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<document id="9D8A11B1BAFCCAF22CC4C0037AC37330" ID-CLB-Dataset="29761" ID-DOI="10.11646/zootaxa.4442.2.8" ID-GBIF-Dataset="d66ed56d-eec7-424a-a7c9-9f2c884269e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303179" ID-ZooBank="A822F33A-0E10-46A4-8630-6944D4A00307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530533711971" checkinUser="plazi" docAuthor="Kopylov, D. S., Spasojevic, T. &amp; Klopfstein, S." docDate="2018" docId="03D9905AFFD73B5D19EC78ADFBCDFB94" docLanguage="en" docName="zootaxa.4442.2.8.pdf" docOrigin="Zootaxa 4442 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crusopimpla Kopylov, Spasojevic &amp; Klopfstein 2018, gen. nov." docType="treatment" docUuid="FBD4FD16-5EED-4D2F-9DDF-A55F7BFC0527" docUuidSource="ZooBank" docVersion="8" lastPageNumber="323" masterDocId="FFE0E822FFD33B59197B7901FFFFFFAB" masterDocTitle="New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East" masterLastPageNumber="330" masterPageNumber="319" pageNumber="323" updateTime="1698552165635" updateUser="ExternalLinkService">
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<mods:title id="0F0485ADE8DADD440490F7A46C49399D">New ichneumonids (Hymenoptera, Ichneumonidae) from the Eocene Tadushi Formation, Russian Far East</mods:title>
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<mods:namePart id="B1C95954495D0B3A5A181586480F909B">Spasojevic, T.</mods:namePart>
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<mods:namePart id="B2668C691F92545766B904C706DACC18">Klopfstein, S.</mods:namePart>
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<treatment id="03D9905AFFD73B5D19EC78ADFBCDFB94" ID-DOI="http://doi.org/10.5281/zenodo.5973708" ID-GBIF-Taxon="146044110" ID-Zenodo-Dep="5973708" LSID="urn:lsid:zoobank.org:act:FBD4FD16-5EED-4D2F-9DDF-A55F7BFC0527" httpUri="http://treatment.plazi.org/id/03D9905AFFD73B5D19EC78ADFBCDFB94" lastPageNumber="323" pageId="4" pageNumber="323">
<subSubSection id="C36A72C7FFD73B5D19EC78ADFC4DFE6D" box="[151,946,428,454]" pageId="4" pageNumber="323" type="nomenclature">
<paragraph id="8BCF214CFFD73B5D19EC78ADFC4DFE6D" blockId="4.[151,946,428,454]" box="[151,946,428,454]" pageId="4" pageNumber="323">
<heading id="D0879620FFD73B5D19EC78ADFC4DFE6D" bold="true" box="[151,946,428,454]" fontSize="11" level="1" pageId="4" pageNumber="323" reason="1">
<emphasis id="B904FD5EFFD73B5D19EC78ADFC4DFE6D" bold="true" box="[151,946,428,454]" pageId="4" pageNumber="323">
Genus
<taxonomicName id="4C705ACFFFD73B5D199578ADFCC8FE6D" authority="Kopylov, Spasojevic &amp; Klopfstein" authorityName="Kopylov, Spasojevic &amp; Klopfstein" authorityYear="2018" box="[238,823,428,454]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B904FD5EFFD73B5D199578ADFE74FE6D" bold="true" box="[238,395,428,454]" italics="true" pageId="4" pageNumber="323">Crusopimpla</emphasis>
Kopylov, Spasojevic &amp; Klopfstein
</taxonomicName>
,
<taxonomicNameLabel id="A2374025FFD73B5D1A3F78ADFC4DFE6D" box="[836,946,428,454]" pageId="4" pageNumber="323" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A72C7FFD73B5D19EC78EEFC16FD82" pageId="4" pageNumber="323" type="materials_examined">
<paragraph id="8BCF214CFFD73B5D19EC78EEFBBAFDA3" blockId="4.[151,1437,495,1087]" box="[151,1093,495,520]" pageId="4" pageNumber="323">
<emphasis id="B904FD5EFFD73B5D19EC78EEFED4FDA3" bold="true" box="[151,299,495,520]" pageId="4" pageNumber="323">Type species</emphasis>
.
<taxonomicName id="4C705ACFFFD73B5D184278EEFC1FFDA3" authority="Kopylov, Spasojevic &amp; Klopfstein" authorityName="Kopylov &amp; Spasojevic &amp; Klopfstein" authorityYear="2018" box="[313,992,495,520]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="tadushensis" status="sp. nov.">
<emphasis id="B904FD5EFFD73B5D184278EEFDAAFDA3" box="[313,597,495,520]" italics="true" pageId="4" pageNumber="323">Crusopimpla tadushensis</emphasis>
Kopylov, Spasojevic &amp; Klopfstein
</taxonomicName>
,
<taxonomicNameLabel id="A2374025FFD73B5D1A9078EEFBBAFDA3" box="[1003,1093,495,520]" pageId="4" pageNumber="323" rank="species">
<emphasis id="B904FD5EFFD73B5D1A9078EEFBBAFDA3" bold="true" box="[1003,1093,495,520]" pageId="4" pageNumber="323">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BCF214CFFD73B5D19BC7B10FC16FD82" blockId="4.[151,1437,495,1087]" box="[199,1001,529,554]" pageId="4" pageNumber="323">
<emphasis id="B904FD5EFFD73B5D19BC7B10FEA0FD81" bold="true" box="[199,351,529,554]" pageId="4" pageNumber="323">Composition</emphasis>
.
<taxonomicName id="4C705ACFFFD73B5D18167B10FCD2FD81" authority=", Crusopimpla" authorityName="Crusopimpla" box="[365,813,529,554]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="tadushensis">
<emphasis id="B904FD5EFFD73B5D18167B10FD76FD81" box="[365,649,529,554]" italics="true" pageId="4" pageNumber="323">Crusopimpla tadushensis</emphasis>
,
<emphasis id="B904FD5EFFD73B5D1BEC7B10FCD2FD81" box="[663,813,529,554]" italics="true" pageId="4" pageNumber="323">Crusopimpla</emphasis>
</taxonomicName>
<emphasis id="B904FD5EFFD73B5D1A567B10FCC8FD81" box="[813,823,529,554]" italics="true" pageId="4" pageNumber="323">?</emphasis>
<taxonomicName id="4C705ACFFFD73B5D1A457B10FC16FD82" authority="Brues." authorityName=": Brues" authorityYear="1910" box="[830,1001,529,554]" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="rediviva">
<emphasis id="B904FD5EFFD73B5D1A457B10FC66FD81" box="[830,921,529,554]" italics="true" pageId="4" pageNumber="323">rediviva</emphasis>
Brues.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36A72C7FFD73B5D19BC7B33FA62FD7B" pageId="4" pageNumber="323" type="diagnosis">
<paragraph id="8BCF214CFFD73B5D19BC7B33FA62FD7B" blockId="4.[151,1437,495,1087]" pageId="4" pageNumber="323">
<emphasis id="B904FD5EFFD73B5D19BC7B33FEC5FDE0" bold="true" box="[199,314,562,587]" pageId="4" pageNumber="323">Diagnosis</emphasis>
. Propodeal carinae well developed, enclosing basal area, areola, petiolar area, and first, second and third lateral areas. Fore wing with pterostigma rather wide, about 3× as long as wide; areolet tetragonal with uneven sides, r-m and 2+3-M long, 4-M short and 3Rs absent; ramulus short or absent. Metasomal tergite 1 wider than long, with strong dorsal carinae converging on basal and almost parallel on apical half. Metasomal tergites 2 and following wider than long. Ovipositor robust, about 0.3–0.8 times as long as metasoma. Hind femur thickened.
</paragraph>
</subSubSection>
<subSubSection id="C36A72C7FFD73B5D19BC7BD8FE19FCB8" pageId="4" pageNumber="323" type="etymology">
<paragraph id="8BCF214CFFD73B5D19BC7BD8FE19FCB8" blockId="4.[151,1437,495,1087]" pageId="4" pageNumber="323">
<emphasis id="B904FD5EFFD73B5D19BC7BD8FEB9FD59" bold="true" box="[199,326,729,754]" pageId="4" pageNumber="323">Etymology</emphasis>
. From the
<emphasis id="B904FD5EFFD73B5D18BD7BDBFDFDFD5A" box="[454,514,730,753]" italics="true" pageId="4" pageNumber="323">Latin</emphasis>
word “crus”, which means stem, and the genus name
<taxonomicName id="4C705ACFFFD73B5D1D0B7BD8FB3EFD59" box="[1136,1217,729,754]" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D1D0B7BD8FB3EFD59" box="[1136,1217,729,754]" italics="true" pageId="4" pageNumber="323">Pimpla</emphasis>
</taxonomicName>
, to reflect its basal position within the subfamily.
</paragraph>
</subSubSection>
<subSubSection id="C36A72C7FFD73B5D19BC7A1DFBCDFB94" pageId="4" pageNumber="323" type="discussion">
<paragraph id="8BCF214CFFD73B5D19BC7A1DFBCDFB94" blockId="4.[151,1437,495,1087]" pageId="4" pageNumber="323">
<emphasis id="B904FD5EFFD73B5D19BC7A1DFECBFC9E" bold="true" box="[199,308,796,821]" pageId="4" pageNumber="323">Remarks</emphasis>
. The genus
<emphasis id="B904FD5EFFD73B5D18BA7A1DFDACFC9E" box="[449,595,796,821]" italics="true" pageId="4" pageNumber="323">Crusopimpla</emphasis>
is similar in general appearance to several extant genera in the subfamily
<taxonomicName id="4C705ACFFFD73B5D19EC7A3CFEF3FCFD" box="[151,268,829,854]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="subFamily" subFamily="Pimplinae">Pimplinae</taxonomicName>
, especially among the
<emphasis id="B904FD5EFFD73B5D1B5F7A3CFCE1FCFD" box="[548,798,829,854]" italics="true" pageId="4" pageNumber="323">
<taxonomicName id="4C705ACFFFD73B5D1B5F7A3CFD44FCFD" authorityName="Momoi" authorityYear="1965" box="[548,699,829,854]" class="Insecta" family="Ichneumonidae" genus="Gregopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">Gregopimpla</taxonomicName>
,
<taxonomicName id="4C705ACFFFD73B5D1BB67A3CFCE1FCFD" box="[717,798,829,854]" class="Insecta" family="Ichneumonidae" genus="Pimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">Pimpla</taxonomicName>
</emphasis>
and
<taxonomicName id="4C705ACFFFD73B5D1A1A7A3CFC3BFCFD" box="[865,964,829,854]" class="Insecta" family="Ichneumonidae" genus="Scambus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D1A1A7A3CFC3BFCFD" box="[865,964,829,854]" italics="true" pageId="4" pageNumber="323">Scambus</emphasis>
</taxonomicName>
genus groups. However, the propodeal carination is always reduced in these groups (
<figureCitation id="134B3DC9FFD73B5D1BE37A5FFD26FCDC" box="[664,729,862,887]" captionStart="FIGURE 1" captionStartId="2.[151,250,1688,1710]" captionTargetBox="[161,1435,198,1667]" captionTargetId="figure@2.[151,1436,195,1667]" captionTargetPageId="2" captionText="FIGURE 1. Phylogeny of Pimplinae and outgroups, redrawn according to Gauld et al. (2002), with examples of propodea, first metasomal tergites and hind wings shown to the left of the genus names. The drawings are modified from Townes (1969), and the species representatives are, from top to bottom: Xorides irrigator Fabricius, Odontocolon ochropus Townes, Poemenia hectica Gravenhorst, Rhyssa persuasoria Linnaeus, Theronia atalantae Poda, Lissopimpla excelsa Costa, Xanthopimpla hova Saussure, Echthromorpha agrestoria Swederus, Itoplectis maculator Fabricius, Pimpla madecassa Saussure, Zatypota percontatoria Müller, Dreisbachia mira Tosquinet, Camptotypus pulchripennis Saussure, Scambus tecumseh Viereck, Flavopimpla nava Tosquinet, Delomerista novita Cresson, Atractogaster semisculptus Kriechbaumer, and Perithous albicinctus Gravenhorst." httpUri="https://zenodo.org/record/1303181/files/figure.png" pageId="4" pageNumber="323">Fig. 1</figureCitation>
). The only extant genera in the subfamily which have a nearly full set of propodeal carinae at least in a few species are
<taxonomicName id="4C705ACFFFD73B5D1A6A7A7EFC88FC33" authorityName="Holmgren" authorityYear="1859" box="[785,887,895,920]" class="Insecta" family="Ichneumonidae" genus="Theronia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D1A6A7A7EFC88FC33" box="[785,887,895,920]" italics="true" pageId="4" pageNumber="323">Theronia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C705ACFFFD73B5D1AD67A7EFBB3FC33" authorityName="Saussure" authorityYear="1892" box="[941,1100,895,920]" class="Insecta" family="Ichneumonidae" genus="Xanthopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D1AD67A7EFBB3FC33" box="[941,1100,895,920]" italics="true" pageId="4" pageNumber="323">Xanthopimpla</emphasis>
</taxonomicName>
from the tribe
<taxonomicName id="4C705ACFFFD73B5D1D8F7A7EFAA7FC33" box="[1268,1368,895,920]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="tribe" tribe="Pimplini">Pimplini</taxonomicName>
. They both have a much more elongate pterostigma and radial cell in the fore wing and the ground color of the body is orange or yellow, while
<taxonomicName id="4C705ACFFFD73B5D18D57AC2FDABFC70" box="[430,596,962,987]" class="Insecta" family="Ichneumonidae" genus="Crusopimpla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="tadushensis">
<emphasis id="B904FD5EFFD73B5D18D57AC2FDABFC70" box="[430,596,962,987]" italics="true" pageId="4" pageNumber="323">C. tadushensis</emphasis>
</taxonomicName>
is dark-colored, and basal area of propodeum not developed. The extinct genus
<taxonomicName id="4C705ACFFFD73B5D19A47AE5FE5BFC57" authority="Brown" authorityName="Brown" authorityYear="1986" box="[223,420,996,1021]" class="Insecta" family="Ichneumonidae" genus="Lithoserix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D19A47AE5FEAEFC56" box="[223,337,996,1021]" italics="true" pageId="4" pageNumber="323">Lithoserix</emphasis>
Brown
</taxonomicName>
also has a similar body shape and the state of its propodeal carinae is somewhat uncertain from the single known fossil; however,
<taxonomicName id="4C705ACFFFD73B5D1B2F7D04FD39FBB5" authorityName="Brown" authorityYear="1986" box="[596,710,1029,1054]" class="Insecta" family="Ichneumonidae" genus="Lithoserix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B904FD5EFFD73B5D1B2F7D04FD39FBB5" box="[596,710,1029,1054]" italics="true" pageId="4" pageNumber="323">Lithoserix</emphasis>
</taxonomicName>
also has an elongate pterostigma and radial cell, and possesses a long ramulus in the discocubital cell (
<bibRefCitation id="EFE15CBDFFD73B5D1B477D26FD33FB94" box="[572,716,1062,1087]" pageId="4" pageNumber="323" refId="ref5040" refString="Brown, M. (1986) Lithosirex williamsi (Siricidae: Hymenoptera) a newly recognized fossil horntail from Florissant, Colorado. Insecta Mundi, 1, 119 - 120." type="journal article">Brown 1986</bibRefCitation>
;
<bibRefCitation id="EFE15CBDFFD73B5D1BA37D26FBD9FB94" box="[728,1062,1062,1087]" pageId="4" pageNumber="323" refId="ref5561" refString="Kasparyan, D. R. &amp; Rasnitsyn, A. P. (1992) The systematic position of the Oligocene Lithoserix williamsi Brown (Hymenoptera, Ichneumonidae) from Colorado (USA). Paleontological Journal, 26, 134 - 135." type="journal article">Kasparyan &amp; Rasnitsyn 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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