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<document id="7E5E5343CEEE40EEA1270BCFE0DC45FA" ID-CLB-Dataset="26834" ID-DOI="10.11646/zootaxa.4604.2.2" ID-GBIF-Dataset="943ac4e5-eee2-409e-a058-eb76866f102a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2806972" ID-ZooBank="3B7FA7A1-3EFC-4AC2-B08C-0A8FD52247A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557812968979" checkinUser="plazi" docAuthor="Tixier, Marie-Stéphane, Dennj, Principato, Douin, Martial, Kreiter, Serge &amp; Haralabos, Tsolakis" docDate="2019" docId="03A10A70FFCDFF92FF55FB18EE6CFDBE" docLanguage="en" docName="zootaxa.4604.2.2.pdf" docOrigin="Zootaxa 4604 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Typhlodromus (Typhlodromus) exhilaratus Ragusa 1977" docType="treatment" docVersion="10" lastPageNumber="265" masterDocId="FF987208FFDBFF85FFC2FFF3EC3BFFDD" masterDocTitle="Mites of the genus Typhlodromus (Acari: Phytoseiidae) from Southern France: combined morphological and molecular approaches for species identification" masterLastPageNumber="280" masterPageNumber="242" pageNumber="264" updateTime="1698728565905" updateUser="ExternalLinkService">
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<mods:title id="6FC0DC54ED4607555729B2BBA50B5A38">Mites of the genus Typhlodromus (Acari: Phytoseiidae) from Southern France: combined morphological and molecular approaches for species identification</mods:title>
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<mods:namePart id="4BCBE29AC5DE2E1B6CE4665D88D8B4D1">Tixier, Marie-Stéphane</mods:namePart>
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<mods:namePart id="29F67536FDDA8C2959F81EAD02B63014">Dennj, Principato</mods:namePart>
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<mods:namePart id="9EF2D23F9040E884597951BD119387F9">Douin, Martial</mods:namePart>
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<mods:namePart id="4CA36DAF527C7427E893F5784A25D6ED">Kreiter, Serge</mods:namePart>
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<mods:namePart id="01075701C661CD72AA185D618E7B1E63">Haralabos, Tsolakis</mods:namePart>
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<mods:title id="922317D97122A49DD70221599AA18066">Zootaxa</mods:title>
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<mods:date id="D71A0EA6DC9811F4AB2D712442757A92">2019</mods:date>
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<mods:number id="8CD2CBD916B23C494E10BB20652FFDBE">4604</mods:number>
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<treatment id="03A10A70FFCDFF92FF55FB18EE6CFDBE" ID-DOI="http://doi.org/10.5281/zenodo.5585259" ID-GBIF-Taxon="157137040" ID-Zenodo-Dep="5585259" LSID="urn:lsid:plazi:treatment:03A10A70FFCDFF92FF55FB18EE6CFDBE" httpUri="http://treatment.plazi.org/id/03A10A70FFCDFF92FF55FB18EE6CFDBE" lastPageId="23" lastPageNumber="265" pageId="22" pageNumber="264">
<subSubSection id="C312E8EDFFCDFF93FF55FB18EF75FAD8" box="[151,846,1259,1286]" pageId="22" pageNumber="264" type="nomenclature">
<paragraph id="8BB7BB66FFCDFF93FF55FB18EF75FAD8" blockId="22.[151,846,1259,1286]" box="[151,846,1259,1286]" pageId="22" pageNumber="264">
<heading id="D0FF0C0AFFCDFF93FF55FB18EF75FAD8" bold="true" box="[151,846,1259,1286]" fontSize="11" level="1" pageId="22" pageNumber="264" reason="1">
<taxonomicName id="4C08C0E5FFCDFF93FF55FB18EF75FAD8" ID-CoL="59WQ2" authority="Ragusa, 1977" authorityName="Ragusa" authorityYear="1977" box="[151,846,1259,1286]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="264" phylum="Arthropoda" rank="species" species="exhilaratus" subGenus="Typhlodromus">
<emphasis id="B97C6774FFCDFF93FF55FB18EF75FAD8" bold="true" box="[151,846,1259,1286]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF55FB18ED7DFADB" bold="true" box="[151,326,1259,1286]" italics="true" pageId="22" pageNumber="264">Typhlodromus</emphasis>
(
<emphasis id="B97C6774FFCDFF93FE95FB18EE3DFADB" bold="true" box="[343,518,1259,1286]" italics="true" pageId="22" pageNumber="264">Typhlodromus</emphasis>
)
<emphasis id="B97C6774FFCDFF93FDD5FB18EE9BFADB" bold="true" box="[535,672,1259,1286]" italics="true" pageId="22" pageNumber="264">exhilaratus</emphasis>
<bibRefCitation id="EF99C697FFCDFF93FD6AFB1FEF75FAD8" author="Ragusa, S." box="[680,846,1259,1286]" pageId="22" pageNumber="264" pagination="379 - 392" refId="ref21723" refString="Ragusa, S. (1977) Notes on phytoseiid mites in Sicily with a description of a new species of Typhlodromus (Acarina: Mesostigmata). Acarologia, 18, 379 - 392." type="journal article" year="1977">Ragusa, 1977</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C312E8EDFFCDFF93FF55FADCEF9CFABB" pageId="22" pageNumber="264" type="reference_group">
<paragraph id="8BB7BB66FFCDFF93FF55FADCEE5FFA95" blockId="22.[151,939,1327,1413]" box="[151,612,1327,1352]" pageId="22" pageNumber="264">
<treatmentCitationGroup id="AB189C48FFCDFF93FF55FADCEE5FFA95" box="[151,612,1327,1352]" pageId="22" pageNumber="264">
<taxonomicName id="4C08C0E5FFCDFF93FF55FADCEE64FA95" ID-CoL="59WQ2" authority="Ragusa 1977: 380" authorityName="Ragusa" authorityPageNumber="380" authorityYear="1977" box="[151,607,1327,1352]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="264" phylum="Arthropoda" rank="species" species="exhilaratus">
<emphasis id="B97C6774FFCDFF93FF55FADCEDA4FA9A" box="[151,415,1327,1351]" italics="true" pageId="22" pageNumber="264">Typhlodromus exhilaratus</emphasis>
<treatmentCitation id="0AA99D77FFCDFF93FE67FAC3EE64FA95" author="Ragusa, S." box="[421,607,1328,1352]" page="380" pageId="22" pageNumber="264" year="1977">
<bibRefCitation id="EF99C697FFCDFF93FE67FAC3EE17FA95" author="Ragusa, S." box="[421,556,1328,1352]" pageId="22" pageNumber="264" pagination="379 - 392" refId="ref21723" refString="Ragusa, S. (1977) Notes on phytoseiid mites in Sicily with a description of a new species of Typhlodromus (Acarina: Mesostigmata). Acarologia, 18, 379 - 392." type="journal article" year="1977">Ragusa 1977</bibRefCitation>
: 380
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB7BB66FFCDFF93FF55FABDEF9CFABB" blockId="22.[151,939,1327,1413]" box="[151,935,1357,1382]" pageId="22" pageNumber="264">
<treatmentCitationGroup id="AB189C48FFCDFF93FF55FABDEF9CFABB" box="[151,935,1357,1382]" pageId="22" pageNumber="264">
<taxonomicName id="4C08C0E5FFCDFF93FF55FABDEE2CFABB" ID-CoL="59WQ2" authority="exhilaratus" box="[151,535,1358,1382]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="264" phylum="Arthropoda" rank="subSpecies" species="exhilaratus" subSpecies="exhilaratus">
<emphasis id="B97C6774FFCDFF93FF55FABDEE2CFABB" box="[151,535,1358,1382]" italics="true" pageId="22" pageNumber="264">Typhlodromus exhilaratus exhilaratus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AA99D77FFCDFF93FDF3FABEEF98FABB" author="Chant, D. A. &amp; Yoshida-Shaul, E." box="[561,931,1357,1382]" page="1795" pageId="22" pageNumber="264" year="1987">
<bibRefCitation id="EF99C697FFCDFF93FDF3FABEEF58FABB" author="Chant, D. A. &amp; Yoshida-Shaul, E." box="[561,867,1357,1382]" pageId="22" pageNumber="264" pagination="1770 - 1804" refId="ref20083" refString="Chant, D. A. &amp; Yoshida-Shaul, E. (1987) A world review of the pyri species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae). Canadian Journal of Zoology, 65, 1770 - 1804. https: // doi. org / 10.1139 / z 87 - 272" type="journal article" year="1987">Chant &amp; Yoshida-Shaul, 1987</bibRefCitation>
: 1795
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C312E8EDFFCDFF92FF55FA9EEE6CFDBE" lastPageId="23" lastPageNumber="265" pageId="22" pageNumber="264" type="description">
<paragraph id="8BB7BB66FFCDFF93FF55FA9EEF90FA58" blockId="22.[151,939,1327,1413]" box="[151,939,1388,1413]" pageId="22" pageNumber="264">
<treatmentCitationGroup id="AB189C48FFCDFF93FF55FA9EEF90FA58" box="[151,939,1388,1413]" pageId="22" pageNumber="264">
<taxonomicName id="4C08C0E5FFCDFF93FF55FA9EEE20FA58" ID-CoL="59WNC" box="[151,539,1389,1413]" class="Arachnida" family="Phytoseiidae" genus="Typhlodromus" kingdom="Animalia" order="Mesostigmata" pageId="22" pageNumber="264" phylum="Arthropoda" rank="subSpecies" species="exhilaratus" subSpecies="americanus">
<emphasis id="B97C6774FFCDFF93FF55FA9EEE20FA58" box="[151,539,1389,1413]" italics="true" pageId="22" pageNumber="264">Typhlodromus exhilaratus americanus</emphasis>
</taxonomicName>
—
<treatmentCitation id="0AA99D77FFCDFF93FDF7FA9FEF93FA58" author="Chant, D. A. &amp; Yoshida-Shaul, E." box="[565,936,1388,1413]" page="1796" pageId="22" pageNumber="264" year="1987">
<bibRefCitation id="EF99C697FFCDFF93FDF7FA9FEF53FA58" author="Chant, D. A. &amp; Yoshida-Shaul, E." box="[565,872,1388,1413]" pageId="22" pageNumber="264" pagination="1770 - 1804" refId="ref20083" refString="Chant, D. A. &amp; Yoshida-Shaul, E. (1987) A world review of the pyri species group in the genus Typhlodromus Scheuten (Acari: Phytoseiidae). Canadian Journal of Zoology, 65, 1770 - 1804. https: // doi. org / 10.1139 / z 87 - 272" type="journal article" year="1987">Chant &amp; Yoshida-Shaul, 1987</bibRefCitation>
: 1796
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB7BB66FFCDFF93FF55FA41EDDDF9E4" blockId="22.[151,1437,1458,2026]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF55FA41EDB4FA10" bold="true" box="[151,399,1458,1485]" pageId="22" pageNumber="264">Specimens collected.</emphasis>
One female on
<taxonomicName id="4C08C0E5FFCDFF93FD8CFA47EF74FA10" box="[590,847,1459,1485]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" kingdom="Plantae" order="Lamiales" pageId="22" pageNumber="264" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="B97C6774FFCDFF93FD8CFA47EF74FA10" box="[590,847,1459,1485]" italics="true" pageId="22" pageNumber="264">Rosmarinus officinalis</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C08C0E5FFCDFF93FC41FA40E839FA13" box="[899,1026,1459,1486]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="22" pageNumber="264" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
),
<specimenCount id="9D0E70EFFFCDFF93FBDAFA40E8BCFA10" box="[1048,1159,1459,1485]" pageId="22" pageNumber="264" type="female">2 females</specimenCount>
on
<taxonomicName id="4C08C0E5FFCDFF93FB7AFA40E941FA10" box="[1208,1402,1459,1485]" class="Magnoliopsida" family="Rosaceae" genus="Malus" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="264" phylum="Tracheophyta" rank="species" species="domestica">
<emphasis id="B97C6774FFCDFF93FB7AFA40E941FA10" box="[1208,1402,1459,1485]" italics="true" pageId="22" pageNumber="264">Malus domestica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C08C0E5FFCDFF93FF5DFA24ED34FA2C" box="[159,271,1495,1521]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="264" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
), Montpellier SupAgro campus,
<collectingCountry id="F31FFBF6FFCDFF93FD4DFA2BEEE0FA2F" box="[655,731,1496,1522]" name="France" pageId="22" pageNumber="264">France</collectingCountry>
(
<geoCoordinate id="EE3CDDA1FFCDFF93FD32FA25EFA9FA2C" box="[752,914,1494,1521]" direction="north" orientation="latitude" pageId="22" pageNumber="264" precision="1" value="43.615387">43°36’55.4”N</geoCoordinate>
,
<geoCoordinate id="EE3CDDA1FFCDFF93FC60FA25E809FA2C" box="[930,1074,1494,1521]" direction="east" orientation="longitude" pageId="22" pageNumber="264" precision="1" value="3.861222">3°51’40.4”E</geoCoordinate>
), on
<date id="FFB69DA6FFCDFF93FBB0FA24E8CFFA2C" box="[1138,1268,1495,1521]" pageId="22" pageNumber="264" value="2017-09-11">11/09/2017</date>
,
<specimenCount id="9D0E70EFFFCDFF93FAC6FA24E94FFA2C" box="[1284,1396,1495,1521]" pageId="22" pageNumber="264" type="female">6 females</specimenCount>
on
<taxonomicName id="4C08C0E5FFCDFF93FF55FA08EEBFF9CB" authority="Miller" authorityName="Miller" box="[151,644,1531,1558]" class="Magnoliopsida" family="Lamiaceae" genus="Lavandula" kingdom="Plantae" order="Lamiales" pageId="22" pageNumber="264" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="B97C6774FFCDFF93FF55FA08EDA4F9C8" box="[151,415,1531,1557]" italics="true" pageId="22" pageNumber="264">Lavandula angustifolia</emphasis>
Miller (Lamiaceae)
</taxonomicName>
in Montferrier-sur-Lez, Cirad de Baillarguet,
<collectingCountry id="F31FFBF6FFCDFF93FB5AFA0FE8DFF9CB" box="[1176,1252,1532,1558]" name="France" pageId="22" pageNumber="264">France</collectingCountry>
(
<geoCoordinate id="EE3CDDA1FFCDFF93FB34FA09E9A3F9C8" box="[1270,1432,1530,1557]" direction="north" orientation="latitude" pageId="22" pageNumber="264" precision="1" value="43.682697">43°40’57.7”N</geoCoordinate>
,
<geoCoordinate id="EE3CDDA1FFCDFF93FF55F9EDED1CF9E4" box="[151,295,1566,1593]" direction="east" orientation="longitude" pageId="22" pageNumber="264" precision="1" value="3.8750558">3°52’30.2”E</geoCoordinate>
), on
<date id="FFB69DA6FFCDFF93FE9DF9ECEDD9F9E4" box="[351,482,1567,1593]" pageId="22" pageNumber="264" value="2017-09-18">18/09/2017</date>
.
</paragraph>
<paragraph id="8BB7BB66FFCDFF93FF05F9B0E9B4F980" blockId="22.[151,1437,1458,2026]" box="[199,1423,1602,1630]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F9B0ED81F980" bold="true" box="[199,442,1602,1629]" pageId="22" pageNumber="264">Known distribution.</emphasis>
<collectingCountry id="F31FFBF6FFCDFF93FE7DF9B0EE2FF980" box="[447,532,1603,1629]" name="Austria" pageId="22" pageNumber="264">Austria</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FDE2F9B0EE4FF980" box="[544,628,1603,1629]" name="Cyprus" pageId="22" pageNumber="264">Cyprus</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FDBDF9B7EEFEF983" box="[639,709,1604,1630]" name="Egypt" pageId="22" pageNumber="264">Egypt</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FD12F9B7EF24F983" box="[720,799,1604,1630]" name="France" pageId="22" pageNumber="264">France</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FCE8F9B0EF47F980" box="[810,892,1603,1629]" name="Greece" pageId="22" pageNumber="264">Greece</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FC45F9B7EFD1F983" box="[903,1002,1604,1630]" name="Hungary" pageId="22" pageNumber="264">Hungary</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FC37F9B0E80DF980" box="[1013,1078,1603,1629]" name="Israel" pageId="22" pageNumber="264">Israel</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FB83F9B0E84EF980" box="[1089,1141,1603,1629]" name="Italy" pageId="22" pageNumber="264">Italy</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FB43F9B7E8D2F983" box="[1153,1257,1604,1630]" name="Morocco" pageId="22" pageNumber="264">Morocco</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FB31F9B0E97FF980" box="[1267,1348,1603,1629]" name="Turkey" pageId="22" pageNumber="264">Turkey</collectingCountry>
,
<collectingCountry id="F31FFBF6FFCDFF93FA8DF9B0E9B0F980" box="[1359,1419,1603,1629]" name="United States of America" pageId="22" pageNumber="264">USA</collectingCountry>
.
</paragraph>
<paragraph id="8BB7BB66FFCDFF93FF05F994EDC7F95F" blockId="22.[151,1437,1458,2026]" box="[199,508,1639,1666]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F994ED4DF95C" bold="true" box="[199,374,1639,1665]" pageId="22" pageNumber="264">Measurements</emphasis>
(
<specimenCount id="9D0E70EFFFCDFF93FE44F994EDCEF95F" box="[390,501,1639,1666]" pageId="22" pageNumber="264" type="female">4 females</specimenCount>
)
</paragraph>
<paragraph id="8BB7BB66FFCDFF93FF05F97FECE3F8A3" blockId="22.[151,1437,1458,2026]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F97FED17F978" bold="true" box="[199,300,1675,1702]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F97FED1FF97B" bold="true" box="[199,292,1676,1702]" italics="true" pageId="22" pageNumber="264">Dorsum</emphasis>
.
</emphasis>
Dorsal shield 318 (311–326) long and 168 (156–174) wide at level of setae
<emphasis id="B97C6774FFCDFF93FBBBF978E8A9F978" box="[1145,1170,1675,1701]" italics="true" pageId="22" pageNumber="264">s4</emphasis>
, 178 (173–183) at level of
<emphasis id="B97C6774FFCDFF93FF75F95CECE8F914" box="[183,211,1711,1737]" italics="true" pageId="22" pageNumber="264">S2</emphasis>
, 171 (168–174) at level of
<emphasis id="B97C6774FFCDFF93FDCEF95CEE13F914" box="[524,552,1711,1737]" italics="true" pageId="22" pageNumber="264">S4</emphasis>
; slightly reticulated, with four pairs of solenostomes (
<emphasis id="B97C6774FFCDFF93FB50F95CE887F914" box="[1170,1212,1711,1737]" italics="true" pageId="22" pageNumber="264">gd2</emphasis>
,
<emphasis id="B97C6774FFCDFF93FB09F95CE8CEF914" box="[1227,1269,1711,1737]" italics="true" pageId="22" pageNumber="264">gd6</emphasis>
,
<emphasis id="B97C6774FFCDFF93FAC6F95CE915F914" box="[1284,1326,1711,1737]" italics="true" pageId="22" pageNumber="264">gd8</emphasis>
,
<emphasis id="B97C6774FFCDFF93FAFCF95CE95CF914" box="[1342,1383,1711,1737]" italics="true" pageId="22" pageNumber="264">gd9</emphasis>
), 17 pairs of dorsal setae and two pairs of sub-lateral setae:
<emphasis id="B97C6774FFCDFF93FCC0F920EF2CF930" box="[770,791,1747,1773]" italics="true" pageId="22" pageNumber="264">j1</emphasis>
23 (19–27);
<emphasis id="B97C6774FFCDFF93FC68F920EFFBF930" box="[938,960,1747,1773]" italics="true" pageId="22" pageNumber="264">j3</emphasis>
18 (13–20);
<emphasis id="B97C6774FFCDFF93FB91F920E852F930" box="[1107,1129,1747,1773]" italics="true" pageId="22" pageNumber="264">j4</emphasis>
13 (10–15);
<emphasis id="B97C6774FFCDFF93FB39F927E92AF933" box="[1275,1297,1748,1774]" italics="true" pageId="22" pageNumber="264">j5</emphasis>
13 (12–13);
<emphasis id="B97C6774FFCDFF93FF55F904EC96F8CC" box="[151,173,1783,1809]" italics="true" pageId="22" pageNumber="264">j6</emphasis>
15;
<emphasis id="B97C6774FFCDFF93FF23F904ECC0F8CC" box="[225,251,1783,1809]" italics="true" pageId="22" pageNumber="264">J2</emphasis>
19 (18–20);
<emphasis id="B97C6774FFCDFF93FE52F90BED90F8CF" box="[400,427,1784,1810]" italics="true" pageId="22" pageNumber="264">J5</emphasis>
6 (6–7);
<emphasis id="B97C6774FFCDFF93FDD7F904EE15F8CC" box="[533,558,1783,1809]" italics="true" pageId="22" pageNumber="264">z2</emphasis>
15 (14–15);
<emphasis id="B97C6774FFCDFF93FD01F904EEE7F8CC" box="[707,732,1783,1809]" italics="true" pageId="22" pageNumber="264">z3</emphasis>
17 (15–17);
<emphasis id="B97C6774FFCDFF93FCB3F904EFB1F8CC" box="[881,906,1783,1809]" italics="true" pageId="22" pageNumber="264">z4</emphasis>
17 (16–18);
<emphasis id="B97C6774FFCDFF93FBDDF90BE803F8CF" box="[1055,1080,1784,1810]" italics="true" pageId="22" pageNumber="264">z5</emphasis>
15 (14–15);
<emphasis id="B97C6774FFCDFF93FB0EF904E8D1F8CC" box="[1228,1258,1783,1809]" italics="true" pageId="22" pageNumber="264">Z4</emphasis>
36 (33–39);
<emphasis id="B97C6774FFCDFF93FABDF90BE9A7F8CF" box="[1407,1436,1784,1810]" italics="true" pageId="22" pageNumber="264">Z5</emphasis>
56 (52–59);
<geoCoordinate id="EE3CDDA1FFCDFF93FEE1F8E8ED5BF8E8" box="[291,352,1819,1845]" direction="south" orientation="latitude" pageId="22" pageNumber="264" precision="925" value="-4.383333">
<emphasis id="B97C6774FFCDFF93FEE1F8E8ED07F8E8" box="[291,316,1819,1845]" italics="true" pageId="22" pageNumber="264">s4</emphasis>
23
</geoCoordinate>
(20–24);
<geoCoordinate id="EE3CDDA1FFCDFF93FE12F8E8EE37F8E8" box="[464,524,1818,1845]" direction="south" orientation="latitude" pageId="22" pageNumber="264" precision="925" value="-6.4166665">
<emphasis id="B97C6774FFCDFF93FE12F8E8EDD2F8E8" box="[464,489,1819,1845]" italics="true" pageId="22" pageNumber="264">s6</emphasis>
25
</geoCoordinate>
(25–26);
<geoCoordinate id="EE3CDDA1FFCDFF93FDBEF8E8EE87F8E8" box="[636,700,1819,1845]" direction="south" orientation="latitude" pageId="22" pageNumber="264" precision="925" value="-2.4833333">
<emphasis id="B97C6774FFCDFF93FDBEF8E8EEA3F8E8" box="[636,664,1819,1845]" italics="true" pageId="22" pageNumber="264">S2</emphasis>
29
</geoCoordinate>
(28–30);
<geoCoordinate id="EE3CDDA1FFCDFF93FCEEF8E8EF57F8E8" box="[812,876,1819,1845]" direction="south" orientation="latitude" pageId="22" pageNumber="264" precision="925" value="-4.5">
<emphasis id="B97C6774FFCDFF93FCEEF8E8EF73F8E8" box="[812,840,1819,1845]" italics="true" pageId="22" pageNumber="264">S4</emphasis>
30
</geoCoordinate>
(28–32);
<emphasis id="B97C6774FFCDFF93FC1EF8E8EFCEF8E8" box="[988,1013,1819,1845]" italics="true" pageId="22" pageNumber="264">r3</emphasis>
23 (23–24);
<emphasis id="B97C6774FFCDFF93FB4AF8E8E893F8E8" box="[1160,1192,1819,1845]" italics="true" pageId="22" pageNumber="264">R1</emphasis>
19 (16–20). All setae smooth, except
<emphasis id="B97C6774FFCDFF93FE88F8CCED53F884" box="[330,360,1855,1881]" italics="true" pageId="22" pageNumber="264">Z4</emphasis>
and
<emphasis id="B97C6774FFCDFF93FE63F8B3ED84F887" box="[417,447,1856,1882]" italics="true" pageId="22" pageNumber="264">Z5</emphasis>
, slightly serrated. Peritreme extending forwards at level between the bases of the setae
<emphasis id="B97C6774FFCDFF93FF55F890ECEAF8A0" box="[151,209,1891,1917]" italics="true" pageId="22" pageNumber="264">j1–j3</emphasis>
.
</paragraph>
<paragraph id="8BB7BB66FFCDFF92FF05F875ECEBFEBA" blockId="22.[151,1437,1458,2026]" lastBlockId="23.[151,1437,152,611]" lastPageId="23" lastPageNumber="265" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F875EDAAF87C" bold="true" box="[199,401,1926,1953]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FF05F875EDB1F87C" bold="true" box="[199,394,1926,1953]" italics="true" pageId="22" pageNumber="264">Ventral idiosoma</emphasis>
.
</emphasis>
Sternal shield slightly reticulated with two pairs of setae (
<emphasis id="B97C6774FFCDFF93FBD9F874E807F87C" box="[1051,1084,1927,1953]" italics="true" pageId="22" pageNumber="264">st1</emphasis>
,
<emphasis id="B97C6774FFCDFF93FB88F874E851F87C" box="[1098,1130,1927,1953]" italics="true" pageId="22" pageNumber="264">st2</emphasis>
) and poroids (
<date id="FFB69DA6FFCDFF93FACCF874E90BF87C" box="[1294,1328,1927,1953]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FACCF874E90BF87C" box="[1294,1328,1927,1953]" italics="true" pageId="22" pageNumber="264">iv1</emphasis>
</date>
,
<date id="FFB69DA6FFCDFF93FAFCF874E95BF87C" box="[1342,1376,1927,1953]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FAFCF874E95BF87C" box="[1342,1376,1927,1953]" italics="true" pageId="22" pageNumber="264">iv2</emphasis>
</date>
). Setae
<emphasis id="B97C6774FFCDFF93FF7DF858ECDBF818" box="[191,224,1963,1989]" italics="true" pageId="22" pageNumber="264">st3</emphasis>
and
<emphasis id="B97C6774FFCDFF93FED7F858ED0DF818" box="[277,310,1963,1989]" italics="true" pageId="22" pageNumber="264">st4</emphasis>
tylochorous (on platelets). Poroids
<date id="FFB69DA6FFCDFF93FD07F858EEDCF818" box="[709,743,1963,1989]" pageId="22" pageNumber="264">
<emphasis id="B97C6774FFCDFF93FD07F858EEDCF818" box="[709,743,1963,1989]" italics="true" pageId="22" pageNumber="264">iv3</emphasis>
</date>
on the interscutal membrane anteriorly to setae
<emphasis id="B97C6774FFCDFF93FAC3F858E91AF818" box="[1281,1313,1963,1989]" italics="true" pageId="22" pageNumber="264">st4</emphasis>
. Distances between setae
<emphasis id="B97C6774FFCDFF93FEFDF83CEDB5F834" box="[319,398,1999,2025]" italics="true" pageId="22" pageNumber="264">st1–st1</emphasis>
50 (48–51),
<emphasis id="B97C6774FFCDFF93FDE7F83CEE4FF834" box="[549,628,1999,2025]" italics="true" pageId="22" pageNumber="264">st2–st2</emphasis>
53 (51–55),
<emphasis id="B97C6774FFCDFF93FCCEF83CEF60F834" box="[780,859,1999,2025]" italics="true" pageId="22" pageNumber="264">st1–st2</emphasis>
32 (31–32). Genital shield almost smooth, genital sigilla visible (1–3 pairs). The posterior margin of the genital shield is straight. The 4
<superScript id="7C7D162EFFCCFF92FB8DFF6BE867FF7B" attach="left" box="[1103,1116,152,166]" fontSize="6" pageId="23" pageNumber="265">th</superScript>
and 5
<superScript id="7C7D162EFFCCFF92FB62FF6BE896FF7B" attach="left" box="[1184,1197,152,166]" fontSize="6" pageId="23" pageNumber="265">th</superScript>
pair of genital sigilla clearly visible anteriorly to the setae
<emphasis id="B97C6774FFCCFF92FD85FF4EEE4DFF0A" box="[583,630,189,215]" italics="true" pageId="23" pageNumber="265">ZV1</emphasis>
. The 6
<superScript id="7C7D162EFFCCFF92FD08FF4FEEECFF17" attach="left" box="[714,727,188,202]" fontSize="6" pageId="23" pageNumber="265">th</superScript>
genital sigilla postero-paraxially to the setae
<emphasis id="B97C6774FFCCFF92FB28FF4EE923FF0A" box="[1258,1304,189,215]" italics="true" pageId="23" pageNumber="265">ZV1</emphasis>
. Ventrianal shield pentagonal, slightly striated, with four pairs of pre-anal setae,
<emphasis id="B97C6774FFCCFF92FC5CFF12EFF2FF26" box="[926,969,225,251]" italics="true" pageId="23" pageNumber="265">JV1</emphasis>
,
<emphasis id="B97C6774FFCCFF92FC1AFF12E838FF26" box="[984,1027,225,251]" italics="true" pageId="23" pageNumber="265">JV2</emphasis>
,
<emphasis id="B97C6774FFCCFF92FBD0FF12E806FF26" box="[1042,1085,225,251]" italics="true" pageId="23" pageNumber="265">JV3</emphasis>
and
<emphasis id="B97C6774FFCCFF92FBB7FF12E89FFF26" box="[1141,1188,225,251]" italics="true" pageId="23" pageNumber="265">ZV2</emphasis>
. Length of the ventrianal shield 104 (101–106), width at level of setae
<emphasis id="B97C6774FFCCFF92FD00FEF6EECAFEC2" box="[706,753,261,287]" italics="true" pageId="23" pageNumber="265">ZV2</emphasis>
89 (83–94), at level of setae
<emphasis id="B97C6774FFCCFF92FBF6FEF6E85BFEC2" box="[1076,1120,261,287]" italics="true" pageId="23" pageNumber="265">JV2</emphasis>
77 (69–87), at level of setae
<emphasis id="B97C6774FFCCFF92FF55FEDAECF8FE9E" box="[151,195,297,323]" italics="true" pageId="23" pageNumber="265">JV3</emphasis>
77 (68–84) and at level of para-anal setae 68 (67–69). Solenostome
<emphasis id="B97C6774FFCCFF92FC06FEDAEFD7FE9E" box="[964,1004,297,323]" italics="true" pageId="23" pageNumber="265">gv3</emphasis>
absent. Setae
<emphasis id="B97C6774FFCCFF92FB4FFEDAE882FE9E" box="[1165,1209,297,323]" italics="true" pageId="23" pageNumber="265">JV5</emphasis>
smooth, 49 (48–50) long.
</paragraph>
<paragraph id="8BB7BB66FFCCFF92FF05FE83ED68FE72" blockId="23.[151,1437,152,611]" pageId="23" pageNumber="265">
<emphasis id="B97C6774FFCCFF92FF05FE83ED0CFE56" bold="true" box="[199,311,368,395]" italics="true" pageId="23" pageNumber="265">Chelicera</emphasis>
. Fixed digit 27 (26–27) long, with three subapical teeth and
<emphasis id="B97C6774FFCCFF92FC1BFE82E853FE56" box="[985,1128,369,395]" italics="true" pageId="23" pageNumber="265">pilus dentilis</emphasis>
. Movable digit, unidentate, 26 (25–27) long.
</paragraph>
<paragraph id="8BB7BB66FFCCFF92FF05FE4AEDB1FE2A" blockId="23.[151,1437,152,611]" pageId="23" pageNumber="265">
<emphasis id="B97C6774FFCCFF92FF05FE4AECC6FE0E" bold="true" box="[199,253,441,467]" italics="true" pageId="23" pageNumber="265">Legs</emphasis>
. Genu II with seven setae, 2 2/0 2/0 1. Legs IV with a slightly bulbous tip macrosetae on the basitarsus,
<emphasis id="B97C6774FFCCFF92FF55FE2EECFCFE2A" box="[151,199,477,503]" italics="true" pageId="23" pageNumber="265">StIV</emphasis>
36 (35–37) long.
</paragraph>
<paragraph id="8BB7BB66FFCCFF92FF05FDF2EE6CFDBE" blockId="23.[151,1437,152,611]" pageId="23" pageNumber="265">
<emphasis id="B97C6774FFCCFF92FF05FDF2EDDEFDC6" bold="true" box="[199,485,513,539]" pageId="23" pageNumber="265">
<emphasis id="B97C6774FFCCFF92FF05FDF2EDE5FDC6" bold="true" box="[199,478,513,539]" italics="true" pageId="23" pageNumber="265">Insemination apparatus</emphasis>
.
</emphasis>
Major duct short 4 (3–4) cylindrical. Atrium slightly bulbous, embolus and minor duct visible. A short neck between the atrium and the calyx. The calyx is almost cylindrical, U-shaped, 12 (10–13) long and 5 (4–5) wide, with thick walls.
</paragraph>
</subSubSection>
</treatment>
</document>
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