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<document id="AF4A5BA012DE6A601EE022C532FBE31B" ID-CLB-Dataset="55625" ID-DOI="10.11646/zootaxa.5133.3.3" ID-GBIF-Dataset="02d6b967-3efb-4c75-96d3-e9ac8541bb3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6522677" ID-ZooBank="44C7B333-7D3E-4235-A158-60E70849ED60" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651756637829" checkinUser="plazi" docAuthor="Haddad, Charles R., Jin, Chi &amp; Platnick, Norman I." docDate="2022" docId="AE2487D1B52AFF4652DF9310C82AFA18" docLanguage="en" docName="zootaxa.5133.3.3.pdf" docOrigin="Zootaxa 5133 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orthobula aethiopica Haddad &amp; Jin &amp; Platnick 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="364" masterDocId="521DFFA9B52CFF4F5248952CCB73FFE9" masterDocTitle="A revision of the spider genus Orthobula Simon, 1897 (Araneae: Trachelidae) in the Afrotropical Region. I. Continental species" masterLastPageNumber="382" masterPageNumber="355" pageNumber="361" updateTime="1699329851161" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8E0DE2DEFD6659C08212A366122E60D4">A revision of the spider genus Orthobula Simon, 1897 (Araneae: Trachelidae) in the Afrotropical Region. I. Continental species</mods:title>
</mods:titleInfo>
<mods:name id="1A6E3F5636F25232EC108F91EC6752F1" type="personal">
<mods:role id="6CD0C71BA8CC4F30E03559C4361F24DE">
<mods:roleTerm id="14C54DA8B8B4647252ED14CA749DE89D">Author</mods:roleTerm>
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<mods:namePart id="507E699EBEAB9331D87CC6C801588083">Haddad, Charles R.</mods:namePart>
<mods:affiliation id="A49498BCEF26E6A84E4234C736D0FAC9">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="8B7D57B0362209227CED116571D2FA04">Jin, Chi</mods:namePart>
<mods:affiliation id="2FBA47CA95E9FB016F2292E4E045686E">School of Landscape and Ecological Engineering, Hebei University of Engineering, Handan, Hebei 056038, P. R. China.</mods:affiliation>
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<mods:namePart id="8197D64FCD226D824A3132A9103903C8">Platnick, Norman I.</mods:namePart>
<mods:nameIdentifier id="5234FD06C8EF4F1BEF81D19E9EDAB118" type="ORCID">0000-0003-4868-7529</mods:nameIdentifier>
<mods:affiliation id="21326085652E4522B27D0EFE7D65A069">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024, U. S. A. https: // orcid. org / 0000 - 0003 - 4868 - 7529</mods:affiliation>
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<mods:title id="0419B91196584222D767D7536A2ABB81">Zootaxa</mods:title>
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<mods:date id="9EF0342DB2B08D5D8052724F11608DB2">2022</mods:date>
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<mods:number id="A646F5661A84FB22A03E10AF5ED01334">2022-05-05</mods:number>
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<mods:number id="14D5D357C72F0E61442A94185B061CAC">5133</mods:number>
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<treatment id="AE2487D1B52AFF4652DF9310C82AFA18" ID-DOI="http://doi.org/10.5281/zenodo.6521600" ID-GBIF-Taxon="195277976" ID-Zenodo-Dep="6521600" LSID="urn:lsid:plazi:treatment:AE2487D1B52AFF4652DF9310C82AFA18" httpUri="http://treatment.plazi.org/id/AE2487D1B52AFF4652DF9310C82AFA18" lastPageId="9" lastPageNumber="364" pageId="6" pageNumber="361">
<subSubSection id="6E97654CB52AFF4952DF9310CA8DF9BF" box="[151,510,1596,1623]" pageId="6" pageNumber="361" type="nomenclature">
<paragraph id="263236C7B52AFF4952DF9310CA8DF9BF" blockId="6.[151,510,1596,1659]" box="[151,510,1596,1623]" pageId="6" pageNumber="361">
<heading id="7D7A81ABB52AFF4952DF9310CA8DF9BF" bold="true" box="[151,510,1596,1623]" fontSize="11" level="1" pageId="6" pageNumber="361" reason="1">
<emphasis id="14F9EAD5B52AFF4952DF9310CA8DF9BF" bold="true" box="[151,510,1596,1623]" pageId="6" pageNumber="361">
<taxonomicName id="E18D4D44B52AFF4952DF9310CAEAF9BE" ID-CoL="992HC" authority="Haddad &amp; Jin &amp; Platnick, 2022" authorityName="Haddad &amp; Jin &amp; Platnick" authorityYear="2022" box="[151,409,1596,1623]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="14F9EAD5B52AFF4952DF9310CAEAF9BE" bold="true" box="[151,409,1596,1623]" italics="true" pageId="6" pageNumber="361">Orthobula aethiopica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0FCA57AEB52AFF4953E99310CA8DF9BF" box="[417,510,1596,1622]" pageId="6" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6E97654CB52AFF4952DF934CCA3FF992" box="[151,332,1632,1659]" pageId="6" pageNumber="361" type="description">
<paragraph id="263236C7B52AFF4952DF934CCA3FF992" blockId="6.[151,510,1596,1659]" box="[151,332,1632,1659]" pageId="6" pageNumber="361">
<figureCitation id="BEB62A42B52AFF4952DF934CCB89F993" box="[151,250,1632,1658]" captionStart="FIGURES 1–12" captionStartId="1.[151,264,1877,1901]" captionTargetBox="[151,1436,489,1854]" captionTargetId="figure-164@1.[151,1436,489,1854]" captionTargetPageId="1" captionText="FIGURES 1–12. Dorsal habitus photographs of Orthobula species: 1. O. aethiopica sp. nov., female; 2. Same, male; 3. O. arca sp. nov., female; 4. Same, male; 5. O. calceata Simon, 1897, female; 6. Same, male; 7. O. marusiki sp. nov., female; 8. Same, male; 9. O. milloti Caporiacco, 1949, female; 10. Same, male; 11. O. radiata Simon, 1897, female; 12. Same, male. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6522679" httpUri="https://zenodo.org/record/6522679/files/figure.png" pageId="6" pageNumber="361">Figs 1, 2</figureCitation>
,
<figureCitation id="BEB62A42B52AFF49534E934CCA3FF992" box="[262,332,1632,1659]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="6" pageNumber="361">54–58</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6E97654CB52AFF4952DF9384CF21F89E" pageId="6" pageNumber="361" type="diagnosis">
<paragraph id="263236C7B52AFF4952DF9384CF21F89E" blockId="6.[151,1437,1704,1911]" pageId="6" pageNumber="361">
<emphasis id="14F9EAD5B52AFF4952DF9384CA61F92B" bold="true" box="[151,274,1704,1730]" pageId="6" pageNumber="361">Diagnosis.</emphasis>
Females of
<taxonomicName id="E18D4D44B52AFF4953EA9385C94EF92B" authority="Haddad &amp; Jin &amp; Platnick, 2022" authorityName="Haddad &amp; Jin &amp; Platnick" authorityYear="2022" box="[418,573,1704,1731]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="14F9EAD5B52AFF4953EA9385C94EF92B" box="[418,573,1704,1731]" italics="true" pageId="6" pageNumber="361">O. aethiopica</emphasis>
</taxonomicName>
<emphasis id="14F9EAD5B52AFF49500F9384C9D0F92B" bold="true" box="[583,675,1704,1730]" pageId="6" pageNumber="361">
<taxonomicNameLabel id="0FCA57AEB52AFF49500F9384C9D0F92B" box="[583,675,1704,1730]" pageId="6" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
share with those of
<taxonomicName id="E18D4D44B52AFF4951D99385C88CF92B" authorityName="Caporiacco" authorityYear="1949" box="[913,1023,1704,1731]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="14F9EAD5B52AFF4951D99385C88CF92B" box="[913,1023,1704,1731]" italics="true" pageId="6" pageNumber="361">O. milloti</emphasis>
</taxonomicName>
a broad, semicircular anterior ridge in the epigyne, but can be distinguished by the CD being about twice as long as the width of the SBB, while approximately equal to the width of the SBB in
<taxonomicName id="E18D4D44B52AFF4950D093DDC870F8E3" authorityName="Caporiacco" authorityYear="1949" box="[664,771,1776,1803]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="milloti">
<emphasis id="14F9EAD5B52AFF4950D093DDC870F8E3" box="[664,771,1776,1803]" italics="true" pageId="6" pageNumber="361">O. milloti</emphasis>
</taxonomicName>
(compare
<figureCitation id="BEB62A42B52AFF49513393DCC8BCF8E2" box="[891,975,1776,1803]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="6" pageNumber="361">Figs 54</figureCitation>
and
<figureCitation id="BEB62A42B52AFF49564D93DCCF56F8E3" box="[1029,1061,1776,1802]" captionStart="FIGURES 90–95" captionStartId="20.[151,264,1862,1886]" captionTargetBox="[159,1423,569,1831]" captionTargetId="figure-169@20.[151,1436,561,1839]" captionTargetPageId="20" captionText="FIGURES 90–95. Orthobula milloti Caporiacco, 1949, genitalic morphology of female from Nairobi (90, 91) and male from Pugu (92–95): epigyne, ventral (90) and dorsal (91) views; right palpal tegulum, retrolateral (92), ventral (93) and prolateral (94) views; left palpal femur, patella and tibia, retrolateral view. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522723" httpUri="https://zenodo.org/record/6522723/files/figure.png" pageId="6" pageNumber="361">90</figureCitation>
). Males of
<taxonomicName id="E18D4D44B52AFF4956EB93DDCE48F8E3" authority="Haddad &amp; Jin &amp; Platnick, 2022" authorityName="Haddad &amp; Jin &amp; Platnick" authorityYear="2022" box="[1187,1339,1776,1803]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="14F9EAD5B52AFF4956EB93DDCE48F8E3" box="[1187,1339,1776,1803]" italics="true" pageId="6" pageNumber="361">O. aethiopica</emphasis>
</taxonomicName>
<emphasis id="14F9EAD5B52AFF49570B93DCCEEFF8E3" bold="true" box="[1347,1436,1776,1802]" pageId="6" pageNumber="361">
<taxonomicNameLabel id="0FCA57AEB52AFF49570B93DCCEEFF8E3" box="[1347,1436,1776,1802]" pageId="6" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most closely resemble those of
<taxonomicName id="E18D4D44B52AFF4953BD9239C905F8C7" authorityName="Simon" authorityYear="1897" box="[501,630,1812,1839]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="14F9EAD5B52AFF4953BD9239C905F8C7" box="[501,630,1812,1839]" italics="true" pageId="6" pageNumber="361">O. calceata</emphasis>
</taxonomicName>
in the slightly undulating embolus in ventral view (
<figureCitation id="BEB62A42B52AFF4956FE9238CE7FF8C6" box="[1206,1292,1812,1839]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="6" pageNumber="361">Figs 57</figureCitation>
,
<figureCitation id="BEB62A42B52AFF4957509238CE44F8C6" box="[1304,1335,1812,1839]" captionStart="FIGURES 65–69" captionStartId="14.[151,264,1064,1088]" captionTargetBox="[168,1424,191,1036]" captionTargetId="figure-26@14.[161,1427,181,1040]" captionTargetPageId="14" captionText="FIGURES 65–69. Orthobula calceata Simon, 1897, genitalic morphology of female (65, 66) and male (67–69) from Bagoé: epigyne, ventral (65) and dorsal (66) views; palp, prolateral (67), ventral (68) and retrolateral (69) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522715" httpUri="https://zenodo.org/record/6522715/files/figure.png" pageId="6" pageNumber="361">68</figureCitation>
), but can be distinguished by the shape of the RFA, which is longer and bifid, with a strongly pronounced distal tooth in
<taxonomicName id="E18D4D44B52AFF4957C99215CA7EF89F" authority="Haddad &amp; Jin &amp; Platnick, 2022" authorityName="Haddad &amp; Jin &amp; Platnick" authorityYear="2022" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="14F9EAD5B52AFF4957C99215CA7EF89F" italics="true" pageId="6" pageNumber="361">O. aethiopica</emphasis>
</taxonomicName>
<emphasis id="14F9EAD5B52AFF49535C9270CA1DF89F" bold="true" box="[276,366,1884,1910]" pageId="6" pageNumber="361">
<taxonomicNameLabel id="0FCA57AEB52AFF49535C9270CA1DF89F" box="[276,366,1884,1910]" pageId="6" pageNumber="361" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="BEB62A42B52AFF4953359270CAA2F89E" box="[381,465,1884,1911]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="6" pageNumber="361">Fig. 58</figureCitation>
), while shorter and hook-shaped in
<taxonomicName id="E18D4D44B52AFF4951299271C891F89F" authorityName="Simon" authorityYear="1897" box="[865,994,1884,1911]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="361" phylum="Arthropoda" rank="species" species="calceata">
<emphasis id="14F9EAD5B52AFF4951299271C891F89F" box="[865,994,1884,1911]" italics="true" pageId="6" pageNumber="361">O. calceata</emphasis>
</taxonomicName>
(
<figureCitation id="BEB62A42B52AFF4951B99270CF35F89E" box="[1009,1094,1884,1911]" captionStart="FIGURES 65–69" captionStartId="14.[151,264,1064,1088]" captionTargetBox="[168,1424,191,1036]" captionTargetId="figure-26@14.[161,1427,181,1040]" captionTargetPageId="14" captionText="FIGURES 65–69. Orthobula calceata Simon, 1897, genitalic morphology of female (65, 66) and male (67–69) from Bagoé: epigyne, ventral (65) and dorsal (66) views; palp, prolateral (67), ventral (68) and retrolateral (69) views. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.6522715" httpUri="https://zenodo.org/record/6522715/files/figure.png" pageId="6" pageNumber="361">Fig. 69</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="72F2664FB52BFF4852DF912EC84FFB6F" ID-DOI="http://doi.org/10.5281/zenodo.6522697" ID-Zenodo-Dep="6522697" httpUri="https://zenodo.org/record/6522697/files/figure.png" pageId="7" pageNumber="362" startId="7.[151,264,1026,1050]" targetBox="[151,1436,151,1008]" targetPageId="7">
<paragraph id="263236C7B52BFF4852DF912EC84FFB6F" blockId="7.[151,1436,1026,1158]" pageId="7" pageNumber="362">
<emphasis id="14F9EAD5B52BFF4852DF912ECA27FBF3" bold="true" box="[151,340,1026,1050]" pageId="7" pageNumber="362">FIGURES 40–44.</emphasis>
Scanning electron microscope photographs of
<taxonomicName id="E18D4D44B52BFF485178912ECF00FBF3" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[816,1139,1026,1050]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="362" phylum="Arthropoda" rank="species" species="radiata">
<emphasis id="14F9EAD5B52BFF485178912EC898FBF3" box="[816,1003,1026,1050]" italics="true" pageId="7" pageNumber="362">Orthobula radiata</emphasis>
Simon, 1897
</taxonomicName>
female (40, 42–44) and male (41): 40, 41. Spinnerets, general view; 42. Anterior lateral spinnerets; 43. Posterior median spinnerets; 44. Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—minor ampullate gland spigot; MAmp—major ampullate gland spigot; Pi—piriform gland spigot.
</paragraph>
</caption>
<subSubSection id="6E97654CB52BFF47528F91F5CE14F872" lastPageId="8" lastPageNumber="363" pageId="7" pageNumber="362" type="etymology">
<paragraph id="263236C7B52BFF48528F91F5CF43FB1B" blockId="7.[151,1437,1240,1915]" box="[199,1072,1240,1267]" pageId="7" pageNumber="362">
<emphasis id="14F9EAD5B52BFF48528F91F5CA3DFB1A" bold="true" box="[199,334,1241,1267]" pageId="7" pageNumber="362">Etymology.</emphasis>
The species name is derived from the country of origin,
<collectingCountry id="5E9A7657B52BFF48518191F4CF5FFB1B" box="[969,1068,1240,1266]" name="Ethiopia" pageId="7" pageNumber="362">Ethiopia</collectingCountry>
.
</paragraph>
<paragraph id="263236C7B52BFF48528F91D1CAA0FA6A" blockId="7.[151,1437,1240,1915]" pageId="7" pageNumber="362">
<emphasis id="14F9EAD5B52BFF48528F91D1C9C9FAFF" bold="true" box="[199,698,1276,1303]" pageId="7" pageNumber="362">
Female (
<typeStatus id="F9368865B52BFF48536791D0CAEAFAFF" box="[303,409,1276,1302]" pageId="7" pageNumber="362" type="holotype">holotype</typeStatus>
,
<collectingRegion id="E449F825B52BFF4853EE91D0C937FAFF" box="[422,580,1276,1302]" country="Ethiopia" name="Adis Abeba" pageId="7" pageNumber="362">Addis Ababa</collectingRegion>
, BMNH).
</emphasis>
Measurements: CL 0.98, CW 0.74, AL 1.03, AW 0.73, TL 1.95, PERW 0.33, MOQAW 0.13, MOQPW 0.15, MOQL 0.14. Length of leg segments: I 0.62 + 0.25 + 0.54 + 0.48 + 0.29 = 2.18; II 0.41 + 0.24 + 0.45 + 0.40 + 0.28 = 1.78; III 0.46 + 0.22 + 0.33 + 0.38 + 0.27 = 1.66; IV 0.62 + 0.23 + 0.54 + 0.57 + 0.31 = 2.27.
</paragraph>
<paragraph id="263236C7B52BFF48528F90A0CB90F996" blockId="7.[151,1437,1240,1915]" pageId="7" pageNumber="362">Colour: carapace dark brown, with reticulated black mottled pattern; chelicerae dark brown; endites and labium slightly paler brown than chelicerae, cream distally; sternum dark brown, with black mottled margins and faint mottling centrally; palps dark yellow-brown; legs with femora, patellae and tibiae orange-brown, with dense black mottling laterally; distal ends of femora and tibiae, and basal end of patellae, with yellow rings; metatarsi and tarsi yellow, metatarsi with faint black mottling; abdomen black dorsally, with pair of cream anterolateral patches and transverse cream band at midpoint; venter cream, with small mottled grey patch in front of spinnerets; spinnerets cream.</paragraph>
<paragraph id="263236C7B52BFF48528F93A5C933F92E" blockId="7.[151,1437,1240,1915]" pageId="7" pageNumber="362">Leg spination: femora and patellae: spineless; tibiae: I plv 5 rlv 4, II plv 4 rlv 4; metatarsi: I plv 4 rlv 4, II plv 4 rlv 4; tarsi: I plv 3 rlv 3, II plv 3 rlv 3.</paragraph>
<paragraph id="263236C7B52BFF48528F93FCC988F892" blockId="7.[151,1437,1240,1915]" pageId="7" pageNumber="362">
Epigyne with lateral CO in broad semi-circular anterior epigynal ridge (
<figureCitation id="BEB62A42B52BFF48564393FCCF10F902" box="[1035,1123,1744,1771]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="7" pageNumber="362">Fig. 54</figureCitation>
); CD initially curving posteromesally, then bending sharply and diverging slightly before passing round SBB at centre of epigyne; SBB separated by approximately half their width; CD approximately twice as long as width of SBB; BU large and oval, with stalk-like accessory gland on their anteromesal surface; SBB connected to ST I by short looping ducts, entering transversely oval ST I on their mesal surface (
<figureCitation id="BEB62A42B52BFF4850D2924CC99DF892" box="[666,750,1888,1915]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="7" pageNumber="362">Fig. 55</figureCitation>
).
</paragraph>
<caption id="72F2664FB524FF4752DF919ECEE7FADF" ID-DOI="http://doi.org/10.5281/zenodo.6522701" ID-Zenodo-Dep="6522701" httpUri="https://zenodo.org/record/6522701/files/figure.png" pageId="8" pageNumber="363" startId="8.[151,264,1202,1226]" targetBox="[151,1436,151,1184]" targetPageId="8">
<paragraph id="263236C7B524FF4752DF919ECEE7FADF" blockId="8.[151,1437,1202,1334]" pageId="8" pageNumber="363">
<emphasis id="14F9EAD5B524FF4752DF919ECA20FB23" bold="true" box="[151,339,1202,1226]" pageId="8" pageNumber="363">FIGURES 45–53.</emphasis>
Scanning electron microscope photographs of
<taxonomicName id="E18D4D44B524FF475160919ECF14FB23" authority="Simon, 1897" authorityName="Simon" authorityYear="1897" box="[808,1127,1202,1226]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="363" phylum="Arthropoda" rank="species" species="radiata">
<emphasis id="14F9EAD5B524FF475160919EC891FB23" box="[808,994,1202,1226]" italics="true" pageId="8" pageNumber="363">Orthobula radiata</emphasis>
Simon, 1897
</taxonomicName>
female (45) and male (46–53): 45. Epigyne, ventral view; 46. Left palpal femur, retrolateral view; 47. Same, detail of ventrodistal apophysis; 48. Right palpal patella, tibia and tarsus, ventrolateral view; 49. Same, detail of patellar lyriform organ; 50. Same, detail of pores in lyriform organ; 51. Left palpal tibia, dorsolateral view; 52. Left embolus, retrolateral view; 53. Right embolus, retrolateral view, detail.
</paragraph>
</caption>
<paragraph id="263236C7B524FF47528F90A4CEE6FA03" blockId="8.[151,1437,1416,2019]" pageId="8" pageNumber="363">
<emphasis id="14F9EAD5B524FF47528F90A4C9EDFA4B" bold="true" box="[199,670,1416,1442]" pageId="8" pageNumber="363">
Male (
<typeStatus id="F9368865B524FF47535B90A4CAF1FA4B" box="[275,386,1416,1442]" pageId="8" pageNumber="363" type="paratype">paratype</typeStatus>
,
<collectingRegion id="E449F825B524FF4753C590A4C959FA4B" box="[397,554,1416,1442]" country="Ethiopia" name="Adis Abeba" pageId="8" pageNumber="363">Addis Ababa</collectingRegion>
, BMNH).
</emphasis>
Measurements: CL 0.86, CW 0.68, AL 0.87, AW 0.60, TL 1.72, PERW 0.28, MOQAW 0.11, MOQPW 0.16, MOQL 0.14. Length of leg segments: I 0.55 + 0.22 + 0.48 + 0.40 + 0.26 = 1.91; II 0.48 + 0.21 + 0.36 + 0.35 + 0.26 = 1.66; III 0.40 + 0.19 + 0.29 + 0.33 + 0.21 = 1.42; IV 0.49 + 0.22 + 0.42 + 0.51 + 0.27 = 1.91.
</paragraph>
<paragraph id="263236C7B524FF47528F90D8C828F90E" blockId="8.[151,1437,1416,2019]" pageId="8" pageNumber="363">Colour: carapace deep orange-brown, with reticulated black mottled pattern; chelicerae orange-brown; endites and labium slightly paler than chelicerae, cream distally; sternum orange-brown, with black mottled margins and mottling centrally; palps deep yellow-brown; anterior legs slightly darker than posteriors; legs with femora, patellae and tibiae yellow-brown, with black mottling laterally; distal ends of femora and tibiae, and basal end of patellae, with yellow rings; metatarsi and tarsi yellow; abdomen black dorsally, with pair of cream anterolateral patches and pair of transverse cream bands at midpoint, separated medially; venter cream, with large grey patch in front of spinnerets and black mottling on epigastric plate; spinnerets cream.</paragraph>
<paragraph id="263236C7B524FF47528F93DDC933F8C7" blockId="8.[151,1437,1416,2019]" pageId="8" pageNumber="363">Leg spination: femora and patellae: spineless; tibiae: I plv 5 rlv 4, II plv 4 rlv 4; metatarsi: I plv 4 rlv 4, II plv 4 rlv 4; tarsi: I plv 3 rlv 3, II plv 3 rlv 3.</paragraph>
<paragraph id="263236C7B524FF47528F9214CE14F872" blockId="8.[151,1437,1416,2019]" pageId="8" pageNumber="363">
Palpal femur with small bifid RFA, proximal branch sharply hook-shaped, distal branch tooth-like (
<figureCitation id="BEB62A42B524FF4757479214CEFDF8BA" box="[1295,1422,1848,1875]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="8" pageNumber="363">Figs 56, 58</figureCitation>
); tibia with small, simple triangular apophysis; retrodistal tegular ridge strongly developed, curving around ventrally to base of embolus (
<figureCitation id="BEB62A42B524FF47533192ACCABDF872" box="[377,462,1920,1947]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="8" pageNumber="363">Fig. 57</figureCitation>
); embolus narrow, undulating slightly, with weak curve at half its length (
<figureCitation id="BEB62A42B524FF47574F92ACCE28F872" box="[1287,1371,1920,1947]" captionStart="FIGURES 54–58" captionStartId="9.[151,264,1124,1148]" captionTargetBox="[159,1428,158,1096]" captionTargetId="figure-316@9.[151,1436,151,1103]" captionTargetPageId="9" captionText="FIGURES 54–58. Orthobula aethiopica sp. nov., genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae." figureDoi="http://doi.org/10.5281/zenodo.6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="8" pageNumber="363">Fig. 57</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6E97654CB524FF46528F9288CE3FFA24" lastPageId="9" lastPageNumber="364" pageId="8" pageNumber="363" type="materials_examined">
<paragraph id="263236C7B524FF47528F9288CF41F80B" blockId="8.[151,1437,1416,2019]" pageId="8" pageNumber="363">
<emphasis id="14F9EAD5B524FF47528F9288CA04F857" bold="true" box="[199,375,1956,1982]" pageId="8" pageNumber="363">Type material.</emphasis>
<materialsCitation id="96E53C9AB524FF4753379288CF41F80B" ID-GBIF-Occurrence="3763890392" collectingDate="1988-06-18" collectionCode="BMNH" collectorName="A. Russell-Smith" country="Ethiopia" latitude="9.0" location="Oromiya" longLatPrecision="1300" longitude="39.333332" pageId="8" pageNumber="363" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Oromiya" typeStatus="holotype">
<typeStatus id="F9368865B524FF4753379288CA94F857" box="[383,487,1956,1982]" pageId="8" pageNumber="363" type="holotype">Holotype</typeStatus>
<specimenCount id="308BFD4EB524FF4753B89289C976F856" box="[496,517,1957,1983]" pageId="8" pageNumber="363" type="female">♀</specimenCount>
and
<typeStatus id="F9368865B524FF4750779289C9ECF856" box="[575,671,1957,1983]" pageId="8" pageNumber="363" type="paratype">paratype</typeStatus>
<specimenCount id="308BFD4EB524FF4750E09289C9B3F856" box="[680,704,1957,1983]" pageId="8" pageNumber="363" type="male">♂</specimenCount>
:
<emphasis id="14F9EAD5B524FF4750859288C811F857" bold="true" box="[717,866,1956,1982]" pageId="8" pageNumber="363">
<collectingCountry id="5E9A7657B524FF4750859288C82EF857" box="[717,861,1956,1982]" name="Ethiopia" pageId="8" pageNumber="363">ETHIOPIA</collectingCountry>
:
</emphasis>
<emphasis id="14F9EAD5B524FF4751239289C8BDF856" box="[875,974,1957,1983]" italics="true" pageId="8" pageNumber="363">
<collectingRegion id="E449F825B524FF4751239289C8BDF856" box="[875,974,1957,1983]" country="Ethiopia" name="Oromiya" pageId="8" pageNumber="363">Oromiya</collectingRegion>
</emphasis>
:
<quantity id="E1759B22B524FF4751969288CF55F857" box="[990,1062,1956,1983]" metricMagnitude="4" metricUnit="m" metricValue="5.1" pageId="8" pageNumber="363" unit="km" value="51.0">51 km</quantity>
E of
<collectingRegion id="E449F825B524FF47562F9288CF89F857" box="[1127,1274,1956,1982]" country="Ethiopia" name="Adis Abeba" pageId="8" pageNumber="363">Addis Ababa</collectingRegion>
[ca.
<geoCoordinate id="43B95000B524FF47577D9288CEEBF857" box="[1333,1432,1956,1982]" degrees="09" direction="north" minutes="00" orientation="latitude" pageId="8" pageNumber="363" precision="925" value="9.0">09°00’N</geoCoordinate>
,
<geoCoordinate id="43B95000B524FF4752DF92E4CB8AF80A" box="[151,249,1992,2019]" degrees="39" direction="east" minutes="20" orientation="longitude" pageId="8" pageNumber="363" precision="925" value="39.333332">39°20’E</geoCoordinate>
],
<date id="52331007B524FF47534492E4CAFDF80B" box="[268,398,1992,2018]" pageId="8" pageNumber="363" value="1988-06-18">
<collectingDate id="4277E9EFB524FF47534492E4CAFDF80B" box="[268,398,1992,2018]" pageId="8" pageNumber="363" value="1988-06-18">18.VI.1988</collectingDate>
</date>
, leg.
<collectorName id="8B785311B524FF47538092E4C9F9F80B" box="[456,650,1992,2018]" pageId="8" pageNumber="363">A. Russell-Smith</collectorName>
(under stones, dry hillside) (
<collectionCode id="409CAE02B524FF47518392E4CF56F80B" box="[971,1061,1992,2018]" pageId="8" pageNumber="363">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<caption id="72F2664FB525FF4652DF9148C9FEFAE5" ID-DOI="http://doi.org/10.5281/zenodo.6522707" ID-Zenodo-Dep="6522707" httpUri="https://zenodo.org/record/6522707/files/figure.png" pageId="9" pageNumber="364" startId="9.[151,264,1124,1148]" targetBox="[159,1428,158,1096]" targetPageId="9">
<paragraph id="263236C7B525FF4652DF9148C9FEFAE5" blockId="9.[151,1437,1124,1293]" pageId="9" pageNumber="364">
<emphasis id="14F9EAD5B525FF4652DF9148CA27FB95" bold="true" box="[151,340,1124,1148]" pageId="9" pageNumber="364">FIGURES 54–58.</emphasis>
<taxonomicName id="E18D4D44B525FF4653119148C940FB95" authority="Haddad &amp; Jin &amp; Platnick, 2022" authorityName="Haddad &amp; Jin &amp; Platnick" authorityYear="2022" box="[345,563,1124,1148]" class="Arachnida" family="Trachelidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="aethiopica" status="sp. nov.">
<emphasis id="14F9EAD5B525FF4653119148C940FB95" box="[345,563,1124,1148]" italics="true" pageId="9" pageNumber="364">Orthobula aethiopica</emphasis>
</taxonomicName>
<emphasis id="14F9EAD5B525FF4650719149C9FAFB94" bold="true" box="[569,649,1125,1149]" pageId="9" pageNumber="364">
<taxonomicNameLabel id="0FCA57AEB525FF4650719149C9FAFB94" box="[569,649,1125,1149]" pageId="9" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, genitalic morphology of holotype female (54, 55) and paratype male (56–58) from Addis Ababa: epigyne, ventral (54) and dorsal (55) views; palp, prolateral (56), ventral (57) and retrolateral (58) views. Scale bars: 0.10 mm. Abbreviations: AG—accessory gland; BU—bursa; CD—copulatory duct; CO—copulatory opening; EM—embolus; FD—fertilisation duct; RFA—retrodistal femoral apophysis; RTA—retrolateral tibial apophysis; SBB—sclerotised base of bursa; ST I—primary spermathecae.
</paragraph>
</caption>
<paragraph id="263236C7B525FF46528F906BC9D8FA40" blockId="9.[151,1437,1350,1521]" pageId="9" pageNumber="364">
<materialsCitation id="96E53C9AB525FF46528F906BCAF7FA6C" ID-GBIF-Occurrence="3763890418" collectingDate="1973-12" collectionCode="MRAC" collectorName="G. de Rougemont" country="Ethiopia" latitude="12.0" location="Lake Tana" longLatPrecision="1294" longitude="37.333332" municipality="Bahir Dar" pageId="9" pageNumber="364" specimenCode="MRAC 158945" specimenCount="1" specimenCount-male="1" stateProvince="Amara" typeStatus="paratype">
<typeStatus id="F9368865B525FF46528F906BCA4BFA88" box="[199,312,1351,1377]" pageId="9" pageNumber="364" type="paratype">Paratypes</typeStatus>
:
<emphasis id="14F9EAD5B525FF46530C906ACAAAFA89" bold="true" box="[324,473,1350,1376]" pageId="9" pageNumber="364">
<collectingCountry id="5E9A7657B525FF46530C906ACAA7FA89" box="[324,468,1350,1376]" name="Ethiopia" pageId="9" pageNumber="364">ETHIOPIA</collectingCountry>
:
</emphasis>
<emphasis id="14F9EAD5B525FF4653A9906BC948FA88" box="[481,571,1351,1377]" italics="true" pageId="9" pageNumber="364">
<collectingRegion id="E449F825B525FF4653A9906BC948FA88" box="[481,571,1351,1377]" country="Ethiopia" name="Amara" pageId="9" pageNumber="364">Amhara</collectingRegion>
</emphasis>
:
<collectingMunicipality id="C656ACBDB525FF465002906BC9CFFA88" box="[586,700,1351,1377]" pageId="9" pageNumber="364">Bahir Dar</collectingMunicipality>
,
<location id="2352601CB525FF465080906BC833FA88" LSID="urn:lsid:plazi:treatment:AE2487D1B52AFF4652DF9310C82AFA18:2352601CB525FF465080906BC833FA88" box="[712,832,1351,1377]" country="Ethiopia" latitude="12.0" longLatPrecision="1294" longitude="37.333332" municipality="Bahir Dar" name="Lake Tana" pageId="9" pageNumber="364" stateProvince="Amara">Lake Tana</location>
,
<geoCoordinate id="43B95000B525FF465104906BC8DCFA88" box="[844,943,1351,1377]" degrees="12" direction="north" minutes="00" orientation="latitude" pageId="9" pageNumber="364" precision="925" value="12.0">12°00’N</geoCoordinate>
,
<geoCoordinate id="43B95000B525FF4651F3906BCF6FFA88" box="[955,1052,1351,1377]" degrees="37" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="364" precision="925" value="37.333332">37°20’E</geoCoordinate>
,
<date id="52331007B525FF465660906BCFE3FA88" box="[1064,1168,1351,1377]" pageId="9" pageNumber="364" value="1973-12">
<collectingDate id="4277E9EFB525FF465660906BCFE3FA88" box="[1064,1168,1351,1377]" pageId="9" pageNumber="364" value="1973-12">XII.1973</collectingDate>
</date>
, leg.
<collectorName id="8B785311B525FF465685906BCEEAFA88" box="[1229,1433,1351,1377]" pageId="9" pageNumber="364">G. de Rougemont</collectorName>
,
<specimenCount id="308BFD4EB525FF4652DF9047CBC9FA6C" box="[151,186,1387,1413]" pageId="9" pageNumber="364" type="male">1♂</specimenCount>
(
<specimenCode id="762B9EBCB525FF4652809047CA0FFA6C" box="[200,380,1386,1413]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="9" pageNumber="364" type="Museum">MRAC 158945</specimenCode>
)
</materialsCitation>
;
<emphasis id="14F9EAD5B525FF4653C79047CA81FA6C" box="[399,498,1387,1413]" italics="true" pageId="9" pageNumber="364">
<collectingRegion id="E449F825B525FF4653C79047CA81FA6C" box="[399,498,1387,1413]" country="Ethiopia" name="Oromiya" pageId="9" pageNumber="364">Oromiya</collectingRegion>
</emphasis>
:
<quantity id="E1759B22B525FF4650489047C934FA6C" box="[512,583,1387,1413]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="9" pageNumber="364" unit="km" value="17.0">17 km</quantity>
E of
<materialsCitation id="96E53C9AB525FF4650CA9047C9D4FA40" ID-GBIF-Occurrence="3763890489" collectingDate="1988-05-28" collectionCode="BMNH" collectorName="A. Russell-Smith" country="Ethiopia" latitude="9.083333" location="Addis Ababa" longLatPrecision="1300" longitude="39.083332" pageId="9" pageNumber="364" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Adis Abeba" typeStatus="paratype">
<collectingRegion id="E449F825B525FF4650CA9047C860FA6C" box="[642,787,1387,1413]" country="Ethiopia" name="Adis Abeba" pageId="9" pageNumber="364">Addis Ababa</collectingRegion>
[ca.
<geoCoordinate id="43B95000B525FF4651029046C8DFFA6C" box="[842,940,1386,1413]" degrees="09" direction="north" minutes="05" orientation="latitude" pageId="9" pageNumber="364" precision="925" value="9.083333">09°05’N</geoCoordinate>
,
<geoCoordinate id="43B95000B525FF4651F09046CF69FA6C" box="[952,1050,1386,1413]" degrees="39" direction="east" minutes="05" orientation="longitude" pageId="9" pageNumber="364" precision="925" value="39.083332">39°05’E</geoCoordinate>
],
<date id="52331007B525FF4656659047CFD1FA6C" box="[1069,1186,1387,1413]" pageId="9" pageNumber="364" value="1988-05-28">
<collectingDate id="4277E9EFB525FF4656659047CFD1FA6C" box="[1069,1186,1387,1413]" pageId="9" pageNumber="364" value="1988-05-28">28.V.1988</collectingDate>
</date>
, leg.
<collectorName id="8B785311B525FF4656949047CEEEFA6C" box="[1244,1437,1387,1413]" pageId="9" pageNumber="364">A. Russell-Smith</collectorName>
(under stones, stream valley),
<specimenCount id="308BFD4EB525FF4653A190A3C97FFA40" box="[489,524,1423,1449]" pageId="9" pageNumber="364" type="male">1♂</specimenCount>
<specimenCount id="308BFD4EB525FF46505B90A3C946FA40" box="[531,565,1423,1449]" pageId="9" pageNumber="364" type="female">2♀</specimenCount>
(
<collectionCode id="409CAE02B525FF46500D90A3C9ECFA40" box="[581,671,1423,1449]" pageId="9" pageNumber="364">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="263236C7B525FF46528F909ECE3FFA24" blockId="9.[151,1437,1350,1521]" box="[199,1356,1458,1485]" pageId="9" pageNumber="364">
<emphasis id="14F9EAD5B525FF46528F909ECAC8FA25" bold="true" box="[199,443,1458,1485]" pageId="9" pageNumber="364">Habitat and biology.</emphasis>
A ground-dwelling species sampled from under rocks in grassy savanna habitats.
</paragraph>
</subSubSection>
<subSubSection id="6E97654CB525FF46528F90FAC82AFA18" box="[199,857,1494,1521]" pageId="9" pageNumber="364" type="distribution">
<paragraph id="263236C7B525FF46528F90FAC82AFA18" blockId="9.[151,1437,1350,1521]" box="[199,857,1494,1521]" pageId="9" pageNumber="364">
<emphasis id="14F9EAD5B525FF46528F90FACA12FA19" bold="true" box="[199,353,1494,1520]" pageId="9" pageNumber="364">Distribution.</emphasis>
Only known from central
<collectingCountry id="5E9A7657B525FF4650C290FBC99AFA18" box="[650,745,1495,1521]" name="Ethiopia" pageId="9" pageNumber="364">Ethiopia</collectingCountry>
(
<figureCitation id="BEB62A42B525FF4650B090FBC83EFA18" box="[760,845,1495,1521]" captionStart="FIGURE 82" captionStartId="17.[151,250,1267,1291]" captionTargetBox="[179,1408,237,1243]" captionTargetId="figure-48@17.[179,1408,237,1243]" captionTargetPageId="17" captionText="FIGURE 82. Distribution of Orthobula aethiopica sp. nov. (circles), O. arca sp. nov. (squares) and O. calceata Simon, 1897 (triangles) in the continental Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.6522719" httpUri="https://zenodo.org/record/6522719/files/figure.png" pageId="9" pageNumber="364">Fig. 82</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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