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<document id="4F56353FEE76056D3D87F944D057EDD8" ID-DOI="10.5852/ejt.2024.962.2683" ID-ISSN="2118-9773" ID-Zenodo-Dep="13947260" ID-ZooBank="40171C5B-90EB-4641-91FF-EA8CA2C60D23" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729183243575" checkinUser="plazi" docAuthor="Ramos-Sánchez, Mariela" docDate="2024" docId="03F2DF38026EFF9FFE61496BFC747D96" docLanguage="en" docName="EJT.2024.962.1-67.pdf" docOrigin="European Journal of Taxonomy 962 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2683/12419" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Theama occidua Sopott-Ehlers &amp; Schmidt 1975" docType="treatment" docVersion="1" lastPageNumber="37" masterDocId="FFCBA740024AFFBBFF994B10FFC27830" masterDocTitle="The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="37" updateTime="1729617757279" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8D27009AC1D8FE0E20652E618892E67D">The polyclads (Platyhelminthes: Polycladida) from the Tropical Eastern Pacific: Commented checklist and description of a new species</mods:title>
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<mods:name id="1B8DBCCB66153C799D83B41E301C6B1D" type="personal">
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<mods:roleTerm id="CD24C659AE12A7A59475F913AAA2EE37">Author</mods:roleTerm>
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<mods:namePart id="9062D37A4BF7A6259C1FD5FCE418592D">Ramos-Sánchez, Mariela</mods:namePart>
<mods:nameIdentifier id="6D5717C965483A4325D8A042EC4BFAC7" type="LSID">E9D05B3F-CF64-48E3-B5ED-8D24E47F9D2A</mods:nameIdentifier>
<mods:affiliation id="3C3931AB41F0BE006E3751800142495A">Doctorado en Ciencias Biológico Agropecuarias, Área Ciencias Biológico Agropecuaria y Pesquera, Ciudad de la cultura “ Amado Nervo ”, Tepic, Nayarit, 63000, Mexico. &amp; Laboratorio de Sistemática de Invertebrados Marinos (LABSIM), Universidad del Mar, campus Puerto Ángel, Ciudad Universitaria, Puerto Ángel, Oaxaca, 70902, Mexico.</mods:affiliation>
<mods:nameIdentifier id="58C89756587043F57B51CA24F465227E" type="email">rmariela220@gmail.com</mods:nameIdentifier>
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<mods:title id="B7BAB74FF6CE23EFB20A7F73E873DB3E">European Journal of Taxonomy</mods:title>
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<mods:number id="E8B690EC87B0501A6FA6081E773943E7">962</mods:number>
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<mods:classification id="CDD5A658EEEE68E5D5A3B27794A9DF4C">journal article</mods:classification>
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<treatment id="03F2DF38026EFF9FFE61496BFC747D96" LSID="urn:lsid:plazi:treatment:03F2DF38026EFF9FFE61496BFC747D96" httpUri="http://treatment.plazi.org/id/03F2DF38026EFF9FFE61496BFC747D96" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection id="C3413DA5026EFF9FFE61496BFBF97AA7" box="[504,1083,635,663]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BE46E2E026EFF9FFE61496BFBF97AA7" blockId="36.[504,1083,635,663]" box="[504,1083,635,663]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFE61496BFBF97AA7" box="[504,1083,635,663]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
<taxonomicName id="4C5B15AD026EFF9FFE61496BFBF97AA7" ID-CoL="565QZ" authority="Sopott-Ehlers &amp; Schmidt, 1975" authorityName="Sopott-Ehlers &amp; Schmidt" authorityYear="1975" box="[504,1083,635,663]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFE61496BFD007AA6" bold="true" box="[504,706,635,662]" italics="true" pageId="36" pageNumber="37">Theama occidua</emphasis>
<bibRefCitation id="EFCA13DF026EFF9FFD50496CFBF97AA7" author="Sopott-Ehlers B. &amp; Schmidt P." box="[713,1083,636,663]" pageId="36" pageNumber="37" pagination="193 - 222" refId="ref32626" refString="Sopott-Ehlers B. &amp; Schmidt P. 1975. Interstitielle Fauna von Galapagos XIV. Polycladida (Turbellaria). Mikrofauna des Meeresbodens 54: 193 - 222." type="journal article" year="1975">Sopott-Ehlers &amp; Schmidt, 1975</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3413DA5026EFF9FFF2449ADFEF97BB2" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BE46E2E026EFF9FFF2449ADFDBD7ACA" blockId="36.[189,1399,701,762]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB4B4900026EFF9FFF2449ADFDBD7ACA" pageId="36" pageNumber="37">
<treatmentCitation id="0AFA483F026EFF9FFF2449ADFC837AE7" author="Sopott-Ehlers B. &amp; Schmidt P." box="[189,833,701,728]" page="11" pageId="36" pageNumber="37" year="1975">
<taxonomicName id="4C5B15AD026EFF9FFF2449ADFC837AE7" ID-CoL="565QZ" authority="Sopott-Ehlers &amp; Schmidt, 1975: 8 - 11" authorityName="Sopott-Ehlers &amp; Schmidt" authorityPageNumber="8 - 11" authorityYear="1975" box="[189,833,701,728]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFF2449ADFEBD7AE7" box="[189,383,701,727]" italics="true" pageId="36" pageNumber="37">Theama occidua</emphasis>
<bibRefCitation id="EFCA13DF026EFF9FFE1F49ADFC837AE7" author="Sopott-Ehlers B. &amp; Schmidt P." box="[390,833,701,728]" pageId="36" pageNumber="37" pagination="193 - 222" refId="ref32626" refString="Sopott-Ehlers B. &amp; Schmidt P. 1975. Interstitielle Fauna von Galapagos XIV. Polycladida (Turbellaria). Mikrofauna des Meeresbodens 54: 193 - 222." type="journal article" year="1975">Sopott-Ehlers &amp; Schmidt, 1975: 8–11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 3, pl. 4 figs a–c, pl. 5 figs a–c (
<typeStatus id="54E0D08C026EFF9FFB4249ADFACD7AE7" box="[1243,1295,701,727]" pageId="36" pageNumber="37">type</typeStatus>
locality:
<collectingRegion id="499FA0CC026EFF9FFF7349F0FEA47ACA" box="[234,358,736,762]" country="Ecuador" name="Galapagos" pageId="36" pageNumber="37">Galapagos</collectingRegion>
Archipelago,
<collectingCountry id="F34C2EBE026EFF9FFD9249F0FDB07ACA" box="[523,626,736,762]" name="Ecuador" pageId="36" pageNumber="37">Ecuador</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF244832FB8A7B0C" blockId="36.[189,1399,802,898]" box="[189,1096,802,828]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB4B4900026EFF9FFF244832FB8A7B0C" box="[189,1096,802,828]" pageId="36" pageNumber="37">
<taxonomicName id="4C5B15AD026EFF9FFF244832FE5A7B0C" ID-CoL="6HJF5" authorityName="Sopott-Ehlers &amp; Schmidt" authorityYear="1975" box="[189,408,802,828]" family="Theamatidae" genus="Eutheama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFF244832FE5A7B0C" box="[189,408,802,828]" italics="true" pageId="36" pageNumber="37">Eutheama occidua</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AFA483F026EFF9FFE2C4832FDB67B0C" author="Faubel A." box="[437,628,802,828]" page="97" pageId="36" pageNumber="37" year="1983">
<bibRefCitation id="EFCA13DF026EFF9FFE2C4832FDB67B0C" author="Faubel A." box="[437,628,802,828]" pageId="36" pageNumber="37" pagination="17 - 121" refId="ref28888" refString="Faubel A. 1983. The Polycladida, Turbellaria proposal and establishment of a new system part I. The Acotylea. Mitteilungen des Hamburgischen Zoologischen Museum und Institut 80: 17 - 121." type="journal article" year="1983">Faubel 1983: 97</bibRefCitation>
</treatmentCitation>
(new combination, identification keys).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF244855FEF97BB2" blockId="36.[189,1399,802,898]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB4B4900026EFF9FFF244855FEF97BB2" pageId="36" pageNumber="37">
<taxonomicName id="4C5B15AD026EFF9FFF244855FE417B6F" ID-CoL="565QZ" authorityName="Sopott-Ehlers &amp; Schmidt" authorityYear="1975" box="[189,387,837,863]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFF244855FE417B6F" box="[189,387,837,863]" italics="true" pageId="36" pageNumber="37">Theama occidua</emphasis>
</taxonomicName>
–
<treatmentCitation id="0AFA483F026EFF9FFE3E4855FD017B6F" author="Westheide W." box="[423,707,837,863]" page="43" pageId="36" pageNumber="37" year="1991">
<bibRefCitation id="EFCA13DF026EFF9FFE3E4855FD017B6F" author="Westheide W." box="[423,707,837,863]" pageId="36" pageNumber="37" pagination="37 - 73" refId="ref33180" refString="Westheide W. 1991. The meiofauna of the Galapagos. In: Matthew J. J. (ed.) Galapagos Marine Invertebrates. Taxonomy, Biogeography, and Evolution in Darwin's Islands: 37 - 73. Topics in Geobiology 8. Springer Science + Business Media New York. https: // doi. org / 10.1007 / 978 - 1 - 4899 - 0646 - 5 _ 3" type="journal article" year="1991">Westheide 1991: 43–44</bibRefCitation>
</treatmentCitation>
(mentioned only). —
<treatmentCitation id="0AFA483F026EFF9FFC4E4855FAB07B6F" author="Curini-Galletti M. &amp; Campus P. &amp; Delogu V." box="[983,1394,837,864]" page="82" pageId="36" pageNumber="37" year="2008">
<bibRefCitation id="EFCA13DF026EFF9FFC4E4855FAB07B6F" author="Curini-Galletti M. &amp; Campus P. &amp; Delogu V." box="[983,1394,837,864]" pageId="36" pageNumber="37" pagination="77 - 83" refId="ref28594" refString="Curini-Galletti M., Campus P. &amp; Delogu V. 2008. Thaema mediterranea sp. nov. (Platyhelminthes, Polycladida), the first interstitial polyclad from the Mediterranean. Italian Journal of Zoology 75 (1): 77 - 83. https: // doi. org / 10.1080 / 11250000701690525" type="journal article" year="2008">
Curini-Galletti
<emphasis id="B92FB23C026EFF9FFB164856FB047B6F" box="[1167,1222,837,863]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2008: 82–83
</bibRefCitation>
</treatmentCitation>
, fig. 1c.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<subSubSection id="C3413DA5026EFF9FFF2448B9FC037C25" pageId="36" pageNumber="37" type="distribution">
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<paragraph id="8BE46E2E026EFF9FFF2448B9FE947BF3" blockId="36.[189,342,937,963]" box="[189,342,937,963]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFF2448B9FE947BF3" bold="true" box="[189,342,937,963]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B92FB23C026EFF9FFF2448B9FE947BF3" bold="true" box="[189,342,937,963]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF2448C9FC037C25" blockId="36.[188,1398,984,1046]" pageId="36" pageNumber="37">
James Bay and Caleta Negra (
<bibRefCitation id="EFCA13DF026EFF9FFD8148C8FC897BC2" author="Curini-Galletti M. &amp; Campus P. &amp; Delogu V." box="[536,843,984,1010]" pageId="36" pageNumber="37" pagination="77 - 83" refId="ref28594" refString="Curini-Galletti M., Campus P. &amp; Delogu V. 2008. Thaema mediterranea sp. nov. (Platyhelminthes, Polycladida), the first interstitial polyclad from the Mediterranean. Italian Journal of Zoology 75 (1): 77 - 83. https: // doi. org / 10.1080 / 11250000701690525" type="journal article" year="2008">
Curini-Galletti
<emphasis id="B92FB23C026EFF9FFD5248C9FCC77BC2" box="[715,773,984,1010]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
2008
</bibRefCitation>
), Fernandina, Isabela, Genovesa, Bartholomew,
<collectingCountry id="F34C2EBE026EFF9FFF2548ECFE9C7C25" box="[188,350,1019,1046]" name="Saint Kitts and Nevis" pageId="36" pageNumber="37">San Cristobal</collectingCountry>
, Hood and Santa Cruz Islands,
<collectingRegion id="499FA0CC026EFF9FFD5748EBFC8F7C25" box="[718,845,1019,1045]" country="Ecuador" name="Galapagos" pageId="36" pageNumber="37">Galapagos</collectingRegion>
,
<collectingCountry id="F34C2EBE026EFF9FFCC048EBFC7F7C25" box="[857,957,1019,1045]" name="Ecuador" pageId="36" pageNumber="37">Ecuador</collectingCountry>
.
</paragraph>
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</subSubSection>
<subSubSection id="C3413DA5026EFF9FFF244F2DFEFF7CB5" box="[189,317,1085,1157]" pageId="36" pageNumber="37" type="biology_ecology">
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<paragraph id="8BE46E2E026EFF9FFF244F2DFEDF7C67" blockId="36.[189,285,1085,1111]" box="[189,285,1085,1111]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFF244F2DFEDF7C67" bold="true" box="[189,285,1085,1111]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B92FB23C026EFF9FFF244F2DFEDF7C67" bold="true" box="[189,285,1085,1111]" pageId="36" pageNumber="37">Habitat</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF244F7BFEFF7CB5" blockId="36.[189,317,1131,1157]" box="[189,317,1131,1157]" pageId="36" pageNumber="37">Interstitial.</paragraph>
</subSubSection>
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<subSubSection id="C3413DA5026EFF9FFF244FBDFC747D96" pageId="36" pageNumber="37" type="discussion">
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<paragraph id="8BE46E2E026EFF9FFF244FBDFEED7CF7" blockId="36.[189,303,1197,1223]" box="[189,303,1197,1223]" pageId="36" pageNumber="37">
<heading id="D0ACD942026EFF9FFF244FBDFEED7CF7" bold="true" box="[189,303,1197,1223]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B92FB23C026EFF9FFF244FBDFEED7CF7" bold="true" box="[189,303,1197,1223]" pageId="36" pageNumber="37">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE46E2E026EFF9FFF244FCBFC747D96" blockId="36.[189,1399,1243,1446]" pageId="36" pageNumber="37">
<taxonomicName id="4C5B15AD026EFF9FFF244FCBFE407CC5" authorityName="Sopott-Ehlers &amp; Schmidt" authorityYear="1975" box="[189,386,1243,1269]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFF244FCBFE407CC5" box="[189,386,1243,1269]" italics="true" pageId="36" pageNumber="37">Theama occidua</emphasis>
</taxonomicName>
was originally described by
<bibRefCitation id="EFCA13DF026EFF9FFD7B4FCBFBA97CC6" author="Sopott-Ehlers B. &amp; Schmidt P." box="[738,1131,1243,1270]" pageId="36" pageNumber="37" pagination="193 - 222" refId="ref32626" refString="Sopott-Ehlers B. &amp; Schmidt P. 1975. Interstitielle Fauna von Galapagos XIV. Polycladida (Turbellaria). Mikrofauna des Meeresbodens 54: 193 - 222." type="journal article" year="1975">Sopott-Ehlers &amp; Schmidt (1975)</bibRefCitation>
. Subsequently,
<bibRefCitation id="EFCA13DF026EFF9FFABE4FCBFECE7D29" author="Faubel A." pageId="36" pageNumber="37" pagination="17 - 121" refId="ref28888" refString="Faubel A. 1983. The Polycladida, Turbellaria proposal and establishment of a new system part I. The Acotylea. Mitteilungen des Hamburgischen Zoologischen Museum und Institut 80: 17 - 121." type="journal article" year="1983">Faubel (1983)</bibRefCitation>
established the genus
<taxonomicName id="4C5B15AD026EFF9FFDB84FEFFD5A7D29" box="[545,664,1279,1305]" family="Theamatidae" genus="Eutheama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026EFF9FFDB84FEFFD5A7D29" box="[545,664,1279,1305]" italics="true" pageId="36" pageNumber="37">Eutheama</emphasis>
</taxonomicName>
and determined
<taxonomicName id="4C5B15AD026EFF9FFCFE4FEFFC267D29" authorityName="Sopott-Ehlers &amp; Schmidt" authorityYear="1975" box="[871,996,1279,1305]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="species" species="occidua">
<emphasis id="B92FB23C026EFF9FFCFE4FEFFC267D29" box="[871,996,1279,1305]" italics="true" pageId="36" pageNumber="37">T. occidua</emphasis>
</taxonomicName>
as the
<typeStatus id="54E0D08C026EFF9FFBD84FEFFBB67D29" box="[1089,1140,1279,1305]" pageId="36" pageNumber="37">type</typeStatus>
species, based on an unarmed penis papilla; however,
<bibRefCitation id="EFCA13DF026EFF9FFDC54E32FC767D0C" author="Curini-Galletti M. &amp; Campus P. &amp; Delogu V." box="[604,948,1314,1340]" pageId="36" pageNumber="37" pagination="77 - 83" refId="ref28594" refString="Curini-Galletti M., Campus P. &amp; Delogu V. 2008. Thaema mediterranea sp. nov. (Platyhelminthes, Polycladida), the first interstitial polyclad from the Mediterranean. Italian Journal of Zoology 75 (1): 77 - 83. https: // doi. org / 10.1080 / 11250000701690525" type="journal article" year="2008">
Curini-Galletti
<emphasis id="B92FB23C026EFF9FFC8E4E33FC9A7D0C" box="[791,856,1314,1340]" italics="true" pageId="36" pageNumber="37">et al.</emphasis>
(2008)
</bibRefCitation>
conducted a detailed review of the reproductive system of the species and observed the presence of a small sclerotized area in the internal ejaculatory duct. Based on this character, they transferred the species back to the genus
<taxonomicName id="4C5B15AD026EFF9FFB7D4E78FA817DB2" authorityName="Marcus" authorityYear="1949" box="[1252,1347,1384,1410]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026EFF9FFB7D4E78FA817DB2" box="[1252,1347,1384,1410]" italics="true" pageId="36" pageNumber="37">Theama</emphasis>
</taxonomicName>
and established
<taxonomicName id="4C5B15AD026EFF9FFEDE4E9BFE7C7D95" box="[327,446,1419,1445]" family="Theamatidae" genus="Eutheama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026EFF9FFEDE4E9BFE7C7D95" box="[327,446,1419,1445]" italics="true" pageId="36" pageNumber="37">Eutheama</emphasis>
</taxonomicName>
as a junior synonym of the genus
<taxonomicName id="4C5B15AD026EFF9FFCC94E9BFC6D7D95" authorityName="Marcus" authorityYear="1949" box="[848,943,1419,1445]" family="Theamatidae" genus="Theama" kingdom="Animalia" order="Polycladida" pageId="36" pageNumber="37" phylum="Platyhelminthes" rank="genus">
<emphasis id="B92FB23C026EFF9FFCC94E9BFC6D7D95" box="[848,943,1419,1445]" italics="true" pageId="36" pageNumber="37">Theama</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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