494 lines
83 KiB
XML
494 lines
83 KiB
XML
|
<document id="D0B71DD754CE693C8104BA6932D30DCE" ID-DOI="10.11646/zootaxa.3722.2.8" ID-GBIF-Dataset="6af4eefc-93cf-49e0-b570-8f50f59e966d" ID-ISSN="1175-5326" ID-Zenodo-Dep="223843" ID-ZooBank="9116C1A3-EA5B-4798-B852-12B44B976A99" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460691703237" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2013" docId="03C887A6D870FFFC0988FF3DFBB9F87E" docLanguage="en" docName="zt03722p282.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alpheus pseudopugnax Banner 1953" docType="treatment" docVersion="7" lastPageNumber="281" masterDocId="FFF1FFDED878FFF2091FFFAAFFF6FFD1" masterDocTitle="Notes on two rare and little-known Indo-Pacific snapping shrimps, Alpheus percyi Coutière, 1908 and A. pseudopugnax (Banner, 1953) (Decapoda, Alpheidae)" masterLastPageNumber="282" masterPageNumber="267" pageNumber="275" updateTime="1698351849637" updateUser="plazi">
|
|||
|
|
<mods:mods id="4A4080E937F00E0D1AC8273E83A5CEAE" xmlns:mods="http://www.loc.gov/mods/v3">
|
|||
|
|
<mods:titleInfo id="2C1321B7517103074DB14BB9A1EE9364">
|
|||
|
|
<mods:title id="D4C81A4F85FD7B1B83A61E92C31E132A">Notes on two rare and little-known Indo-Pacific snapping shrimps, Alpheus percyi Coutière, 1908 and A. pseudopugnax (Banner, 1953) (Decapoda, Alpheidae)</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:name id="1914B817D6AA63C7D02982E799CFF863" type="personal">
|
|||
|
|
<mods:role id="862A2E3F7B963E13483C0C9A4DE9608D">
|
|||
|
|
<mods:roleTerm id="F5113FEA6FBE9E4A5B35F920F6C724C9">Author</mods:roleTerm>
|
|||
|
|
</mods:role>
|
|||
|
|
<mods:namePart id="B1A6CE0BB6F6332591DFC6E153E6C6DA">Anker, Arthur</mods:namePart>
|
|||
|
|
</mods:name>
|
|||
|
|
<mods:typeOfResource id="476DF3392812BD949C9A3782D5C6AAE4">text</mods:typeOfResource>
|
|||
|
|
<mods:relatedItem id="7D5E9E03EEA7B2674467F3CDC6025A48" type="host">
|
|||
|
|
<mods:titleInfo id="2953687A48E7576EA6017A0582D7E85C">
|
|||
|
|
<mods:title id="4ACCC1FF3F8E0DE5197579DA4F9235ED">Zootaxa</mods:title>
|
|||
|
|
</mods:titleInfo>
|
|||
|
|
<mods:part id="4C205E8F36E424394BDC434E30A4BCBE">
|
|||
|
|
<mods:date id="60634510413AF9795C2B783EA74CC071">2013</mods:date>
|
|||
|
|
<mods:detail id="B383C4680C7975093F37C260E61A3E93" type="volume">
|
|||
|
|
<mods:number id="896F0D521D97D5644A5FE9E944C9BBBD">3722</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:detail id="78522D48CE16A10087874B304F1617F0" type="issue">
|
|||
|
|
<mods:number id="A11FACE728E414AC604053F4C7A8E8B4">2</mods:number>
|
|||
|
|
</mods:detail>
|
|||
|
|
<mods:extent id="291122B1552EC2EF416363285D2D7393" unit="page">
|
|||
|
|
<mods:start id="3DCAA111B70C38431367206A9D3647F2">267</mods:start>
|
|||
|
|
<mods:end id="9D3304F24F17603BC737227DE584A37D">282</mods:end>
|
|||
|
|
</mods:extent>
|
|||
|
|
</mods:part>
|
|||
|
|
</mods:relatedItem>
|
|||
|
|
<mods:classification id="F162D0712B4A17E8C2678679B1BA4086">journal article</mods:classification>
|
|||
|
|
<mods:identifier id="50046B57ECBED206A8EAEB79A8C1EA1A" type="DOI">10.11646/zootaxa.3722.2.8</mods:identifier>
|
|||
|
|
<mods:identifier id="D42459707F2771D4DDCC210A3A4435B9" type="GBIF-Dataset">6af4eefc-93cf-49e0-b570-8f50f59e966d</mods:identifier>
|
|||
|
|
<mods:identifier id="985034FE202589981775B89E72090EDF" type="ISSN">1175-5326</mods:identifier>
|
|||
|
|
<mods:identifier id="4E6E01C0BDC3FA2A8F8C94B9E42054D4" type="Zenodo-Dep">223843</mods:identifier>
|
|||
|
|
<mods:identifier id="51BDBB5884C18915A97382B33A1BA778" type="ZooBank">9116C1A3-EA5B-4798-B852-12B44B976A99</mods:identifier>
|
|||
|
|
</mods:mods>
|
|||
|
|
<treatment id="03C887A6D870FFFC0988FF3DFBB9F87E" ID-DOI="http://doi.org/10.5281/zenodo.6495798" ID-GBIF-Taxon="119580100" ID-Zenodo-Dep="6495798" LSID="urn:lsid:plazi:treatment:03C887A6D870FFFC0988FF3DFBB9F87E" httpUri="http://treatment.plazi.org/id/03C887A6D870FFFC0988FF3DFBB9F87E" lastPageId="14" lastPageNumber="281" pageId="8" pageNumber="275">
|
|||
|
|
<subSubSection id="C37B653BD870FFFA0988FF3DFD6DFEC9" pageId="8" pageNumber="275" type="nomenclature">
|
|||
|
|
<paragraph id="8BDE36B0D870FFFA0988FF3DFD99FF63" blockId="8.[151,623,151,211]" box="[151,623,151,178]" pageId="8" pageNumber="275">
|
|||
|
|
<heading id="D09681DCD870FFFA0988FF3DFD99FF63" bold="true" box="[151,623,151,178]" fontSize="11" level="1" pageId="8" pageNumber="275" reason="1">
|
|||
|
|
<taxonomicName id="4C614D33D870FFFA0988FF3DFD99FF63" ID-CoL="84W93" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[151,623,151,178]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FF3DFD99FF63" bold="true" box="[151,623,151,178]" pageId="8" pageNumber="275">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FF3DFE47FF60" bold="true" box="[151,433,151,177]" italics="true" pageId="8" pageNumber="275">Alpheus pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDE36B0D870FFFA0988FF10FE8AFF02" blockId="8.[151,623,151,211]" box="[151,380,186,211]" pageId="8" pageNumber="275">
|
|||
|
|
(
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0980FF10FEEFFF02" box="[159,281,186,211]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="9.[151,250,1868,1890]" captionStartId-1="10.[151,250,1433,1455]" captionStartId-2="11.[151,250,1861,1883]" captionStartId-3="12.[151,250,1165,1187]" captionStartId-4="13.[151,250,1828,1850]" captionTargetBox-0="[169,1427,310,1793]" captionTargetBox-1="[163,1429,199,1401]" captionTargetBox-2="[155,1429,449,1820]" captionTargetBox-3="[184,1414,185,1138]" captionTargetBox-4="[151,1436,193,1805]" captionTargetId-0="figure@9.[151,1436,301,1846]" captionTargetId-1="figure@10.[151,1436,193,1412]" captionTargetId-2="figure@11.[151,1436,445,1840]" captionTargetId-3="figure@12.[167,1417,181,1156]" captionTargetId-4="figure@13.[151,1436,193,1806]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." captionText-1="FIGURE 7. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—minor cheliped, lateral view; B—same, distal carpus and chela, mesial view; C—major cheliped, lateral view; D—same, basis, ischium, merus and carpus, mesial view; E—major chela, mesial view; F—major chela, lateral view; G—distal major chela, fingers opened, lateral view." captionText-2="FIGURE 8. Alpheus pseudopugnax (Banner, 1953), male from Reunion Island (La Réunion), UF 12873: A—frontal region, dorsal view; B—same, lateral view; C—third maxilliped, lateral view; D—second pereiopod, lateral view; E—third pereiopod, lateral view; F—same, distal carpus, propodus and dactylus, lateral view; G—second pleopod, appendices masculina and interna, mesial view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." captionText-3="FIGURE 9. Alpheus pseudopugnax (Banner, 1953), male from Reunion Island (La Réunion), UF 12873: A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—major cheliped, ischium, merus and carpus, mesial view; D – major chela, mesial view; E—major chela, lateral view; F—distal major chela, fingers opened, lateral view." captionText-4="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri-0="https://zenodo.org/record/223849/files/figure.png" httpUri-1="https://zenodo.org/record/223850/files/figure.png" httpUri-2="https://zenodo.org/record/223851/files/figure.png" httpUri-3="https://zenodo.org/record/223852/files/figure.png" httpUri-4="https://zenodo.org/record/223853/files/figure.png" pageId="8" pageNumber="275">Figs. 6–10</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="135A2A35D870FFFA083BFF11FECAFF02" box="[292,316,187,211]" captionStart="FIGURE 11" captionStartId="14.[151,250,997,1019]" captionTargetBox="[173,1419,205,957]" captionTargetId="figure@14.[150,1436,193,976]" captionTargetPageId="14" captionText="FIGURE 11. Variation in the shape of the postrostral region in Alpheus percyi Coutière, 1908 (A – D) and Alpheus pseudopugnax (Banner, 1953) (E – G): A—holotype female from Amirante, MNHN 3188; B—female from Tabuaeran, Line Islands, UF 10662; C—male from Oahu, Hawaiian Archipelago, UF 15237; D—non-type male of Alpheus percyi (?) from Amirante, MNHN 3486; E—holotype female from Maui, Hawaiian Archipelago, USNM 93513; F—ovigerous female from Reunion Island (La Réunion), UF 12880; G—female from Palmyra Atoll, Line Islands, UF 10622." httpUri="https://zenodo.org/record/223854/files/figure.png" pageId="8" pageNumber="275">11</figureCitation>
|
|||
|
|
E–G)
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDE36B0D870FFFA0988FEA9FD6DFEC9" blockId="8.[151,747,258,311]" box="[151,667,258,280]" pageId="8" pageNumber="275">
|
|||
|
|
<taxonomicName id="4C614D33D870FFFA0988FEA9FDB1FEC9" authority="Banner 1953: 122" authorityName="Banner" authorityPageNumber="122" authorityYear="1953" box="[151,583,258,280]" class="Malacostraca" family="Crangonidae" genus="Crangon" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FEA9FE7FFEC9" box="[151,393,258,280]" italics="true" pageId="8" pageNumber="275">Crangon pseudopugnax</emphasis>
|
|||
|
|
Banner 1953: 122
|
|||
|
|
</taxonomicName>
|
|||
|
|
, fig. 44.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B653BD870FFFA0988FE8BFD1DFEE6" box="[151,747,289,311]" pageId="8" pageNumber="275" type="description">
|
|||
|
|
<paragraph id="8BDE36B0D870FFFA0988FE8BFD1DFEE6" blockId="8.[151,747,258,311]" box="[151,747,289,311]" pageId="8" pageNumber="275">
|
|||
|
|
<taxonomicName id="4C614D33D870FFFA0988FE8BFE77FEE6" box="[151,385,289,311]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FE8BFE77FEE6" box="[151,385,289,311]" italics="true" pageId="8" pageNumber="275">Alpheus pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
—
|
|||
|
|
<collectingCountry id="F3767620D870FFFA08B9FE88FE32FEE6" box="[422,452,290,311]" name="South Georgia and the South Sandwich Islands" pageId="8" pageNumber="275">De</collectingCountry>
|
|||
|
|
Grave & Fransen 2011: 388.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B653BD870FFFA0988FECEFAD5FCFC" pageId="8" pageNumber="275" type="materials_examined">
|
|||
|
|
<paragraph id="8BDE36B0D870FFFA0988FECEFAD5FCFC" blockId="8.[151,1436,356,2037]" pageId="8" pageNumber="275">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FECEFE8DFEAC" bold="true" box="[151,379,356,381]" pageId="8" pageNumber="275">Material examined</emphasis>
|
|||
|
|
.
|
|||
|
|
<specimenCount id="9D67FD39D870FFFA0894FECEFE06FEAC" box="[395,496,356,381]" pageId="8" pageNumber="275" type="female">1 female</specimenCount>
|
|||
|
|
,
|
|||
|
|
<typeStatus id="54DA8812D870FFFA08E2FECEFDA9FEAC" box="[509,607,356,381]" pageId="8" pageNumber="275" type="holotype">holotype</typeStatus>
|
|||
|
|
(cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0B91FECEFD1CFEAD" box="[654,746,356,381]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="275" unit="mm" value="4.6">4.6 mm</quantity>
|
|||
|
|
), USNM 93513, Hawaiian Islands, Maui, southeast Maalaea Bay, Kalama Park, old coral, ~
|
|||
|
|
<quantity id="4C999B55D870FFFA08E8FE22FDCAFE4E" box="[503,572,392,416]" metricMagnitude="0" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="275" unit="m" value="2.7">2.7 m</quantity>
|
|||
|
|
; 1 ovig. female (cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0A02FE2DFC8EFE71" box="[797,888,391,416]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="275" unit="mm" value="6.8">6.8 mm</quantity>
|
|||
|
|
), UF 12880, Mascarene Islands,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0DEFFE2DFAB9FE71" box="[1264,1359,391,416]" name="Réunion" pageId="8" pageNumber="275">Reunion</collectingCountry>
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0C48FE2DFA6AFE71" box="[1367,1436,391,416]" name="Iceland" pageId="8" pageNumber="275">Island</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0980FE06FEDDFE14" box="[159,299,428,453]" name="Réunion" pageId="8" pageNumber="275">La Réunion</collectingCountry>
|
|||
|
|
), Saint-Leu, Sec Jaune, -21.1533° -55.2811°, rocky slope, basalt blocks, fore reef, depth
|
|||
|
|
<quantity id="4C999B55D870FFFA0C23FE06FA61FE15" box="[1340,1431,428,453]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="8" pageNumber="275" unit="m" value="10.5" valueMax="15.0" valueMin="6.0">6–15 m</quantity>
|
|||
|
|
, rubble extraction, leg. H. Bruggemann, N. Hubert, F. Michonneau, G. Paulay,
|
|||
|
|
<date id="FFDF1070D870FFFA0D0CFE7AFB5FFE39" box="[1043,1193,464,488]" pageId="8" pageNumber="275" value="2007-08-17">17 Aug 2007</date>
|
|||
|
|
[fcn BREU-1359];
|
|||
|
|
<specimenCount id="9D67FD39D870FFFA0C91FE7AFF3BFDDC" pageId="8" pageNumber="275" type="male">1 male</specimenCount>
|
|||
|
|
(cl
|
|||
|
|
<quantity id="4C999B55D870FFFA09E3FE5EFEAEFDDD" box="[252,344,500,525]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="275" unit="mm" value="6.5">6.5 mm</quantity>
|
|||
|
|
), UF 12873, Mascarene Islands,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0BCEFE5EFCC6FDDC" box="[721,816,500,525]" name="Réunion" pageId="8" pageNumber="275">Reunion</collectingCountry>
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0A27FE5EFC8AFDDC" box="[824,892,500,525]" name="Iceland" pageId="8" pageNumber="275">Island</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0A92FE5EFBEFFDDC" box="[909,1049,500,525]" name="Réunion" pageId="8" pageNumber="275">La Réunion</collectingCountry>
|
|||
|
|
), Saint-Leu, Sec Jaune, -21.1533° -55.2811°, rocky slope, basalt blocks, fore reef, depth
|
|||
|
|
<quantity id="4C999B55D870FFFA0BE2FDBDFCA3FDFE" box="[765,853,535,560]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="8" pageNumber="275" unit="m" value="10.5" valueMax="15.0" valueMin="6.0">6–15 m</quantity>
|
|||
|
|
, rubble extraction, leg. H. Bruggemann, N. Hubert, F. Michonneau, G. Paulay,
|
|||
|
|
<date id="FFDF1070D870FFFA08D3FD96FD95FD85" box="[460,611,572,596]" pageId="8" pageNumber="275" value="2007-08-17">17 Aug 2007</date>
|
|||
|
|
[fcn BREU-1350];
|
|||
|
|
<specimenCount id="9D67FD39D870FFFA0A56FD96FC5AFD84" box="[841,940,572,597]" pageId="8" pageNumber="275" type="female">1 female</specimenCount>
|
|||
|
|
(cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0AC3FD96FBCFFD85" box="[988,1081,572,597]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="275" unit="mm" value="4.6">4.6 mm</quantity>
|
|||
|
|
), UF 15443,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0DCEFD96FA6EFD84" box="[1233,1432,572,597]" name="French Polynesia" pageId="8" pageNumber="275">French Polynesia</collectingCountry>
|
|||
|
|
, Society Islands, Moorea, of northern coast between Pihaena and Opunohu Bay, outer reef slope, depth ~
|
|||
|
|
<quantity id="4C999B55D870FFFA0C46FDCAFA61FDA6" box="[1369,1431,608,632]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="275" unit="m" value="12.0">12 m</quantity>
|
|||
|
|
, rubble cracking, leg. S. McKeon, J. Moore, G. Paulay,
|
|||
|
|
<date id="FFDF1070D870FFFA0BE6FD2EFC75FD4D" box="[761,899,644,668]" pageId="8" pageNumber="275" value="2008-10-12">12 Oct 2008</date>
|
|||
|
|
[fcn BMOO-2258]; 1 ovig. female (cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0C29FD2EFA66FD4D" box="[1334,1424,644,669]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="8" pageNumber="275" unit="mm" value="6.8">6.8 mm</quantity>
|
|||
|
|
), UF 28992,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0802FD0DFE12FD11" box="[285,484,679,704]" name="French Polynesia" pageId="8" pageNumber="275">French Polynesia</collectingCountry>
|
|||
|
|
, Society Islands, Moorea, south end of island, -17.60085° -149.84128°, reef flat, depth
|
|||
|
|
<quantity id="4C999B55D870FFFA09C2FD66FEECFD35" box="[221,282,716,740]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="275" unit="m" value="20.0">20 m</quantity>
|
|||
|
|
, leg. D. Uyeno,
|
|||
|
|
<date id="FFDF1070D870FFFA08C9FD66FD9DFD35" box="[470,619,716,740]" pageId="8" pageNumber="275" value="2010-11-17">17 Nov 2010</date>
|
|||
|
|
[fcn BMOO-15118];
|
|||
|
|
<specimenCount id="9D67FD39D870FFFA0A7BFD66FC46FD34" box="[868,944,716,741]" pageId="8" pageNumber="275" type="male">1 male</specimenCount>
|
|||
|
|
(cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0AC0FD66FBCCFD35" box="[991,1082,716,741]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="275" unit="mm" value="6.2">6.2 mm</quantity>
|
|||
|
|
), 1 ov. female (cl
|
|||
|
|
<quantity id="4C999B55D870FFFA0C16FD66FA92FD35" box="[1289,1380,716,741]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="8" pageNumber="275" unit="mm" value="6.3">6.3 mm</quantity>
|
|||
|
|
), UF 10622,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA09F1FD45FEBEFCD9" box="[238,328,751,776]" name="Kiribati" pageId="8" pageNumber="275">Kiribati</collectingCountry>
|
|||
|
|
, Line Islands,
|
|||
|
|
<collectingCountry id="F3767620D870FFFA08E9FD45FD6CFCD9" box="[502,666,751,776]" name="United States Pacific Island Wildlife Refuges" pageId="8" pageNumber="275">Palmyra Atoll</collectingCountry>
|
|||
|
|
, north side of atoll, 5.8958° -162.0815°, outer reef slope, in dead
|
|||
|
|
<taxonomicName id="4C614D33D870FFFA0988FCBEFEEBFCFD" box="[151,285,788,812]" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="8" pageNumber="275" phylum="Cnidaria" rank="genus">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0988FCBEFEEBFCFD" box="[151,285,788,812]" italics="true" pageId="8" pageNumber="275">Pocillopora</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
head, depth ~
|
|||
|
|
<quantity id="4C999B55D870FFFA08A4FCBEFE03FCFD" box="[443,501,788,812]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="275" unit="m" value="11.0">11 m</quantity>
|
|||
|
|
, leg. G. Paulay and N. Knowlton,
|
|||
|
|
<date id="FFDF1070D870FFFA0A6DFCBEFBF2FCFC" box="[882,1028,788,813]" pageId="8" pageNumber="275" value="2005-08-19">19 Aug 2005</date>
|
|||
|
|
[fcn BPALM-956, -957].
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B653BD870FFF909D8FC9DFC72FECD" lastPageId="11" lastPageNumber="278" pageId="8" pageNumber="275" type="description">
|
|||
|
|
<paragraph id="8BDE36B0D870FFFB09D8FC9DFAB8FF29" blockId="8.[151,1436,356,2037]" lastBlockId="9.[151,1436,151,248]" lastPageId="9" lastPageNumber="276" pageId="8" pageNumber="275">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA09D8FC9DFEA7FC81" bold="true" box="[199,337,823,848]" pageId="8" pageNumber="275">Description</emphasis>
|
|||
|
|
. Small-sized species of
|
|||
|
|
<taxonomicName id="4C614D33D870FFFA0B7CFC92FD4BFC81" box="[611,701,824,848]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B915EAA2D870FFFA0B7CFC92FD4BFC81" box="[611,701,824,848]" italics="true" pageId="8" pageNumber="275">Alpheus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(cl range
|
|||
|
|
<quantity id="4C999B55D870FFFA0A31FC9DFC4CFC81" box="[814,954,823,848]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.8" metricValueMin="4.8" pageId="8" pageNumber="275" unit="mm" value="5.8" valueMax="6.8" valueMin="4.8">4.8–6.8 mm</quantity>
|
|||
|
|
). Carapace with small post-rostral tubercle on mid-dorsal line, at about 0.25 carapace length, separating more or less descendant frontal area from horizontal or slightly domed posterior area; post-rostral tubercle continued by low, sometimes barely discernable, mid-dorsal crest flattening completely before reaching posterior margin of carapace (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0AE9FC0EFBBEFC6C" box="[1014,1096,932,957]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
A, B, 8A, B). Rostrum welldeveloped, abruptly rising above deep orbito-rostral furrows, anterior portion narrow, carina-like, not overhanging orbito-rostral furrows, posterior portion flattened, broadening, overhanging orbito-rostral furrows; faint median carina running from rostral tip to post-rostral tubercle (better developed in younger individuals); rostral tip almost reaching distal margin of first article of antennular peduncle, pointing straight-foward or slightly descendant (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0C7BFB9EFF53FBA1" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
A, B, 8A, B). Orbital hoods unarmed, rounded anteriorly (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0A21FBFDFC7AFBA1" box="[830,908,1111,1136]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
A, 8A). Pterygostomial angle broadly rounded (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0980FBD6FF19FB44" box="[159,239,1148,1173]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
B, 8B). Telson moderately broad, tapering distally; dorsal spiniform setae stout, inserted at some distance from lateral margins; posterior margin broadly rounded, with two pairs of distolateral spiniform setae, mesial much longer than lateral, and row of smaller spiniform setae (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0A37FB6EFC8CFB0C" box="[808,890,1220,1245]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
J, K, 8J, K). Antennular peduncle with slender stylocerite, its acute tip overreaching distal margin of first article; second article about 1.3-1.5 times as long as wide (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0980FAA6FF02FAF4" box="[159,244,1292,1317]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
A, 8A). Antenna with stout basicerite armed with strong, sharp distoventral tooth; carpocerite not exceeding scaphocerite; scaphocerite with broadly concave lateral margin, strong distolateral tooth overreaching end of antennular peduncle, blade moderately broad (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0BE5FAFEFCB1FABC" box="[762,839,1364,1389]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
A-C, 8A, B). Third maxilliped with antepenultimate article not expanded; penultimate article at least 2.5 times as long as wide (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0D0AFADDFB90FA41" box="[1045,1126,1399,1424]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
D, E, 8C). Minor cheliped significantly smaller and more slender than major cheliped; merus with dorsal margin ending in stout distodorsal tooth, mesial margin armed with one or several spiniform seta and ending in small, sharp distomesial tooth; carpus cup-shaped; chela with palm subcylindrical, rounded in cross-section, smooth, distomesial and distolateral areas on each side of dactylar articulation blunt; fingers slightly shorter than palm, conical, non-gaping, dactylus without balaeniceps setae in both sexes (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0B0FF986FD97F995" box="[528,609,1580,1605]" captionStart="FIGURE 7" captionStartId="10.[151,250,1433,1455]" captionTargetBox="[163,1429,199,1401]" captionTargetId="figure@10.[151,1436,193,1412]" captionTargetPageId="10" captionText="FIGURE 7. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—minor cheliped, lateral view; B—same, distal carpus and chela, mesial view; C—major cheliped, lateral view; D—same, basis, ischium, merus and carpus, mesial view; E—major chela, mesial view; F—major chela, lateral view; G—distal major chela, fingers opened, lateral view." httpUri="https://zenodo.org/record/223850/files/figure.png" pageId="8" pageNumber="275">Figs. 7</figureCitation>
|
|||
|
|
A, B, 9A, B). Major cheliped with short, stout merus; dorsal margin of merus slightly rugose, ending in strong, curved distodorsal tooth; mesial margin of merus rugose, with widely spaced, stout spiniform setae, ending in strong, sharp distomesial tooth; carpus very short, bowl-shaped; chela with palm ovoid, rounded in cross-section, smooth except for deep transverse groove on distodorsal surface and shallow longitudinal depression on distolateral surface, adjacent to transverse groove; fingers less than half-length of palm, dactylus with short, truncate plunger (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0B20F975FD7CF929" box="[575,650,1759,1784]" captionStart="FIGURE 7" captionStartId="10.[151,250,1433,1455]" captionTargetBox="[163,1429,199,1401]" captionTargetId="figure@10.[151,1436,193,1412]" captionTargetPageId="10" captionText="FIGURE 7. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—minor cheliped, lateral view; B—same, distal carpus and chela, mesial view; C—major cheliped, lateral view; D—same, basis, ischium, merus and carpus, mesial view; E—major chela, mesial view; F—major chela, lateral view; G—distal major chela, fingers opened, lateral view." httpUri="https://zenodo.org/record/223850/files/figure.png" pageId="8" pageNumber="275">Figs. 7</figureCitation>
|
|||
|
|
C-G, 9C-F). Second pereiopod with first carpal article 1.0-1.3 times as long as second (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA084DF8AEFE57F8CC" box="[338,417,1796,1821]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
F, 8D). Third and fourth pereiopods similar, slender; ischium with spiniform seta; merus ~5.0 times as long as wide, with stout, sharp distoventral tooth; carpus with sharp distoventral tooth; propodus with row of slender spiniform setae on ventral margin (some inserted in pairs), as well as one pair of distoventral and one distodorsal spiniform setae; dactylus simple, conical, slightly curving, about 0.3 length of propodus (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0C28F8C5FA75F859" box="[1335,1411,1903,1928]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="8" pageNumber="275">Figs. 6</figureCitation>
|
|||
|
|
G, 8E). Fifth pereiopod more slender than third and fourth pereiopods; ischium with small spiniform seta; propodus with row of slender spiniform setae on ventral margin, cleaning brush poorly developed.
|
|||
|
|
<collectingCountry id="F3767620D870FFFA0D6AF812FB59F800" box="[1141,1199,1976,2001]" name="Malta" pageId="8" pageNumber="275">Male</collectingCountry>
|
|||
|
|
second pleopod with appendix masculina reaching far beyond appendix interna (
|
|||
|
|
<figureCitation id="135A2A35D870FFFA0A55F876FC66F825" box="[842,912,2012,2037]" captionStart="FIGURE 8" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[155,1429,449,1820]" captionTargetId="figure@11.[151,1436,445,1840]" captionTargetPageId="11" captionText="FIGURE 8. Alpheus pseudopugnax (Banner, 1953), male from Reunion Island (La Réunion), UF 12873: A—frontal region, dorsal view; B—same, lateral view; C—third maxilliped, lateral view; D—second pereiopod, lateral view; E—third pereiopod, lateral view; F—same, distal carpus, propodus and dactylus, lateral view; G—second pleopod, appendices masculina and interna, mesial view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223851/files/figure.png" pageId="8" pageNumber="275">Fig. 8</figureCitation>
|
|||
|
|
G). Uropod with exopod slightly longer than endopod; protopod with lateral lobe ending in strong, sharp tooth; exopod with diaeresis sinuous, ending in strong sublateral tooth, distolateral tooth strong; distolateral spiniform seta long, slender; endopod with row of slender spiniform setae on distal margin (
|
|||
|
|
<figureCitation id="135A2A35D871FFFB0B11FF75FDAAFF29" box="[526,604,223,248]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="9" pageNumber="276">Figs. 6</figureCitation>
|
|||
|
|
H, I, 8H, I). See also description and illustrations in Banner (1953).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1E6638D871FFFB0988F8E6FEE4F80D" httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="9" pageNumber="276" targetBox="[169,1427,310,1793]" targetPageId="9">
|
|||
|
|
<paragraph id="8BDE36B0D871FFFB0988F8E6FEE4F80D" blockId="9.[151,1436,1868,2012]" pageId="9" pageNumber="276">
|
|||
|
|
<emphasis id="B915EAA2D871FFFB0988F8E6FEF8F8B0" bold="true" box="[151,270,1868,1890]" pageId="9" pageNumber="276">FIGURE 6</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C614D33D871FFFB0801F8E6FD47F8B3" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[286,689,1868,1890]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="276" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D871FFFB0801F8E6FE86F8B3" box="[286,368,1868,1890]" italics="true" pageId="9" pageNumber="276">Alpheus</emphasis>
|
|||
|
|
<emphasis id="B915EAA2D871FFFB0866F8E6FDFDF8B3" box="[377,523,1868,1890]" italics="true" pageId="9" pageNumber="276">pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF1E6638D872FFF80988FA33FEE5F9DA" httpUri="https://zenodo.org/record/223850/files/figure.png" pageId="10" pageNumber="277" targetBox="[163,1429,199,1401]" targetPageId="10">
|
|||
|
|
<paragraph id="8BDE36B0D872FFF80988FA33FEE5F9DA" blockId="10.[151,1436,1433,1547]" pageId="10" pageNumber="277">
|
|||
|
|
<emphasis id="B915EAA2D872FFF80988FA33FEFBFA7F" bold="true" box="[151,269,1433,1455]" pageId="10" pageNumber="277">FIGURE 7</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C614D33D872FFF80803FA33FD5AFA7E" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[284,684,1433,1455]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="277" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D872FFF80803FA33FDF1FA7E" box="[284,519,1433,1455]" italics="true" pageId="10" pageNumber="277">Alpheus pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, female from Moorea, French Polynesia, UF 15443: A—minor cheliped, lateral view; B—same, distal carpus and chela, mesial view; C—major cheliped, lateral view; D—same, basis, ischium, merus and carpus, mesial view; E—major chela, mesial view; F—major chela, lateral view; G—distal
|
|||
|
|
<taxonomicName id="4C614D33D872FFF80D68FA7CFB44FA3D" box="[1143,1202,1494,1516]" pageId="10" pageNumber="277" rank="species" species="major">major</taxonomicName>
|
|||
|
|
chela, fingers opened, lateral view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDE36B0D872FFF909D8F998FC72FECD" blockId="10.[151,1437,1586,2007]" lastBlockId="11.[151,1436,151,392]" lastPageId="11" lastPageNumber="278" pageId="10" pageNumber="277">
|
|||
|
|
<emphasis id="B915EAA2D872FFF809D8F998FE8DF99A" bold="true" box="[199,379,1586,1611]" pageId="10" pageNumber="277">Colour pattern</emphasis>
|
|||
|
|
. Two colour pattern
|
|||
|
|
<typeStatus id="54DA8812D872FFF80B78F999FD54F99A" box="[615,674,1587,1611]" pageId="10" pageNumber="277">types</typeStatus>
|
|||
|
|
differing mainly in the dorsal colour of the fifth abdominal somite.
|
|||
|
|
<typeStatus id="54DA8812D872FFF80988F9FDFF39F9BE" box="[151,207,1623,1647]" pageId="10" pageNumber="277">Type</typeStatus>
|
|||
|
|
A (
|
|||
|
|
<figureCitation id="135A2A35D872FFF809E7F9FCFEBEF9BE" box="[248,328,1622,1647]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="10" pageNumber="277">Fig. 10</figureCitation>
|
|||
|
|
A–C): body with semitransparent-white background and broad, wine-red transverse bands arranged as following, from anterior to posterior: first band crossing anterior margin of carapace, encompassing rostroorbital area; second band crossing carapace at about mid-length; third band crossing first and second abdominal somites, tangentially touching posterior margin of carapace; fourth band crossing posterior third of third and most of fourth abdominal somites; fifth band occupying most of sixth abdominal somite; fifth abdominal somite mostly white except for deep-red ventral portions of pleura; antennular and antennal peduncles dark red with some whitish areas, flagella pale reddish; minor chela dark red; major chela mostly white, with some red-orange on mesialproximal portion of palm; dactylus orange-brown with hyaline-white tip; second pereiopods and walking legs dark red; pleopod with whitish protopods, exopods and endopods red; tail fan dark red distally, mainly white proximally.
|
|||
|
|
<typeStatus id="54DA8812D872FFF80988F831FF39F862" box="[151,207,1947,1971]" pageId="10" pageNumber="277">Type</typeStatus>
|
|||
|
|
B (
|
|||
|
|
<figureCitation id="135A2A35D872FFF8081DF830FEAEF862" box="[258,344,1946,1971]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="10" pageNumber="277">Fig. 10</figureCitation>
|
|||
|
|
D–F): generally similar to
|
|||
|
|
<typeStatus id="54DA8812D872FFF80B86F831FD3FF862" box="[665,713,1947,1971]" pageId="10" pageNumber="277">type</typeStatus>
|
|||
|
|
A, but first and second bands of carapace with respectively posterior and anterior margins more irregular, white band between them narrower, more crescent-shape; major chela sometimes with orange rash spots on mesial face of palm; posterior white band on carapace sometimes with yellow-orange spots dorsally; fourth band of abdomen crossing posterior third of third, fourth and anterior half of fifth abdominal somites, leaving either a narrow transverse white band (
|
|||
|
|
<figureCitation id="135A2A35D873FFF90AA1FF75FBFBFF29" box="[958,1037,223,248]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="11" pageNumber="278">Fig. 10</figureCitation>
|
|||
|
|
D–F) or a more irregular, U-shaped band (
|
|||
|
|
<figureCitation id="135A2A35D873FFF909C2FEAEFEDAFECD" box="[221,300,260,285]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="11" pageNumber="278">Fig. 10</figureCitation>
|
|||
|
|
G, H) on fifth somite. Fresh eggs dark red (
|
|||
|
|
<figureCitation id="135A2A35D873FFF90A11FEAEFC96FECD" box="[782,864,260,285]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="11" pageNumber="278">Fig. 10</figureCitation>
|
|||
|
|
H).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B653BD873FFFE09D8FE8DFBD9FA95" lastPageId="12" lastPageNumber="279" pageId="11" pageNumber="278" type="distribution">
|
|||
|
|
<paragraph id="8BDE36B0D873FFF909D8FE8DFCCAFE59" blockId="11.[151,1436,151,392]" pageId="11" pageNumber="278">
|
|||
|
|
<emphasis id="B915EAA2D873FFF909D8FE8DFEAFFE91" bold="true" box="[199,345,295,320]" pageId="11" pageNumber="278">Distribution</emphasis>
|
|||
|
|
. Indo-West Pacific from southwestern Indian Ocean to Hawaii and
|
|||
|
|
<collectingCountry id="F3767620D873FFF90D63FE8DFAB0FE91" box="[1148,1350,295,320]" name="French Polynesia" pageId="11" pageNumber="278">French Polynesia</collectingCountry>
|
|||
|
|
. Indian Ocean:
|
|||
|
|
<collectingCountry id="F3767620D873FFF909F2FEE6FEBAFEB4" box="[237,332,332,357]" name="Réunion" pageId="11" pageNumber="278">Reunion</collectingCountry>
|
|||
|
|
<collectingCountry id="F3767620D873FFF9084BFEE6FE6FFEB4" box="[340,409,332,357]" name="Iceland" pageId="11" pageNumber="278">Island</collectingCountry>
|
|||
|
|
(present study). Pacific Ocean: Maui and Oahu, Hawaiian Islands (Banner, 1953);
|
|||
|
|
<collectingCountry id="F3767620D873FFF90C20FEE6FF25FE59" name="United States Pacific Island Wildlife Refuges" pageId="11" pageNumber="278">Palmyra Atoll</collectingCountry>
|
|||
|
|
, Line Islands; Moorea, Society Islands (present study).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1E6638D873FFF90988F8EFFD43F807" httpUri="https://zenodo.org/record/223851/files/figure.png" pageId="11" pageNumber="278" targetBox="[155,1429,449,1820]" targetPageId="11">
|
|||
|
|
<paragraph id="8BDE36B0D873FFF90988F8EFFD43F807" blockId="11.[151,1436,1861,2006]" pageId="11" pageNumber="278">
|
|||
|
|
<emphasis id="B915EAA2D873FFF90988F8EFFEFBF88B" bold="true" box="[151,269,1861,1883]" pageId="11" pageNumber="278">FIGURE 8</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C614D33D873FFF90803F8EFFD5BF88A" authority="(Banner, 1953)" authorityName="Banner" authorityYear="1953" baseAuthorityName="Banner" baseAuthorityYear="1953" box="[284,685,1861,1883]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="278" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D873FFF90803F8EFFDFEF88A" box="[284,520,1861,1883]" italics="true" pageId="11" pageNumber="278">Alpheus pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male from Reunion Island (La Réunion), UF 12873: A—frontal region, dorsal view; B—same, lateral view; C—third maxilliped, lateral view; D—second pereiopod, lateral view; E—third pereiopod, lateral view; F—same, distal carpus, propodus and dactylus, lateral view; G—second pleopod, appendices masculina and interna, mesial view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF1E6638D874FFFE0988FB27FB61FB31" httpUri="https://zenodo.org/record/223852/files/figure.png" pageId="12" pageNumber="279" targetBox="[184,1414,185,1138]" targetPageId="12">
|
|||
|
|
<paragraph id="8BDE36B0D874FFFE0988FB27FB61FB31" blockId="12.[151,1436,1165,1248]" pageId="12" pageNumber="279">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0988FB27FEFBFB73" bold="true" box="[151,269,1165,1187]" pageId="12" pageNumber="279">FIGURE 9</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0804FB27FD5EFB72" authority="(Banner, 1953)" authorityName="Banner" authorityYear="1953" baseAuthorityName="Banner" baseAuthorityYear="1953" box="[283,680,1165,1187]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0804FB27FDF3FB72" box="[283,517,1165,1187]" italics="true" pageId="12" pageNumber="279">Alpheus pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male from Reunion Island (La Réunion), UF 12873: A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—major cheliped, ischium, merus and carpus, mesial view; D –major chela, mesial view; E—major chela, lateral view; F—distal major chela, fingers opened, lateral view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDE36B0D874FFFE09D8FAADFBD9FA95" blockId="12.[151,1436,1287,2032]" pageId="12" pageNumber="279">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE09D8FAADFED2FAF1" bold="true" box="[199,292,1287,1312]" pageId="12" pageNumber="279">Ecology</emphasis>
|
|||
|
|
. Coral reef and associated habitats, in dead coral heads, in crevices of coral rubble; depth range:
|
|||
|
|
<quantity id="4C999B55D874FFFE0C7BFAA2FF5BFA92" metricMagnitude="1" metricUnit="m" metricValue="1.1" metricValueMax="2.0" metricValueMin="0.2" pageId="12" pageNumber="279" unit="m" value="11.0" valueMax="20.0" valueMin="2.0">2–20 m</quantity>
|
|||
|
|
[e.g.,
|
|||
|
|
<quantity id="4C999B55D874FFFE09ECFA81FECCFA92" box="[243,314,1323,1348]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="12" pageNumber="279" unit="m" value="4.0" valueMax="6.0" valueMin="2.0">2–6 m</quantity>
|
|||
|
|
(Hawaii),
|
|||
|
|
<quantity id="4C999B55D874FFFE08ADFA86FE1CFA92" box="[434,490,1324,1348]" metricMagnitude="1" metricUnit="m" metricValue="1.1" pageId="12" pageNumber="279" unit="m" value="11.0">11 m</quantity>
|
|||
|
|
(Palmyra),
|
|||
|
|
<quantity id="4C999B55D874FFFE0B70FA86FD24FA92" box="[623,722,1324,1348]" metricMagnitude="1" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="12" pageNumber="279" unit="m" value="16.0" valueMax="20.0" valueMin="12.0">12–20 m</quantity>
|
|||
|
|
(Moorea),
|
|||
|
|
<quantity id="4C999B55D874FFFE0A4EFA81FC50FA92" box="[849,934,1323,1348]" metricMagnitude="1" metricUnit="m" metricValue="1.05" metricValueMax="1.5" metricValueMin="0.6" pageId="12" pageNumber="279" unit="m" value="10.5" valueMax="15.0" valueMin="6.0">6–15 m</quantity>
|
|||
|
|
(
|
|||
|
|
<collectingCountry id="F3767620D874FFFE0AABFA81FBEAFA95" box="[948,1052,1323,1348]" name="Réunion" pageId="12" pageNumber="279">Reunion</collectingCountry>
|
|||
|
|
)].
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C37B653BD874FFFC09D8FAE5FBB9F87E" lastPageId="14" lastPageNumber="281" pageId="12" pageNumber="279" type="discussion">
|
|||
|
|
<paragraph id="8BDE36B0D874FFFE09D8FAE5FD97F991" blockId="12.[151,1436,1287,2032]" pageId="12" pageNumber="279">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE09D8FAE5FEC2FAB9" bold="true" box="[199,308,1359,1384]" pageId="12" pageNumber="279">Remarks</emphasis>
|
|||
|
|
. The specimens from Moorea,
|
|||
|
|
<collectingCountry id="F3767620D874FFFE0BA3FAE5FC95FAB9" box="[700,867,1359,1384]" name="United States Pacific Island Wildlife Refuges" pageId="12" pageNumber="279">Palmyra Atoll</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F3767620D874FFFE0AB6FAE5FBFEFAB9" box="[937,1032,1359,1384]" name="Réunion" pageId="12" pageNumber="279">Reunion</collectingCountry>
|
|||
|
|
represent the first records of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0C9BFAFAFECFFA5D" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0C9BFAFAFECFFA5D" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
since the original description of the species from Hawaii. The new records significantly extend the range of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE09E2FA32FE36FA61" box="[253,448,1432,1456]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE09E2FA32FE36FA61" box="[253,448,1432,1456]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
from north-central Pacific Ocean (Hawaii) to south-central and western Pacific Ocean (
|
|||
|
|
<collectingCountry id="F3767620D874FFFE0981FA11FE97FA05" box="[158,353,1467,1492]" name="French Polynesia" pageId="12" pageNumber="279">French Polynesia</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F3767620D874FFFE0888FA11FE06FA05" box="[407,496,1467,1492]" name="Kiribati" pageId="12" pageNumber="279">Kiribati</collectingCountry>
|
|||
|
|
) and south-western Indian Ocean (Mascarene Islands). Thus,
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0DBBFA16FA93FA05" box="[1188,1381,1468,1492]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0DBBFA16FA93FA05" box="[1188,1381,1468,1492]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, like its relative
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE080BFA4AFE8FFA26" box="[276,377,1504,1527]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE080BFA4AFE8FFA26" box="[276,377,1504,1527]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, appears to be widely distributed in the Indo-West and Central Pacific. However, both species are relatively uncommon and appear to be more common around oceanic or offshore islands with healthy coral reef stands and biologically rich coral rubble.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDE36B0D874FFFE09D8F9E6FC31F821" blockId="12.[151,1436,1287,2032]" pageId="12" pageNumber="279">
|
|||
|
|
Banner (1953) listed three morphological differences between
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0A8DF9E6FBA3F9B5" box="[914,1109,1612,1636]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0A8DF9E6FBA3F9B5" box="[914,1109,1612,1636]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0D90F9E6FB03F9B2" box="[1167,1269,1612,1635]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0D90F9E6FB03F9B2" box="[1167,1269,1612,1635]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, involving the orbito-rostral region and major and minor chelipeds. In
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0A27F9DAFBF6F959" box="[824,1024,1648,1672]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0A27F9DAFBF6F959" box="[824,1024,1648,1672]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the rostral carina is not flattened posterior to the rostral tip, running as a flattened ridge, before broadening, becoming more abrupt and overhanging the orbito-rostral grooves between or posterior to the eyes; in addition, a faint median carina is noticeable in the flattened portion of the post-rostral plate (
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0B6CF971FD37F925" box="[627,705,1755,1780]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="12" pageNumber="279">Figs. 6</figureCitation>
|
|||
|
|
A, 8A). In
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0A20F976FC52F922" box="[831,932,1756,1779]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0A20F976FC52F922" box="[831,932,1756,1779]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the rostral carina flattens close to the tip of the rostrum, forming a narrowly triangular post-rostral plate without a trace of a median carina (
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0DD8F955FAE7F8C9" box="[1223,1297,1791,1816]" captionStart="FIGURE 1" captionStartId="3.[151,250,1768,1790]" captionTargetBox="[156,1422,383,1734]" captionTargetId="figure@3.[151,1436,373,1746]" captionTargetPageId="3" captionText="FIGURE 1. Alpheus percyi Coutière, 1908, male from French Frigate Shoals, Hawaiian Archipelago (UF 12372): A—frontal region, dorsal view; B—same, lateral view; C—second pereiopod, lateral view; D—third pereiopod, lateral view; E—same, distal carpus, propodus and dactylus, lateral view; F—second pleopod, appendices masculina and interna, mesial view; Guropod, dorsal view; H—same, spiniform setae on distal margin of endopod, dorsal view; I—telson, dorsal view; J—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223844/files/figure.png" pageId="12" pageNumber="279">Figs. 1</figureCitation>
|
|||
|
|
A, 3A). The major chela of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE085FF88EFDF7F8ED" box="[320,513,1828,1852]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE085FF88EFDF7F8ED" box="[320,513,1828,1852]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
has a slight longitudinal depression on the lateral face of the palm, adjacent to the deep transverse groove (
|
|||
|
|
<figureCitation id="135A2A35D874FFFE08B6F8EDFE00F88E" box="[425,502,1863,1888]" captionStart="FIGURE 7" captionStartId="10.[151,250,1433,1455]" captionTargetBox="[163,1429,199,1401]" captionTargetId="figure@10.[151,1436,193,1412]" captionTargetPageId="10" captionText="FIGURE 7. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—minor cheliped, lateral view; B—same, distal carpus and chela, mesial view; C—major cheliped, lateral view; D—same, basis, ischium, merus and carpus, mesial view; E—major chela, mesial view; F—major chela, lateral view; G—distal major chela, fingers opened, lateral view." httpUri="https://zenodo.org/record/223850/files/figure.png" pageId="12" pageNumber="279">Figs. 7</figureCitation>
|
|||
|
|
F, 9E); such a depression is not discernable on the palm of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0D90F8E2FB05F88E" box="[1167,1267,1864,1887]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0D90F8E2FB05F88E" box="[1167,1267,1864,1887]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0C1CF8EDFAA7F8B1" box="[1283,1361,1863,1888]" captionStart="FIGURE 2" captionStartId="4.[151,250,1238,1260]" captionTargetBox="[151,1425,204,1209]" captionTargetId="figure@4.[151,1436,193,1217]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus percyi Coutière, 1908, male from French Frigate Shoals, Hawaiian Archipelago (UF 12372): A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—same, ischium, merus and carpus, mesial view; D—major cheliped, lateral view; E—major chela, mesial view; F—major chela, lateral view." httpUri="https://zenodo.org/record/223845/files/figure.png" pageId="12" pageNumber="279">Figs. 2</figureCitation>
|
|||
|
|
F, 4E). The palm of the minor chela of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0B11F8C6FD25F855" box="[526,723,1900,1924]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0B11F8C6FD25F855" box="[526,723,1900,1924]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
does not have strong, sharp distolateral and distomesial teeth characteristic of
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE084EF83AFE43F876" box="[337,437,1936,1959]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE084EF83AFE43F876" box="[337,437,1936,1959]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(compare
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0B34F825FD8EF879" box="[555,632,1935,1960]" captionStart="FIGURE 2" captionStartId="4.[151,250,1238,1260]" captionTargetBox="[151,1425,204,1209]" captionTargetId="figure@4.[151,1436,193,1217]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus percyi Coutière, 1908, male from French Frigate Shoals, Hawaiian Archipelago (UF 12372): A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—same, ischium, merus and carpus, mesial view; D—major cheliped, lateral view; E—major chela, mesial view; F—major chela, lateral view." httpUri="https://zenodo.org/record/223845/files/figure.png" pageId="12" pageNumber="279">Figs. 2</figureCitation>
|
|||
|
|
A, B and 9A, B). One additional differentiating character not mentioned by Banner (1953) is the presence of balaeniceps setae on the male minor chela in
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0D43F81EFB35F81A" box="[1116,1219,1972,1995]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0D43F81EFB35F81A" box="[1116,1219,1972,1995]" italics="true" pageId="12" pageNumber="279">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0DC9F819FAEAF81D" box="[1238,1308,1971,1996]" captionStart="FIGURE 2" captionStartId="4.[151,250,1238,1260]" captionTargetBox="[151,1425,204,1209]" captionTargetId="figure@4.[151,1436,193,1217]" captionTargetPageId="4" captionText="FIGURE 2. Alpheus percyi Coutière, 1908, male from French Frigate Shoals, Hawaiian Archipelago (UF 12372): A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—same, ischium, merus and carpus, mesial view; D—major cheliped, lateral view; E—major chela, mesial view; F—major chela, lateral view." httpUri="https://zenodo.org/record/223845/files/figure.png" pageId="12" pageNumber="279">Fig. 2</figureCitation>
|
|||
|
|
B; see also Coutière 1921, fig. 22b) and their absence in
|
|||
|
|
<taxonomicName id="4C614D33D874FFFE0B8DF872FCA5F821" box="[658,851,2008,2032]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D874FFFE0B8DF872FCA5F821" box="[658,851,2008,2032]" italics="true" pageId="12" pageNumber="279">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="135A2A35D874FFFE0A7DF87DFC53F821" box="[866,933,2007,2032]" captionStart="FIGURE 9" captionStartId="12.[151,250,1165,1187]" captionTargetBox="[184,1414,185,1138]" captionTargetId="figure@12.[167,1417,181,1156]" captionTargetPageId="12" captionText="FIGURE 9. Alpheus pseudopugnax (Banner, 1953), male from Reunion Island (La Réunion), UF 12873: A—minor cheliped, lateral view; B—same, carpus and chela, mesial view; C—major cheliped, ischium, merus and carpus, mesial view; D – major chela, mesial view; E—major chela, lateral view; F—distal major chela, fingers opened, lateral view." httpUri="https://zenodo.org/record/223852/files/figure.png" pageId="12" pageNumber="279">Fig. 9</figureCitation>
|
|||
|
|
B).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF1E6638D875FFFF0988F88EFBD6F865" httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="13" pageNumber="280" targetBox="[151,1436,193,1805]" targetPageId="13">
|
|||
|
|
<paragraph id="8BDE36B0D875FFFF0988F88EFBD6F865" blockId="13.[151,1436,1828,1972]" pageId="13" pageNumber="280">
|
|||
|
|
<emphasis id="B915EAA2D875FFFF0988F88EFEECF8E8" bold="true" box="[151,282,1828,1850]" pageId="13" pageNumber="280">FIGURE 10</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C614D33D875FFFF0837F88EFD41F8EB" authority="Banner, 1953" authorityName="Banner" authorityYear="1953" box="[296,695,1828,1850]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="280" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D875FFFF0837F88EFE8CF8EB" box="[296,378,1828,1850]" italics="true" pageId="13" pageNumber="280">Alpheus</emphasis>
|
|||
|
|
<emphasis id="B915EAA2D875FFFF089EF88EFDE5F8EB" box="[385,531,1828,1850]" italics="true" pageId="13" pageNumber="280">pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
: A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF1E6638D876FFFC0988FC4FFB88FBA7" httpUri="https://zenodo.org/record/223854/files/figure.png" pageId="14" pageNumber="281" targetBox="[173,1419,205,957]" targetPageId="14">
|
|||
|
|
<paragraph id="8BDE36B0D876FFFC0988FC4FFB88FBA7" blockId="14.[151,1436,997,1142]" pageId="14" pageNumber="281">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0988FC4FFEE9FC2A" bold="true" box="[151,287,997,1019]" pageId="14" pageNumber="281">FIGURE 11</emphasis>
|
|||
|
|
. Variation in the shape of the postrostral region in
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0A70FC4FFB4CFC2A" authority="Coutiere, 1908" authorityName="Coutiere" authorityYear="1908" box="[879,1210,997,1020]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0A70FC4FFBFBFC2D" box="[879,1037,997,1020]" italics="true" pageId="14" pageNumber="281">Alpheus percyi</emphasis>
|
|||
|
|
Coutière, 1908
|
|||
|
|
</taxonomicName>
|
|||
|
|
(A–D) and
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0C55FC4FFE3BFBCB" authority="(Banner, 1953)" authorityName="Banner" authorityYear="1953" baseAuthorityName="Banner" baseAuthorityYear="1953" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0C55FC4FFEDFFBCB" italics="true" pageId="14" pageNumber="281">Alpheus pseudopugnax</emphasis>
|
|||
|
|
(Banner, 1953)
|
|||
|
|
</taxonomicName>
|
|||
|
|
(E–G): A—holotype female from Amirante, MNHN 3188; B—female from Tabuaeran, Line Islands, UF 10662; C—male from Oahu, Hawaiian Archipelago, UF 15237; D—non-type male of
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0DBCFB89FACAFBE8" box="[1187,1340,1059,1081]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0DBCFB89FACAFBE8" box="[1187,1340,1059,1081]" italics="true" pageId="14" pageNumber="281">Alpheus percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(?) from Amirante, MNHN 3486; E—holotype female from Maui, Hawaiian Archipelago, USNM 93513; F—ovigerous female from Reunion Island (La Réunion), UF 12880; G—female from Palmyra Atoll, Line Islands, UF 10622.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BDE36B0D876FFFC09D8FB09FE9DF95E" blockId="14.[151,1436,1187,1968]" pageId="14" pageNumber="281">
|
|||
|
|
Interestingly,
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC087EFB09FDD5FB6A" box="[353,547,1187,1211]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC087EFB09FDD5FB6A" box="[353,547,1187,1211]" italics="true" pageId="14" pageNumber="281">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
appears to display two
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0A2EFB0EFC9AFB6D" box="[817,876,1188,1212]" pageId="14" pageNumber="281">types</typeStatus>
|
|||
|
|
of colour pattern, one with the fifth somite white dorsally (
|
|||
|
|
<typeStatus id="54DA8812D876FFFC081CFB6DFEC3FB0E" box="[259,309,1223,1247]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
A) and one with the fifth somite dark red dorsally (
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0A82FB6DFC39FB0E" box="[925,975,1223,1247]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
B).
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0D18FB62FBC9FB31" box="[1031,1087,1224,1248]" pageId="14" pageNumber="281">Type</typeStatus>
|
|||
|
|
A pattern is displayed by the smaller Moorea specimen and one of the
|
|||
|
|
<collectingCountry id="F3767620D876FFFC0B7DFB41FD37FAD5" box="[610,705,1259,1284]" name="Réunion" pageId="14" pageNumber="281">Reunion</collectingCountry>
|
|||
|
|
specimens (
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0A54FB41FC6DFAD2" box="[843,923,1259,1284]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="14" pageNumber="281">Fig. 10</figureCitation>
|
|||
|
|
A–C), whereas
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0D56FB46FB8FFAD5" box="[1097,1145,1260,1284]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
B pattern is present in the Palmyra specimens and the second
|
|||
|
|
<collectingCountry id="F3767620D876FFFC0B5BFAA4FD55FAF6" box="[580,675,1294,1319]" name="Réunion" pageId="14" pageNumber="281">Reunion</collectingCountry>
|
|||
|
|
specimen (
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0A2FFAA4FC71FAF6" box="[816,903,1294,1319]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="14" pageNumber="281">Fig. 10</figureCitation>
|
|||
|
|
D–F). However, the larger Moorea specimen displays a colour pattern that is intermediate between
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0A0CFA9EFCB5FA9D" box="[787,835,1332,1356]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
A and
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0ABDFA9EFC24FA9D" box="[930,978,1332,1356]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
B (
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0D1CFA99FBA1FA9A" box="[1027,1111,1331,1356]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="14" pageNumber="281">Fig. 10</figureCitation>
|
|||
|
|
G, H), suggesting that these patterns are not consistent and may vary individually and/or with sex or age. Both colour pattern
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0DFCFAF2FAE8FAA1" box="[1251,1310,1368,1392]" pageId="14" pageNumber="281">types</typeStatus>
|
|||
|
|
agree with Banner’s (1953) short description of the pattern of
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0BEBFAD1FC4FFA42" box="[756,953,1403,1427]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0BEBFAD1FC4FFA42" box="[756,953,1403,1427]" italics="true" pageId="14" pageNumber="281">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: “in life,
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0D2EFAD6FB97FA45" box="[1073,1121,1404,1428]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
specimen with broad, red, transverse bands”. A side-by-side morphological comparison between the specimens of
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0D8BFA0AFB32FA69" box="[1172,1220,1440,1464]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
A and B revealed some differences, for instance, in the post-rostral region (which is generally quite variable), but also in the length of the second article of the antennular peduncle (cf.
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0BDCFA4CFCF9FA2E" box="[707,783,1510,1535]" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="14" pageNumber="281">Figs. 6</figureCitation>
|
|||
|
|
A, 8A) and the proportions of the second pereiopod (
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0C7BFA4CFF52F9F5" captionStart="FIGURE 6" captionStartId="9.[151,250,1868,1890]" captionTargetBox="[169,1427,310,1793]" captionTargetId="figure@9.[151,1436,301,1846]" captionTargetPageId="9" captionText="FIGURE 6. Alpheus pseudopugnax (Banner, 1953), female from Moorea, French Polynesia, UF 15443: A—frontal region, dorsal view; B—same, lateral view; C—antennal scaphocerite, dorsal view; D—third maxilliped, lateral view; E—same, lateral plate and mastigobranch, lateral view; F—second pereiopod, lateral view; G—third pereiopod, lateral view; H—uropod, dorsal view; I—same, spiniform setae on distal margin of endopod, dorsal view; J—telson, dorsal view; K—same, posterior margin, dorsal view." httpUri="https://zenodo.org/record/223849/files/figure.png" pageId="14" pageNumber="281">Figs. 6</figureCitation>
|
|||
|
|
G, 8E), which appears to be more slender in the
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0BA0F9A6FD19F9F5" box="[703,751,1548,1572]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
B specimen from
|
|||
|
|
<collectingCountry id="F3767620D876FFFC0AA0F9A1FBE8F9F5" box="[959,1054,1547,1572]" name="Réunion" pageId="14" pageNumber="281">Reunion</collectingCountry>
|
|||
|
|
than in the
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0DBDF9A6FB24F9F5" box="[1186,1234,1548,1572]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
A specimen from Moorea. However, a preliminary sequencing of the barcoding segment of the COI gene (C. Hurt, pers. comm.) revealed only a very insignificant difference between the smaller Moorea specimen (
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0D67F9F9FB5CF9BA" box="[1144,1194,1619,1643]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
A) and the Palmyra specimen (
|
|||
|
|
<typeStatus id="54DA8812D876FFFC0810F9DDFEB7F95E" box="[271,321,1655,1679]" pageId="14" pageNumber="281">type</typeStatus>
|
|||
|
|
B).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BDE36B0D876FFFC09D8F931FBB9F87E" blockId="14.[151,1436,1187,1968]" pageId="14" pageNumber="281">
|
|||
|
|
A remarkable and unusual feature of the colour pattern of
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0A47F931FBECF962" box="[856,1050,1691,1715]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0A47F931FBECF962" box="[856,1050,1691,1715]" italics="true" pageId="14" pageNumber="281">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the colour contrast between the major and minor cheliped. The major chela is conspicuously white, only with brown-red base and occasionally with orange, rash-like pattern, whilst the minor chela is dark red (
|
|||
|
|
<figureCitation id="135A2A35D876FFFC0AD9F949FBE8F92A" box="[966,1054,1763,1788]" captionStart="FIGURE 10" captionStartId="13.[151,250,1828,1850]" captionTargetBox="[151,1436,193,1805]" captionTargetId="figure@13.[151,1436,193,1806]" captionTargetPageId="13" captionText="FIGURE 10. Alpheus pseudopugnax (Banner, 1953): A, B—ovigerous female from Reunion Island (La Réunion), UF 12880 [type A colour pattern]; C—female from Moorea, French Polynesia, UF 15443 [type A colour pattern]; D, E—male from Palmyra Atoll, Line Islands, UF 10622 [type B colour pattern]; F—male from Reunion Island (La Réunion), UF 12873 [type B colour pattern]; G, H—ovigerous female (missing major cheliped) from Moorea, French Polynesia, UF 28992 [intermediate A- B colour pattern]. Photographers: Gustav Paulay (A, B, D, E, F); Arthur Anker (C, G, H)." httpUri="https://zenodo.org/record/223853/files/figure.png" pageId="14" pageNumber="281">Fig. 10</figureCitation>
|
|||
|
|
A–F). Because of this disparate colouration of the chelipeds and the presence of broad red bands on the carapace and abdomen, the colour pattern of
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC09AAF881FE80F892" box="[181,374,1835,1859]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="pseudopugnax">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC09AAF881FE80F892" box="[181,374,1835,1859]" italics="true" pageId="14" pageNumber="281">A. pseudopugnax</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
greatly differs from that of
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0BB4F881FCF9F892" box="[683,783,1835,1859]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="percyi">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0BB4F881FCF9F892" box="[683,783,1835,1859]" italics="true" pageId="14" pageNumber="281">A. percyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which shows more resemblance to some species of the
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0C9BF881FEF1F8B6" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="gracilipes">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0C9BF881FEF1F8B6" italics="true" pageId="14" pageNumber="281">A. gracilipes</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species complex (see above). A strikingly dissimilar colouration of the chelipeds is rare in alpheid shrimps and decapods in general, although it is a common character of many fiddler crabs (
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0DDCF8DEFB04F85A" box="[1219,1266,1908,1931]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0DDCF8DEFB04F85A" box="[1219,1266,1908,1931]" italics="true" pageId="14" pageNumber="281">Uca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
spp.) and also occurs in some hermit crabs, e.g.,
|
|||
|
|
<taxonomicName id="4C614D33D876FFFC0B0AF83DFBBAF87E" authority="Asakura & Paulay, 2003" authorityName="Asakura & Paulay" authorityYear="2003" box="[533,1100,1942,1967]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="lemaitrei">
|
|||
|
|
<emphasis id="B915EAA2D876FFFC0B0AF83DFCD9F87E" box="[533,815,1943,1967]" italics="true" pageId="14" pageNumber="281">Pylopaguropsis lemaitrei</emphasis>
|
|||
|
|
Asakura & Paulay, 2003
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|