360 lines
46 KiB
XML
360 lines
46 KiB
XML
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<document id="B4CAACE793B9275EC9730FC97EB29A26" ID-CLB-Dataset="27716" ID-DOI="10.5852/ejt.2019.510" ID-GBIF-Dataset="08fe4f6b-5422-45af-bc2e-29ad35fed42f" ID-Zenodo-Dep="2616378" ID-ZooBank="BF58E108-2C6C-4A80-BD7D-47FEF5805B06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553845906407" checkinUser="plazi" docAuthor="Fujimoto, Shinta & Hansen, Jesper Guldberg" docDate="2019" docId="03D3878DFF9F58191C3F65CFD9C5FCD1" docLanguage="en" docName="ejt-510_fujimoto_hansen.pdf.imf" docOrigin="European Journal of Taxonomy 510" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Angursa seisuimaruae Fujimoto & Hansen 2019, sp. nov." docType="treatment" docUuid="208985F1-BBB3-40B4-A2E4-26E8662277DD" docUuidSource="ZooBank" docVersion="9" lastPageNumber="13" masterDocId="FFEAFFF5FF9658151E5F6411DB46FFE9" masterDocTitle="Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan" masterLastPageNumber="19" masterPageNumber="1" pageNumber="10" updateTime="1698715651624" updateUser="ExternalLinkService">
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<mods:mods id="AA253F87C6AECDD80F18B0434F9AC04D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="13AB34AF827106B714B6687927C4D858">
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<mods:title id="F046275B42D2BBF248C4F04846D78FDF">Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan</mods:title>
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</mods:titleInfo>
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<mods:name id="2FBF9C1A545E8A228E24C3697F571098" type="personal">
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<mods:namePart id="A099C6F2075276D54B81E223FD2D8D0D">Fujimoto, Shinta</mods:namePart>
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</mods:name>
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<mods:name id="977BD10125447FC72C8F5BEAA97C5407" type="personal">
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<mods:roleTerm id="602A4CEFC31278BC0F466605A5B97F11">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="13A7CBFC6E85894EA412829CCF096479">Hansen, Jesper Guldberg</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1465D4EBDE987E7745F5193ECEC55C36">text</mods:typeOfResource>
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<mods:title id="58A5389DABEB529C5998A91AAB83C35A">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="04C200367F74EC2457720B97F5FB5B2D">
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<mods:date id="BC0B4857E7AC47CDE8186B7F76B5555E">2019</mods:date>
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<mods:number id="682E0BB50788CE97838DE4883A224D55">2019-03-28</mods:number>
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</mods:detail>
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<mods:detail id="2E5A5BAF309F63BF1C4768E08F61E38B" type="volume">
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<mods:number id="38961DA6915A5F106F86199AD52E0A3C">510</mods:number>
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<mods:start id="2FA75908B05DCFA161CEB3991AF14387">1</mods:start>
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<mods:end id="0D27DAF22C437B6BDD22CB46DE0E9FD7">19</mods:end>
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<mods:classification id="78D0D5850883C329B46650A6EF57118A">journal article</mods:classification>
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<mods:identifier id="7085DE987671149171FFD7BD1E30392D" type="CLB-Dataset">27716</mods:identifier>
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<mods:identifier id="99AF1A5953E4199A41AA06073B9F1C9E" type="DOI">10.5852/ejt.2019.510</mods:identifier>
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<mods:identifier id="BEF86FD10AA88BAA2C6CD8B8164C1AD1" type="GBIF-Dataset">08fe4f6b-5422-45af-bc2e-29ad35fed42f</mods:identifier>
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<mods:identifier id="772BC7F7773D0C21DFB1632A37292C14" type="Zenodo-Dep">2616378</mods:identifier>
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<mods:identifier id="AD10E1F769F132A00C1F581EE768F46D" type="ZooBank">BF58E108-2C6C-4A80-BD7D-47FEF5805B06</mods:identifier>
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</mods:mods>
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<treatment id="03D3878DFF9F58191C3F65CFD9C5FCD1" ID-DOI="http://doi.org/10.5281/zenodo.5941840" ID-GBIF-Taxon="154522279" ID-Zenodo-Dep="5941840" LSID="urn:lsid:zoobank.org:act:208985F1-BBB3-40B4-A2E4-26E8662277DD" httpUri="http://treatment.plazi.org/id/03D3878DFF9F58191C3F65CFD9C5FCD1" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10">
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<subSubSection id="C3606510FF9F581C1C3F65CFDF85FDF5" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BC5369BFF9F581C1C3F65CFD895FE10" blockId="9.[369,1219,478,575]" box="[608,979,478,505]" pageId="9" pageNumber="10">
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<heading id="D08D81F7FF9F581C1C3F65CFD895FE10" box="[608,979,478,505]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="10" reason="2">
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<taxonomicName id="4C7A4D18FF9F581C1C3F65CFD837FE11" ID-CoL="9bec59d6-dfd9-4ff8-b1e4-3c5bf21491ef" authority="Fujimoto & Hansen, 2019" authorityName="Fujimoto & Hansen" authorityYear="2019" box="[608,881,478,504]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="species" species="seisuimaruae" status="sp. nov.">
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<emphasis id="B90EEA89FF9F581C1C3F65CFD837FE11" bold="true" box="[608,881,478,504]" italics="true" pageId="9" pageNumber="10">Angursa seisuimaruae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D57F2FF9F581C1D2665CED895FE10" box="[889,979,479,505]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BC5369BFF9F581C1F2E6610DF85FDF5" blockId="9.[369,1219,478,575]" box="[369,1219,513,540]" pageId="9" pageNumber="10">
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<uri id="FFEB3A99FF9F581C1F2E6610DF85FDF5" box="[369,1219,513,540]" pageId="9" pageNumber="10">
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urn:lsid:zoobank.org:act:
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<uuid id="FFDC0C4EFF9F581C1CCD6610DF85FDF5" box="[658,1219,513,540]" pageId="9" pageNumber="10">208985F1-BBB3-40B4-A2E4-26E8662277DD</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9F581C1CB86634D80BFDD6" box="[743,845,549,575]" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BC5369BFF9F581C1CB86634D80BFDD6" blockId="9.[369,1219,478,575]" box="[743,845,549,575]" pageId="9" pageNumber="10">
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<figureCitation id="13412A1EFF9F581C1CB86634D80BFDD6" box="[743,845,549,575]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[189,232,2000,2026]" captionStartId-1="11.[189,232,1923,1949]" captionTargetBox-1="[238,1344,814,1876]" captionTargetId-1="figure@11.[238,1352,814,1876]" captionTargetPageId-1="11" captionText-0="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." captionText-1="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri-1="https://zenodo.org/record/2616390/files/figure.png" pageId="9" pageNumber="10">Figs 6–7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9F581C1EE26676D9FFFD68" box="[189,697,614,641]" pageId="9" pageNumber="10" type="reference_group">
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<paragraph id="8BC5369BFF9F581C1EE26676D9FFFD68" blockId="9.[189,697,614,641]" box="[189,697,614,641]" pageId="9" pageNumber="10">
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<taxonomicName id="4C7A4D18FF9F581C1EE26676DA66FD68" ID-CoL="7443d693-6738-42b4-9e43-1b41918cea81" authorityName="Pollock" authorityYear="1979" box="[189,288,615,641]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="genus">
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<emphasis id="B90EEA89FF9F581C1EE26676DA66FD68" box="[189,288,615,641]" italics="true" pageId="9" pageNumber="10">Angursa</emphasis>
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</taxonomicName>
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sp. –
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<bibRefCitation id="EFEB4B6AFF9F581C1F396677D90EFD68" author="Kimura T. & Kimura S. & Jimi N. & Kakui K. & Tomioka S. & Oya Y. & Matsumoto Y. & Tanabe Y. & Hasegawa N. & Hookabe N. & Homma R. & Hosoda Y. & Fujimoto S. & Kuramochi T. & Fujita T. & Ogawa A. & Kobayashi I. & Ishida Y. & Tanaka H. & Onishi H. & Shimetsugu M. & Yoshikawa A. & Tanaka M. & Kushida Y. & Maekawa Y. & Nakamura T. & Okumura J. & Tanaka K." box="[358,584,614,641]" pageId="9" pageNumber="10" pagination="1 - 32" refId="ref8854" refString="Kimura T., Kimura S., Jimi N., Kakui K., Tomioka S., Oya Y., Matsumoto Y., Tanabe Y., Hasegawa N., Hookabe N., Homma R., Hosoda Y., Fujimoto S., Kuramochi T., Fujita T., Ogawa A., Kobayashi I., Ishida Y., Tanaka H., Onishi H., Shimetsugu M., Yoshikawa A., Tanaka M., Kushida Y., Maekawa Y., Nakamura T., Okumura J. & Tanaka K. 2018. Benthic deep-sea fauna in the Sea of Kumano, Mie Prefecture, Japan. Annals of Field Research and Technology Mie University 16: 1 - 32. [In Japanese.]" type="journal article" year="2018">
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Kimura
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<emphasis id="B90EEA89FF9F581C1F986676DABDFD68" box="[455,507,615,641]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
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. 2018
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</bibRefCitation>
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: table 23.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9F581C1EE266B9D9E3FC7B" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="8BC5369BFF9F581C1EE266B9DA70FD2B" blockId="9.[189,310,680,706]" box="[189,310,680,706]" pageId="9" pageNumber="10">
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<heading id="D08D81F7FF9F581C1EE266B9DA70FD2B" bold="true" box="[189,310,680,706]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="B90EEA89FF9F581C1EE266B9DA70FD2B" bold="true" box="[189,310,680,706]" pageId="9" pageNumber="10">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC5369BFF9F581C1EE266FBD9E3FC7B" blockId="9.[189,1399,746,914]" pageId="9" pageNumber="10">
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<taxonomicName id="4C7A4D18FF9F581C1EE266FBDA66FCED" authorityName="Pollock" authorityYear="1979" box="[189,288,746,772]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="9" pageNumber="10" phylum="Tardigrada" rank="genus">
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<emphasis id="B90EEA89FF9F581C1EE266FBDA66FCED" box="[189,288,746,772]" italics="true" pageId="9" pageNumber="10">Angursa</emphasis>
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</taxonomicName>
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with club-shaped primary clavae longer than lateral cirri; pedestals of primary clavae and lateral cirri absent; D-shaped secondary clavae; three-lobed tertiary clavae; leg I sensory organs as subdivided spines; legs II and III sensory organs absent; leg IV sensory organs each as papilla enveloped in hemispherical cuticular sheath with short, apical spine; anal papillae absent; seminal receptacles open latero-ventrally with short, sinuous ducts.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9F581C1EE267ABD891FBEB" box="[189,983,954,1027]" pageId="9" pageNumber="10" type="etymology">
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<paragraph id="8BC5369BFF9F581C1EE267ABDA02FC3D" blockId="9.[189,324,954,980]" box="[189,324,954,980]" pageId="9" pageNumber="10">
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<heading id="D08D81F7FF9F581C1EE267ABDA02FC3D" bold="true" box="[189,324,954,980]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="B90EEA89FF9F581C1EE267ABDA02FC3D" bold="true" box="[189,324,954,980]" pageId="9" pageNumber="10">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC5369BFF9F581C1EE267F9D891FBEB" blockId="9.[189,983,1000,1027]" box="[189,983,1000,1027]" pageId="9" pageNumber="10">
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The specific epithet refers to the T/V
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<emphasis id="B90EEA89FF9F581C1C2C67F8D9B1FBEA" box="[627,759,1001,1027]" italics="true" pageId="9" pageNumber="10">Seisuimaru</emphasis>
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of
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<collectingRegion id="49BEF879FF9F581C1D4167F9D80BFBEB" box="[798,845,1000,1026]" country="Japan" name="Mie" pageId="9" pageNumber="10">Mie</collectingRegion>
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University.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606510FF9F581C1EE2603BD934FB34" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BC5369BFF9F581C1EE2603BDA2AFBAD" blockId="9.[189,364,1066,1092]" box="[189,364,1066,1092]" pageId="9" pageNumber="10">
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<heading id="D08D81F7FF9F581C1EE2603BDA2AFBAD" bold="true" box="[189,364,1066,1092]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
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<emphasis id="B90EEA89FF9F581C1EE2603BDA2AFBAD" bold="true" box="[189,364,1066,1092]" pageId="9" pageNumber="10">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC5369BFF9F581C1EB56048DA1AFB9A" blockId="9.[189,1399,1113,1246]" box="[234,348,1113,1139]" pageId="9" pageNumber="10">
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<typeStatus id="54C18839FF9F581C1EB56048DA1AFB9A" box="[234,348,1113,1139]" pageId="9" pageNumber="10" type="holotype">
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<emphasis id="B90EEA89FF9F581C1EB56048DA1AFB9A" bold="true" box="[234,348,1113,1139]" pageId="9" pageNumber="10">Holotype</emphasis>
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</typeStatus>
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</paragraph>
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<paragraph id="8BC5369BFF9F581C1EE2606DD934FB34" blockId="9.[189,1399,1113,1246]" pageId="9" pageNumber="10">
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<materialsCitation id="3B123CC6FF9F581C1EE2606DD934FB34" ID-GBIF-Occurrence="2147483701" collectingDate="2017-11-10" collectionCode="KUZ" collectorName="Fujimoto" country="Japan" elevation="782" latitude="33.995" location="Northwestern Pacific" longLatPrecision="119" longitude="136.945" municipality="Sea of Kumano" pageId="9" pageNumber="10" specimenCode="Z2066" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
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<collectionCode id="ED6BAE5EFF9F581C1EE2606DDADBFB7F" box="[189,413,1148,1174]" pageId="9" pageNumber="10">PACIFIC OCEAN</collectionCode>
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• adult
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<specimenCount id="9D7CFD12FF9F581C1C54606CD964FB7E" box="[523,546,1149,1175]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
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;
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<location id="8EA56040FF9F581C1C68606DD87EFB7F" LSID="urn:lsid:plazi:treatment:03D3878DFF9F58191C3F65CFD9C5FCD1:8EA56040FF9F581C1C68606DD87EFB7F" box="[567,824,1148,1174]" country="Japan" latitude="33.995" longLatPrecision="119" longitude="136.945" municipality="Sea of Kumano" name="Northwestern Pacific" pageId="9" pageNumber="10">Northwestern Pacific</location>
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,
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<collectingCountry id="F36D760BFF9F581C1D16606CD8C9FB7E" box="[841,911,1149,1175]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
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,
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<collectingMunicipality id="6BA1ACE1FF9F581C1DFE606DDF2EFB7E" box="[929,1128,1148,1175]" pageId="9" pageNumber="10">Sea of Kumano</collectingMunicipality>
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; between
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<geoCoordinate id="EE4E505CFF9F581C1AB6606BDE34FB7E" box="[1257,1394,1146,1175]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="33.991665">33°59.5′ N</geoCoordinate>
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,
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<geoCoordinate id="EE4E505CFF9F581C1EE2608FDA0AFB53" box="[189,332,1182,1210]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="136.945">136°56.7′ E</geoCoordinate>
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and
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<geoCoordinate id="EE4E505CFF9F581C1FD060B1D953FB53" box="[399,533,1182,1210]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="33.995">33°59.7′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE4E505CFF9F581C1C79608FD9FEFB53" box="[550,696,1182,1210]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="136.945">136°56.7′ E</geoCoordinate>
|
|||
|
|
;
|
|||
|
|
<elevation id="0057D1A8FF9F581C1C96608EDF67FB50" box="[713,1057,1183,1210]" metricMagnitude="2" metricUnit="m" metricValue="7.82" metricValueMax="7.959999999999999" metricValueMin="7.68" pageId="9" pageNumber="10" unit="m" value="782.0" valueMax="796.0" valueMin="768.0">
|
|||
|
|
<quantity id="4C829B7EFF9F581C1C96608ED815FB53" box="[713,851,1183,1210]" metricMagnitude="2" metricUnit="m" metricValue="7.82" metricValueMax="7.959999999999999" metricValueMin="7.68" pageId="9" pageNumber="10" unit="m" value="782.0" valueMax="796.0" valueMin="768.0">768–796 m</quantity>
|
|||
|
|
below sea level
|
|||
|
|
</elevation>
|
|||
|
|
;
|
|||
|
|
<collectingDate id="EF80E9B3FF9F581C1A6E60B1DF9AFB53" box="[1073,1244,1184,1210]" pageId="9" pageNumber="10" value="2017-11-10">
|
|||
|
|
<date id="FFC4105BFF9F581C1A6E60B1DF9AFB53" box="[1073,1244,1184,1210]" pageId="9" pageNumber="10" value="2017-11-10">10 Nov. 2017</date>
|
|||
|
|
</collectingDate>
|
|||
|
|
; mounted in glycerol;
|
|||
|
|
<collectorName id="268F534DFF9F581C1F7260D2DADFFB34" box="[301,409,1219,1245]" pageId="9" pageNumber="10">Fujimoto</collectorName>
|
|||
|
|
leg.;
|
|||
|
|
<collectionCode id="ED6BAE5EFF9F581C1F8560D5D950FB37" box="[474,534,1220,1246]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="9" pageNumber="10">KUZ</collectionCode>
|
|||
|
|
<specimenCode id="DBDC9EE0FF9F581C1C4160D2D929FB34" box="[542,623,1219,1245]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="9" pageNumber="10">Z2066</specimenCode>
|
|||
|
|
.
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606510FF9F581C1EB56114DF47FAAB" pageId="9" pageNumber="10" type="description">
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EB56114DA1AFAF6" blockId="9.[189,1025,1285,1347]" box="[234,348,1285,1311]" pageId="9" pageNumber="10">
|
|||
|
|
<typeStatus id="54C18839FF9F581C1EB56114DA1AFAF6" box="[234,348,1285,1311]" pageId="9" pageNumber="10" type="paratype">
|
|||
|
|
<emphasis id="B90EEA89FF9F581C1EB56114DA1AFAF6" bold="true" box="[234,348,1285,1311]" pageId="9" pageNumber="10">Paratype</emphasis>
|
|||
|
|
</typeStatus>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EE26139DF47FAAB" blockId="9.[189,1025,1285,1347]" box="[189,1025,1320,1347]" pageId="9" pageNumber="10">
|
|||
|
|
PACIFIC OCEAN •
|
|||
|
|
<materialsCitation id="3B123CC6FF9F581C1FEF6139D8BBFAAB" ID-GBIF-Occurrence="2147483702" box="[432,1021,1320,1347]" collectingDate="2017-11-10" collectionCode="KUZ" collectorName="Fujimoto" country="Japan" elevation="782" latitude="33.995" location="Northwestern Pacific" longLatPrecision="119" longitude="136.945" municipality="Sea of Kumano" pageId="9" pageNumber="10" specimenCode="Z2067" specimenCount="2" specimenCount-adult="1" specimenCount-female="1" typeStatus="paratype">
|
|||
|
|
<specimenCount id="9D7CFD12FF9F581C1FEF6139D947FAAB" box="[432,513,1320,1346]" pageId="9" pageNumber="10" type="adult">1 adult</specimenCount>
|
|||
|
|
<specimenCount id="9D7CFD12FF9F581C1C576138D959FAAA" box="[520,543,1321,1347]" pageId="9" pageNumber="10" type="female">♀</specimenCount>
|
|||
|
|
; same data as for holotype;
|
|||
|
|
<collectionCode id="ED6BAE5EFF9F581C1D366138D8E3FAAA" box="[873,933,1321,1347]" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="9" pageNumber="10">KUZ</collectionCode>
|
|||
|
|
<specimenCode id="DBDC9EE0FF9F581C1DF36139D8BBFAAB" box="[940,1021,1320,1346]" collectionCode="KUZ" country="Japan" httpUri="http://grbio.org/cool/n11d-rfwg" name="Zoological Collection of the Kyoto University" pageId="9" pageNumber="10">Z2067</specimenCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606510FF9F581C1EE2617BD9A2FA3F" pageId="9" pageNumber="10" type="materials_examined">
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EE2617BDA1BFA6D" blockId="9.[189,349,1386,1412]" box="[189,349,1386,1412]" pageId="9" pageNumber="10">
|
|||
|
|
<heading id="D08D81F7FF9F581C1EE2617BDA1BFA6D" bold="true" box="[189,349,1386,1412]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
|||
|
|
<emphasis id="B90EEA89FF9F581C1EE2617BDA1BFA6D" bold="true" box="[189,349,1386,1412]" pageId="9" pageNumber="10">Type locality</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EE26188D9A2FA3F" blockId="9.[189,1398,1431,1495]" pageId="9" pageNumber="10">
|
|||
|
|
Sea of Kumano, Northwestern Pacific (between
|
|||
|
|
<geoCoordinate id="EE4E505CFF9F581C1CA56186D838FA5A" box="[762,894,1431,1459]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="33.991665">33°59.5′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE4E505CFF9F581C1DD46186DF51FA5A" box="[907,1047,1431,1459]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="136.945">136°56.7′ E</geoCoordinate>
|
|||
|
|
and
|
|||
|
|
<geoCoordinate id="EE4E505CFF9F581C1A0D6186DF93FA5A" box="[1106,1237,1431,1459]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="92" value="33.995">33°59.7′ N</geoCoordinate>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="EE4E505CFF9F581C1ABD6186DE37FA5A" box="[1250,1393,1431,1459]" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="92" value="136.945">136°56.7′ E</geoCoordinate>
|
|||
|
|
; depth range,
|
|||
|
|
<quantity id="4C829B7EFF9F581C1F0A61ADDA9DFA3E" box="[341,475,1468,1495]" metricMagnitude="2" metricUnit="m" metricValue="7.82" metricValueMax="7.959999999999999" metricValueMin="7.68" pageId="9" pageNumber="10" unit="m" value="782.0" valueMax="796.0" valueMin="768.0">768–796 m</quantity>
|
|||
|
|
) (
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9F581C1FAC61ADD99EFA3F" author="Kimura T. & Kimura S. & Jimi N. & Kakui K. & Tomioka S. & Oya Y. & Matsumoto Y. & Tanabe Y. & Hasegawa N. & Hookabe N. & Homma R. & Hosoda Y. & Fujimoto S. & Kuramochi T. & Fujita T. & Ogawa A. & Kobayashi I. & Ishida Y. & Tanaka H. & Onishi H. & Shimetsugu M. & Yoshikawa A. & Tanaka M. & Kushida Y. & Maekawa Y. & Nakamura T. & Okumura J. & Tanaka K." box="[499,728,1468,1494]" pageId="9" pageNumber="10" pagination="1 - 32" refId="ref8854" refString="Kimura T., Kimura S., Jimi N., Kakui K., Tomioka S., Oya Y., Matsumoto Y., Tanabe Y., Hasegawa N., Hookabe N., Homma R., Hosoda Y., Fujimoto S., Kuramochi T., Fujita T., Ogawa A., Kobayashi I., Ishida Y., Tanaka H., Onishi H., Shimetsugu M., Yoshikawa A., Tanaka M., Kushida Y., Maekawa Y., Nakamura T., Okumura J. & Tanaka K. 2018. Benthic deep-sea fauna in the Sea of Kumano, Mie Prefecture, Japan. Annals of Field Research and Technology Mie University 16: 1 - 32. [In Japanese.]" type="journal article" year="2018">
|
|||
|
|
Kimura
|
|||
|
|
<emphasis id="B90EEA89FF9F581C1C0961ACD9D7FA3F" box="[598,657,1468,1494]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
|||
|
|
2018
|
|||
|
|
</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606510FF9F58191EE261EFDFD2FDE4" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EE261EFDAA7F9F1" blockId="9.[189,481,1534,1560]" box="[189,481,1534,1560]" pageId="9" pageNumber="10">
|
|||
|
|
<heading id="D08D81F7FF9F581C1EE261EFDAA7F9F1" bold="true" box="[189,481,1534,1560]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
|||
|
|
<emphasis id="B90EEA89FF9F581C1EE261EFDAA7F9F1" bold="true" box="[189,481,1534,1560]" pageId="9" pageNumber="10">
|
|||
|
|
Description of
|
|||
|
|
<typeStatus id="54C18839FF9F581C1F2A61EFDAA7F9F1" box="[373,481,1534,1560]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9F581C1EE2623DD971F983" blockId="9.[189,1398,1580,1642]" pageId="9" pageNumber="10">
|
|||
|
|
Adult female with unarmoured, slender body (143 μm long; 35 μm wide between legs II and III) (
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1B79623DDE37F9AF" box="[1318,1393,1580,1606]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="9" pageNumber="10">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1EE26241DBA3F983" box="[189,229,1616,1642]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="9" pageNumber="10">7A</figureCitation>
|
|||
|
|
). Epicuticular pillars visible.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9F581F1EE26283DE50FE85" blockId="9.[189,1399,1682,2027]" lastBlockId="10.[189,1399,267,364]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
|||
|
|
Cephalic region with unpaired median cirrus, paired internal cirri, paired external cirri, paired lateral cirri, paired primary clavae, paired secondary clavae, paired tertiary clavae, and mouth opening. Median cirrus (1 μm) positioned dorsally. Internal cirri (5 μm) inserted on anterior margin of body each consisting of scapus (2 μm) and distal flagellar part with blunt tip (2 μm). External cirri (9 μm) inserted ventrally each consisting of scapus (5 μm) and distal flagellar part with blunt tip (5 μm). Lateral cirri (4 μm) positioned dorso-laterally. Primary clavae (10 μm) positioned immediately ventral to lateral cirri and both organs inserted directly on body with no apparent pedestal (only very slight rise). Primary clavae club-shaped and distal half slightly bent posteriorly. Secondary clavae (
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1A44639BDF2CF84D" box="[1051,1130,1930,1956]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="9" pageNumber="10">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1A26639BDFDAF84D" box="[1145,1180,1930,1956]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="9" pageNumber="10">7B</figureCitation>
|
|||
|
|
) present posterior to median cirrus as flat, D-shaped structures without epicuticular pillars. Tertiary clavae (
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1AB263BCDE7AF82E" box="[1261,1340,1965,1991]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="9" pageNumber="10">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9F581C1B1563BCDE28F82E" box="[1354,1390,1965,1991]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="9" pageNumber="10">7C</figureCitation>
|
|||
|
|
) indented around external cirri, giving impression of three-lobed structures (one lobe directed anteriorly, interior to external cirri; second lobe developed interior to first lobe; third lobe developed posterior to external cirri) without epicuticular pillars. Anteriorly directed mouth at antero-most position of body. Buccal apparatus dissolved and only pharyngeal bulb recognised (length 14 μm, width 11 μm).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9C581F1EE26582D917FE38" blockId="10.[189,1399,403,465]" pageId="10" pageNumber="11">
|
|||
|
|
Paired cirri E (18 μm) (
|
|||
|
|
<figureCitation id="13412A1EFF9C581F1F906582D95AFE44" box="[463,540,403,429]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="10" pageNumber="11">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9C581F1C776585D908FE47" box="[552,590,404,430]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="10" pageNumber="11">7D</figureCitation>
|
|||
|
|
) as simple, tapering spines without recognisable annulation, inserted dorsally slightly anterior to leg IV.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF056613FF9C581F1EE263C1D8FDF805" box="[189,955,2000,2028]" pageId="10" pageNumber="11">
|
|||
|
|
<paragraph id="8BC5369BFF9C581F1EE263C1D8FDF805" blockId="10.[189,955,2000,2028]" box="[189,955,2000,2028]" pageId="10" pageNumber="11">
|
|||
|
|
<emphasis id="B90EEA89FF9C581F1EE263C1DA4CF802" bold="true" box="[189,266,2000,2027]" pageId="10" pageNumber="11">Fig. 6.</emphasis>
|
|||
|
|
Drawing of
|
|||
|
|
<taxonomicName id="4C7A4D18FF9C581F1FC063C0D9E0F802" authority="Fujimoto & Hansen, 2019" authorityName="Fujimoto & Hansen" authorityYear="2019" box="[415,678,2001,2027]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="10" pageNumber="11" phylum="Tardigrada" rank="species" species="seisuimaruae" status="sp. nov.">
|
|||
|
|
<emphasis id="B90EEA89FF9C581F1FC063C0D9E0F802" box="[415,678,2001,2027]" italics="true" pageId="10" pageNumber="11">Angursa seisuimaruae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23D57F2FF9C581F1CF163C0D84FF802" box="[686,777,2001,2027]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(KUZ Z2066).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC5369BFF9D581E1EE2651ADE58FDF5" blockId="11.[189,1399,267,540]" pageId="11" pageNumber="12">
|
|||
|
|
Each leg consists of indistinct coxa, femur, indistinct tibia, and tarsus bearing four digits terminating in claws with two divergent points. Leg I sensory organ (10 μm) (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1DB1653FDF78FEA1" box="[1006,1086,302,328]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1A10653EDF36FEA0" box="[1103,1136,303,329]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7E</figureCitation>
|
|||
|
|
) present on femur as spine consisting of proximal portion and distal portion with blunt tip. Legs II and III lack sensory organs. Leg IV sensory organs (4 μm) (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1CD06564D99AFE66" box="[655,732,373,399]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1CB66564D848FE66" box="[745,782,373,399]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7D</figureCitation>
|
|||
|
|
) present at base of leg IV each as papilla enveloped in hemispherical cuticular sheath with short, apical spine. Internal digits have proximal pads and lack peduncles (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1F1A65AADAD2FE3C" box="[325,404,443,469]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1FFC65ADDA81FE3F" box="[419,455,444,470]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7E</figureCitation>
|
|||
|
|
). External digits have arched peduncles at their base which never reach claws (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1E9A65CFDA54FE11" box="[197,274,478,504]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1F4165CEDA04FE10" box="[286,322,479,505]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7E</figureCitation>
|
|||
|
|
). Internal digits always longer than external digits and leg IV has longer digits than those of legs I–III (leg I, internal digits 9 μm/external digits 6 μm; leg II, 9/6; leg III, 10/6; leg IV, 13/7).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9D581E1EE26652DFFAFD2E" blockId="11.[189,1399,579,711]" pageId="11" pageNumber="12">
|
|||
|
|
Rosette-like gonopore (4 μm in diameter) (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1C956652D85CFDB4" box="[714,794,579,605]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1D756655D80CFDB7" box="[810,842,580,606]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7F</figureCitation>
|
|||
|
|
) situated 2 μm anterior to anus. Longitudinal cuticluar fold of anus reach dorsal side (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1CCA6677D9A4FD69" box="[661,738,614,640]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1CB16676D850FD68" box="[750,790,615,641]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7D</figureCitation>
|
|||
|
|
). Paired seminal receptacles (
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1A2C6677DFF9FD69" box="[1139,1215,614,640]" captionStart="Fig" captionStartId="10.[189,232,2000,2026]" captionText="Fig. 6. Drawing of Angursa seisuimaruae sp. nov. (KUZ Z2066)." pageId="11" pageNumber="12">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="13412A1EFF9D581E1A936676DFAAFD68" box="[1228,1260,615,641]" captionStart="Fig" captionStartId="11.[189,232,1923,1949]" captionTargetBox="[238,1344,814,1876]" captionTargetId="figure@11.[238,1352,814,1876]" captionTargetPageId="11" captionText="Fig. 7. Micrographs of Angursa seisuimaruae sp. nov. (KUZ Z2066). A. Habitus. B. Cephalic region (dorsal). C. Cephalic region (ventral). D. Caudal region (dorsal). E. Leg I. F. Caudal region (ventral)." httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12">7F</figureCitation>
|
|||
|
|
) positioned near latero-ventral margin of body, at level slightly anterior to gonopore. Each seminal receptacle duct runs interiorly, antero-dorsally, and then postero-ventrally terminating in round vesicle.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF056613FF9D581E1EE26392DE36F828" httpUri="https://zenodo.org/record/2616390/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1923,1949]" targetBox="[238,1344,814,1876]" targetPageId="11">
|
|||
|
|
<paragraph id="8BC5369BFF9D581E1EE26392DE36F828" blockId="11.[189,1399,1923,1985]" pageId="11" pageNumber="12">
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1EE26392DA4AF874" bold="true" box="[189,268,1923,1949]" pageId="11" pageNumber="12">Fig. 7.</emphasis>
|
|||
|
|
Micrographs of
|
|||
|
|
<taxonomicName id="4C7A4D18FF9D581E1F896392D9A7F874" authority="Fujimoto & Hansen, 2019" authorityName="Fujimoto & Hansen" authorityYear="2019" box="[470,737,1923,1949]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="11" pageNumber="12" phylum="Tardigrada" rank="species" species="seisuimaruae" status="sp. nov.">
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1F896392D9A7F874" box="[470,737,1923,1949]" italics="true" pageId="11" pageNumber="12">Angursa seisuimaruae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23D57F2FF9D581E1CB66395D800F877" box="[745,838,1924,1950]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
(KUZ Z2066).
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1A5B6392DF5FF874" bold="true" box="[1028,1049,1923,1949]" pageId="11" pageNumber="12">A</emphasis>
|
|||
|
|
. Habitus.
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1ACA6392DFEFF874" bold="true" box="[1173,1193,1923,1949]" pageId="11" pageNumber="12">B</emphasis>
|
|||
|
|
. Cephalic region (dorsal).
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1F7763B7DA7BF829" bold="true" box="[296,317,1958,1984]" pageId="11" pageNumber="12">C</emphasis>
|
|||
|
|
. Cephalic region (ventral).
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1CDA63B6D9DCF828" bold="true" box="[645,666,1959,1985]" pageId="11" pageNumber="12">D</emphasis>
|
|||
|
|
. Caudal region (dorsal).
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1D9D63B6D890F828" bold="true" box="[962,982,1959,1985]" pageId="11" pageNumber="12">E</emphasis>
|
|||
|
|
. Leg I.
|
|||
|
|
<emphasis id="B90EEA89FF9D581E1A6C63B6DF03F828" bold="true" box="[1075,1093,1959,1985]" pageId="11" pageNumber="12">F</emphasis>
|
|||
|
|
. Caudal region (ventral).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BC5369BFF9A58191EE2651ADAA3FECC" blockId="12.[189,485,267,293]" box="[189,485,267,293]" pageId="12" pageNumber="13">
|
|||
|
|
<heading id="D08D81F7FF9A58191EE2651ADAA3FECC" bold="true" box="[189,485,267,293]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191EE2651ADAA3FECC" bold="true" box="[189,485,267,293]" pageId="12" pageNumber="13">
|
|||
|
|
Description of
|
|||
|
|
<typeStatus id="54C18839FF9A58191F2A651ADAA3FECC" box="[373,485,267,293]" pageId="12" pageNumber="13" type="paratype">paratype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC5369BFF9A58191EE2652ADFD2FDE4" blockId="12.[189,1399,315,525]" pageId="12" pageNumber="13">
|
|||
|
|
Cephalic region accidentally destroyed during cleaning of specimen before observation and mouth opening, buccal apparatus, median cirrus, secondary clavae, and tertiary clavae not recognized. Primary clava (11 μm) and scapi of internal, external, and lateral cirri recognized on torn cuticle. Presence of scapi suggests subdivision of lateral cirri as in the internal and external cirri. Gonopore and anus recognised before permanent preparation in lateral positioned body. Other characters confirm that of
|
|||
|
|
<typeStatus id="54C18839FF9A58191EE265E3DA62FDE5" box="[189,292,498,524]" pageId="12" pageNumber="13" type="holotype">holotype</typeStatus>
|
|||
|
|
(cirrus E, 17 μm; leg I sensory organ, 10 μm; leg IV sensory organ, 4 μm).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606510FF9A58191EE26627DA69FDB9" box="[189,303,566,592]" pageId="12" pageNumber="13" type="discussion">
|
|||
|
|
<paragraph id="8BC5369BFF9A58191EE26627DA69FDB9" blockId="12.[189,303,566,592]" box="[189,303,566,592]" pageId="12" pageNumber="13">
|
|||
|
|
<heading id="D08D81F7FF9A58191EE26627DA69FDB9" bold="true" box="[189,303,566,592]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191EE26627DA69FDB9" bold="true" box="[189,303,566,592]" pageId="12" pageNumber="13">Remarks</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3606510FF9A58191EE26677D9C5FCD1" pageId="12" pageNumber="13" type="nomenclature">
|
|||
|
|
<paragraph id="8BC5369BFF9A58191EE26677D9C5FCD1" blockId="12.[189,1399,613,824]" pageId="12" pageNumber="13">
|
|||
|
|
<taxonomicName id="4C7A4D18FF9A58191EE26677DA82FD96" authority="Fujimoto & Hansen, 2019" authorityName="Fujimoto & Hansen" authorityYear="2019" box="[189,452,614,640]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="seisuimaruae" status="sp. nov.">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191EE26677DA82FD96" box="[189,452,614,640]" italics="true" pageId="12" pageNumber="13">Angursa seisuimaruae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23D57F2FF9A58191F926677D96EFD69" box="[461,552,614,640]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
resembles
|
|||
|
|
<taxonomicName id="4C7A4D18FF9A58191CF26677D87CFD96" authorityName="Renaud-Mornant" authorityYear="1981" box="[685,826,613,640]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="abyssalis">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191CF26677D87CFD96" box="[685,826,613,640]" italics="true" pageId="12" pageNumber="13">A. abyssalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C7A4D18FF9A58191D176677D894FD96" authorityName="Noda" authorityYear="1985" box="[840,978,613,640]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="clavifera">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191D176677D894FD96" box="[840,978,613,640]" italics="true" pageId="12" pageNumber="13">A. clavifera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C7A4D18FF9A58191A4C6677DFD6FD96" authorityName="Bussau" authorityYear="1992" box="[1043,1168,613,640]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="capsula">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191A4C6677DFD6FD96" box="[1043,1168,613,640]" italics="true" pageId="12" pageNumber="13">A. capsula</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by the club-shaped primary clavae longer than the lateral cirri, the leg IV sensory organs in hemispherical shape, and the lack of the anal papillae (
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191FB966BED9B0FD20" author="Renaud-Mornant J." box="[486,758,687,713]" pageId="12" pageNumber="13" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant 1981</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191D5C66BED8CEFD23" author="Noda H." box="[771,904,687,714]" pageId="12" pageNumber="13" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda 1985</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191DCB66BEDF76FD20" author="Bussau C." box="[916,1072,687,713]" pageId="12" pageNumber="13" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
|||
|
|
). However, the new species is distinguished from all three species by its D-shaped secondary clavae and its three-lobed tertiary clavae (
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191F4766E9DAD9FCFA" author="Noda H." box="[280,415,760,787]" pageId="12" pageNumber="13" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda 1985</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191FF366E8D90EFCFB" author="Bussau C." box="[428,584,760,787]" pageId="12" pageNumber="13" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau 1992</bibRefCitation>
|
|||
|
|
). Further, the new species’s seminal receptacle ducts are shorter than
|
|||
|
|
<taxonomicName id="4C7A4D18FF9A58191EE2670CDA00FCDE" authorityName="Noda" authorityYear="1985" box="[189,326,797,823]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="clavifera">
|
|||
|
|
<emphasis id="B90EEA89FF9A58191EE2670CDA00FCDE" box="[189,326,797,823]" italics="true" pageId="12" pageNumber="13">A. clavifera</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
’s spiral ones (
|
|||
|
|
<bibRefCitation id="EFEB4B6AFF9A58191FB1670CD930FCD1" author="Noda H." box="[494,630,797,824]" pageId="12" pageNumber="13" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda 1985</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|