2721E4BAFDBFDBC" box="[542,661,546,565]" gridcol="2" gridrow="15" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11E4BAE7DFDBC" box="[701,819,546,565]" gridcol="3" gridrow="15" pageId="11" pageNumber="798">16 (39.5)</td>
<td id="76BF6846FFDC0045B3371E4BAE9CFDBC" box="[859,978,546,565]" gridcol="4" gridrow="15" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B39B1E4BA920FDBC" box="[1015,1134,546,565]" gridcol="5" gridrow="15" pageId="11" pageNumber="798">–</td>
</tr>
<tr id="356E013AFFDC0045B0C71E53A920FDC4" box="[171,1134,570,589]" gridrow="16" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71E53AC1BFDC4" box="[171,341,570,589]" gridcol="0" gridrow="16" pageId="11" pageNumber="798">Posteriormost testes</th>
<td id="76BF6846FFDC0045B11C1E53ACB0FDC4" box="[368,510,570,589]" gridcol="1" gridrow="16" pageId="11" pageNumber="798">34.5 (66)</td>
<td id="76BF6846FFDC0045B2721E53AFDBFDC4" box="[542,661,570,589]" gridcol="2" gridrow="16" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B2D11E53AE7DFDC4" box="[701,819,570,589]" gridcol="3" gridrow="16" pageId="11" pageNumber="798">30 (55.5)</td>
<td id="76BF6846FFDC0045B3371E53AE9CFDC4" box="[859,978,570,589]" gridcol="4" gridrow="16" pageId="11" pageNumber="798">–</td>
<td id="76BF6846FFDC0045B39B1E53A920FDC4" box="[1015,1134,570,589]" gridcol="5" gridrow="16" pageId="11" pageNumber="798">22.5 (62.5)</td>
</tr>
<tr id="356E013AFFDC0045B0C71E3BA920FDEC" box="[171,1134,594,613]" gridrow="17" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71E3BAC1BFDEC" box="[171,341,594,613]" gridcol="0" gridrow="17" pageId="11" pageNumber="798">Prostatic vesicle</th>
<td id="76BF6846FFDC0045B11C1E3BACB0FDEC" box="[368,510,594,613]" gridcol="1" gridrow="17" pageId="11" pageNumber="798">0.3</td>
<td id="76BF6846FFDC0045B2721E3BAFDBFDEC" box="[542,661,594,613]" gridcol="2" gridrow="17" pageId="11" pageNumber="798">0.5</td>
<td id="76BF6846FFDC0045B2D11E3BAE7DFDEC" box="[701,819,594,613]" gridcol="3" gridrow="17" pageId="11" pageNumber="798">0.5</td>
<td id="76BF6846FFDC0045B3371E3BAE9CFDEC" box="[859,978,594,613]" gridcol="4" gridrow="17" pageId="11" pageNumber="798">0.7</td>
<td id="76BF6846FFDC0045B39B1E3BA920FDEC" box="[1015,1134,594,613]" gridcol="5" gridrow="17" pageId="11" pageNumber="798">0.2</td>
</tr>
<tr id="356E013AFFDC0045B0C71E03A920FDF4" box="[171,1134,618,637]" gridrow="18" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71E03AC1BFDF4" box="[171,341,618,637]" gridcol="0" gridrow="18" pageId="11" pageNumber="798">Penis papilla</th>
<td id="76BF6846FFDC0045B11C1E03ACB0FDF4" box="[368,510,618,637]" gridcol="1" gridrow="18" pageId="11" pageNumber="798">1.1</td>
<td id="76BF6846FFDC0045B2721E03AFDBFDF4" box="[542,661,618,637]" gridcol="2" gridrow="18" pageId="11" pageNumber="798">0.6</td>
<td id="76BF6846FFDC0045B2D11E03AE7DFDF4" box="[701,819,618,637]" gridcol="3" gridrow="18" pageId="11" pageNumber="798">1</td>
<td id="76BF6846FFDC0045B3371E03AE9CFDF4" box="[859,978,618,637]" gridcol="4" gridrow="18" pageId="11" pageNumber="798">0.9</td>
<td id="76BF6846FFDC0045B39B1E03A920FDF4" box="[1015,1134,618,637]" gridcol="5" gridrow="18" pageId="11" pageNumber="798">0.4</td>
</tr>
<tr id="356E013AFFDC0045B0C71EEBA920FD1C" box="[171,1134,642,661]" gridrow="19" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71EEBAC1BFD1C" box="[171,341,642,661]" gridcol="0" gridrow="19" pageId="11" pageNumber="798">Male atrium</th>
<td id="76BF6846FFDC0045B11C1EEBACB0FD1C" box="[368,510,642,661]" gridcol="1" gridrow="19" pageId="11" pageNumber="798">1.1</td>
<td id="76BF6846FFDC0045B2721EEBAFDBFD1C" box="[542,661,642,661]" gridcol="2" gridrow="19" pageId="11" pageNumber="798">0.7</td>
<td id="76BF6846FFDC0045B2D11EEBAE7DFD1C" box="[701,819,642,661]" gridcol="3" gridrow="19" pageId="11" pageNumber="798">1.3</td>
<td id="76BF6846FFDC0045B3371EEBAE9CFD1C" box="[859,978,642,661]" gridcol="4" gridrow="19" pageId="11" pageNumber="798">0.95</td>
<td id="76BF6846FFDC0045B39B1EEBA920FD1C" box="[1015,1134,642,661]" gridcol="5" gridrow="19" pageId="11" pageNumber="798">0.5</td>
</tr>
<tr id="356E013AFFDC0045B0C71EF3A920FD24" box="[171,1134,666,685]" gridrow="20" pageId="11" pageNumber="798">
<th id="76BF6846FFDC0045B0C71EF3AC1BFD24" box="[171,341,666,685]" gridcol="0" gridrow="20" pageId="11" pageNumber="798">Female atrium</th>
<td id="76BF6846FFDC0045B11C1EF3ACB0FD24" box="[368,510,666,685]" gridcol="1" gridrow="20" pageId="11" pageNumber="798">0.8</td>
<td id="76BF6846FFDC0045B2721EF3AFDBFD24" box="[542,661,666,685]" gridcol="2" gridrow="20" pageId="11" pageNumber="798">0.2</td>
<td id="76BF6846FFDC0045B2D11EF3AE7DFD24" box="[701,819,666,685]" gridcol="3" gridrow="20" pageId="11" pageNumber="798">0.5</td>
<td id="76BF6846FFDC0045B3371EF3AE9CFD24" box="[859,978,666,685]" gridcol="4" gridrow="20" pageId="11" pageNumber="798">0.8</td>
<td id="76BF6846FFDC0045B39B1EF3A920FD24" box="[1015,1134,666,685]" gridcol="5" gridrow="20" pageId="11" pageNumber="798">0.2</td>
</tr>
</table>
</paragraph>
<tableNote id="76B802F6FFDCFFB1B0CC1ED2AE26FD76" pageId="11" pageNumber="798" targetBox="[171,1134,178,685]" targetPageId="11">
<paragraph id="8BE10378FFDCFFB1B0CC1ED2A9CDFD47" blockId="11.[160,1157,699,767]" box="[160,1155,699,718]" pageId="11" pageNumber="798">–, not measured; *, after fixation; DG, distance of gonopore from anterior end; DM, distance of mouth from anterior</paragraph>
<paragraph id="8BE10378FFDCFFB1B0DC1EBAA9CBFD6F" blockId="11.[160,1157,699,767]" box="[176,1157,723,742]" pageId="11" pageNumber="798">end; DMG, distance between mouth and gonopore; DPVP, distance between prostatic vesicle and pharyngeal pouch.</paragraph>
<paragraph id="8BE10378FFDCFFB1B0DC1E85AE26FD76" blockId="11.[160,1157,699,767]" box="[176,872,748,767]" pageId="11" pageNumber="798">The numbers given in parentheses represent the position relative to body length.</paragraph>
</tableNote>
<paragraph id="8BE10378FFDCFFB1B0AC1814A950FACB" blockId="11.[160,1156,837,1346]" pageId="11" pageNumber="798">
Eyes monolobated. They are initially uniserial, with pigment cups of about 15 μm in diameter, surrounding anterior tip (
<figureCitation id="13651FFDFFDCFFB1B25D18F6AFD2FB3E" box="[561,668,1183,1207]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">Figure 3C</figureCitation>
). After the second millimetre, the monolobated eyes become larger (with pigment cups of about
<quantity id="4CA6AE9DFFDCFFB1B34C18ABAE24FB53" box="[800,874,1218,1242]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="798" unit="mm" value="30.0">30 µm</quantity>
in diameter) and spread onto the dorsal surface, occupying the maximum width of about 40% of the body width on each side of the body. Eyes remain dorsal, but become less numerous towards posterior tip (
<figureCitation id="13651FFDFFDCFFB1B0CB1943AC5AFACB" box="[167,276,1322,1346]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">Figure 3D</figureCitation>
). Inconspicuous clear halos may occur around dorsal eyes (
<figureCitation id="13651FFDFFDCFFB1B3C51943A95CFACB" box="[937,1042,1322,1346]" captionStart="Figure 3" captionStartId="6.[160,225,1613,1636]" captionTargetBox="[268,1047,142,1582]" captionTargetId="figure@6.[268,1048,142,1582]" captionTargetPageId="6" captionText="Figure 3. Cratera ochra sp. nov. in dorsal view. (A) Detail of the anterior extremity of the holotype. (B) Detail of the median third of the body of the holotype. (C) Detail of eye pattern at the anterior extremity of the holotype. (D) Eye pattern of a fixed specimen (paratype MZU PL.00187)." figureDoi="http://doi.org/10.5281/zenodo.3990055" httpUri="https://zenodo.org/record/3990055/files/figure.png" pageId="11" pageNumber="798">Figure 3B</figureCitation>
).
</paragraph>
<paragraph id="8BE10378FFDCFFB1B0CC1907A91BFA66" blockId="11.[160,1156,1390,1727]" pageId="11" pageNumber="798">
<emphasis id="B92ADF6AFFDCFFB1B0CC1907AE42FA0E" bold="true" box="[160,780,1390,1415]" inLineHeading="true" italics="true" pageId="11" pageNumber="798" reason="3">Sensory organs, epidermis and body musculature.</emphasis>
Sensory pits (
<figureCitation id="13651FFDFFDCFFB1B3D21906A91FFA0E" box="[958,1105,1391,1415]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="11" pageNumber="798">Figure 4A, B</figureCitation>
), as simple invaginations, contour anterior tip and occur ventromarginally in an irregular, single row in the anterior eighth of the body. Initially they occur at intervals of approximately
<quantity id="4CA6AE9DFFDCFFB1B12619BEACDBFA66" box="[330,405,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" pageNumber="798" unit="mm" value="18.0">18 µm</quantity>
, becoming gradually sparser. Their depth is about
<quantity id="4CA6AE9DFFDCFFB1B3B119BEA91FFA66" box="[989,1105,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="11" pageNumber="798" unit="mm" value="37.5" valueMax="50.0" valueMin="25.0">
25
<emphasis id="B92ADF6AFFDCFFB1B39419BEA94BFA66" box="[1016,1029,1495,1519]" italics="true" pageId="11" pageNumber="798">–</emphasis>
50 µm
</quantity>
.
</paragraph>
<paragraph id="8BE10378FFDCFFB6B0AF1993ACBEFD36" blockId="11.[160,1156,1390,1727]" lastBlockId="12.[160,1156,436,1188]" lastPageId="12" lastPageNumber="799" pageId="11" pageNumber="798">
Creeping sole occupying the whole body width. Two
<typeStatus id="54E5BDDAFFDCFFB1B33C1993AEC2F99B" box="[848,908,1530,1554]" pageId="11" pageNumber="798">types</typeStatus>
of glands discharge through dorsal epidermis and body margins of the pre-pharyngeal region: numerous rhabditogen cells with xanthophil secretion (rhammites) and cells with amorphous, cyanophil secretion (
<figureCitation id="13651FFDFFDCFFB1B1FC1A0BACB2F9F3" box="[400,508,1634,1658]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="11" pageNumber="798">Figure 4E</figureCitation>
). Creeping sole receives the secretion from amorphous, cyanophil glands, as well as few rhabditogen cells with small, xanthophil rhabdites and cells with coarse granular, xanthophil secretion (
<figureCitation id="13651FFDFFDCFFB1B29E1ACEAE2DF936" box="[754,867,1703,1727]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="11" pageNumber="798">Figure 4F</figureCitation>
). The glandular margin (
<figureCitation id="13651FFDFFDBFFB6B0CB1DDDAC7FFE45" box="[167,305,436,460]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4C, D</figureCitation>
), which is visible after the second millimetre of the body, receives openings of at least four
<typeStatus id="54E5BDDAFFDBFFB6B13E1DBEACC0FE66" box="[338,398,471,495]" pageId="12" pageNumber="799">types</typeStatus>
of glands. Two of them are more numerous: glands with erythrophil, coarse granules, and glands with xanthophil, coarse granules. Other two
<typeStatus id="54E5BDDAFFDBFFB6B4771D90A919FD98" box="[1051,1111,505,529]" pageId="12" pageNumber="799">types</typeStatus>
are scarcer: cyanophil glands with amorphous secretion and erythrophil glands with fine granules. On anterior tip, glands with cyanophil, amorphous secretion and rhabditogen glands with xanthophil secretion open through the whole surface of the body and glands with coarse granular, xanthophil secretion have numerous openings through the ventral epidermis (
<figureCitation id="13651FFDFFDBFFB6B11A1ECEACAAFD36" box="[374,484,679,703]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4A</figureCitation>
).
</paragraph>
<caption id="DF2153F0FFDBFFB6B0CC1CE7AE31FF69" pageId="12" pageNumber="799">
<paragraph id="8BE10378FFDBFFB6B0CC1CE7AE31FF69" blockId="12.[160,1155,142,224]" pageId="12" pageNumber="799">
<emphasis id="B92ADF6AFFDBFFB6B0CC1CE7ADBFFF2C" bold="true" box="[160,241,142,165]" pageId="12" pageNumber="799">Table 2.</emphasis>
Body height and cutaneous musculature in the median region of a transverse section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc: h index) of specimens of
<taxonomicName id="4C5E78FBFFDBFFB6B2CF1CA0AE66FF69" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[675,808,201,224]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="12" pageNumber="799" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFDBFFB6B2CF1CA0AE66FF69" box="[675,808,201,224]" italics="true" pageId="12" pageNumber="799">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFDBFFB6B35C1CA0AE31FF69" box="[816,895,201,224]" pageId="12" pageNumber="799" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BE10378FFDBFFB6B1FF1C84A96DFEE1" pageId="12" pageNumber="799">
<table id="F95EF1D8FFDB0045B0C71C84A937FEE1" box="[171,1145,237,360]" gridcols="4" gridrows="5" pageId="12" pageNumber="799">
<tr id="356E013AFFDB0045B0C71C84A937FE89" box="[171,1145,237,256]" gridrow="0" pageId="12" pageNumber="799" rowspan-0="1">
<th id="76BF6846FFDB0045B1FF1C84AF26FE89" box="[403,616,237,256]" gridcol="1" gridrow="0" pageId="12" pageNumber="799">Holotype MZUSP PL.1564</th>
<th id="76BF6846FFDB0045B2C41C84AE3FFE89" box="[680,881,237,256]" gridcol="2" gridrow="0" pageId="12" pageNumber="799">Paratype MZU PL.00188</th>
<th id="76BF6846FFDB0045B3DC1C84A937FE89" box="[944,1145,237,256]" gridcol="3" gridrow="0" pageId="12" pageNumber="799">Paratype MZU PL.00190</th>
</tr>
<tr id="356E013AFFDB0045B0C71D64A937FEA9" box="[171,1145,269,288]" gridrow="1" pageId="12" pageNumber="799">
<th id="76BF6846FFDB0045B0C71D64AC1DFEA9" box="[171,339,269,288]" gridcol="0" gridrow="1" pageId="12" pageNumber="799">Dorsal musculature</th>
<td id="76BF6846FFDB0045B1FF1D64AF26FEA9" box="[403,616,269,288]" gridcol="1" gridrow="1" pageId="12" pageNumber="799">56</td>
<td id="76BF6846FFDB0045B2C41D64AE3FFEA9" box="[680,881,269,288]" gridcol="2" gridrow="1" pageId="12" pageNumber="799">71</td>
<td id="76BF6846FFDB0045B3DC1D64A937FEA9" box="[944,1145,269,288]" gridcol="3" gridrow="1" pageId="12" pageNumber="799">40</td>
</tr>
<tr id="356E013AFFDB0045B0C71D4CA937FEB1" box="[171,1145,293,312]" gridrow="2" pageId="12" pageNumber="799">
<th id="76BF6846FFDB0045B0C71D4CAC1DFEB1" box="[171,339,293,312]" gridcol="0" gridrow="2" pageId="12" pageNumber="799">Ventral musculature</th>
<td id="76BF6846FFDB0045B1FF1D4CAF26FEB1" box="[403,616,293,312]" gridcol="1" gridrow="2" pageId="12" pageNumber="799">65</td>
<td id="76BF6846FFDB0045B2C41D4CAE3FFEB1" box="[680,881,293,312]" gridcol="2" gridrow="2" pageId="12" pageNumber="799">108</td>
<td id="76BF6846FFDB0045B3DC1D4CA937FEB1" box="[944,1145,293,312]" gridcol="3" gridrow="2" pageId="12" pageNumber="799">50</td>
</tr>
<tr id="356E013AFFDB0045B0C71D54A937FED9" box="[171,1145,317,336]" gridrow="3" pageId="12" pageNumber="799">
<th id="76BF6846FFDB0045B0C71D54AC1DFED9" box="[171,339,317,336]" gridcol="0" gridrow="3" pageId="12" pageNumber="799">Body height</th>
<td id="76BF6846FFDB0045B1FF1D54AF26FED9" box="[403,616,317,336]" gridcol="1" gridrow="3" pageId="12" pageNumber="799">992</td>
<td id="76BF6846FFDB0045B2C41D54AE3FFED9" box="[680,881,317,336]" gridcol="2" gridrow="3" pageId="12" pageNumber="799">1500</td>
<td id="76BF6846FFDB0045B3DC1D54A937FED9" box="[944,1145,317,336]" gridcol="3" gridrow="3" pageId="12" pageNumber="799">806</td>
</tr>
<tr id="356E013AFFDB0045B0C71D3CA937FEE1" box="[171,1145,341,360]" gridrow="4" pageId="12" pageNumber="799">
<th id="76BF6846FFDB0045B0C71D3CAC1DFEE1" box="[171,339,341,360]" gridcol="0" gridrow="4" pageId="12" pageNumber="799">mc: h (%)</th>
<td id="76BF6846FFDB0045B1FF1D3CAF26FEE1" box="[403,616,341,360]" gridcol="1" gridrow="4" pageId="12" pageNumber="799">12</td>
<td id="76BF6846FFDB0045B2C41D3CAE3FFEE1" box="[680,881,341,360]" gridcol="2" gridrow="4" pageId="12" pageNumber="799">12</td>
<td id="76BF6846FFDB0045B3DC1D3CA937FEE1" box="[944,1145,341,360]" gridcol="3" gridrow="4" pageId="12" pageNumber="799">11</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE10378FFDBFFB6B0B21EA0AEA4FC06" blockId="12.[160,1156,436,1188]" pageId="12" pageNumber="799">
Cutaneous musculature with the usual three layers (circular, oblique and longitudinal layers), longitudinal layer with thick bundles (
<figureCitation id="13651FFDFFDBFFB6B3661E85AEDCFC8D" box="[778,914,748,772]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">
Figure 4C
<emphasis id="B92ADF6AFFDBFFB6B3141E85AECBFC8D" box="[888,901,748,772]" italics="true" pageId="12" pageNumber="799">–</emphasis>
F
</figureCitation>
,
<tableCitation id="C6DC36C3FFDBFFB6B3CD1E85AEBBFC8D" box="[929,1013,748,772]" captionStart="Table 2" captionStartId="12.[160,216,142,165]" captionText="Table 2. Body height and cutaneous musculature in the median region of a transverse section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc: h index) of specimens of Cratera ochra sp. nov." pageId="12" pageNumber="799">Table 2</tableCitation>
). Cutaneous musculature thinner in the pre-pharyngeal region than in the anterior region of the body, gradually diminishing its thickness towards anterior tip (
<figureCitation id="13651FFDFFDBFFB6B31F1F58AEACFCC0" box="[883,994,817,841]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4A</figureCitation>
). Musculature becoming progressively lower towards body margins. Ventral musculature slightly higher than dorsal at the sagittal plane. Ratio of mc: h 11
<emphasis id="B92ADF6AFFDBFFB6B3571F1FAE06FC07" box="[827,840,886,910]" italics="true" pageId="12" pageNumber="799">–</emphasis>
12% (
<tableCitation id="C6DC36C3FFDBFFB6B3E51F1EAE93FC06" box="[905,989,887,911]" captionStart="Table 2" captionStartId="12.[160,216,142,165]" captionText="Table 2. Body height and cutaneous musculature in the median region of a transverse section of the pre-pharyngeal region, in micrometers, and ratio of the height of cutaneous musculature to the height of the body (mc: h index) of specimens of Cratera ochra sp. nov." pageId="12" pageNumber="799">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BE10378FFDBFFB6B0AC1FF0AC38FB2D" blockId="12.[160,1156,436,1188]" pageId="12" pageNumber="799">
Mesenchymal musculature (
<figureCitation id="13651FFDFFDBFFB6B1961FF0AFD9FC38" box="[506,663,921,945]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4C, E, F</figureCitation>
) well developed, mainly composed of three layers: (1) dorsal subcutaneous, located mainly close to the cutaneous musculature, with oblique fibres variously oriented (about 5
<emphasis id="B92ADF6AFFDBFFB6B2191FB7AFCCFC7F" box="[629,642,990,1014]" italics="true" pageId="12" pageNumber="799">–</emphasis>
14 fibres thick); (2) supra-intestinal transverse (about 7
<emphasis id="B92ADF6AFFDBFFB6B16A1868AC5DFB90" box="[262,275,1025,1049]" italics="true" pageId="12" pageNumber="799">–</emphasis>
14 fibres thick); (3) subintestinal transverse (about 14
<emphasis id="B92ADF6AFFDBFFB6B3F51868AEE8FB90" box="[921,934,1025,1049]" italics="true" pageId="12" pageNumber="799">–</emphasis>
22 fibres thick). In addition, there are scattered transverse subneural fibres and ventral subcutaneous oblique fibres, as well as numerous dorso-ventral fibres. On the anterior region of the body (
<figureCitation id="13651FFDFFDBFFB6B0871800AC36FB08" box="[235,376,1129,1153]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4A, B</figureCitation>
), the mesenchymal musculature is less developed than in the prepharyngeal region.
</paragraph>
<paragraph id="8BE10378FFDBFFB6B0CC188DAEA2FACB" blockId="12.[160,1156,1252,1727]" pageId="12" pageNumber="799">
<emphasis id="B92ADF6AFFDBFFB6B0CC188DAC49FB74" bold="true" box="[160,263,1252,1277]" inLineHeading="true" italics="true" pageId="12" pageNumber="799" reason="3">Pharynx.</emphasis>
Pharynx cylindrical, about 5
<emphasis id="B92ADF6AFFDBFFB6B23E188DAF11FB75" box="[594,607,1252,1276]" italics="true" pageId="12" pageNumber="799">–</emphasis>
6% of body length, with dorsal insertion posteriorly shifted, but located at the anterior third of pharyngeal pouch. Mouth posterior to the dorsal insertion, in the median third of pharyngeal pouch (
<figureCitation id="13651FFDFFDBFFB6B31D1943AE91FACB" box="[881,991,1322,1346]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="12" pageNumber="799">Figure 4G</figureCitation>
).
</paragraph>
<paragraph id="8BE10378FFDBFFB6B0D21924ACF1FA66" blockId="12.[160,1156,1252,1727]" pageId="12" pageNumber="799">
Oesophagus short; oesophagus: pharynx ratio varying from 14% in
<typeStatus id="54E5BDDAFFDBFFB6B3B31924A90DFAEC" box="[991,1091,1357,1381]" pageId="12" pageNumber="799" type="paratype">paratype</typeStatus>
MZU PL.00190 to 18.5% in the
<typeStatus id="54E5BDDAFFDBFFB6B1A51906AF7EFA0E" box="[457,560,1391,1415]" pageId="12" pageNumber="799" type="holotype">holotype</typeStatus>
. It is lined by ciliated, cuboidal to columnar epithelium with some insunk nuclei. The oesophagus is coated with a thick subepithelial muscle layer with circular fibres, followed by a thin muscle layer with longitudinal fibres (about
<quantity id="4CA6AE9DFFDBFFB6B09E19BEAC3CFA66" box="[242,370,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="12" pageNumber="799" unit="mm" value="80.0" valueMax="100.0" valueMin="60.0">
60
<emphasis id="B92ADF6AFFDBFFB6B16219BEAC55FA66" box="[270,283,1495,1519]" italics="true" pageId="12" pageNumber="799">–</emphasis>
100 µm
</quantity>
thick).
</paragraph>
<paragraph id="8BE10378FFDBFFB7B0AC1993AF8AFEB8" blockId="12.[160,1156,1252,1727]" lastBlockId="13.[160,1155,142,305]" lastPageId="13" lastPageNumber="800" pageId="12" pageNumber="799">
Pharynx and pharyngeal lumen lined by ciliated, cuboidal epithelium with insunk nuclei. Pharyngeal glands constituted by three secretory cell
<typeStatus id="54E5BDDAFFDBFFB6B3E51A74AE86F9BC" box="[905,968,1565,1589]" pageId="12" pageNumber="799">types</typeStatus>
: cells with fine granular erythrophil secretion; cells with fine granular xanthophil secretion and cells with amorphous cyanophil secretion. Cell bodies of pharyngeal glands located in the mesenchyme. Outer pharyngeal musculature (about
<quantity id="4CA6AE9DFFDBFFB6B37A1AECAEC3F914" box="[790,909,1668,1693]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.4" metricValueMin="1.0" pageId="12" pageNumber="799" unit="mm" value="17.0" valueMax="24.0" valueMin="10.0">
10
<emphasis id="B92ADF6AFFDBFFB6B35D1AEDAE70F915" box="[817,830,1668,1692]" italics="true" pageId="12" pageNumber="799">–</emphasis>
24 µm
</quantity>
thick) comprised of subepithelial layer of longitudinal muscles, followed by a circular layer and more internally by longitudinal fibres. All layers become thinner towards pharyngeal tip and the longitudinal internal fibres become mixed with the circular fibres. Inner pharyngeal musculature (about
<quantity id="4CA6AE9DFFDAFFB7B1EF1CBDAF41FF65" box="[387,527,211,236]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="13" pageNumber="800" unit="mm" value="125.0" valueMax="150.0" valueMin="100.0">
100
<emphasis id="B92ADF6AFFDAFFB7B1C01CBAACF7FF62" box="[428,441,211,235]" italics="true" pageId="13" pageNumber="800">–</emphasis>
150 µm
</quantity>
thick) comprises a thick subepithelial layer with circular fibres, mixed internally and externally with longitudinal fibres. Inner musculature gradually becomes thinner towards pharyngeal tip.
</paragraph>
<paragraph id="8BE10378FFDAFFB7B0CC1D37AC07FCF6" blockId="13.[160,1156,350,1727]" pageId="13" pageNumber="800">
<emphasis id="B92ADF6AFFDAFFB7B0CC1D37ACD3FEFE" bold="true" box="[160,413,350,375]" inLineHeading="true" italics="true" pageId="13" pageNumber="800" reason="3">Reproductive organs.</emphasis>
Testes in one irregular row on either side of the body, located beneath the dorsal transverse mesenchymal muscles (
<figureCitation id="13651FFDFFDAFFB7B3671DEBAEDAFE13" box="[779,916,386,410]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="13" pageNumber="800">Figure 4C, E</figureCitation>
). They begin slightly posterior to the ovaries, in the anterior third of the body, and extend to near the root of the pharynx (
<tableCitation id="C6DC36C3FFDAFFB7B1531DAEACD8FE56" box="[319,406,455,479]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="13" pageNumber="800">Table 1</tableCitation>
). Sperm ducts medial to ovovitelline ducts, above fibres of the subintestinal transverse mesenchymal musculature, in pre-pharyngeal region. They form spermiducal vesicles posterior to pharynx. Distally, spermiducal vesicles enter laterally into the proximal expanded portion of the prostatic vesicle (
<figureCitation id="13651FFDFFDAFFB7B3F11E46A90FFDCE" box="[925,1089,559,583]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="13" pageNumber="800">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFDAFFB7B4341E46A938FDCE" box="[1112,1142,559,583]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">7A</figureCitation>
). Extrabulbar prostatic vesicle, unpaired, located near the common muscle coat, with ample proximal portion and tubular distal portion. The proximal portion is laterally expanded and T-shaped (
<figureCitation id="13651FFDFFDAFFB7B1A61EFEAF0DFD26" box="[458,579,663,687]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="13" pageNumber="800">Figures 5B</figureCitation>
,
<figureCitation id="13651FFDFFDAFFB7B23F1EFEAF3FFD26" box="[595,625,663,687]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">7A</figureCitation>
); it is located closer to the ventral epidermis than to the dorsal epidermis. The prostatic vesicle penetrates the common muscle coat, becoming sinuous, and continues inside the penis papilla as an ejaculatory duct (
<figureCitation id="13651FFDFFDAFFB7B0CB1E96AC58FC9E" box="[167,278,767,791]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="13" pageNumber="800">Figure 5A</figureCitation>
). Ejaculatory duct slightly sinuous, opening through an expansion at the tip of the penis papilla (
<figureCitation id="13651FFDFFDAFFB7B1E21F4BACB4FCB3" box="[398,506,802,826]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="13" pageNumber="800">Figure 5B</figureCitation>
,
<figureCitation id="13651FFDFFDAFFB7B2641F4BAF6AFCB3" box="[520,548,802,826]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">7B</figureCitation>
). Male atrium without folds, occupied by the conical and symmetric penis papilla with ventral insertion posteriorly displaced (
<figureCitation id="13651FFDFFDAFFB7B38B1F2DA933FCD5" box="[999,1149,836,860]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="13" pageNumber="800">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFDAFFB7B0CC1F0EAD94FCF6" box="[160,218,871,895]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="13" pageNumber="800">6A, B</figureCitation>
,
<tableCitation id="C6DC36C3FFDAFFB7B0851F0EAC73FCF6" box="[233,317,871,895]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="13" pageNumber="800">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BE10378FFDAFFB7B0D21FE3AF76FB75" blockId="13.[160,1156,350,1727]" pageId="13" pageNumber="800">
Lining epithelium of sperm ducts cuboidal and ciliated; thin muscularis (about
<quantity id="4CA6AE9DFFDAFFB7B4041FE3AD93FC4D" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="13" pageNumber="800" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">
2
<emphasis id="B92ADF6AFFDAFFB7B41A1FE3A9CDFC2B" box="[1142,1155,906,930]" italics="true" pageId="13" pageNumber="800">–</emphasis>
4 µm
</quantity>
) constituted of interwoven circular and longitudinal fibres. Prostatic vesicle lined with ciliated, tall columnar or pseudostratified epithelium. Muscularis of prostatic vesicle (about
<quantity id="4CA6AE9DFFDAFFB7B0981F9BAC26FB83" box="[244,360,1010,1034]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="800" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">
10
<emphasis id="B92ADF6AFFDAFFB7B17C1F9BAC53FB83" box="[272,285,1010,1034]" italics="true" pageId="13" pageNumber="800">–</emphasis>
20 µm
</quantity>
thick) comprises circular fibres mixed with longitudinal and oblique fibres. Ejaculatory duct lined with ciliated, columnar epithelium, showing a pseudostratified appearance at the expanded portion (
<figureCitation id="13651FFDFFDAFFB7B2F5185EAE4BFBC6" box="[665,773,1079,1103]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">Figure 7B</figureCitation>
). Muscle coat of ejaculatory duct thin (about
<quantity id="4CA6AE9DFFDAFFB7B1421833ACC5FBFB" box="[302,395,1114,1138]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="800" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">
3
<emphasis id="B92ADF6AFFDAFFB7B1501833AC07FBFB" box="[316,329,1114,1138]" italics="true" pageId="13" pageNumber="800">–</emphasis>
5 µm
</quantity>
), mainly constituted of circular fibres. Both prostatic vesicle and ejaculatory duct receive openings from glands producing a fine granular, mixed secretion (
<figureCitation id="13651FFDFFDAFFB7B0B118F6AC04FB3E" box="[221,330,1183,1207]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">Figure 7A</figureCitation>
) as well as glands with amorphous cyanophil secretion. Granules with a mixed secretion have a cyanophil external part and an erythrophil internal core. Both glands with extrabulbar cell bodies.
</paragraph>
<paragraph id="8BE10378FFDAFFB7B0AC196EAE10FA66" blockId="13.[160,1156,350,1727]" pageId="13" pageNumber="800">
Penis papilla lined with non-ciliated, pseudostratified epithelium. Penis glands produce numerous fine granular, xanthophil and erythrophil secretions as well as less numerous cyanophil secretions of two
<typeStatus id="54E5BDDAFFDAFFB7B2091925AFEFFAED" box="[613,673,1356,1380]" pageId="13" pageNumber="800">types</typeStatus>
(amorphous and fine granular secretion). All penis glands with cell bodies external to common muscle coat; their long necks run longitudinally through the papilla (
<figureCitation id="13651FFDFFDAFFB7B24319FBAF8DFA23" box="[559,707,1426,1450]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="13" pageNumber="800">Figures 6A, B</figureCitation>
,
<figureCitation id="13651FFDFFDAFFB7B2BD19FBAFA3FA23" box="[721,749,1426,1450]" captionStart="Figure 7" captionStartId="10.[160,225,614,637]" captionTargetBox="[162,1152,142,583]" captionTargetId="figure@10.[162,1153,142,583]" captionTargetPageId="10" captionText="Figure 7. Cratera ochra sp. nov. (A) Detail of prostatic vesicle of paratype MZU PL.00189 in horizontal section. (B) Detail of tip of the penis papilla of the holotype in sagittal section." figureDoi="http://doi.org/10.5281/zenodo.3990063" httpUri="https://zenodo.org/record/3990063/files/figure.png" pageId="13" pageNumber="800">7B</figureCitation>
), with numerous openings through its lining epithelium. Muscularis of the penis papilla (about
<quantity id="4CA6AE9DFFDAFFB7B35319DDAEECFA45" box="[831,930,1460,1484]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="13" pageNumber="800" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">
6
<emphasis id="B92ADF6AFFDAFFB7B32119DDAE14FA45" box="[845,858,1460,1484]" italics="true" pageId="13" pageNumber="800">–</emphasis>
10 µm
</quantity>
thick) composed of a subepithelial circular layer, followed by a longitudinal layer.
</paragraph>
<paragraph id="8BE10378FFDAFFB4B0AC1993AD92FEDD" blockId="13.[160,1156,350,1727]" lastBlockId="14.[160,1156,142,1414]" lastPageId="14" lastPageNumber="801" pageId="13" pageNumber="800">
Epithelial lining of male atrium columnar to pseudostratified (about
<quantity id="4CA6AE9DFFDAFFB7B3D71993A977F99B" box="[955,1081,1529,1554]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="15.0" metricValueMin="2.0" pageId="13" pageNumber="800" unit="mm" value="85.0" valueMax="150.0" valueMin="20.0">
20
<emphasis id="B92ADF6AFFDAFFB7B3BB1990AEAAF998" box="[983,996,1529,1553]" italics="true" pageId="13" pageNumber="800">–</emphasis>
150 µm
</quantity>
thick), non-ciliated. Four
<typeStatus id="54E5BDDAFFDAFFB7B11E1A75ACE0F9BD" box="[370,430,1564,1588]" pageId="13" pageNumber="800">types</typeStatus>
of glands empty through this epithelium: abundant cells with fine granular, cyanophil secretion and cells with cyanophil amorphous secretion; and few glands with fine granular, mixed secretion (cyanophil peripheral portion and erythrophil core) as well as scarce xanthophil glands. These glands have their cell bodies located in the mesenchyme, mainly anterior and laterally to the copulatory apparatus, or between the fibres of the common muscle coat. Numerous necks of both
<typeStatus id="54E5BDDAFFD9FFB4B44F1CE7A911FF2F" box="[1059,1119,142,166]" pageId="14" pageNumber="801">types</typeStatus>
of cyanophil glands concentrate their openings at the dorso-lateral wall of the male atrium (
<figureCitation id="13651FFDFFD9FFB4B0CB1CBDAC7FFF65" box="[167,305,212,236]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">Figure 6A, B</figureCitation>
). Their numerous gland necks are located between fibres of the stroma in the dorso-lateral walls of the male atrium. Muscularis of male atrium (about 4
<emphasis id="B92ADF6AFFD9FFB4B4541C9FA90BFE87" box="[1080,1093,246,270]" italics="true" pageId="14" pageNumber="801">–</emphasis>
8 μm thick) comprised of a subepithelial layer with circular fibres, followed by a longitudinal layer.
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0AC1D37AC28FD58" blockId="14.[160,1156,142,1414]" pageId="14" pageNumber="801">
Vitellaria (
<figureCitation id="13651FFDFFD9FFB4B15B1D37AC83FEFF" box="[311,461,350,374]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="14" pageNumber="801">Figure 4C, F</figureCitation>
), situated between intestinal branches, opening into the ovovitelline ducts. Ovaries oval-elongate (
<figureCitation id="13651FFDFFD9FFB4B2EE1DE8AFBCFE10" box="[642,754,385,409]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="14" pageNumber="801">Figure 4H</figureCitation>
), measuring about
<quantity id="4CA6AE9DFFD9FFB4B3BA1DE8A92AFE10" box="[982,1124,385,409]" metricMagnitude="-1" metricUnit="m" metricValue="4.1" metricValueMax="5.2" metricValueMin="3.0" pageId="14" pageNumber="801" unit="mm" value="410.0" valueMax="520.0" valueMin="300.0">
300
<emphasis id="B92ADF6AFFD9FFB4B3931DE8A942FE10" box="[1023,1036,385,409]" italics="true" pageId="14" pageNumber="801">–</emphasis>
520 µm
</quantity>
in diameter in the
<typeStatus id="54E5BDDAFFD9FFB4B10C1DCDAC8AFE35" box="[352,452,420,444]" pageId="14" pageNumber="801" type="holotype">holotype</typeStatus>
and
<typeStatus id="54E5BDDAFFD9FFB4B26A1DCDAF24FE35" box="[518,618,420,444]" pageId="14" pageNumber="801" type="paratype">paratype</typeStatus>
MZU PL.00188, respectively. They are located dorsal to the ventral nerve plate, in the anterior third of the body (
<tableCitation id="C6DC36C3FFD9FFB4B3FA1DAFAEA6FE57" box="[918,1000,454,478]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="14" pageNumber="801">Table 1</tableCitation>
). Ovovitelline ducts emerge dorsally from the median third of the ovaries (
<figureCitation id="13651FFDFFD9FFB4B3381D80AE8DFD88" box="[852,963,489,513]" captionStart="Figure 4" captionStartId="7.[160,225,1516,1539]" captionTargetBox="[163,1153,145,1485]" captionTargetId="figure@7.[163,1153,142,1485]" captionTargetPageId="7" captionText="Figure 4. Cratera ochra sp. nov., holotype, in transverse (A–F) or sagittal (G, H) sections. (A) Anterior region of the body. (B) Detail of the anterior region of the body. (C) Pre-pharyngeal region. (D) Detail of body margin of the pre-pharyngeal region. (E) Detail of a dorsal portion of the pre-pharyngeal region. (F) Detail of a ventral portion of the pre-pharyngeal region. (G) Pharynx. (H) Ovary." figureDoi="http://doi.org/10.5281/zenodo.3990057" httpUri="https://zenodo.org/record/3990057/files/figure.png" pageId="14" pageNumber="801">Figure 4H</figureCitation>
) and run posteriorly immediately above the nerve plate. Lateral to the female atrium, ovovitelline ducts ascend postero-medially, to unite dorsal to the end of the median third or dorsal to the posterior third of the female atrium, thus forming the common glandular ovovitelline duct (
<figureCitation id="13651FFDFFD9FFB4B08D1E1AAC3FFD02" box="[225,369,627,651]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="14" pageNumber="801">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFD9FFB4B1121E1AACD2FD02" box="[382,412,627,651]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">6A</figureCitation>
). The female genital duct is dorso-anteriorly curved (
<figureCitation id="13651FFDFFD9FFB4B3811E1AA933FD02" box="[1005,1149,627,651]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="14" pageNumber="801">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFD9FFB4B0CC1EFFADF0FD27" box="[160,190,662,686]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">6A</figureCitation>
). Female atrium, shorter than male atrium, funnel-shaped without folds (
<figureCitation id="13651FFDFFD9FFB4B4691EFFADE0FD58" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="14" pageNumber="801">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFD9FFB4B0D11ED0ADB9FD58" box="[189,247,697,721]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">6A, B</figureCitation>
,
<tableCitation id="C6DC36C3FFD9FFB4B16A1ED0AC14FD58" box="[262,346,697,721]" captionStart="Table 1" captionStartId="11.[160,216,142,165]" captionText="Table 1. Measurements, in mm, of specimens of Cratera ochra sp. nov." pageId="14" pageNumber="801">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0AC1EB2AE43FCF7" blockId="14.[160,1156,142,1414]" pageId="14" pageNumber="801">
Ovovitelline ducts and common ovovitelline duct lined with ciliated, cuboidal to columnar epithelium and covered with intermingled circular and longitudinal muscle fibres (about 3
<emphasis id="B92ADF6AFFD9FFB4B1211F48AC14FCB0" box="[333,346,801,825]" italics="true" pageId="14" pageNumber="801">–</emphasis>
5 μm). Numerous shell glands with erythrophil secretion empty into common glandular ovovitelline duct as well as into the distal half of the ascending portion of the ovovitelline ducts (
<figureCitation id="13651FFDFFD9FFB4B24D1F0FAFF8FCF7" box="[545,694,870,894]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="14" pageNumber="801">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFD9FFB4B2A91F0FAFB1FCF7" box="[709,767,870,894]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">6A, B</figureCitation>
).
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0AC1FE0ACF4FBF8" blockId="14.[160,1156,142,1414]" pageId="14" pageNumber="801">
Female genital duct and atrium lined by tall columnar to pseudostratified epithelium (
<figureCitation id="13651FFDFFD9FFB4B0C41FC2AC5BFC4A" box="[168,277,939,963]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">Figure 6A</figureCitation>
) containing some lacunae. Female duct and atrium receive abundant cyanophil secretion of two
<typeStatus id="54E5BDDAFFD9FFB4B1F91FA7AC9FFC6F" box="[405,465,974,998]" pageId="14" pageNumber="801">types</typeStatus>
(amorphous and fine granular secretion), as well as erythrophil glands and few xanthophil glands. Their cell bodies are located between fibres of the atrial stroma or external to the common muscle coat. Muscularis of vagina and female atrium (about
<quantity id="4CA6AE9DFFD9FFB4B1F1185FACA9FBC7" box="[413,487,1078,1102]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="801" unit="mm" value="15.0">15 µm</quantity>
thick) composed mainly of circular fibres interposed with some longitudinal fibres.
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0AC1812AEE0FA97" blockId="14.[160,1156,142,1414]" pageId="14" pageNumber="801">
Gonoduct vertical at the sagittal plane. Male and female atria with ample communication, without separating folds (
<figureCitation id="13651FFDFFD9FFB4B25418F7AF9CFB3F" box="[568,722,1182,1206]" captionStart="Figure 5" captionStartId="8.[160,225,1407,1430]" captionTargetBox="[163,1153,142,1376]" captionTargetId="figure@8.[163,1153,142,1376]" captionTargetPageId="8" captionText="Figure 5. Cratera ochra sp. nov. (A) Sagittal composite reconstruction of the copulatory apparatus of the holotype. (B) Diagrammatic horizontal composite reconstruction of the copulatory apparatus of paratype MZU PL.00189." figureDoi="http://doi.org/10.5281/zenodo.3990059" httpUri="https://zenodo.org/record/3990059/files/figure.png" pageId="14" pageNumber="801">Figures 5A, B</figureCitation>
,
<figureCitation id="13651FFDFFD9FFB4B28F18F7AE51FB3F" box="[739,799,1182,1206]" captionStart="Figure 6" captionStartId="9.[160,225,1375,1398]" captionTargetBox="[163,1153,152,1344]" captionTargetId="figure@9.[163,1153,142,1344]" captionTargetPageId="9" captionText="Figure 6. Cratera ochra sp. nov. (A) Copulatory apparatus of the holotype in sagittal section. (B) Copulatory apparatus of paratype MZU PL.00189 in horizontal section." figureDoi="http://doi.org/10.5281/zenodo.3990061" httpUri="https://zenodo.org/record/3990061/files/figure.png" pageId="14" pageNumber="801">6A, B</figureCitation>
). Gonoduct lined with ciliated columnar epithelium, receiving the openings of numerous rhabditogen glands and cyanophil glands with amorphous secretion. Muscularis of gonoduct comprised of a subepithelial layer of circular fibres, followed by a longitudinal layer.
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0AC1940AFF0FA0F" blockId="14.[160,1156,142,1414]" pageId="14" pageNumber="801">Common muscle coat with circular, longitudinal and oblique fibres; it is thin along both male and female atria. A stroma with sparse intermingled muscle fibres separates the atrial muscularis and common muscle coat.</paragraph>
<paragraph id="8BE10378FFD9FFB4B0CC19A1AC54FA68" blockId="14.[160,1155,1480,1713]" box="[160,282,1480,1505]" pageId="14" pageNumber="801">
<heading id="D0A9B414FFD9FFB4B0CC19A1AC54FA68" box="[160,282,1480,1505]" fontSize="11" level="3" pageId="14" pageNumber="801" reason="5">
<emphasis id="B92ADF6AFFD9FFB4B0CC19A1AC54FA68" bold="true" box="[160,282,1480,1505]" inLineHeading="true" italics="true" pageId="14" pageNumber="801" reason="3">Variability</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFD9FFB4B0CC1982AF93F938" blockId="14.[160,1155,1480,1713]" pageId="14" pageNumber="801">
<materialsCitation id="3B360925FFD9FFB4B0CC1982AC24F98A" ID-GBIF-Occurrence="2831860310" box="[160,362,1515,1539]" collectionCode="PL" pageId="14" pageNumber="801" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E5BDDAFFD9FFB4B0CC1982AC4DF98A" box="[160,259,1515,1539]" pageId="14" pageNumber="801" type="paratype">Paratype</typeStatus>
MZU
<collectionCode id="ED4F9BBDFFD9FFB4B1221982AC24F98A" box="[334,362,1515,1539]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" pageNumber="801" type="Museum">PL</collectionCode>
</materialsCitation>
.000190 is at an initial stage of maturity, showing poorly developed testes and vitellaria, few shell glands and short female atrium.
<materialsCitation id="3B360925FFD9FFB4B3D91A67ADF3F9C0" ID-GBIF-Occurrence="2831860317" collectionCode="PL" pageId="14" pageNumber="801" specimenCount="1" typeStatus="paratype">
In
<typeStatus id="54E5BDDAFFD9FFB4B3B61A67A970F9AF" box="[986,1086,1550,1574]" pageId="14" pageNumber="801" type="paratype">paratype</typeStatus>
MZU
<collectionCode id="ED4F9BBDFFD9FFB4B0CC1A58ADF3F9C0" box="[160,189,1585,1609]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" pageNumber="801" type="Museum">PL</collectionCode>
</materialsCitation>
.00187, the penis papilla is protruded towards the gonoduct and there is abundant cyanophil secretion in the male atrium and in the gonoduct; the epithelial lining of the female atrium is taller laterally than medially.
<materialsCitation id="3B360925FFD9FFB4B2AB1A1FAFC1F938" ID-GBIF-Occurrence="2831860319" collectionCode="PL" pageId="14" pageNumber="801" specimenCount="1" typeStatus="paratype">
The proximal portion of the prostatic vesicle contains sperm in
<typeStatus id="54E5BDDAFFD9FFB4B1AB1AF0AF65F938" box="[455,555,1689,1713]" pageId="14" pageNumber="801" type="paratype">paratype</typeStatus>
MZU
<collectionCode id="ED4F9BBDFFD9FFB4B21E1AF0AFC1F938" box="[626,655,1689,1713]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="14" pageNumber="801" type="Museum">PL</collectionCode>
</materialsCitation>
.00189.
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0CC1CE7ACF4FF2E" blockId="15.[160,1156,142,1380]" box="[160,442,142,167]" pageId="15" pageNumber="802">
<heading id="D0A9B414FFD8FFB5B0CC1CE7ACF4FF2E" box="[160,442,142,167]" fontSize="11" level="3" pageId="15" pageNumber="802" reason="5">
<emphasis id="B92ADF6AFFD8FFB5B0CC1CE7ACF4FF2E" bold="true" box="[160,442,142,167]" inLineHeading="true" italics="true" pageId="15" pageNumber="802" reason="3">Comparative discussion</emphasis>
</heading>
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0CC1CDBAF6DFEDD" blockId="15.[160,1156,142,1380]" pageId="15" pageNumber="802">
In accordance with the phylogenetic analyses, the external and internal characters of the new species herein described as
<taxonomicName id="4C5E78FBFFD8FFB5B27C1CBDAFEBFF65" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[528,677,212,236]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B27C1CBDAFEBFF65" box="[528,677,212,236]" italics="true" pageId="15" pageNumber="802">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B2C01CBDAE4DFF65" box="[684,771,212,236]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
strongly support its inclusion into the genus
<taxonomicName id="4C5E78FBFFD8FFB5B1731C9EAF00FE86" authority="Carbayo et al. The" authorityName="Carbayo et al. The" authorityYear="2013" box="[287,590,247,271]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFD8FFB5B1731C9EAC21FE86" box="[287,367,247,271]" italics="true" pageId="15" pageNumber="802">Cratera</emphasis>
Carbayo et al. The
</taxonomicName>
morphological characters concern the following features: monolobated eyes, ejaculatory duct forming a distal cavity in the penis papilla and funnel-shaped female atrium.
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0AC1D36A962FD7D" blockId="15.[160,1156,142,1380]" pageId="15" pageNumber="802">
<taxonomicName id="4C5E78FBFFD8FFB5B0AC1D36AC16FEFE" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[192,344,351,375]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B0AC1D36AC16FEFE" box="[192,344,351,375]" italics="true" pageId="15" pageNumber="802">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B10F1D36ACF1FEFE" box="[355,447,351,375]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
, with dorsal eyes, can be differentiated from
<taxonomicName id="4C5E78FBFFD8FFB5B3BF1D36AC95FE13" authority="(Riester, 1938)" baseAuthorityName="Riester" baseAuthorityYear="1938" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="pseudovaginuloides">
<emphasis id="B92ADF6AFFD8FFB5B3BF1D36AC67FE13" italics="true" pageId="15" pageNumber="802">Cratera pseudovaginuloides</emphasis>
(
<bibRefCitation id="EFCF7E89FFD8FFB5B1571DEBAC9CFE13" author="Riester A" box="[315,466,386,410]" pageId="15" pageNumber="802" pagination="1 - 88" refId="ref13409" refString="Riester A. 1938. Beitrage zur Geoplaniden-Fauna Brasiliens. Abh Senckenberg Naturf Ges. 441: 1 - 88." type="journal article" year="1938">Riester, 1938</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C5E78FBFFD8FFB5B1801DEBAECCFE13" authority="(E. M. Froehlich, 1955)" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[492,898,386,410]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="yara">
<emphasis id="B92ADF6AFFD8FFB5B1801DEBAF37FE13" box="[492,633,386,410]" italics="true" pageId="15" pageNumber="802">Cratera yara</emphasis>
(E.M. Froehlich, 1955)
</taxonomicName>
and
<taxonomicName id="4C5E78FBFFD8FFB5B3AA1DEBAC88FE35" authority="Carbayo &amp; Almeida, 2015" authorityName="Carbayo &amp; Almeida" authorityYear="2015" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="cuarassu">
<emphasis id="B92ADF6AFFD8FFB5B3AA1DEBA9CAFE13" box="[966,1156,386,410]" italics="true" pageId="15" pageNumber="802">Cratera cuarassu</emphasis>
<bibRefCitation id="EFCF7E89FFD8FFB5B0CC1DCDAC88FE35" author="Carbayo F &amp; Almeida AL" box="[160,454,420,444]" pageId="15" pageNumber="802" pagination="27 - 40" refId="ref12355" refString="Carbayo F, Almeida AL. 2015. Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa. 3931: 27 - 40." type="journal article" year="2015">Carbayo &amp; Almeida, 2015</bibRefCitation>
</taxonomicName>
, which have exclusively marginal eyes (E.M. Froehlich 1955;
<bibRefCitation id="EFCF7E89FFD8FFB5B0CC1DAEAC06FE56" author="Froehlich CG" box="[160,328,455,479]" pageId="15" pageNumber="802" pagination="173 - 191" refId="ref12666" refString="Froehlich CG. 1956. Planarias terrestres do Parana. Dusenia. 7: 173 - 191." type="journal article" year="1956">Froehlich 1956</bibRefCitation>
;
<bibRefCitation id="EFCF7E89FFD8FFB5B13B1DAEAFC0FE56" author="Carbayo F &amp; Almeida AL" box="[343,654,455,479]" pageId="15" pageNumber="802" pagination="27 - 40" refId="ref12355" refString="Carbayo F, Almeida AL. 2015. Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa. 3931: 27 - 40." type="journal article" year="2015">Carbayo and Almeida 2015</bibRefCitation>
). It is also easily distinguished from
<taxonomicName id="4C5E78FBFFD8FFB5B45F1DAEACE3FD8B" authority="Rossi et al., 2014" authorityName="Rossi" authorityYear="2014" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="steffeni">
<emphasis id="B92ADF6AFFD8FFB5B45F1DAEADA3FD8B" italics="true" pageId="15" pageNumber="802">Cratera steffeni</emphasis>
<bibRefCitation id="EFCF7E89FFD8FFB5B0991D83ACE3FD8B" author="Rossi I &amp; Fontoura M &amp; Amaral S &amp; Leal-Zanchet AM" box="[245,429,490,514]" pageId="15" pageNumber="802" pagination="514 - 524" refId="ref13470" refString="Rossi I, Fontoura M, Amaral S, Leal-Zanchet AM. 2014. A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil. Zootaxa. 3794: 514 - 524." type="journal article" year="2014">Rossi et al., 2014</bibRefCitation>
</taxonomicName>
, in which the eyes are restricted to either side of the body (
<bibRefCitation id="EFCF7E89FFD8FFB5B4251D83AC5AFDAD" author="Rossi I &amp; Fontoura M &amp; Amaral S &amp; Leal-Zanchet AM" pageId="15" pageNumber="802" pagination="514 - 524" refId="ref13470" refString="Rossi I, Fontoura M, Amaral S, Leal-Zanchet AM. 2014. A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil. Zootaxa. 3794: 514 - 524." type="journal article" year="2014">Rossi et al. 2014</bibRefCitation>
). These four species and also
<taxonomicName id="4C5E78FBFFD8FFB5B20D1E65AEB7FDAD" authority="(E. M. Froehlich, 1955)" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[609,1017,524,548]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="crioula">
<emphasis id="B92ADF6AFFD8FFB5B20D1E65AE4CFDAD" box="[609,770,524,548]" italics="true" pageId="15" pageNumber="802">Cratera crioula</emphasis>
(E.M. Froehlich, 1955)
</taxonomicName>
,
<taxonomicName id="4C5E78FBFFD8FFB5B4681E65AC12FDCE" authority="(Froehlich, 1956)" baseAuthorityName="Froehlich" baseAuthorityYear="1956" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="joia">
<emphasis id="B92ADF6AFFD8FFB5B4681E65A9CDFDAD" box="[1028,1155,524,548]" italics="true" pageId="15" pageNumber="802">Cratera joia</emphasis>
(
<bibRefCitation id="EFCF7E89FFD8FFB5B0CA1E46AC1AFDCE" author="Froehlich CG" box="[166,340,559,583]" pageId="15" pageNumber="802" pagination="173 - 191" refId="ref12666" refString="Froehlich CG. 1956. Planarias terrestres do Parana. Dusenia. 7: 173 - 191." type="journal article" year="1956">Froehlich, 1956</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C5E78FBFFD8FFB5B1FB1E46AEC3FDCE" authority="Carbayo &amp; Almeida, 2015" authorityName="Carbayo &amp; Almeida" authorityYear="2015" box="[407,909,559,583]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="anamariae">
<emphasis id="B92ADF6AFFD8FFB5B1FB1E46AF28FDCE" box="[407,614,559,583]" italics="true" pageId="15" pageNumber="802">Cratera anamariae</emphasis>
<bibRefCitation id="EFCF7E89FFD8FFB5B2031E46AEC3FDCE" author="Carbayo F &amp; Almeida AL" box="[623,909,559,583]" pageId="15" pageNumber="802" pagination="27 - 40" refId="ref12355" refString="Carbayo F, Almeida AL. 2015. Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa. 3931: 27 - 40." type="journal article" year="2015">Carbayo &amp; Almeida, 2015</bibRefCitation>
</taxonomicName>
have different colour patterns in comparison to the new species herein described (
<bibRefCitation id="EFCF7E89FFD8FFB5B3221E3BAE94FDE3" author="Riester A" box="[846,986,594,618]" pageId="15" pageNumber="802" pagination="1 - 88" refId="ref13409" refString="Riester A. 1938. Beitrage zur Geoplaniden-Fauna Brasiliens. Abh Senckenberg Naturf Ges. 441: 1 - 88." type="journal article" year="1938">Riester 1938</bibRefCitation>
; E.M. Froehlich 1955;
<bibRefCitation id="EFCF7E89FFD8FFB5B08B1E1DACDEFD05" author="Froehlich CG" box="[231,400,628,652]" pageId="15" pageNumber="802" pagination="173 - 191" refId="ref12666" refString="Froehlich CG. 1956. Planarias terrestres do Parana. Dusenia. 7: 173 - 191." type="journal article" year="1956">Froehlich 1956</bibRefCitation>
;
<bibRefCitation id="EFCF7E89FFD8FFB5B1CC1E1DAF11FD05" author="Rossi I &amp; Fontoura M &amp; Amaral S &amp; Leal-Zanchet AM" box="[416,607,628,652]" pageId="15" pageNumber="802" pagination="514 - 524" refId="ref13470" refString="Rossi I, Fontoura M, Amaral S, Leal-Zanchet AM. 2014. A new species of land flatworm (Platyhelminthes: Continenticola) from areas of Araucaria Forest in southern Brazil. Zootaxa. 3794: 514 - 524." type="journal article" year="2014">Rossi et al. 2014</bibRefCitation>
;
<bibRefCitation id="EFCF7E89FFD8FFB5B2031E1DAEE7FD05" author="Carbayo F &amp; Almeida AL" box="[623,937,628,652]" pageId="15" pageNumber="802" pagination="27 - 40" refId="ref12355" refString="Carbayo F, Almeida AL. 2015. Anatomical deviation of male organs of land planarians from Rio de Janeiro, Brazil, with description of two new species of Cratera (Platyhelminthes, Tricladida). Zootaxa. 3931: 27 - 40." type="journal article" year="2015">Carbayo and Almeida 2015</bibRefCitation>
).
<taxonomicName id="4C5E78FBFFD8FFB5B3AD1E1DA917FD05" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[961,1113,628,652]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B3AD1E1DA917FD05" box="[961,1113,628,652]" italics="true" pageId="15" pageNumber="802">Cratera ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B4081E1DAD9FFD26" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
has colour and eye patterns similar to those of
<taxonomicName id="4C5E78FBFFD8FFB5B2881EFEA931FD26" authority="(E. M. Froehlich, 1955)" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[740,1151,663,687]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="tamoia">
<emphasis id="B92ADF6AFFD8FFB5B2881EFEAEC7FD26" box="[740,905,663,687]" italics="true" pageId="15" pageNumber="802">Cratera tamoia</emphasis>
(E.M. Froehlich, 1955)
</taxonomicName>
, but the new species can be mainly distinguished from the latter by its cylindrical pharynx, whereas the pharynx of
<taxonomicName id="4C5E78FBFFD8FFB5B24D1EB5AFC2FD7D" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[545,652,732,756]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="tamoia">
<emphasis id="B92ADF6AFFD8FFB5B24D1EB5AFC2FD7D" box="[545,652,732,756]" italics="true" pageId="15" pageNumber="802">C. tamoia</emphasis>
</taxonomicName>
is bell-shaped (E.M. Froehlich 1955).
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0AC1E96AE2EFBC6" blockId="15.[160,1156,142,1380]" pageId="15" pageNumber="802">
With respect to the morphology of the copulatory apparatus, species of the genus
<taxonomicName id="4C5E78FBFFD8FFB5B0CC1F4BADBEFCB3" authorityName="Carbayo et al. The" authorityYear="2013" box="[160,240,802,826]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92ADF6AFFD8FFB5B0CC1F4BADBEFCB3" box="[160,240,802,826]" italics="true" pageId="15" pageNumber="802">Cratera</emphasis>
</taxonomicName>
have an extrabulbar prostatic vesicle, a penis papilla with a distal cavity and a funnel-shaped female atrium, among other characteristics. The prostatic vesicle has two portions, a proximal expanded section and a distal tubular and sinuous portion. In
<taxonomicName id="4C5E78FBFFD8FFB5B0CC1FE3AC4CFC2B" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[160,258,906,930]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B0CC1FE3AC4CFC2B" box="[160,258,906,930]" italics="true" pageId="15" pageNumber="802">C. ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B17E1FE3AC3DFC2B" box="[274,371,906,930]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
, similarly to
<taxonomicName id="4C5E78FBFFD8FFB5B2741FE3AFC4FC2B" box="[536,650,905,930]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="steffeni">
<emphasis id="B92ADF6AFFD8FFB5B2741FE3AFC4FC2B" box="[536,650,905,930]" italics="true" pageId="15" pageNumber="802">C. steffeni</emphasis>
</taxonomicName>
, the proximal portion is characteristically T-shaped with the sperm ducts opening through its lateral walls. This form is particularly evident in horizontal sections of the copulatory apparatus. Other species of the genus, such as
<emphasis id="B92ADF6AFFD8FFB5B0901F9BAFF9FB83" box="[252,695,1010,1034]" italics="true" pageId="15" pageNumber="802">
<taxonomicName id="4C5E78FBFFD8FFB5B0901F9BAC2AFB83" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[252,356,1010,1034]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="crioula">C. crioula</taxonomicName>
,
<taxonomicName id="4C5E78FBFFD8FFB5B11D1F9BACF6FB83" baseAuthorityName="Froehlich" baseAuthorityYear="1956" box="[369,440,1010,1034]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="joia">C. joia</taxonomicName>
,
<taxonomicName id="4C5E78FBFFD8FFB5B1A81F9BAFF9FB83" baseAuthorityName="Riester" baseAuthorityYear="1938" box="[452,695,1010,1034]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="pseudovaginuloides">C. pseudovaginuloides</taxonomicName>
</emphasis>
and
<taxonomicName id="4C5E78FBFFD8FFB5B29E1F9BAE13FB83" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[754,861,1010,1034]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="tamoia">
<emphasis id="B92ADF6AFFD8FFB5B29E1F9BAE13FB83" box="[754,861,1010,1034]" italics="true" pageId="15" pageNumber="802">C. tamoia</emphasis>
</taxonomicName>
, which were described as possessing a prostatic vesicle with proximal diverticula (
<bibRefCitation id="EFCF7E89FFD8FFB5B351187DAE80FBA5" author="Riester A" box="[829,974,1044,1068]" pageId="15" pageNumber="802" pagination="1 - 88" refId="ref13409" refString="Riester A. 1938. Beitrage zur Geoplaniden-Fauna Brasiliens. Abh Senckenberg Naturf Ges. 441: 1 - 88." type="journal article" year="1938">Riester 1938</bibRefCitation>
; E.M. Froehlich 1955;
<bibRefCitation id="EFCF7E89FFD8FFB5B089185EACC3FBC6" author="Froehlich CG" box="[229,397,1079,1103]" pageId="15" pageNumber="802" pagination="173 - 191" refId="ref12666" refString="Froehlich CG. 1956. Planarias terrestres do Parana. Dusenia. 7: 173 - 191." type="journal article" year="1956">Froehlich 1956</bibRefCitation>
), may have a vesicle with a similar form.
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0AC1833A9CAFB3E" blockId="15.[160,1156,142,1380]" pageId="15" pageNumber="802">
In summary,
<taxonomicName id="4C5E78FBFFD8FFB5B1271833ACECFBFB" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[331,418,1114,1138]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B1271833ACECFBFB" box="[331,418,1114,1138]" italics="true" pageId="15" pageNumber="802">C. ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B1CB1833ACB5FBFB" box="[423,507,1114,1138]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
is mainly differentiated from its congeners by the combination of the following morphological characters: dorsal body surface with dispersed pigmentation forming two broad bands, eyes spreading over the dorsal surface, cylindrical pharynx.
</paragraph>
<paragraph id="8BE10378FFD8FFB5B0AC18ABAEB9FAED" blockId="15.[160,1156,142,1380]" pageId="15" pageNumber="802">
Measures of intraspecific and interspecific variation of the COI gene, as well as the ABGD algorithm applied to the COI data set, supported
<taxonomicName id="4C5E78FBFFD8FFB5B343188DAEC2FB75" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[815,908,1252,1276]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B343188DAEC2FB75" box="[815,908,1252,1276]" italics="true" pageId="15" pageNumber="802">C. ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B3FA188DAEBEFB75" box="[918,1008,1252,1276]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
as a species different from its congeners. In addition, both phylogenetic methods (maximum likelihood and Bayesian inference) revealed similar topologies, placing
<taxonomicName id="4C5E78FBFFD8FFB5B3CA1943A94FFACB" authority="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet, 2015" authorityName="Rossi &amp; Amaral &amp; Ribeiro &amp; Cauduro &amp; Fick &amp; Valiati &amp; Leal-Zanchet" authorityYear="2015" box="[934,1025,1322,1346]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="ochra" status="sp. nov.">
<emphasis id="B92ADF6AFFD8FFB5B3CA1943A94FFACB" box="[934,1025,1322,1346]" italics="true" pageId="15" pageNumber="802">C. ochra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2196211FFD8FFB5B4661943A92DFACB" box="[1034,1123,1322,1346]" pageId="15" pageNumber="802" rank="species">sp. nov.</taxonomicNameLabel>
as the sister group of the clade formed by
<taxonomicName id="4C5E78FBFFD8FFB5B2021925AF97FAED" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[622,729,1356,1380]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="tamoia">
<emphasis id="B92ADF6AFFD8FFB5B2021925AF97FAED" box="[622,729,1356,1380]" italics="true" pageId="15" pageNumber="802">C. tamoia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5E78FBFFD8FFB5B37A1925AE33FAED" baseAuthorityName="E. M. Froehlich" baseAuthorityYear="1955" box="[790,893,1356,1380]" class="Rhabditophora" family="Geoplanidae" genus="Cratera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="15" pageNumber="802" phylum="Platyhelminthes" rank="species" species="crioula">
<emphasis id="B92ADF6AFFD8FFB5B37A1925AE33FAED" box="[790,893,1356,1380]" italics="true" pageId="15" pageNumber="802">C. crioula</emphasis>
</taxonomicName>
(
<figureCitation id="13651FFDFFD8FFB5B3E11925AEA5FAED" box="[909,1003,1356,1380]" captionStart="Figure 1" captionStartId="4.[160,225,1585,1608]" captionTargetBox="[163,1116,142,1553]" captionTargetId="figure@4.[163,1153,142,1554]" captionTargetPageId="4" captionText="Figure 1. Bayesian phylogenetic tree inferred from the 640-bp of cytochrome c oxidase subunit I gene under GTR + I + G model of sequence evolution. The two new species are highlighted in light grey (Cratera ochra sp. nov.) and dark grey (Obama maculipunctata sp. nov.). Values indicate support for each node according to the maximum posterior probabilities&gt;70% and bootstrap support values&gt; 70%, respectively." figureDoi="http://doi.org/10.5281/zenodo.3990051" httpUri="https://zenodo.org/record/3990051/files/figure.png" pageId="15" pageNumber="802">Figure 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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