674 lines
123 KiB
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674 lines
123 KiB
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<document id="511AC409A4042603F2FE83F990D0C4F6" ID-DOI="10.11646/zootaxa.5318.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="8158357" ID-ZooBank="161E8842-5DB1-40CA-A4B7-2287462D86E1" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1689663496356" checkinUser="plazi" docAuthor="Bennett, Robb, Copley, Claudia & Copley, Darren" docDate="2023" docId="03BF87ABE428071409BEF00A60B5F98B" docLanguage="en" docName="zootaxa.5318.1.5.pdf" docOrigin="Zootaxa 5318 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5318.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybaeina Chamberlin and Ivie" docType="treatment" docVersion="1" lastPageNumber="100" masterDocId="FF86FFD3E42907170929F7706528FFC9" masterDocTitle="Revision of the western Nearctic spider genus Cybaeina including the description of Neocybaeina gen. nov. and Rothaeina gen. nov. (Araneae: Cybaeidae: Cybaeinae)" masterLastPageNumber="129" masterPageNumber="97" pageNumber="98" updateTime="1689879812679" updateUser="carolina" zenodo-license-document="CLOSED">
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<mods:titleInfo id="13C6F20B6545B09360CD5F09A628257D">
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<mods:title id="D34645C5BAFB41C73389EBA72DF60EF5">Revision of the western Nearctic spider genus Cybaeina including the description of Neocybaeina gen. nov. and Rothaeina gen. nov. (Araneae: Cybaeidae: Cybaeinae)</mods:title>
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<mods:namePart id="DC02731B58E9276551B1730A5FE86845">Bennett, Robb</mods:namePart>
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<mods:nameIdentifier id="AD45696726B938E5D1738C491056903D" type="ORCID">0000-0002-6587-7079</mods:nameIdentifier>
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<mods:affiliation id="528963CBFBF4257378635A92DCD2FF02">robb. bennett @ shaw. ca; https: // orcid. org / 0000 - 0002 - 6587 - 7079</mods:affiliation>
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<mods:nameIdentifier id="1E1CFCF95D864733431381035E960340" type="email">robb.bennett@shaw.ca</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="E1D36545291EAAB02C52856DEA93E5C0">Copley, Claudia</mods:namePart>
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<mods:nameIdentifier id="74A8DE300A44DC533DE8BC48B1AC190B" type="ORCID">0000-0002-8184-5819</mods:nameIdentifier>
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<mods:affiliation id="528B991CD40B894DBDAF81038572C675">ccopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 8184 - 5819</mods:affiliation>
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<mods:nameIdentifier id="B751B6ECAA5F366952229B8F259764B6" type="email">ccopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:namePart id="7EBB37DC1EF67F4885954C6A5D9CFA92">Copley, Darren</mods:namePart>
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<mods:nameIdentifier id="3AA282E235B94CCFA516389AB9C543E4" type="ORCID">0000-0002-1944-4272</mods:nameIdentifier>
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<mods:affiliation id="8F0D3C6D4E31BBA657B5A3D4739618F3">dcopley @ royalbcmuseum. bc. ca; https: // orcid. org / 0000 - 0002 - 1944 - 4272</mods:affiliation>
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<mods:nameIdentifier id="37B441AC37C51D673562604569B75AB5" type="email">dcopley@royalbcmuseum.bc.ca</mods:nameIdentifier>
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<mods:typeOfResource id="D0F053A729AC4F0D7A557183466348E4">text</mods:typeOfResource>
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<mods:title id="DE8A025044051EADC13817153618275E">Zootaxa</mods:title>
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<mods:part id="00390B7CB0F7B023CA836A769E6876E2">
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<mods:date id="FD3177ED9AF99702F3D5FF7D22369A93">2023</mods:date>
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<mods:number id="2FAEB81988A81D4669564C9393C02C83">2023-07-18</mods:number>
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<mods:number id="E8A59208471EE13BED7F624445E13DB1">5318</mods:number>
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<mods:start id="2CFAB1D6F597EA68E03BD20EC02AF9E4">97</mods:start>
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<mods:end id="DC8DF67D7C455DA8B394497FACF340F8">129</mods:end>
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</mods:location>
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<mods:classification id="0F2D9CEC93ED0BAE4BC4F78BE3FA9EF2">journal article</mods:classification>
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<mods:identifier id="E70CFAF50EF2DB637731D4968FC5F666" type="DOI">10.11646/zootaxa.5318.1.5</mods:identifier>
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<mods:identifier id="BDA63678FC46268EE66BFF5B5CEB702A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BF87ABE428071409BEF00A60B5F98B" LSID="urn:lsid:plazi:treatment:03BF87ABE428071409BEF00A60B5F98B" httpUri="http://treatment.plazi.org/id/03BF87ABE428071409BEF00A60B5F98B" lastPageId="3" lastPageNumber="100" pageId="1" pageNumber="98">
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<subSubSection id="C30C6536E428071609BEF00A6744F85D" box="[151,620,1914,1941]" pageId="1" pageNumber="98" type="nomenclature">
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<paragraph id="8BA936BDE428071609BEF00A6744F85D" blockId="1.[151,620,1914,1941]" box="[151,620,1914,1941]" pageId="1" pageNumber="98">
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<heading id="D0E181D1E428071609BEF00A6744F85D" bold="true" box="[151,620,1914,1941]" fontSize="11" level="1" pageId="1" pageNumber="98" reason="1">
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<emphasis id="B962EAAFE428071609BEF00A6744F85D" bold="true" box="[151,620,1914,1941]" pageId="1" pageNumber="98">
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Genus
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<taxonomicName id="4C164D3EE428071609C7F00A6744F85D" ID-CoL="3XS8" ID-ENA="880878" authority="Chamberlin and Ivie" authorityName="Chamberlin and Ivie" box="[238,620,1914,1941]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EAAFE428071609C7F00A6448F85C" bold="true" box="[238,352,1914,1941]" italics="true" pageId="1" pageNumber="98">Cybaeina</emphasis>
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Chamberlin and Ivie
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30C6536E428071509BEF0CF667BFEC2" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98" type="reference_group">
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<paragraph id="8BA936BDE428071609BEF0CF64DBF81E" blockId="1.[151,499,1983,2007]" box="[151,499,1983,2007]" pageId="1" pageNumber="98">
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<treatmentCitationGroup id="AB061193E428071609BEF0CF64DBF81E" box="[151,499,1983,2007]" pageId="1" pageNumber="98">
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<taxonomicName id="4C164D3EE428071609BEF0CF65C6F81E" ID-CoL="3XSF" box="[151,238,1983,2007]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="98" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EAAFE428071609BEF0CF65C6F81E" box="[151,238,1983,2007]" italics="true" pageId="1" pageNumber="98">Cybaeus</emphasis>
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</taxonomicName>
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(in part)
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<treatmentCitation id="0AB710ACE42807160864F0CF64C7F81E" author="Banks, N." box="[333,495,1983,2007]" page="95" pageId="1" pageNumber="98" year="1906">
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<bibRefCitation id="EF874B4CE42807160864F0CF64C7F81E" author="Banks, N." box="[333,495,1983,2007]" pageId="1" pageNumber="98" pagination="94 - 100" refId="ref20707" refString="Banks, N. (1906) Descriptions of new American spiders. Proceedings of the Entomological Society of Washington, 7, 94 - 100." type="journal article" year="1906">Banks 1906: 95</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA936BDE42B071509BEF7E7667BFEC2" blockId="2.[151,1437,150,267]" pageId="2" pageNumber="99">
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<treatmentCitationGroup id="AB061193E42B071509BEF7E7667BFEC2" pageId="2" pageNumber="99">
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<treatmentCitation id="0AB710ACE42B071509BEF7E76736FF66" author="Chamberlin, R. V. & Ivie, W." box="[151,542,151,175]" page="28" pageId="2" pageNumber="99" year="1932">
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<taxonomicName id="4C164D3EE42B071509BEF7E76736FF66" ID-CoL="3XS8" authority="Chamberlin & Ivie 1932: 28" authorityName="Chamberlin & Ivie" authorityPageNumber="28" authorityYear="1932" box="[151,542,151,175]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
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<emphasis id="B962EAAFE42B071509BEF7E765D0FF66" box="[151,248,151,175]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
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<bibRefCitation id="EF874B4CE42B071509D7F7E76736FF66" author="Chamberlin, R. V. & Ivie, W." box="[254,542,151,175]" pageId="2" pageNumber="99" pagination="1 - 43" refId="ref21835" refString="Chamberlin, R. V. & Ivie, W. (1932) A review of the North American spiders of the genera Cybaeus and Cybaeina. Bulletin of the University of Utah, 23 (2), 1 - 43." type="journal article" year="1932">Chamberlin & Ivie 1932: 28</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figs 5–14, 28–30;
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<treatmentCitation id="0AB710ACE42B07150BC8F7E76129FF66" author="Chamberlin, R. V. & Ivie, W." box="[737,1025,151,175]" page="19" pageId="2" pageNumber="99" year="1942">
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<bibRefCitation id="EF874B4CE42B07150BC8F7E76129FF66" author="Chamberlin, R. V. & Ivie, W." box="[737,1025,151,175]" pageId="2" pageNumber="99" pagination="1 - 117" refId="ref21933" refString="Chamberlin, R. V. & Ivie, W. (1942) A hundred new species of American spiders. Bulletin of the University of Utah, 32 (13), 1 - 117." type="journal article" year="1942">Chamberlin & Ivie 1942: 19</bibRefCitation>
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</treatmentCitation>
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, fig. 38;
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<bibRefCitation id="EF874B4CE42B07150D72F7E761EEFF67" author="Roth, V. D." box="[1115,1222,150,175]" pageId="2" pageNumber="99" pagination="195 - 201" refId="ref22719" refString="Roth, V. D. (1952) Notes and a new species in Cybaeina. Pan-Pacific Entomologist, 28, 195 - 201." type="journal article" year="1952">Roth 1952</bibRefCitation>
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(in part): 195, figs 1, 3, 6;
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<treatmentCitation id="0AB710ACE42B071509D3F7C66480FF04" author="Roewer, C. F." box="[250,424,181,205]" page="86" pageId="2" pageNumber="99" year="1954">
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<bibRefCitation id="EF874B4CE42B071509D3F7C66480FF04" author="Roewer, C. F." box="[250,424,181,205]" pageId="2" pageNumber="99" refId="ref22691" refString="Roewer, C. F. (1954) Katalog der Araneae 2 a. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp." type="book" year="1954">Roewer 1954: 86</bibRefCitation>
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</treatmentCitation>
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(in part);
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<bibRefCitation id="EF874B4CE42B07150B27F7C667B8FF04" author="Bonnet, P." box="[526,656,181,205]" pageId="2" pageNumber="99" pagination="919 - 1926" refId="ref21567" refString="Bonnet, P. (1956) Bibliographia Araneorum. Douladoure, Toulouse, Tome 2, pp. 919 - 1926." type="journal article" year="1956">Bonnet 1956</bibRefCitation>
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(in part): 1296;
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<bibRefCitation id="EF874B4CE42B07150A1CF7C5612BFF04" author="Roth, V. D. & Brame, P. L." box="[821,1027,181,205]" pageId="2" pageNumber="99" pagination="1 - 52" refId="ref22745" refString="Roth, V. D. & Brame, P. L. (1972) Nearctic genera of the spider family Agelenidae (Arachnida, Araneida). American Museum Novitates, 2505, 1 - 52." type="journal article" year="1972">Roth & Brame 1972</bibRefCitation>
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(in part): 16, figs 21–22;
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<bibRefCitation id="EF874B4CE42B07150C2FF7C565D2FF25" author="Roth, V. D. & Brown, W. L." pageId="2" pageNumber="99" pagination="1 - 21" refId="ref22784" refString="Roth, V. D. & Brown, W. L. (1986) Catalog of Nearctic Agelenidae. The Museum Texas Tech University Occasional Papers, 99, 1 - 21. https: // doi. org / 10.5962 / bhl. title. 142809" type="journal article" year="1986">Roth & Brown 1986</bibRefCitation>
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(in part): 2;
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<treatmentCitation id="0AB710ACE42B071508AFF7A46714FF25" author="Bennett, R. G." box="[390,572,212,236]" page="88" pageId="2" pageNumber="99" year="2005">
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<bibRefCitation id="EF874B4CE42B071508AFF7A46714FF25" author="Bennett, R. G." box="[390,572,212,236]" pageId="2" pageNumber="99" pagination="85 - 90" refId="ref20845" refString="Bennett, R. G. (2005) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. & Roth, V. D. (Eds.), Spiders of North America: an Identification Manual. American Arachnological Society, pp. 85 - 90." type="book chapter" year="2005">Bennett 2005: 88</bibRefCitation>
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</treatmentCitation>
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, figs 22.19–22.21, 22.24–22.25, 22.27–22.34;
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<treatmentCitation id="0AB710ACE42B07150D0DF7A461F3FF25" author="Bennett, R. G." box="[1060,1243,212,236]" page="99" pageId="2" pageNumber="99" year="2017">
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|
<bibRefCitation id="EF874B4CE42B07150D0DF7A461F3FF25" author="Bennett, R. G." box="[1060,1243,212,236]" pageId="2" pageNumber="99" pagination="96 - 101" refId="ref20957" refString="Bennett, R. G. (2017) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. & Roth, V. D. (Eds.), Spiders of North America: an Identification Manual, 2 nd ed. American Arachnological Society, pp. 96 - 101." type="book chapter" year="2017">Bennett 2017: 99</bibRefCitation>
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</treatmentCitation>
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, figs 23.19–23.21, 23.24–23.25, 23.27–23.34;
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<bibRefCitation id="EF874B4CE42B071508F7F78367DCFEC2" author="World Spider Catalog" box="[478,756,243,267]" pageId="2" pageNumber="99" refId="ref23160" refString="World Spider Catalog (2023) World Spider Catalog. Version 24. Natural History Museum, Bern. Available from: https: // wsc. nmbe. ch / (accessed 5 March 2023) https: // doi. org / 10.24436 / 2" type="book" year="2023">World Spider Catalog 2023</bibRefCitation>
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(in part).
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</treatmentCitationGroup>
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</paragraph>
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|
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</subSubSection>
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|
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<paragraph id="8BA936BDE42B071509B1F47B66A9FC59" blockId="2.[151,1436,779,912]" pageId="2" pageNumber="99">
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<emphasis id="B962EAAFE42B071509B1F47B6469FCED" bold="true" box="[152,321,779,804]" pageId="2" pageNumber="99">FIGURES 1–2.</emphasis>
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<taxonomicName id="4C164D3EE42B0715086EF47C6480FCED" authorityName="Chamberlin & Ivie" authorityYear="1932" box="[327,424,780,804]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
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|
|
<emphasis id="B962EAAFE42B0715086EF47C6480FCED" box="[327,424,780,804]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C164D3EE42B071508F0F47C6769FCED" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[473,577,780,804]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B071508F0F47C6769FCED" box="[473,577,780,804]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150B6EF47C678EFCED" bold="true" box="[583,678,780,804]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150B6EF47C678EFCED" box="[583,678,780,804]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
species (1
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A3EF47C6652FCEA" authorityName="Chamberlin and Ivie" authorityYear="1932" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[791,890,780,803]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="minuta">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A3EF47C6652FCEA" box="[791,890,780,803]" italics="true" pageId="2" pageNumber="99">C. minuta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
female from Tofino, British Columbia; 2
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150C0BF47C60B4FCEA" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[1314,1436,780,803]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="petersoni" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150C0BF47C60B4FCEA" box="[1314,1436,780,803]" italics="true" pageId="2" pageNumber="99">R. petersoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B071509BEF44065D7FC81" bold="true" box="[151,255,816,840]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B071509BEF44065D7FC81" box="[151,255,816,840]" pageId="2" pageNumber="99" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
male from vicinity of Yuba Pass, California). 1 Eye group, anterior. 2 Patellae, tibiae, and metatarsi of legs I and II, ventral showing ventral tibial macrosetal pattern of 2(+1p)-2-2-2-2-0 on leg I and 2(+1p)-2-2-2-1-2-0 on leg II. AME—anterior median eyes, I—leg 1, II—leg II, Mt—metatarsus, Pt—patella, Ti—tibia.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C30C6536E42B071509BEF49261FDFC35" box="[151,1237,994,1020]" pageId="2" pageNumber="99" type="type_taxon">
|
|||
|
|
<paragraph id="8BA936BDE42B071509BEF49261FDFC35" blockId="2.[151,1437,994,2028]" box="[151,1237,994,1020]" pageId="2" pageNumber="99">
|
|||
|
|
<emphasis id="B962EAAFE42B071509BEF4926403FC35" bold="true" box="[151,299,994,1020]" pageId="2" pageNumber="99">
|
|||
|
|
<typeStatus id="54AD881FE42B071509BEF49265FAFC35" box="[151,210,994,1020]" pageId="2" pageNumber="99">Type</typeStatus>
|
|||
|
|
species
|
|||
|
|
</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150810F49267ADFC35" authority="Banks 1906" authorityName="Banks" authorityYear="1906" box="[313,645,994,1020]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="minutus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150810F49264D2FC35" box="[313,506,994,1020]" italics="true" pageId="2" pageNumber="99">Cybaeus minutus</emphasis>
|
|||
|
|
<bibRefCitation id="EF874B4CE42B07150B29F49267ADFC35" author="Banks, N." box="[512,645,994,1020]" pageId="2" pageNumber="99" pagination="94 - 100" refId="ref20707" refString="Banks, N. (1906) Descriptions of new American spiders. Proceedings of the Entomological Society of Washington, 7, 94 - 100." type="journal article" year="1906">Banks 1906</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by original designation (
|
|||
|
|
<bibRefCitation id="EF874B4CE42B07150AB4F49261E0FC35" author="Chamberlin, R. V. & Ivie, W." box="[925,1224,994,1020]" pageId="2" pageNumber="99" pagination="1 - 43" refId="ref21835" refString="Chamberlin, R. V. & Ivie, W. (1932) A review of the North American spiders of the genera Cybaeus and Cybaeina. Bulletin of the University of Utah, 23 (2), 1 - 43." type="journal article" year="1932">Chamberlin and Ivie 1932</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30C6536E42B071509EEF376600CFBE9" box="[199,1316,1030,1056]" pageId="2" pageNumber="99" type="etymology">
|
|||
|
|
<paragraph id="8BA936BDE42B071509EEF376600CFBE9" blockId="2.[151,1437,994,2028]" box="[199,1316,1030,1056]" pageId="2" pageNumber="99">
|
|||
|
|
<emphasis id="B962EAAFE42B071509EEF3766466FBE9" bold="true" box="[199,334,1030,1056]" pageId="2" pageNumber="99">Etymology.</emphasis>
|
|||
|
|
The genus name
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150B3AF3766756FBE9" authorityName="Chamberlin & Ivie" authorityYear="1932" box="[531,638,1030,1056]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150B3AF3766756FBE9" box="[531,638,1030,1056]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is derived from
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A1EF37666D3FBE9" authority="L. Koch" authorityName="L. Koch" box="[823,1019,1030,1056]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A1EF37666BFFBE9" box="[823,919,1030,1056]" italics="true" pageId="2" pageNumber="99">Cybaeus</emphasis>
|
|||
|
|
L. Koch
|
|||
|
|
</taxonomicName>
|
|||
|
|
and is feminine in gender.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30C6536E42B071409EEF35A61C3FF30" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99" type="diagnosis">
|
|||
|
|
<paragraph id="8BA936BDE42B071509EEF35A616DFB31" blockId="2.[151,1437,994,2028]" pageId="2" pageNumber="99">
|
|||
|
|
<emphasis id="B962EAAFE42B071509EEF35A646AFB8D" bold="true" box="[199,322,1066,1092]" pageId="2" pageNumber="99">Diagnosis.</emphasis>
|
|||
|
|
The males and females of the Nearctic genera of Cybaeinae are readily separated into two groups on the basis of tibia I and II ventral macrosetal characters: members of
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150AB6F33E61D5FBA1" authority="Chamberlin and Ivie" authorityName="Chamberlin and Ivie" box="[927,1277,1102,1128]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150AB6F33E6122FBA1" box="[927,1034,1102,1128]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
Chamberlin and Ivie
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150C23F33E65C6FB45" authority="Bennett" authorityName="Bennett & Copley & Copley" authorityYear="2023" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150C23F33E60B4FBA1" box="[1290,1436,1102,1128]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
|||
|
|
Bennett
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B071509D3F302644FFB45" bold="true" box="[250,359,1138,1164]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B071509D3F302644FFB45" box="[250,359,1138,1164]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150853F3026778FB45" authority="Bennett" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[378,592,1138,1164]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150853F30264C5FB45" box="[378,493,1138,1164]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
Bennett
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150B72F30267E1FB45" bold="true" box="[603,713,1138,1164]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150B72F30267E1FB45" box="[603,713,1138,1164]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A26F302614BFB45" authority="Chamberlin & Ivie" authorityName="Chamberlin & Ivie" authorityYear="1933" box="[783,1123,1138,1164]" class="Arachnida" family="Cybaeidae" genus="Cybaeota" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A26F3026652FB45" box="[783,890,1138,1164]" italics="true" pageId="2" pageNumber="99">Cybaeota</emphasis>
|
|||
|
|
Chamberlin & Ivie
|
|||
|
|
</taxonomicName>
|
|||
|
|
have four or five pairs of conspicuous elongate, often overlapping macrosetae (
|
|||
|
|
<figureCitation id="132D2A38E42B07150BC5F3E66607FB79" box="[748,815,1174,1200]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,779,803]" captionTargetBox="[151,1436,291,756]" captionTargetId="figure-108@2.[151,1436,291,756]" captionTargetPageId="2" captionText="FIGURES 1–2. Cybaeina and Rothaeina gen. nov. species (1 C. minuta female from Tofino, British Columbia; 2 R. petersoni spec. nov. male from vicinity of Yuba Pass, California). 1 Eye group, anterior. 2 Patellae, tibiae, and metatarsi of legs I and II, ventral showing ventral tibial macrosetal pattern of 2(+1p)-2-2-2-2-0 on leg I and 2(+1p)-2-2-2-1-2-0 on leg II. AME—anterior median eyes, I—leg 1, II—leg II, Mt—metatarsus, Pt—patella, Ti—tibia." figureDoi="http://doi.org/10.5281/zenodo.8158359" httpUri="https://zenodo.org/record/8158359/files/figure.png" pageId="2" pageNumber="99">Fig. 2</figureCitation>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B962EAAFE42B07150A14F3E766ADFB79" box="[829,901,1175,1200]" italics="true" pageId="2" pageNumber="99">versus</emphasis>
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150AA2F3E66157FB79" authority="Bennett" authorityName="Bennett and Pseudocybaeota Bennett" authorityYear="2020" box="[907,1151,1174,1200]" class="Arachnida" family="Cybaeidae" genus="Allocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150AA2F3E66137FB79" box="[907,1055,1174,1200]" italics="true" pageId="2" pageNumber="99">Allocybaeina</emphasis>
|
|||
|
|
Bennett
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150DA0F3E665CDFB1D" authority="Chamberlin & Ivie" authorityName="Chamberlin & Ivie" class="Arachnida" family="Cybaeidae" genus="Cybaeozyga" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150DA0F3E6603AFB79" box="[1161,1298,1174,1200]" italics="true" pageId="2" pageNumber="99">Cybaeozyga</emphasis>
|
|||
|
|
Chamberlin & Ivie
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B071509DAF3CA663EFB1D" authority="L. Koch, and Pseudocybaeota Bennett" authorityName="L. Koch, and Pseudocybaeota Bennett" box="[243,790,1210,1236]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B071509DAF3CA647BFB1D" box="[243,339,1210,1236]" italics="true" pageId="2" pageNumber="99">Cybaeus</emphasis>
|
|||
|
|
L. Koch, and
|
|||
|
|
<emphasis id="B962EAAFE42B071508D7F3CA679EFB1D" box="[510,694,1210,1236]" italics="true" pageId="2" pageNumber="99">Pseudocybaeota</emphasis>
|
|||
|
|
Bennett
|
|||
|
|
</taxonomicName>
|
|||
|
|
which have only two or three pairs of shorter and nonoverlapping macrosetae (as in
|
|||
|
|
<bibRefCitation id="EF874B4CE42B071508C4F3AE67E8FB31" author="Copley, C. R. & Bennett, R. & Perlman, S. J." box="[493,704,1246,1272]" pageId="2" pageNumber="99" pagination="367 - 401" refId="ref22032" refString="Copley, C. R., Bennett, R. & Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
|
|||
|
|
Copley
|
|||
|
|
<emphasis id="B962EAAFE42B07150B6CF3AF6755FB31" box="[581,637,1246,1272]" italics="true" pageId="2" pageNumber="99">et al.</emphasis>
|
|||
|
|
2009
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: fig. 14;
|
|||
|
|
<bibRefCitation id="EF874B4CE42B07150A0AF3AE6695FB31" author="Bennett, R. G." box="[803,957,1246,1272]" pageId="2" pageNumber="99" pagination="96 - 101" refId="ref20957" refString="Bennett, R. G. (2017) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. & Roth, V. D. (Eds.), Spiders of North America: an Identification Manual, 2 nd ed. American Arachnological Society, pp. 96 - 101." type="book chapter" year="2017">Bennett 2017</bibRefCitation>
|
|||
|
|
: fig. 23.14).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42B071509EEF27267CFFA3D" blockId="2.[151,1437,994,2028]" pageId="2" pageNumber="99">
|
|||
|
|
Both sexes of
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150841F27264FBFAD5" authorityName="Chamberlin & Ivie" authorityYear="1933" box="[360,467,1282,1308]" class="Arachnida" family="Cybaeidae" genus="Cybaeota" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150841F27264FBFAD5" box="[360,467,1282,1308]" italics="true" pageId="2" pageNumber="99">Cybaeota</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be separated from those of
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A15F272668FFAD5" box="[828,935,1282,1308]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A15F272668FFAD5" box="[828,935,1282,1308]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A9FF2726160FAD5" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[950,1096,1282,1308]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A9FF2726160FAD5" box="[950,1096,1282,1308]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150D79F2726191FAD5" bold="true" box="[1104,1209,1282,1308]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150D79F2726191FAD5" box="[1104,1209,1282,1308]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150DD8F272604CFAD5" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[1265,1380,1282,1308]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150DD8F272604CFAD5" box="[1265,1380,1282,1308]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150C42F27265E0FA89" bold="true" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150C42F27265E0FA89" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
by the absence of a retrolateral patellar apophysis and associated peg setae on the male pedipalp and the relatively simple vulva dominated by very large, rounded spermathecal bases (
|
|||
|
|
<bibRefCitation id="EF874B4CE42B07150D3EF23A619DFAAD" author="Bennett, R. G." box="[1047,1205,1354,1380]" pageId="2" pageNumber="99" pagination="103 - 119" refId="ref20734" refString="Bennett, R. G. (1988) The spider genus Cybaeota (Araneae, Agelenidae). Journal of Arachnology, 16, 103 - 119." type="journal article" year="1988">Bennett 1988</bibRefCitation>
|
|||
|
|
: figs 25, 28, 35, 38). In
|
|||
|
|
<taxonomicName id="4C164D3EE42B071509E1F21E641BFA41" box="[200,307,1390,1416]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B071509E1F21E641BFA41" box="[200,307,1390,1416]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B0715086DF21E64FEFA41" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[324,470,1390,1416]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B0715086DF21E64FEFA41" box="[324,470,1390,1416]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B071508C9F21E6763FA41" bold="true" box="[480,587,1390,1416]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B071508C9F21E6763FA41" box="[480,587,1390,1416]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
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|
|
</emphasis>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150BA7F21E6629FA41" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[654,769,1390,1416]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150BA7F21E6629FA41" box="[654,769,1390,1416]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150A23F21E665EFA41" bold="true" box="[778,886,1390,1416]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150A23F21E665EFA41" box="[778,886,1390,1416]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, a retrolateral patellar apophysis and associated peg setae are present (
|
|||
|
|
<figureCitation id="132D2A38E42B071508B4F2E264C2FA65" box="[413,490,1426,1452]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="2" pageNumber="99">Figs 9</figureCitation>
|
|||
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,
|
|||
|
|
<figureCitation id="132D2A38E42B071508D0F2E2673FFA65" box="[505,535,1426,1452]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="2" pageNumber="99">18</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150B0EF2E2676DFA65" box="[551,581,1426,1452]" captionStart="FIGURES 29–35" captionStartId="13.[152,270,1250,1274]" captionTargetBox="[151,1433,181,1227]" captionTargetId="figure-21@13.[151,1436,181,1227]" captionTargetPageId="13" captionText="FIGURES 29–35. Neocybaeina xantha comb. nov., males from Oregon (29, 33–35 from near Glide; 30 from unknown locale; 31–32 from near Bridge), left pedipalp. 29 Genital bulb, ventral. 30 Tegular apophysis and embolus, retrolateral. 31–34 Tibia and patella (31, 33 retrolateral; 32, 34 dorsal). 35 medial component and distal tip of distal component of retrolateral tibial apophysis, dorsal. Cy—cymbium, DA—distal arm of tegular apophysis, dRTA—distal component of retrolateral tibial apophysis, E—embolus, mRTA—medial component of retrolateral tibial apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, Tr—trichobothrium. Note: in Figs 31, 33–35 the distal end of the mRTA is a short rigid spine-like process; in Fig. 32 the mRTA terminates distally with a short basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158375" httpUri="https://zenodo.org/record/8158375/files/figure.png" pageId="2" pageNumber="99">33</figureCitation>
|
|||
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,
|
|||
|
|
<figureCitation id="132D2A38E42B07150B7CF2E2675BFA64" box="[597,627,1426,1453]" captionStart="FIGURES 53–59" captionStartId="18.[152,270,1632,1656]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-37@18.[151,1436,181,1608]" captionTargetPageId="18" captionText="FIGURES 53–59. Rothaeina gen. nov. species, males (53 R. jamesi spec. nov. holotype; 54–55 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 56–57 R. petersoni spec. nov. from near Kyburz, California; 58–59 R. sequoia comb. nov. holotype), left pedipalp. 53–54, 56, 58 Patellar apophysis and tibia, retrolateral. 55 Patella, dorsal. 57, 59 Tibia and patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis. Unlabelled arrow indicates basal hump of patellar apophysis (59). Note: in Figs 53, 58–59 the distal end of the mRTA is a rigid spine-like process; in Figs 56–57 the mRTA terminates distally in a basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158381" httpUri="https://zenodo.org/record/8158381/files/figure.png" pageId="2" pageNumber="99">59</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150BABF2E267B7FA65" box="[642,671,1426,1452]" captionStart="FIGURES 60–65" captionStartId="19.[152,270,1253,1277]" captionTargetBox="[151,1436,181,1229]" captionTargetId="figure-24@19.[151,1436,181,1229]" captionTargetPageId="19" captionText="FIGURES 60–65. Rothaeina gen. nov. species, males (60–62 R. jamesi spec. nov. holotype; 63 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 64–65 R. petersoni spec. nov. from near Kyburz, California), left pedipalp. 60 Proximal arm of tegular apophysis, ventral. 61 Medial component of retrolateral tibial apophysis, retrolateral. 62, 64–65 Patellar apophysis, dorsal. 63 Tip of proximal arm of tegular apophysis, retrolateral. Unlabelled arrows indicate diagnostic characters: basal projection of proximal arm of tegular apophysis (60); dorso-basal group of peg setae (62); dorsal keel of proximal arm of tegular apophysis (63); large and small peg setae (64–65)." figureDoi="http://doi.org/10.5281/zenodo.8158432" httpUri="https://zenodo.org/record/8158432/files/figure.png" pageId="2" pageNumber="99">64</figureCitation>
|
|||
|
|
) and the vulva is relatively complex with the spermathecal bases undifferentiated from the spermathecal heads and stalks (
|
|||
|
|
<figureCitation id="132D2A38E42B07150A05F2C666AEFA19" box="[812,902,1462,1488]" captionStart="FIGURES 19–21" captionStartId="9.[152,270,1259,1283]" captionTargetBox="[153,1433,181,1235]" captionTargetId="figure-19@9.[151,1436,181,1235]" captionTargetPageId="9" captionText="FIGURES 19–21. Cybaeina confusa, females (19 from Rose Harbour, British Columbia; 20–21 holotype), copulatory organ. 19 Epigyne, ventral. 20–21 Vulva (20 ventral, 21 dorsal). BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (19–21)." figureDoi="http://doi.org/10.5281/zenodo.8158369" httpUri="https://zenodo.org/record/8158369/files/figure.png" pageId="2" pageNumber="99">Figs 20</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150ABDF2C6669AFA19" box="[916,946,1462,1488]" captionStart="FIGURES 66–69" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 66–69. Rothaeina beaudini spec. nov., females from California (66–67 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 67–69 Vulva (67–68 dorsal, 69 ventral). BG—Bennett’s gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (66–67, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="2" pageNumber="99">67</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150AE8F2C666F6FA19" box="[961,990,1462,1488]" captionStart="FIGURES 75–79" captionStartId="25.[152,270,1849,1873]" captionTargetBox="[151,1436,418,1825]" captionTargetId="figure-112@25.[151,1436,417,1825]" captionTargetPageId="25" captionText="FIGURES 75–79. Rothaeina petersoni spec. nov., females from California (75 from Kyburz; 76 from Sierra City; 77 from Tamarack Flat; 78 from Forest Hill; 79 from near Yuba Pass), copulatory organ. 75 Epigyne, ventral. 76–79 Vulva, dorsal). BG—Bennett’s gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (75–76, 78 single shafted arrows), and ring-like loops of spermathecae (75–76, 79 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158389" httpUri="https://zenodo.org/record/8158389/files/figure.png" pageId="2" pageNumber="99">77</figureCitation>
|
|||
|
|
) or the bases are relatively small, less rounded, and not dominating the vulva (
|
|||
|
|
<figureCitation id="132D2A38E42B07150B71F2AA6787FA3D" box="[600,687,1498,1524]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="2" pageNumber="99">Figs 23</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150B92F2AA67F3FA3D" box="[699,731,1498,1524]" captionStart="FIGURES 36–42" captionStartId="15.[152,270,1834,1858]" captionTargetBox="[151,1436,182,1810]" captionTargetId="figure-19@15.[151,1436,181,1810]" captionTargetPageId="15" captionText="FIGURES 36–42. Neocybaeina gen. nov. species, females from Oregon (36–39 N. xantha comb. nov., 36 from Bandon, 37–39 from Roseburg; 40–42 N. burnetti spec. nov., 40–41 from Carpenterville, 42 from Gold Beach), copulatory organ. 36, 40 Epigyne, ventral. 37–39, 41–42 Vulva (37 dorsal; 38 ventral; 39, 41 atrial openings, copulatory ducts, and heads, stalks, and proximal portion of bases of spermathecae, ventral; 42 left half of vulva, dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (36, 38, 40)." figureDoi="http://doi.org/10.5281/zenodo.8158377" httpUri="https://zenodo.org/record/8158377/files/figure.png" pageId="2" pageNumber="99">42</figureCitation>
|
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|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42B071509EEF28E606AF94D" blockId="2.[151,1437,994,2028]" pageId="2" pageNumber="99">
|
|||
|
|
Females of
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150879F28E6493F9D1" box="[336,443,1534,1560]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150879F28E6493F9D1" box="[336,443,1534,1560]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B071508E4F28E6777F9D1" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[461,607,1534,1560]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B071508E4F28E6777F9D1" box="[461,607,1534,1560]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150B40F28E67FDF9D1" bold="true" box="[617,725,1534,1560]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150B40F28E67FDF9D1" box="[617,725,1534,1560]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A30F28E66A4F9D1" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[793,908,1534,1560]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150A30F28E66A4F9D1" box="[793,908,1534,1560]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150ABFF28E612AF9D1" bold="true" box="[918,1026,1534,1560]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150ABFF28E612AF9D1" box="[918,1026,1534,1560]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
are readily distinguished by vulval morphology. In females of
|
|||
|
|
<taxonomicName id="4C164D3EE42B071508F6F1526762F9F5" box="[479,586,1570,1596]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B071508F6F1526762F9F5" box="[479,586,1570,1596]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the copulatory ducts are elongate, lightly sclerotized, and encircle the spermathecal stalks (
|
|||
|
|
<figureCitation id="132D2A38E42B07150856F13664D7F9A9" box="[383,511,1606,1632]" captionStart="FIGURES 19–21" captionStartId="9.[152,270,1259,1283]" captionTargetBox="[153,1433,181,1235]" captionTargetId="figure-19@9.[151,1436,181,1235]" captionTargetPageId="9" captionText="FIGURES 19–21. Cybaeina confusa, females (19 from Rose Harbour, British Columbia; 20–21 holotype), copulatory organ. 19 Epigyne, ventral. 20–21 Vulva (20 ventral, 21 dorsal). BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (19–21)." figureDoi="http://doi.org/10.5281/zenodo.8158369" httpUri="https://zenodo.org/record/8158369/files/figure.png" pageId="2" pageNumber="99">Figs 20–21</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150B23F1366780F9A9" box="[522,680,1606,1632]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="2" pageNumber="99">23–24, 26–27</figureCitation>
|
|||
|
|
). In
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150BF1F1366642F9A9" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[728,874,1606,1632]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150BF1F1366642F9A9" box="[728,874,1606,1632]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150A59F13666F0F9A9" bold="true" box="[880,984,1606,1632]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150A59F13666F0F9A9" box="[880,984,1606,1632]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150D24F13661A8F9A9" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[1037,1152,1606,1632]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150D24F13661A8F9A9" box="[1037,1152,1606,1632]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150DAEF13661C7F9A9" bold="true" box="[1159,1263,1606,1632]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150DAEF13661C7F9A9" box="[1159,1263,1606,1632]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, the copulatory ducts are very short, heavily sclerotized, and do not encircle the stalks (
|
|||
|
|
<figureCitation id="132D2A38E42B07150A9EF11A611FF94D" box="[951,1079,1642,1668]" captionStart="FIGURES 36–42" captionStartId="15.[152,270,1834,1858]" captionTargetBox="[151,1436,182,1810]" captionTargetId="figure-19@15.[151,1436,181,1810]" captionTargetPageId="15" captionText="FIGURES 36–42. Neocybaeina gen. nov. species, females from Oregon (36–39 N. xantha comb. nov., 36 from Bandon, 37–39 from Roseburg; 40–42 N. burnetti spec. nov., 40–41 from Carpenterville, 42 from Gold Beach), copulatory organ. 36, 40 Epigyne, ventral. 37–39, 41–42 Vulva (37 dorsal; 38 ventral; 39, 41 atrial openings, copulatory ducts, and heads, stalks, and proximal portion of bases of spermathecae, ventral; 42 left half of vulva, dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (36, 38, 40)." figureDoi="http://doi.org/10.5281/zenodo.8158377" httpUri="https://zenodo.org/record/8158377/files/figure.png" pageId="2" pageNumber="99">Figs 39, 41</figureCitation>
|
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,
|
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|
|
<figureCitation id="132D2A38E42B07150D6AF11A61A4F94D" box="[1091,1164,1642,1668]" captionStart="FIGURES 66–69" captionStartId="20.[152,270,1168,1192]" captionTargetBox="[151,1435,182,1144]" captionTargetId="figure-37@20.[151,1436,181,1144]" captionTargetPageId="20" captionText="FIGURES 66–69. Rothaeina beaudini spec. nov., females from California (66–67 from Delta; 68 from Manzanita Lake; 69 from Lake Almanor), copulatory organ. 66 Epigyne, ventral. 67–69 Vulva (67–68 dorsal, 69 ventral). BG—Bennett’s gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial opening (66–67, 69)." figureDoi="http://doi.org/10.5281/zenodo.8158434" httpUri="https://zenodo.org/record/8158434/files/figure.png" pageId="2" pageNumber="99">68–69</figureCitation>
|
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,
|
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|
|
<figureCitation id="132D2A38E42B07150DBEF11A61C8F94D" box="[1175,1248,1642,1668]" captionStart="FIGURES 70–74" captionStartId="22.[152,270,1556,1580]" captionTargetBox="[158,1426,182,1532]" captionTargetId="figure-37@22.[158,1430,181,1532]" captionTargetPageId="22" captionText="FIGURES 70–74. Rothaeina gen. nov. species, females (70–72 R. jamesi spec. nov. from Klamath, California; 73–74 R. mackinleyi spec. nov. from Charleston, Oregon), copulatory organ. 70 Epigyne, ventral. 71–74 Vulva (71, 73 ventral; 72, 74 dorsal). BG—Bennett’s gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (70–71, 74 single shafted arrows), additional loop of spermatheca (72 double shafted arrow), and medial transverse sections of spermathecae (72, 74 dashed arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158438" httpUri="https://zenodo.org/record/8158438/files/figure.png" pageId="2" pageNumber="99">73–74</figureCitation>
|
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,
|
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|
|
<figureCitation id="132D2A38E42B07150DC2F11A601DF94D" box="[1259,1333,1642,1668]" captionStart="FIGURES 75–79" captionStartId="25.[152,270,1849,1873]" captionTargetBox="[151,1436,418,1825]" captionTargetId="figure-112@25.[151,1436,417,1825]" captionTargetPageId="25" captionText="FIGURES 75–79. Rothaeina petersoni spec. nov., females from California (75 from Kyburz; 76 from Sierra City; 77 from Tamarack Flat; 78 from Forest Hill; 79 from near Yuba Pass), copulatory organ. 75 Epigyne, ventral. 76–79 Vulva, dorsal). BG—Bennett’s gland, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca. Unlabelled arrows indicate atrial openings (75–76, 78 single shafted arrows), and ring-like loops of spermathecae (75–76, 79 double shafted arrows)." figureDoi="http://doi.org/10.5281/zenodo.8158389" httpUri="https://zenodo.org/record/8158389/files/figure.png" pageId="2" pageNumber="99">76, 78</figureCitation>
|
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).
|
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|
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</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42B071409EEF1FE61C3FF30" blockId="2.[151,1437,994,2028]" lastBlockId="3.[151,1437,151,1077]" lastPageId="3" lastPageNumber="100" pageId="2" pageNumber="99">
|
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|
|
The males of these three genera, however, are not easily diagnosed; separation of them requires careful referral to a combination of characters of the pedipalp and comparison with published illustrations. In addition, the male of only one of the two species of
|
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|
|
<taxonomicName id="4C164D3EE42B07150B32F1A66785F939" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[539,685,1750,1776]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
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|
|
<emphasis id="B962EAAFE42B07150B32F1A66785F939" box="[539,685,1750,1776]" italics="true" pageId="2" pageNumber="99">Neocybaeina</emphasis>
|
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|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150B9FF1A66609F939" bold="true" box="[694,801,1750,1776]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150B9FF1A66609F939" box="[694,801,1750,1776]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
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|
|
</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150A18F1A761BAF939" authority="(Chamberlin & Ivie)" authorityName="Bennett & Copley & Copley" authorityYear="2023" baseAuthorityName="Bennett & Copley & Copley" baseAuthorityYear="2023" box="[817,1170,1750,1776]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="xantha" status="comb. nov.">
|
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|
|
<emphasis id="B962EAAFE42B07150A18F1A766B7F939" box="[817,927,1750,1776]" italics="true" pageId="2" pageNumber="99">N. xantha</emphasis>
|
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|
|
(Chamberlin & Ivie)
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150DB2F1A66035F939" bold="true" box="[1179,1309,1750,1776]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150DB2F1A66035F939" box="[1179,1309,1750,1776]" pageId="2" pageNumber="99" rank="species">comb. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, is known; the unknown male of
|
|||
|
|
<taxonomicName id="4C164D3EE42B071508A4F18B672FF8DD" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[397,519,1786,1812]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="burnetti" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B071508A4F18B672FF8DD" box="[397,519,1786,1812]" italics="true" pageId="2" pageNumber="99">N. burnetti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150B27F18A67A8F8DD" bold="true" box="[526,640,1786,1812]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150B27F18A67A8F8DD" box="[526,640,1786,1812]" pageId="2" pageNumber="99" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
may not fit the genus diagnoses presented in this paper. The following characteristics may be helpful in distinguishing the males of these three genera. In the males of all three, the patellar apophysis is quite short (
|
|||
|
|
<figureCitation id="132D2A38E42B07150891F0326712F895" box="[440,570,1858,1884]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="2" pageNumber="99">Figs 11, 13</figureCitation>
|
|||
|
|
) to approximately as long as the width of the patella (
|
|||
|
|
<figureCitation id="132D2A38E42B07150D9BF032601EF895" box="[1202,1334,1858,1884]" captionStart="FIGURES 29–35" captionStartId="13.[152,270,1250,1274]" captionTargetBox="[151,1433,181,1227]" captionTargetId="figure-21@13.[151,1436,181,1227]" captionTargetPageId="13" captionText="FIGURES 29–35. Neocybaeina xantha comb. nov., males from Oregon (29, 33–35 from near Glide; 30 from unknown locale; 31–32 from near Bridge), left pedipalp. 29 Genital bulb, ventral. 30 Tegular apophysis and embolus, retrolateral. 31–34 Tibia and patella (31, 33 retrolateral; 32, 34 dorsal). 35 medial component and distal tip of distal component of retrolateral tibial apophysis, dorsal. Cy—cymbium, DA—distal arm of tegular apophysis, dRTA—distal component of retrolateral tibial apophysis, E—embolus, mRTA—medial component of retrolateral tibial apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, Tr—trichobothrium. Note: in Figs 31, 33–35 the distal end of the mRTA is a short rigid spine-like process; in Fig. 32 the mRTA terminates distally with a short basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158375" httpUri="https://zenodo.org/record/8158375/files/figure.png" pageId="2" pageNumber="99">Figs 31–34</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150C6AF0326049F894" box="[1347,1377,1858,1885]" captionStart="FIGURES 53–59" captionStartId="18.[152,270,1632,1656]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-37@18.[151,1436,181,1608]" captionTargetPageId="18" captionText="FIGURES 53–59. Rothaeina gen. nov. species, males (53 R. jamesi spec. nov. holotype; 54–55 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 56–57 R. petersoni spec. nov. from near Kyburz, California; 58–59 R. sequoia comb. nov. holotype), left pedipalp. 53–54, 56, 58 Patellar apophysis and tibia, retrolateral. 55 Patella, dorsal. 57, 59 Tibia and patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis. Unlabelled arrow indicates basal hump of patellar apophysis (59). Note: in Figs 53, 58–59 the distal end of the mRTA is a rigid spine-like process; in Figs 56–57 the mRTA terminates distally in a basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158381" httpUri="https://zenodo.org/record/8158381/files/figure.png" pageId="2" pageNumber="99">53</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150C59F03260B8F895" box="[1392,1424,1858,1884]" captionStart="FIGURES 60–65" captionStartId="19.[152,270,1253,1277]" captionTargetBox="[151,1436,181,1229]" captionTargetId="figure-24@19.[151,1436,181,1229]" captionTargetPageId="19" captionText="FIGURES 60–65. Rothaeina gen. nov. species, males (60–62 R. jamesi spec. nov. holotype; 63 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 64–65 R. petersoni spec. nov. from near Kyburz, California), left pedipalp. 60 Proximal arm of tegular apophysis, ventral. 61 Medial component of retrolateral tibial apophysis, retrolateral. 62, 64–65 Patellar apophysis, dorsal. 63 Tip of proximal arm of tegular apophysis, retrolateral. Unlabelled arrows indicate diagnostic characters: basal projection of proximal arm of tegular apophysis (60); dorso-basal group of peg setae (62); dorsal keel of proximal arm of tegular apophysis (63); large and small peg setae (64–65)." figureDoi="http://doi.org/10.5281/zenodo.8158432" httpUri="https://zenodo.org/record/8158432/files/figure.png" pageId="2" pageNumber="99">62</figureCitation>
|
|||
|
|
). When elongate, however, the tip of the patellar apophysis is retrolaterad in
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150AFDF0166117F849" box="[980,1087,1894,1920]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B07150AFDF0166117F849" box="[980,1087,1894,1920]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132D2A38E42B07150D67F01661B8F849" box="[1102,1168,1894,1920]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="2" pageNumber="99">Fig. 9</figureCitation>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B962EAAFE42B07150DB7F01761CEF849" box="[1182,1254,1895,1920]" italics="true" pageId="2" pageNumber="99">versus</emphasis>
|
|||
|
|
dorsad or distad in
|
|||
|
|
<taxonomicName id="4C164D3EE42B0715099CF0FA64B4F86D" authorityName="Bennett & Copley & Copley" authorityYear="2023" baseAuthorityName="Bennett & Copley & Copley" baseAuthorityYear="2023" box="[181,412,1930,1956]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="species" species="xantha" status="comb. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B0715099CF0FA64B4F86D" box="[181,412,1930,1956]" italics="true" pageId="2" pageNumber="99">Neocybaeina xantha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B0715088CF0FA670FF86D" bold="true" box="[421,551,1930,1956]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B0715088CF0FA670FF86D" box="[421,551,1930,1956]" pageId="2" pageNumber="99" rank="species">comb. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132D2A38E42B07150B1EF0FA6791F86D" box="[567,697,1930,1956]" captionStart="FIGURES 29–35" captionStartId="13.[152,270,1250,1274]" captionTargetBox="[151,1433,181,1227]" captionTargetId="figure-21@13.[151,1436,181,1227]" captionTargetPageId="13" captionText="FIGURES 29–35. Neocybaeina xantha comb. nov., males from Oregon (29, 33–35 from near Glide; 30 from unknown locale; 31–32 from near Bridge), left pedipalp. 29 Genital bulb, ventral. 30 Tegular apophysis and embolus, retrolateral. 31–34 Tibia and patella (31, 33 retrolateral; 32, 34 dorsal). 35 medial component and distal tip of distal component of retrolateral tibial apophysis, dorsal. Cy—cymbium, DA—distal arm of tegular apophysis, dRTA—distal component of retrolateral tibial apophysis, E—embolus, mRTA—medial component of retrolateral tibial apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, Tr—trichobothrium. Note: in Figs 31, 33–35 the distal end of the mRTA is a short rigid spine-like process; in Fig. 32 the mRTA terminates distally with a short basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158375" httpUri="https://zenodo.org/record/8158375/files/figure.png" pageId="2" pageNumber="99">Figs 31, 33</figureCitation>
|
|||
|
|
) and
|
|||
|
|
<taxonomicName id="4C164D3EE42B07150BD2F0FA6646F86D" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[763,878,1930,1956]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42B07150BD2F0FA6646F86D" box="[763,878,1930,1956]" italics="true" pageId="2" pageNumber="99">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42B07150A5EF0FA66C9F86D" bold="true" box="[887,993,1930,1956]" pageId="2" pageNumber="99">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42B07150A5EF0FA66C9F86D" box="[887,993,1930,1956]" pageId="2" pageNumber="99" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132D2A38E42B07150AD8F0FA618AF86C" box="[1009,1186,1930,1957]" captionStart="FIGURES 53–59" captionStartId="18.[152,270,1632,1656]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-37@18.[151,1436,181,1608]" captionTargetPageId="18" captionText="FIGURES 53–59. Rothaeina gen. nov. species, males (53 R. jamesi spec. nov. holotype; 54–55 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 56–57 R. petersoni spec. nov. from near Kyburz, California; 58–59 R. sequoia comb. nov. holotype), left pedipalp. 53–54, 56, 58 Patellar apophysis and tibia, retrolateral. 55 Patella, dorsal. 57, 59 Tibia and patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis. Unlabelled arrow indicates basal hump of patellar apophysis (59). Note: in Figs 53, 58–59 the distal end of the mRTA is a rigid spine-like process; in Figs 56–57 the mRTA terminates distally in a basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158381" httpUri="https://zenodo.org/record/8158381/files/figure.png" pageId="2" pageNumber="99">Figs 53, 57, 59</figureCitation>
|
|||
|
|
). As well, in males of
|
|||
|
|
<taxonomicName id="4C164D3EE42B071509BEF0DE642AF801" box="[151,258,1966,1992]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="99" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42B071509BEF0DE642AF801" box="[151,258,1966,1992]" italics="true" pageId="2" pageNumber="99">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the medial component of the retrolateral tibial apophysis is blade- or ledge-like and distally acuminate (
|
|||
|
|
<figureCitation id="132D2A38E42B071509B6F0A26409F825" box="[159,289,2002,2028]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="2" pageNumber="99">Figs 10–12</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42B07150804F0A2645CF825" box="[301,372,2002,2028]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="2" pageNumber="99">14–17</figureCitation>
|
|||
|
|
) or terminating in a single elongate basally articulated macroseta (
|
|||
|
|
<figureCitation id="132D2A38E42B07150D4CF0A261E2F825" box="[1125,1226,2002,2028]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="2" pageNumber="99">Figs 8–9</figureCitation>
|
|||
|
|
) whereas in males of
|
|||
|
|
<taxonomicName id="4C164D3EE42A0714099FF7E764B3FF78" authorityName="Bennett & Copley & Copley" authorityYear="2023" baseAuthorityName="Bennett & Copley & Copley" baseAuthorityYear="2023" box="[182,411,151,177]" class="Arachnida" family="Cybaeidae" genus="Neocybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="xantha" status="comb. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A0714099FF7E764B3FF78" box="[182,411,151,177]" italics="true" pageId="3" pageNumber="100">Neocybaeina xantha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A0714088BF7E7670BFF78" bold="true" box="[418,547,151,177]" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A0714088BF7E7670BFF78" box="[418,547,151,177]" pageId="3" pageNumber="100" rank="species">comb. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
and most species of
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140A27F7E766A9FF78" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[782,897,151,177]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A07140A27F7E766A9FF78" box="[782,897,151,177]" italics="true" pageId="3" pageNumber="100">Rothaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A07140AA1F7E766DAFF78" bold="true" box="[904,1010,151,177]" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A07140AA1F7E766DAFF78" box="[904,1010,151,177]" pageId="3" pageNumber="100" rank="genus">gen. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
the medial component is reduced to a single rigid spine-like process or basally articulated macroseta (
|
|||
|
|
<figureCitation id="132D2A38E42A07140A7FF7CB66FEFF1C" box="[854,982,186,213]" captionStart="FIGURES 29–35" captionStartId="13.[152,270,1250,1274]" captionTargetBox="[151,1433,181,1227]" captionTargetId="figure-21@13.[151,1436,181,1227]" captionTargetPageId="13" captionText="FIGURES 29–35. Neocybaeina xantha comb. nov., males from Oregon (29, 33–35 from near Glide; 30 from unknown locale; 31–32 from near Bridge), left pedipalp. 29 Genital bulb, ventral. 30 Tegular apophysis and embolus, retrolateral. 31–34 Tibia and patella (31, 33 retrolateral; 32, 34 dorsal). 35 medial component and distal tip of distal component of retrolateral tibial apophysis, dorsal. Cy—cymbium, DA—distal arm of tegular apophysis, dRTA—distal component of retrolateral tibial apophysis, E—embolus, mRTA—medial component of retrolateral tibial apophysis, PA—proximal arm of tegular apophysis, PTA—patellar apophysis, Tr—trichobothrium. Note: in Figs 31, 33–35 the distal end of the mRTA is a short rigid spine-like process; in Fig. 32 the mRTA terminates distally with a short basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158375" httpUri="https://zenodo.org/record/8158375/files/figure.png" pageId="3" pageNumber="100">Figs 31–35</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140AC9F7CA617AFF1C" box="[992,1106,186,213]" captionStart="FIGURES 53–59" captionStartId="18.[152,270,1632,1656]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-37@18.[151,1436,181,1608]" captionTargetPageId="18" captionText="FIGURES 53–59. Rothaeina gen. nov. species, males (53 R. jamesi spec. nov. holotype; 54–55 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 56–57 R. petersoni spec. nov. from near Kyburz, California; 58–59 R. sequoia comb. nov. holotype), left pedipalp. 53–54, 56, 58 Patellar apophysis and tibia, retrolateral. 55 Patella, dorsal. 57, 59 Tibia and patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis. Unlabelled arrow indicates basal hump of patellar apophysis (59). Note: in Figs 53, 58–59 the distal end of the mRTA is a rigid spine-like process; in Figs 56–57 the mRTA terminates distally in a basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158381" httpUri="https://zenodo.org/record/8158381/files/figure.png" pageId="3" pageNumber="100">53, 56–59</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140D72F7CB6150FF1C" box="[1115,1144,187,213]" captionStart="FIGURES 60–65" captionStartId="19.[152,270,1253,1277]" captionTargetBox="[151,1436,181,1229]" captionTargetId="figure-24@19.[151,1436,181,1229]" captionTargetPageId="19" captionText="FIGURES 60–65. Rothaeina gen. nov. species, males (60–62 R. jamesi spec. nov. holotype; 63 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 64–65 R. petersoni spec. nov. from near Kyburz, California), left pedipalp. 60 Proximal arm of tegular apophysis, ventral. 61 Medial component of retrolateral tibial apophysis, retrolateral. 62, 64–65 Patellar apophysis, dorsal. 63 Tip of proximal arm of tegular apophysis, retrolateral. Unlabelled arrows indicate diagnostic characters: basal projection of proximal arm of tegular apophysis (60); dorso-basal group of peg setae (62); dorsal keel of proximal arm of tegular apophysis (63); large and small peg setae (64–65)." figureDoi="http://doi.org/10.5281/zenodo.8158432" httpUri="https://zenodo.org/record/8158432/files/figure.png" pageId="3" pageNumber="100">61</figureCitation>
|
|||
|
|
) or, in
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140DECF7CB6074FF1C" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[1221,1372,187,213]" class="Arachnida" family="Cybaeidae" genus="Rothaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="mackinleyi" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A07140DECF7CB6074FF1C" box="[1221,1372,187,213]" italics="true" pageId="3" pageNumber="100">R. mackinleyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A07140C4BF7CB65E0FF30" bold="true" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A07140C4BF7CB65E0FF30" pageId="3" pageNumber="100" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, is a simple elongate ledge not terminating anteriorly in such a process or macroseta (
|
|||
|
|
<figureCitation id="132D2A38E42A07140DA3F7AF61F6FF30" box="[1162,1246,222,249]" captionStart="FIGURES 53–59" captionStartId="18.[152,270,1632,1656]" captionTargetBox="[151,1436,181,1605]" captionTargetId="figure-37@18.[151,1436,181,1608]" captionTargetPageId="18" captionText="FIGURES 53–59. Rothaeina gen. nov. species, males (53 R. jamesi spec. nov. holotype; 54–55 R. mackinleyi spec. nov. from Beaver Sulphur Forest Camp, Oregon; 56–57 R. petersoni spec. nov. from near Kyburz, California; 58–59 R. sequoia comb. nov. holotype), left pedipalp. 53–54, 56, 58 Patellar apophysis and tibia, retrolateral. 55 Patella, dorsal. 57, 59 Tibia and patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis. Unlabelled arrow indicates basal hump of patellar apophysis (59). Note: in Figs 53, 58–59 the distal end of the mRTA is a rigid spine-like process; in Figs 56–57 the mRTA terminates distally in a basally articulated macroseta." figureDoi="http://doi.org/10.5281/zenodo.8158381" httpUri="https://zenodo.org/record/8158381/files/figure.png" pageId="3" pageNumber="100">Fig. 54</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30C6536E42A071409EEF6726601FBFC" pageId="3" pageNumber="100" type="description">
|
|||
|
|
<paragraph id="8BA936BDE42A071409EEF672674EFE3C" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="100">
|
|||
|
|
<emphasis id="B962EAAFE42A071409EEF6726471FED5" bold="true" box="[199,345,258,284]" pageId="3" pageNumber="100">Description.</emphasis>
|
|||
|
|
As for
|
|||
|
|
<taxonomicName id="4C164D3EE42A0714088FF673672EFED4" box="[422,518,259,285]" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42A0714088FF673672EFED4" box="[422,518,259,285]" italics="true" pageId="3" pageNumber="100">Cybaeus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation id="EF874B4CE42A07140B17F6736622FED4" author="Copley, C. R. & Bennett, R. & Perlman, S. J." box="[574,778,259,285]" pageId="3" pageNumber="100" pagination="367 - 401" refId="ref22032" refString="Copley, C. R., Bennett, R. & Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
|
|||
|
|
Copley
|
|||
|
|
<emphasis id="B962EAAFE42A07140BBDF67367E3FED4" box="[660,715,259,285]" italics="true" pageId="3" pageNumber="100">et al.</emphasis>
|
|||
|
|
2009
|
|||
|
|
</bibRefCitation>
|
|||
|
|
) and in genus diagnosis in this paper.Additional descriptive characters presented here. Small- to medium-sized spiders, carapace lengths averaging
|
|||
|
|
<quantity id="4CEE9B58E42A07140D47F6576039FE88" box="[1134,1297,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="2.015" metricValueMax="2.23" metricValueMin="1.8" pageId="3" pageNumber="100" unit="mm" value="2.015" valueMax="2.23" valueMin="1.8">1.80–2.23 mm</quantity>
|
|||
|
|
(males) and
|
|||
|
|
<quantity id="4CEE9B58E42A071409BEF63A6411FEAC" box="[151,313,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.935" metricValueMax="2.02" metricValueMin="1.85" pageId="3" pageNumber="100" unit="mm" value="1.935" valueMax="2.02" valueMin="1.85">1.85–2.02 mm</quantity>
|
|||
|
|
(females); females usually slightly smaller than males. Eight eyes (
|
|||
|
|
<figureCitation id="132D2A38E42A07140D37F63B6175FEAC" box="[1054,1117,331,357]" captionStart="FIGURES 1–2" captionStartId="2.[152,270,779,803]" captionTargetBox="[151,1436,291,756]" captionTargetId="figure-108@2.[151,1436,291,756]" captionTargetPageId="2" captionText="FIGURES 1–2. Cybaeina and Rothaeina gen. nov. species (1 C. minuta female from Tofino, British Columbia; 2 R. petersoni spec. nov. male from vicinity of Yuba Pass, California). 1 Eye group, anterior. 2 Patellae, tibiae, and metatarsi of legs I and II, ventral showing ventral tibial macrosetal pattern of 2(+1p)-2-2-2-2-0 on leg I and 2(+1p)-2-2-2-1-2-0 on leg II. AME—anterior median eyes, I—leg 1, II—leg II, Mt—metatarsus, Pt—patella, Ti—tibia." figureDoi="http://doi.org/10.5281/zenodo.8158359" httpUri="https://zenodo.org/record/8158359/files/figure.png" pageId="3" pageNumber="100">Fig. 1</figureCitation>
|
|||
|
|
) similar to
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140DF5F63B6437FE40" authority="(Copley et al. 2009)" baseAuthorityName="Copley" baseAuthorityYear="2009" class="Arachnida" family="Cybaeidae" genus="Cybaeus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42A07140DF5F63B6014FEAC" box="[1244,1340,331,357]" italics="true" pageId="3" pageNumber="100">Cybaeus</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EF874B4CE42A07140C63F63B643FFE40" author="Copley, C. R. & Bennett, R. & Perlman, S. J." pageId="3" pageNumber="100" pagination="367 - 401" refId="ref22032" refString="Copley, C. R., Bennett, R. & Perlman, S. J. (2009) Systematics of Nearctic Cybaeus (Araneae: Cybaeidae). Invertebrate Systematics, 23, 367 - 401. https: // doi. org / 10.1071 / IS 09001" type="journal article" year="2009">
|
|||
|
|
Copley
|
|||
|
|
<emphasis id="B962EAAFE42A071409BEF61F65F9FE40" box="[151,209,367,393]" italics="true" pageId="3" pageNumber="100">et al.</emphasis>
|
|||
|
|
2009
|
|||
|
|
</bibRefCitation>
|
|||
|
|
)
|
|||
|
|
</taxonomicName>
|
|||
|
|
; anterior median eyes smallest, about 1/2 diameter of anterior lateral eyes. Legs pale yellow to light reddish brown, unmarked. Four (
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140B38F6E367A3FE64" authorityName="Chamberlin & Ivie" authorityYear="1942" box="[529,651,403,429]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="confusa">
|
|||
|
|
<emphasis id="B962EAAFE42A07140B38F6E367A3FE64" box="[529,651,403,429]" italics="true" pageId="3" pageNumber="100">C. confusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) or five complete pairs of ventral tibia I macrosetae; pattern usually 2(+1p)-2-1p-2-2-2-0 or 2(+1p)-2-1p-2-2-0. Abdomen pale to light gray without markings. Colulus a low swelling marked by two clumps of very few setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42A071409EEF68E6499FD60" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="100">
|
|||
|
|
<emphasis id="B962EAAFE42A071409EEF68E642AFDD0" bold="true" box="[199,258,510,537]" italics="true" pageId="3" pageNumber="100">Male</emphasis>
|
|||
|
|
: Patellar apophysis (
|
|||
|
|
<figureCitation id="132D2A38E42A071408CFF68F677DFDD0" box="[486,597,511,537]" captionStart-0="FIGURES 8–13" captionStart-1="FIGURES 14–18" captionStartId-0="7.[151,264,1603,1627]" captionStartId-1="8.[152,270,1515,1539]" captionTargetBox-0="[151,1435,181,1579]" captionTargetBox-1="[151,1436,448,1492]" captionTargetId-0="figure-19@7.[151,1436,181,1579]" captionTargetId-1="figure-169@8.[151,1436,448,1492]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." captionText-1="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi-0="http://doi.org/10.5281/zenodo.8158365" figureDoi-1="http://doi.org/10.5281/zenodo.8158367" httpUri-0="https://zenodo.org/record/8158365/files/figure.png" httpUri-1="https://zenodo.org/record/8158367/files/figure.png" pageId="3" pageNumber="100">Figs 8–14</figureCitation>
|
|||
|
|
) with two to nine peg setae. Distal component of retrolateral tibial apophysis (
|
|||
|
|
<figureCitation id="132D2A38E42A071409B6F5536415FDF4" box="[159,317,547,573]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 8, 10, 12</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140860F55364B8FDF4" box="[329,400,547,573]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="3" pageNumber="100">14, 16</figureCitation>
|
|||
|
|
) nearly as long as pedipalpal tibia, usually slightly extended distally. Genital bulb with very long, thin, simple embolus (
|
|||
|
|
<figureCitation id="132D2A38E42A071408E4F537671BFDA8" box="[461,563,583,609]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Figs 3–7</figureCitation>
|
|||
|
|
); distal arm of tegular apophysis well developed with narrow (
|
|||
|
|
<figureCitation id="132D2A38E42A07140DC1F5376004FDA8" box="[1256,1324,583,609]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Fig. 6</figureCitation>
|
|||
|
|
), tapering (
|
|||
|
|
<figureCitation id="132D2A38E42A071409B6F51B65CAFD4C" box="[159,226,618,645]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Fig. 5</figureCitation>
|
|||
|
|
), or slightly expanded tip (
|
|||
|
|
<figureCitation id="132D2A38E42A07140B23F51B6747FD4C" box="[522,623,619,645]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Figs 3–4</figureCitation>
|
|||
|
|
); proximal arm well developed but simple: elongate, slightly twisted with acuminate tip (
|
|||
|
|
<figureCitation id="132D2A38E42A07140817F5FF648CFD60" box="[318,420,655,681]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Figs 3–7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42A071409EEF5C260BCFC94" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="100">
|
|||
|
|
<emphasis id="B962EAAFE42A071409EEF5C26436FD04" bold="true" box="[199,286,690,717]" italics="true" pageId="3" pageNumber="100">Female</emphasis>
|
|||
|
|
: Epigyne (
|
|||
|
|
<figureCitation id="132D2A38E42A071408B5F5C364DCFD04" box="[412,500,691,717]" captionStart="FIGURES 19–21" captionStartId="9.[152,270,1259,1283]" captionTargetBox="[153,1433,181,1235]" captionTargetId="figure-19@9.[151,1436,181,1235]" captionTargetPageId="9" captionText="FIGURES 19–21. Cybaeina confusa, females (19 from Rose Harbour, British Columbia; 20–21 holotype), copulatory organ. 19 Epigyne, ventral. 20–21 Vulva (20 ventral, 21 dorsal). BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (19–21)." figureDoi="http://doi.org/10.5281/zenodo.8158369" httpUri="https://zenodo.org/record/8158369/files/figure.png" pageId="3" pageNumber="100">Figs 19</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140B28F5C36762FD04" box="[513,586,690,717]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="3" pageNumber="100">22, 25</figureCitation>
|
|||
|
|
) simple with single, transverse atrium medially or postero-medially. Vulva (
|
|||
|
|
<figureCitation id="132D2A38E42A071409B6F5A76437FD38" box="[159,287,727,753]" captionStart="FIGURES 19–21" captionStartId="9.[152,270,1259,1283]" captionTargetBox="[153,1433,181,1235]" captionTargetId="figure-19@9.[151,1436,181,1235]" captionTargetPageId="9" captionText="FIGURES 19–21. Cybaeina confusa, females (19 from Rose Harbour, British Columbia; 20–21 holotype), copulatory organ. 19 Epigyne, ventral. 20–21 Vulva (20 ventral, 21 dorsal). BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (19–21)." figureDoi="http://doi.org/10.5281/zenodo.8158369" httpUri="https://zenodo.org/record/8158369/files/figure.png" pageId="3" pageNumber="100">Figs 20–21</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140800F5A764ECFD38" box="[297,452,727,753]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="3" pageNumber="100">23–24, 26–27</figureCitation>
|
|||
|
|
) with long, slender, weakly sclerotized copulatory ducts, broadly or narrowly contiguous at atrium, looped around spermathecal stalks once to up to 2.5 times; spermathecal heads at anterior margin of vulva; spermathecal stalks relatively elongate, convoluted, narrow; spermathecal bases small, rounded or undifferentiated from stalks; fertilization ducts exit bases posteriorly; Bennett’s glands hidden by copulatory ducts or conspicuous.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42A071409EEF4166601FBFC" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="100">
|
|||
|
|
<emphasis id="B962EAAFE42A071409EEF4166702FC49" bold="true" box="[199,554,870,897]" pageId="3" pageNumber="100">Composition and distribution</emphasis>
|
|||
|
|
. (
|
|||
|
|
<figureCitation id="132D2A38E42A07140B69F41767BDFC48" box="[576,661,871,897]" captionStart="FIGURE 28" captionStartId="12.[152,255,1946,1970]" captionTargetBox="[171,1421,197,1914]" captionTargetId="figure-37@12.[157,1431,181,1922]" captionTargetPageId="12" captionText="FIGURE 28. Distributions of Cybaeina species. Cybaeina confusa, C dixoni spec. nov., and C. minuta. AB—Alberta, AZ— Arizona, BC—British Columbia, CA—California, ID—Idaho, MO—Montana, OR—Oregon, NV—Nevada, UT—Utah, WA— Washington." figureDoi="http://doi.org/10.5281/zenodo.8158373" httpUri="https://zenodo.org/record/8158373/files/figure.png" pageId="3" pageNumber="100">Fig. 28</figureCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140B80F417663CFC48" box="[681,788,871,897]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42A07140B80F417663CFC48" box="[681,788,871,897]" italics="true" pageId="3" pageNumber="100">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is endemic to coastal western North America from Haida Gwaii and the adjacent mainland islands of
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140BB8F4FB667FFC6C" box="[657,855,907,933]" country="Canada" name="British Columbia" pageId="3" pageNumber="100">British Columbia</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F301762DE42A07140A4CF4FB6691FC6C" box="[869,953,907,933]" name="Canada" pageId="3" pageNumber="100">Canada</collectingCountry>
|
|||
|
|
to
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140AC9F4FB614DFC6C" box="[992,1125,907,933]" country="United States of America" name="Washington" pageId="3" pageNumber="100">Washington</collectingRegion>
|
|||
|
|
and
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140D89F4FB61DFFC6C" box="[1184,1271,907,933]" country="United States of America" name="Oregon" pageId="3" pageNumber="100">Oregon</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F301762DE42A07140C2DF4FB6433FC00" name="United States of America" pageId="3" pageNumber="100">United States of America</collectingCountry>
|
|||
|
|
. We recognize three species:
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140B5AF4DF6668FC00" authorityName="Chamberlin & Ivie" authorityYear="1942" box="[627,832,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="confusa">
|
|||
|
|
<emphasis id="B962EAAFE42A07140B5AF4DF6668FC00" box="[627,832,943,969]" italics="true" pageId="3" pageNumber="100">Cybaeina confusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140A78F4DF6693FC00" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[849,955,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="dixoni" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A07140A78F4DF6693FC00" box="[849,955,943,969]" italics="true" pageId="3" pageNumber="100">C. dixoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A07140AECF4DF6113FC00" bold="true" box="[965,1083,943,969]" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A07140AECF4DF6113FC00" box="[965,1083,943,969]" pageId="3" pageNumber="100" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140D56F4DF61D9FC01" authorityName="Chamberlin and Ivie" authorityYear="1932" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1151,1265,943,969]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="minuta">
|
|||
|
|
<emphasis id="B962EAAFE42A07140D56F4DF61D9FC01" box="[1151,1265,943,969]" italics="true" pageId="3" pageNumber="100">C. minuta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. All are rarely encountered;
|
|||
|
|
<taxonomicName id="4C164D3EE42A0714081BF4A364B4FC24" authority="Bennett & Copley & Copley, 2023" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[306,412,979,1005]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="dixoni" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A0714081BF4A364B4FC24" box="[306,412,979,1005]" italics="true" pageId="3" pageNumber="100">C. dixoni</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A0714088EF4A36735FC24" bold="true" box="[423,541,979,1005]" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A0714088EF4A36735FC24" box="[423,541,979,1005]" pageId="3" pageNumber="100" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
is known only from southwestern Curry County,
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140D4DF4A36194FC24" box="[1124,1212,979,1005]" country="United States of America" name="Oregon" pageId="3" pageNumber="100">Oregon</collectingRegion>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140DE3F4A3606DFC24" authorityName="Chamberlin & Ivie" authorityYear="1942" box="[1226,1349,979,1005]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="confusa">
|
|||
|
|
<emphasis id="B962EAAFE42A07140DE3F4A3606DFC24" box="[1226,1349,979,1005]" italics="true" pageId="3" pageNumber="100">C. confusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140CAAF4A365CDFBD9" authorityName="Chamberlin and Ivie" authorityYear="1932" baseAuthorityName="Banks" baseAuthorityYear="1906" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="minuta">
|
|||
|
|
<emphasis id="B962EAAFE42A07140CAAF4A365CDFBD9" italics="true" pageId="3" pageNumber="100">C. minuta</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are relatively widespread with records from west coastal North America from
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140D5AF487601FFBD8" box="[1139,1335,1015,1041]" inferredBySuffix="true" pageId="3" pageNumber="100">British Columbia</collectingRegion>
|
|||
|
|
south to
|
|||
|
|
<collectingRegion id="49D2F85FE42A071409BEF36B65C3FBFC" box="[151,235,1051,1077]" country="United States of America" name="Oregon" pageId="3" pageNumber="100">Oregon</collectingRegion>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="4C164D3EE42A071409D2F36B645BFBFC" authorityName="Chamberlin & Ivie" authorityYear="1942" box="[251,371,1051,1077]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="confusa">
|
|||
|
|
<emphasis id="B962EAAFE42A071409D2F36B645BFBFC" box="[251,371,1051,1077]" italics="true" pageId="3" pageNumber="100">C. confusa</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has not been recorded in
|
|||
|
|
<collectingRegion id="49D2F85FE42A07140BBBF36B6635FBFC" box="[658,797,1051,1077]" country="United States of America" name="Washington" pageId="3" pageNumber="100">Washington</collectingRegion>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C30C6536E42A071409BEF3F760B5F98B" pageId="3" pageNumber="100" type="key">
|
|||
|
|
<paragraph id="8BA936BDE42A071409BEF3F76727FB68" blockId="3.[151,527,1158,1185]" box="[151,527,1158,1185]" pageId="3" pageNumber="100">
|
|||
|
|
<heading id="D0E181D1E42A071409BEF3F76727FB68" bold="true" box="[151,527,1158,1185]" fontSize="11" level="1" pageId="3" pageNumber="100" reason="1">
|
|||
|
|
<emphasis id="B962EAAFE42A071409BEF3F76727FB68" bold="true" box="[151,527,1158,1185]" pageId="3" pageNumber="100">
|
|||
|
|
Key to the species of
|
|||
|
|
<taxonomicName id="4C164D3EE42A071408B4F3F66727FB68" ID-CoL="3XS8" ID-ENA="880878" box="[413,527,1158,1185]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B962EAAFE42A071408B4F3F66727FB68" bold="true" box="[413,527,1158,1185]" italics="true" pageId="3" pageNumber="100">Cybaeina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<keyStep id="30E22E18E42A071409BEF3BC60B4FA58" pageId="3" pageNumber="100">
|
|||
|
|
<paragraph id="8BA936BDE42A071409BEF3BC60B4FA9E" blockId="3.[151,1437,1228,1602]" pageId="3" pageNumber="100">
|
|||
|
|
<keyLead id="30E79588E42A071409BEF3BC60B4FA9E" pageId="3" pageNumber="100">
|
|||
|
|
1. Four pairs of elongate, overlapping ventral tibia I macrosetae. Patellar apophysis about 1/2 width of patella with three or four peg setae (
|
|||
|
|
<figureCitation id="132D2A38E42A07140837F399645DFB37" box="[286,373,1257,1279]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 8–9</figureCitation>
|
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|
|
); medial component of retrolateral tibial apophysis distally terminated with single basally articulated macroseta (
|
|||
|
|
<figureCitation id="132D2A38E42A071409C0F276646CFAD2" box="[233,324,1286,1308]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 8–9</figureCitation>
|
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); distal arm of tegular apophysis dorsad, tip slightly expanded (
|
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|
|
<figureCitation id="132D2A38E42A07140AECF2766108FAD2" box="[965,1056,1286,1308]" captionStart="FIGURES 3–7" captionStartId="4.[152,270,1569,1593]" captionTargetBox="[151,1436,181,1545]" captionTargetId="figure-41@4.[151,1436,181,1545]" captionTargetPageId="4" captionText="FIGURES 3–7. Cybaeina species, males (3 C. confusa from Cascadia, Oregon; 4 C. confusa from unknown locality; 5 C. dixoni spec. nov. from Brookings, Oregon; 6–7 C. minuta from Kyuquot, British Columbia), left pedipalp. 3, 5–6 Genital bulb, ventral. 4, 7 Tegular apophysis and embolus, retrolateral. Cy—cymbium, DA—distal arm of tegular apophysis, E—embolus, PA—proximal arm of tegular apophysis, Ti—tibia, TR—tegular ridge." figureDoi="http://doi.org/10.5281/zenodo.8158363" httpUri="https://zenodo.org/record/8158363/files/figure.png" pageId="3" pageNumber="100">Figs 3–4</figureCitation>
|
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). Spermathecal bases undifferentiated, indistinguishable from spermathecal stalks (
|
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<figureCitation id="132D2A38E42A07140BA0F25467D1FAF0" box="[649,761,1316,1338]" captionStart="FIGURES 19–21" captionStartId="9.[152,270,1259,1283]" captionTargetBox="[153,1433,181,1235]" captionTargetId="figure-19@9.[151,1436,181,1235]" captionTargetPageId="9" captionText="FIGURES 19–21. Cybaeina confusa, females (19 from Rose Harbour, British Columbia; 20–21 holotype), copulatory organ. 19 Epigyne, ventral. 20–21 Vulva (20 ventral, 21 dorsal). BS—base of spermatheca, CD—copulatory duct, FD—fertilization duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (19–21)." figureDoi="http://doi.org/10.5281/zenodo.8158369" httpUri="https://zenodo.org/record/8158369/files/figure.png" pageId="3" pageNumber="100">Figs 20–21</figureCitation>
|
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); Bennett’s glands hidden within coils of vulval ducts...........................................................................................
|
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<taxonomicName id="4C164D3EE42A07140DBFF23160B4FA9E" authority="Chamberlin & Ivie" authorityName="Chamberlin & Ivie" authorityYear="1942" box="[1174,1436,1345,1367]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="confusa">
|
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|
|
<emphasis id="B962EAAFE42A07140DBFF23161C9FA9F" box="[1174,1249,1345,1366]" italics="true" pageId="3" pageNumber="100">confusa</emphasis>
|
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Chamberlin & Ivie
|
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</taxonomicName>
|
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</keyLead>
|
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|
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</paragraph>
|
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|
|
<paragraph id="8BA936BDE42A071409BEF22F60B4FA58" blockId="3.[151,1437,1228,1602]" pageId="3" pageNumber="100">
|
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|
|
<keyLead id="30E79588E42A071409BEF22F60B4FA58" pageId="3" pageNumber="100">
|
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- Usually five pairs of elongate, overlapping ventral tibia I macrosetae (as in
|
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|
<bibRefCitation id="EF874B4CE42A07140A94F22F616BFABA" author="Bennett, R. G." box="[957,1091,1374,1396]" pageId="3" pageNumber="100" pagination="96 - 101" refId="ref20957" refString="Bennett, R. G. (2017) Cybaeidae. In: Ubick, D., Paquin, P., Cushing, P. E. & Roth, V. D. (Eds.), Spiders of North America: an Identification Manual, 2 nd ed. American Arachnological Society, pp. 96 - 101." type="book chapter" year="2017">Bennett 2017</bibRefCitation>
|
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|
|
: fig. 23.13). Other characters not as above............................................................................................... 2
|
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</keyLead>
|
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|
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</paragraph>
|
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|
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</keyStep>
|
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|
|
<keyStep id="30E22E18E42A071409BEF2E960B5F98B" pageId="3" pageNumber="100">
|
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|
|
<paragraph id="8BA936BDE42A071409BEF2E960B4FA20" blockId="3.[151,1437,1228,1602]" pageId="3" pageNumber="100">
|
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|
|
<keyLead id="30E79588E42A071409BEF2E960B4FA20" pageId="3" pageNumber="100">
|
|||
|
|
2(1). Patellar apophysis short, dorsoventrally broad with about nine peg setae (
|
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|
|
<figureCitation id="132D2A38E42A07140A9BF2E9610AFA67" box="[946,1058,1433,1455]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 10–11</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140D07F2E96161FA67" box="[1070,1097,1433,1454]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="3" pageNumber="100">14</figureCitation>
|
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|
|
); medial component of retrolateral tibial apophysis a distinct angular, distally acuminate ledge (
|
|||
|
|
<figureCitation id="132D2A38E42A07140A1BF2C66689FA02" box="[818,929,1462,1484]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 10–11</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140A84F2C666C4FA05" box="[941,1004,1462,1484]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="3" pageNumber="100">14–15</figureCitation>
|
|||
|
|
). Copulatory ducts make one complete loop around spermathecal stalks (
|
|||
|
|
<figureCitation id="132D2A38E42A071408DBF2A4674BFA23" box="[498,611,1492,1514]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="3" pageNumber="100">Figs 26–27</figureCitation>
|
|||
|
|
); Bennett’s glands prominent in dorsal view (
|
|||
|
|
<figureCitation id="132D2A38E42A07140D3EF2A46175FA20" box="[1047,1117,1492,1514]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="3" pageNumber="100">Fig. 27</figureCitation>
|
|||
|
|
).....
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140D81F2A4601DFA20" authority="Bennett" authorityName="Bennett & Copley & Copley" authorityYear="2023" box="[1192,1333,1492,1513]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="dixoni" status="spec. nov.">
|
|||
|
|
<emphasis id="B962EAAFE42A07140D81F2A461CCFA20" box="[1192,1252,1492,1513]" italics="true" pageId="3" pageNumber="100">dixoni</emphasis>
|
|||
|
|
Bennett
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B962EAAFE42A07140C13F2A460B4FA20" bold="true" box="[1338,1436,1492,1513]" pageId="3" pageNumber="100">
|
|||
|
|
<taxonomicNameLabel id="A25157D4E42A07140C13F2A460B4FA20" box="[1338,1436,1492,1513]" pageId="3" pageNumber="100" rank="species">spec. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BA936BDE42A071409BEF28160B5F98B" blockId="3.[151,1437,1228,1602]" pageId="3" pageNumber="100">
|
|||
|
|
<keyLead id="30E79588E42A071409BEF28160B5F98B" pageId="3" pageNumber="100">
|
|||
|
|
- Patellar apophysis small, with two peg setae (
|
|||
|
|
<figureCitation id="132D2A38E42A07140B8DF281663EF9CF" box="[676,790,1521,1543]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 12–13</figureCitation>
|
|||
|
|
); medial component of retrolateral tibial apophysis inconspicuous except for an elongate acuminate process distally (
|
|||
|
|
<figureCitation id="132D2A38E42A07140B97F17E6620F9EA" box="[702,776,1550,1572]" captionStart="FIGURES 8–13" captionStartId="7.[151,264,1603,1627]" captionTargetBox="[151,1435,181,1579]" captionTargetId="figure-19@7.[151,1436,181,1579]" captionTargetPageId="7" captionText="FIGURES 8–13. Cybaeina species, males (8–9 C. confusa from Cascadia, Oregon; 10–11 C. dixoni spec. nov. holotype; 12–13 C. minuta from Kyuquot, British Columbia), left pedipalp. 8, 12 Tibia and distal portion of patella, retrolateral. 10 Tibia and patella, retrolateral. 9, 11 Tibia and patella, dorsal. 13 left half of patella, dorsal. Cy—cymbium, dRTA—distal component of retrolateral tibial apophysis, mRTA—medial component of retrolateral tibial apophysis, PTA—patellar apophysis, Ti—tibia. Note: in Figs 8–9 the mRTA terminates distally in a basally articulated macroseta; in Figs 10–12 the distal end is a short rigid spine-like process." figureDoi="http://doi.org/10.5281/zenodo.8158365" httpUri="https://zenodo.org/record/8158365/files/figure.png" pageId="3" pageNumber="100">Figs 12</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="132D2A38E42A07140A39F17E6678F9ED" box="[784,848,1550,1572]" captionStart="FIGURES 14–18" captionStartId="8.[152,270,1515,1539]" captionTargetBox="[151,1436,448,1492]" captionTargetId="figure-169@8.[151,1436,448,1492]" captionTargetPageId="8" captionText="FIGURES 14–18. Cybaeina and Neocybaeina gen. nov. species, males (14–15 C. dixoni spec. nov. holotype; 16–17 C. minuta from Kyuquot, British Columbia; 18 N. xantha comb. nov. from near Bridge, Oregon), left pedipalp retrolateral. 14, 16 tibia and distal tip of patellar apophysis.15, 17 medial component of retrolateral tibial apophysis.18 distal tip of patellar apophysis.Arrows indicate medial component of retrolateral tibial apophysis (14–17 single shafted arrows), distal component of retrolateral tibial apophysis (14, 16 double shafted arrows), unknown contaminant on dorsal surface of patellar apophysis (18 dashed arrow)." figureDoi="http://doi.org/10.5281/zenodo.8158367" httpUri="https://zenodo.org/record/8158367/files/figure.png" pageId="3" pageNumber="100">16–17</figureCitation>
|
|||
|
|
). Copulatory ducts make about 2.5 loops around spermathecal stalks (
|
|||
|
|
<figureCitation id="132D2A38E42A0714080CF15C64BEF988" box="[293,406,1580,1602]" captionStart="FIGURES 22–27" captionStartId="11.[152,270,1811,1835]" captionTargetBox="[151,1436,182,1787]" captionTargetId="figure-19@11.[151,1436,181,1787]" captionTargetPageId="11" captionText="FIGURES 22–27. Cybaeina species, females (22–24 C. minuta from Tofino, British Columbia; 25–27 C. dixoni spec. nov. from Brookings, Oregon), copulatory organ. 22, 25 Epigyne, ventral. 23–24, 26–27 Vulva (23, 26 ventral; 24, 27 dorsal). BG—Bennett’s gland, BS—base of spermatheca, CD—copulatory duct, HS—head of spermatheca, SS—stalk of spermatheca. Unlabelled arrows indicate atrial openings (22, 23, 25–26)." figureDoi="http://doi.org/10.5281/zenodo.8158371" httpUri="https://zenodo.org/record/8158371/files/figure.png" pageId="3" pageNumber="100">Figs 23–24</figureCitation>
|
|||
|
|
); Bennett’s glands inconspicuous...............................................
|
|||
|
|
<taxonomicName id="4C164D3EE42A07140C21F15C60B5F98B" authority="(Banks)" authorityName="Chamberlin and Ivie" authorityYear="1932" baseAuthorityName="Banks" baseAuthorityYear="1906" box="[1288,1437,1580,1602]" class="Arachnida" family="Cybaeidae" genus="Cybaeina" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="100" phylum="Arthropoda" rank="species" species="minuta">
|
|||
|
|
<emphasis id="B962EAAFE42A07140C21F15C6063F988" box="[1288,1355,1580,1601]" italics="true" pageId="3" pageNumber="100">minuta</emphasis>
|
|||
|
|
(Banks)
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|