treatments-xml/data/3E/7A/87/3E7A87DDD973165AFEB5C108FE1DFBAC.xml

<document id="77291C2920C3AD8086755B88F0C432F6" ID-DOI="10.35249/rche.47.2.21.19" ID-ISSN="0718-8994" ID-Zenodo-Dep="13205388" ID-ZooBank="14DEE684-1721-43B7-85C3-2A57525CE1A8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722661749589" checkinUser="felipe" docAuthor="González, Guillermo &amp; Větrovec, Jaroslav" docDate="2021" docId="3E7A87DDD973165AFEB5C108FE1DFBAC" docLanguage="en" docName="RevChilEntomol.47.2.331-374.pdf" docOrigin="Revista Chilena de Entomología (Rev. Chil. Entomol.) 47 (2)" docSource="http://dx.doi.org/10.35249/rche.47.2.21.19" docStyle="DocumentStyle:451321A1F3358405F3D5797D4FC35865.3:RevChilEntomol.2020-.journal_article" docStyleId="451321A1F3358405F3D5797D4FC35865" docStyleName="RevChilEntomol.2020-.journal_article" docStyleVersion="3" docTitle="Toxotoma aguascalientes Gonzalez &amp; Vetrovec 2021, new species" docType="treatment" docUuid="FF8D2755-0159-43FA-B6D6-9C49064BA07F" docUuidSource="ZooBank" docVersion="2" lastPageNumber="365" masterDocId="C243FFA5D9521678FFA1C175FFFFFF98" masterDocTitle="New species and records of Neotropical ladybirds (Coleoptera: Coccinellidae)" masterLastPageNumber="374" masterPageNumber="331" pageNumber="364" updateTime="1722701365769" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="9F01E3ECC4EA8BE9C76AB9A4496002EF">New species and records of Neotropical ladybirds (Coleoptera: Coccinellidae)</mods:title>
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<mods:namePart id="1A8410649C0E90D36CB796B1B2E0DBAA">González, Guillermo</mods:namePart>
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<paragraph id="B66C36CBD9731659FEB5C108FC3BFF0C" blockId="33.[276,964,124,181]" box="[276,964,124,149]" pageId="33" pageNumber="364">
<heading id="ED2481A7D9731659FEB5C108FC3BFF0C" box="[276,964,124,149]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="364" reason="4">
<taxonomicName id="71D34D48D9731659FEB5C108FCD9FF0D" ID-CoL="CGJ3Q" authority="Gonzalez &amp; Vetrovec" authorityName="Gonzalez &amp; Vetrovec" authorityYear="2021" box="[276,806,125,149]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="aguascalientes" status="sp. nov.">
<emphasis id="84A7EAD9D9731659FEB5C108FDE0FF0D" box="[276,543,125,149]" italics="true" pageId="33" pageNumber="364">Toxotoma aguascalientes</emphasis>
González &amp; Větrovec
</taxonomicName>
, 
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<taxonomicNameLabel id="9F9457A2D9731659FC90C108FC3BFF0C" box="[817,964,124,149]" pageId="33" pageNumber="364" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="FEC96540D9731659FD82C1E8FD4AFF2D" box="[547,693,157,181]" pageId="33" pageNumber="364" type="description">
<paragraph id="B66C36CBD9731659FD82C1E8FD4AFF2D" blockId="33.[276,964,124,181]" box="[547,693,157,181]" pageId="33" pageNumber="364">
(
<figureCitation id="2EE82A4ED9731659FD8DC1E8FD87FF2D" box="[556,632,157,181]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Figs. 8</figureCitation>
a-8o)
</paragraph>
</subSubSection>
<subSubSection id="FEC96540D9731659FFFAC1A8FC2BFEAD" pageId="33" pageNumber="364" type="materials_examined">
<paragraph id="B66C36CBD9731659FFFAC1A8FC2BFEAD" blockId="33.[89,1149,221,309]" pageId="33" pageNumber="364">
<materialsCitation id="06BB3C96D9731659FFFAC1A8FF07FE8D" collectingDate="2009-01-18" collectionCode="NMP" collectorName="M. Krejcir" pageId="33" pageNumber="364" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="84A7EAD9D9731659FFFAC1A8FF31FF6D" bold="true" box="[91,206,221,245]" pageId="33" pageNumber="364">
<typeStatus id="69688869D9731659FFFAC1A8FF31FF6D" box="[91,206,221,245]" pageId="33" pageNumber="364" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="A0D5FD42D9731659FF79C1A8FF13FF6C" box="[216,236,221,244]" count="1" pageId="33" pageNumber="364" type="male">♂</specimenCount>
“Perú-Cusco / Agnas [Aguas] Calientes / 
<date id="C26D100BD9731659FCA6C1A8FC8CFF6D" box="[775,883,221,245]" pageId="33" pageNumber="364" value="2009-01-18">
<collectingDate id="D229E9E3D9731659FCA6C1A8FC8CFF6D" box="[775,883,221,245]" pageId="33" pageNumber="364" value="2009-01-18">18.1.2009</collectingDate>
</date>
/ lgt. 
<collectorName id="1B26531DD9731659FC68C1A8FBB8FF6D" box="[969,1095,221,245]" pageId="33" pageNumber="364">M. Krejčíř</collectorName>
” “ 
<specimenCount id="A0D5FD42D9731659FBC8C1A8FB82FF6C" box="[1129,1149,221,244]" count="1" pageId="33" pageNumber="364" type="male">♂</specimenCount>
1954” (
<collectionCode id="D0C2AE0ED9731659FF0DC188FF0EFE8D" box="[172,241,253,277]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="33" pageNumber="364">NMP</collectionCode>
)
</materialsCitation>
. 
<emphasis id="84A7EAD9D9731659FEA1C188FE87FE8D" bold="true" box="[256,376,253,277]" pageId="33" pageNumber="364">
<typeStatus id="69688869D9731659FEA1C188FE87FE8D" box="[256,376,253,277]" pageId="33" pageNumber="364" type="paratype">Paratypes</typeStatus>
</emphasis>
(total 
<specimenCount id="A0D5FD42D9731659FE62C188FDA6FE8D" box="[451,601,253,277]" count="5" pageId="33" pageNumber="364" type="generic">5 specimens</specimenCount>
.): 
<materialsCitation id="06BB3C96D9731659FDCFC188FF52FEAD" collectingDate="2009-01-18" collectionCode="NMP" collectorName="M. Krejcir" pageId="33" pageNumber="364" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="A0D5FD42D9731659FDCFC188FD6CFE8C" box="[622,659,253,277]" count="2" pageId="33" pageNumber="364" type="female">2 ♀</specimenCount>
same data as holotype except “ 
<date id="C26D100BD9731659FBA1C188FB8CFE8D" box="[1024,1139,253,277]" pageId="33" pageNumber="364" value="2009-01-18">
<collectingDate id="D229E9E3D9731659FBA1C188FB8CFE8D" box="[1024,1139,253,277]" pageId="33" pageNumber="364" value="2009-01-18">18.1.2009</collectingDate>
</date>
” (
<collectionCode id="D0C2AE0ED9731659FFC3C068FF59FEAD" box="[98,166,285,309]" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="33" pageNumber="364">NMP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="06BB3C96D9731659FF18C068FC2FFEAD" box="[185,976,285,309]" collectingDate="2009-12-01" collectionCode="JVC" collectorName="M. Krejcir" pageId="33" pageNumber="364" specimenCount="3" typeStatus="paratype">
<specimenCount id="A0D5FD42D9731659FF18C068FEB3FEAD" box="[185,332,285,309]" count="3" pageId="33" pageNumber="364" type="generic">3 specimens</specimenCount>
. same data as holotype except “ 
<date id="C26D100BD9731659FD65C068FCC8FEAD" box="[708,823,285,309]" pageId="33" pageNumber="364" value="2009-12-01">
<collectingDate id="D229E9E3D9731659FD65C068FCC8FEAD" box="[708,823,285,309]" pageId="33" pageNumber="364" value="2009-12-01">12.1.2009</collectingDate>
</date>
” (s/g) (
<collectionCode id="D0C2AE0ED9731659FC32C068FC37FEAD" box="[915,968,285,309]" country="Spain" lsid="urn:lsid:biocol.org:col:13378" name="Jardín Botánico Canario Viera y Clavijo" pageId="33" pageNumber="364" type="Herbarium">JVC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEC96540D9731659FFFAC029FDE8FD2D" pageId="33" pageNumber="364" type="diagnosis">
<paragraph id="B66C36CBD9731659FFFAC029FDE8FD2D" blockId="33.[89,1150,348,693]" pageId="33" pageNumber="364">
<emphasis id="84A7EAD9D9731659FFFAC029FF22FEEC" bold="true" box="[91,221,348,372]" pageId="33" pageNumber="364">Diagnosis.</emphasis>
This species can be distinguished by the male genitalia which presents a long penis guide exceeding the paramere and with a tuft of hairs in the middle of the lateral border, with a small hook at the apex (
<figureCitation id="2EE82A4ED9731659FDDCC0E8FD35FE2D" box="[637,714,413,437]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Figs. 8</figureCitation>
g-8h), the genital plate of the female with transverse coxites and the stylus not visible (
<figureCitation id="2EE82A4ED9731659FD11C0C8FCFDFE4D" box="[688,770,445,469]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Fig. 8o</figureCitation>
), the feebly bilobed labrum, the mandibles with the teeth grouped in the distal half, and a minor projection below the third mandible tooth, all characters which correspond to the 
<taxonomicName id="71D34D48D9731659FD44C088FC4BFD8C" baseAuthorityName="Crotch" baseAuthorityYear="1874" box="[741,948,509,533]" class="Insecta" family="Coccinellidae" genus="Epilachna" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="vittigera">
<emphasis id="84A7EAD9D9731659FD44C088FC4BFD8C" box="[741,948,509,533]" italics="true" pageId="33" pageNumber="364">Epilachna vittigera</emphasis>
</taxonomicName>
species group as defined by 
<bibRefCitation id="D2424B3AD9731659FF43C368FE6FFDAD" author="Gordon, R. D." box="[226,400,541,565]" pageId="33" pageNumber="364" pagination="1 - 409" refId="ref26293" refString="Gordon, R. D. (1975) A revision of Epilachninae of the Western Hemisphere (Coleoptera: Coccinellidae). US Department of Agriculture Technical Bulletin, 1493: 1 - 409." type="journal article" year="1975">Gordon (1975)</bibRefCitation>
. In this species group, the habitus with two elytral yellow and well separated spots, allows recognizing the new species, since other species in the group have two spots which cover most of the elytron, except for 
<taxonomicName id="71D34D48D9731659FCABC328FBC6FDED" authority="Gordon, 1975" authorityName="Gordon" authorityYear="1975" box="[778,1081,605,629]" class="Insecta" family="Coccinellidae" genus="Epilachna" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="84A7EAD9D9731659FCABC328FC6DFDEC" box="[778,914,605,628]" italics="true" pageId="33" pageNumber="364">E. conjuncta</emphasis>
<bibRefCitation id="D2424B3AD9731659FC38C328FBC6FDED" author="Gordon, R. D." box="[921,1081,605,629]" pageId="33" pageNumber="364" pagination="1 - 409" refId="ref26293" refString="Gordon, R. D. (1975) A revision of Epilachninae of the Western Hemisphere (Coleoptera: Coccinellidae). US Department of Agriculture Technical Bulletin, 1493: 1 - 409." type="journal article" year="1975">Gordon, 1975</bibRefCitation>
</taxonomicName>
, from 
<collectingCountry id="CEC4765BD9731659FFFBC308FF4FFD0D" box="[90,176,637,661]" name="Bolivia" pageId="33" pageNumber="364">Bolivia</collectingCountry>
, whose genitalia has a penis guide with an acute apex, while 
<taxonomicName id="71D34D48D9731659FCDEC308FBC9FD0D" authorityName="González &amp; Větrovec" authorityYear="2021" box="[895,1078,637,661]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="364" phylum="Arthropoda" rank="species" species="aguascalientes" status="n. sp.">
<emphasis id="84A7EAD9D9731659FCDEC308FBC9FD0D" box="[895,1078,637,661]" italics="true" pageId="33" pageNumber="364">T. aguascalientes</emphasis>
</taxonomicName>
<emphasis id="84A7EAD9D9731659FB9CC308FB82FD0D" bold="true" box="[1085,1149,637,661]" pageId="33" pageNumber="364">
<taxonomicNameLabel id="9F9457A2D9731659FB9CC308FB82FD0D" box="[1085,1149,637,661]" pageId="33" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has a penis guide with a rounded tip.
</paragraph>
</subSubSection>
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<emphasis id="84A7EAD9D9731659FFFAC3A9FE66FD6D" bold="true" box="[91,409,732,757]" pageId="33" pageNumber="364">Description. Color pattern</emphasis>
(
<figureCitation id="2EE82A4ED9731659FE05C3A8FE11FD6D" box="[420,494,733,757]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Figs. 8</figureCitation>
a-8c). Head black. Antennae and maxillary palpi yellow, four last antennomeres and last palpomere brown. Mouthparts reddish brown with black areas. Pronotum black, with a narrow yellow border at anterior angles. Scutellar shield black. Elytra black, each with two yellow oval spots (1:1) on disc, the first very close to the scutellar shield, occupying one-quarter of the length of the elytra and slightly shorter than half the width, the second about the same size but somewhat more transverse, close to lateral border, on the elytral declivity. Epipleuron black. Ventral side black, except anterior angles of hypomeron yellow. Legs black. Pubescence white. 
<emphasis id="84A7EAD9D9731659FC82C2C8FC3EFC4D" bold="true" box="[803,961,957,981]" pageId="33" pageNumber="364">Morphology.</emphasis>
Body oval with shield-shaped elytra, widest at 1/3 the length of the elytra, narrowing to the apex (
<figureCitation id="2EE82A4ED9731659FBEEC2A8FF85FB8D" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Fig. 8a</figureCitation>
). Frons more than three times width of an eye. Eyes oval, 1/3 longer than wide, inner border of the eye somewhat concave around the antennal insertion, without eye canthus. Clypeus apex concave with rounded corners (
<figureCitation id="2EE82A4ED9731659FDCDC548FD45FBCD" box="[620,698,1085,1109]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Fig. 8c</figureCitation>
). Antennae with eleven antennomeres, the last three forming a club. Labrum feebly bilobed. Apical maxillary palpomere weakly securiform. Mandibles with three major teeth grouped in apical one-third, first tooth bidentate, surfaces without tubercles. Prosternum Y-shaped, short and wide, without carinae. Metaventrite postcoxal lines transverse, slightly curved, not descending to lateral border. Abdominal postcoxal lines complete, evenly rounded, extended 2/3 of the distance to apical margin of ventrite (
<figureCitation id="2EE82A4ED9731659FDE6C588FD67FA8D" box="[583,664,1277,1301]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Fig. 8e</figureCitation>
). Head punctures notorious, disordered, space between the punctures about 1.5 times the diameter; pronotum punctures greater than those on head and thick, separated by a diameter on average; elytra punctures of 
<specimenCount id="A0D5FD42D9731659FFDCC428FF07FAED" box="[125,248,1373,1397]" count="2" pageId="33" pageNumber="364" type="generic" typeStatus="types">two types</specimenCount>
, the smaller similar to those of the pronotum, the largest twice as large, separated by half a diameter on average; ventral side punctures tiny, more abundant on the prosternum, separated by four times their diameter on the metaventrite; abdominal punctures notorious, smaller towards the borders and posterior ventrites. Pubescence at each puncture abundant, decumbent; pronotum hairs long, 3/4 the length of the scutellar shield, elytra pubescence short, 1/3 the size of the scutellar shield, abdominal hairs short and quite dense on the lateral borders. 
<emphasis id="84A7EAD9D9731659FD81C768FD15F9AC" bold="true" box="[544,746,1564,1589]" pageId="33" pageNumber="364">Male terminalia.</emphasis>
Ventrite 5 with a regularly convex apex. Ventrite 6 with semicircular apex, with a strong semicircular notch medially (
<figureCitation id="2EE82A4ED9731659FBE2C748FF96F9ED" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="33" pageNumber="364">Figs. 8</figureCitation>
e-8f). Tegmen three times as long as wide, phallobase as long as wide. Tegminal strut 2/3 as long as the rest of the tegmen. Penis guide symmetrical, four times as long as wide, sides slightly divergent in the basal two fifths, then slightly convergent until ending in a rounded wide apex, a tuft of erect, long hairs on each side at half of the length, in lateral view regularly curved towards the parameres and slightly thinning from the base to 3/4 of the length, then abruptly in the apical 1/10 where it presents a very small hook towards the outer side. Parameres 9/10 the length of the penis guide, slightly curved towards it, widened at the base and at the tip where they are paddle-shaped, with erect hairs on the borders of the distal half, which exceeds the length of the paramere by one quarter at the apex (
<figureCitation id="2EE82A4ED970165AFF41C068FED2FEAD" box="[224,301,285,309]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">Figs. 8</figureCitation>
g-8h). Penis curved in a semicircle in the basal 2/3, then slightly wavy up to the apex where it has a very small hook towards the outer side, where it also has a small membranous area; penis capsule double-walled, the outer arm oval, in the direction of the penis tube, short inner arm, somewhat trapezoidal with rounded tip, basal margin fairly emarginate, accessory piece present (
<figureCitation id="2EE82A4ED970165AFD00C0E8FD10FE2D" box="[673,751,413,437]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">Figs. 8</figureCitation>
i-j). 
<emphasis id="84A7EAD9D970165AFC81C0E8FBF8FE2C" bold="true" box="[800,1031,412,437]" pageId="34" pageNumber="365">Female terminalia.</emphasis>
Tergite 6 apex barely notched in the central 1/3 (
<figureCitation id="2EE82A4ED970165AFD94C0C8FD77FE4D" box="[565,648,445,469]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">Fig. 8k</figureCitation>
), ventrite 6 deeply notched in a triangle in the center (
<figureCitation id="2EE82A4ED970165AFF53C0A8FEC0FE6D" box="[242,319,477,501]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">Fig. 8l</figureCitation>
), tergite 10 trapezoidal, posterior border slightly convex (
<figureCitation id="2EE82A4ED970165AFC56C0A8FBABFE6D" box="[1015,1108,477,501]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">
Fig. 
<quantity id="712B9B2ED970165AFB8DC0A8FBABFE6D" box="[1068,1108,477,501]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="34" pageNumber="365" unit="m" value="8.0">8m</quantity>
</figureCitation>
), genital plate with slightly transverse oval coxites, somewhat concave posterior border, slightly rounded anterior border, very hairy, little apparent stylus, very reduced spermatheca (
<figureCitation id="2EE82A4ED970165AFF01C348FF12FDCD" box="[160,237,573,597]" captionStart="Figures 8" captionStartId="37.[91,170,1462,1484]" captionTargetBox="[143,1103,132,1449]" captionTargetId="figure-187@37.[131,1110,121,1459]" captionTargetPageId="37" captionText="Figures 8a-8ac. a-o: Toxotoma aguascalientes n. sp.; a-d: habitus male (dorsal, lateral, frontal, posterior); e: abdomen; f:ventrite 6; g-h: tegmen (ventral, lateral); i-j:penis and detail; k-o: female terminalia:k: tergite 6; l: ventrite 6; m: ventrite 10; n: genital plate; o: coxite. p-ac: Nexophallus panamensis n. sp.; p-s: habitus (dorsal, lateral, frontal, posterior); t: maxillary palp; u: abdomen; v: ventrite 6; w-x: tegmen; y: penis guide; z-aa: penis and detail; ab-ac: female terminalia; ab: coxitos; ac: spermatheca. Scale bars in mm. / a-o: Toxotoma aguascalientes sp. n.; a-d: habitus del macho (dorsal, lateral, frontal, posterior); e: abdomen; f: ventrito 6; g-h: tegmen (ventral, lateral); i-j: penis y detalle; k-o: terminalia de la hembra: k: tergito 6; l: ventrito 6; m: ventrito 10; n: placa genital; o: coxito. p-ac: Nexophallus panamensis sp. n.; p-s: habitus (dorsal, lateral, frontal, posterior); t: palpo maxilar; u: abdomen; v: ventrito 6; w-x: tegmen; y: guÍa del penis; z-aa: penis y detalle; ab-ac: terminalia de la hembra; ab: coxitos; ac: espermateca. Escalas en mm." figureDoi="http://doi.org/10.5281/zenodo.13205404" httpUri="https://zenodo.org/record/13205404/files/figure.png" pageId="34" pageNumber="365">Figs. 8</figureCitation>
n-8o).
</paragraph>
<paragraph id="B66C36CBD970165AFF36C308FB45FD0D" blockId="34.[151,1210,637,661]" box="[151,1210,637,661]" pageId="34" pageNumber="365">
<emphasis id="84A7EAD9D970165AFF36C308FEB3FD0D" bold="true" box="[151,332,637,661]" pageId="34" pageNumber="365">Measurements</emphasis>
(mm): TL 6.5-7.0; PL 1.0-1.1; PW 2.7-3.0; EL 5.4-5.7; EW 4.8-5.2; GD 2.8-3.0.
</paragraph>
<paragraph id="B66C36CBD970165AFF36C3C9FCE4FD4D" blockId="34.[151,795,700,725]" box="[151,795,700,725]" pageId="34" pageNumber="365">
<emphasis id="84A7EAD9D970165AFF36C3C9FE37FD4C" bold="true" box="[151,456,700,724]" pageId="34" pageNumber="365">Geographic distribution.</emphasis>
<collectingCountry id="CEC4765BD970165AFE6EC3C8FDF6FD4C" box="[463,521,701,724]" name="Peru" pageId="34" pageNumber="365">Peru</collectingCountry>
, Department of 
<collectingRegion id="7417F829D970165AFD6BC3C8FCE9FD4D" box="[714,790,701,725]" country="Peru" name="Cusco" pageId="34" pageNumber="365">Cusco</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FEC96540D970165AFF36C388FBC2FC4D" pageId="34" pageNumber="365" type="discussion">
<paragraph id="B66C36CBD970165AFF36C388FBC2FC4D" blockId="34.[151,1212,765,981]" pageId="34" pageNumber="365">
<emphasis id="84A7EAD9D970165AFF36C388FEF4FC8D" bold="true" box="[151,267,765,789]" pageId="34" pageNumber="365">Remarks.</emphasis>
For an analysis of the taxonomic status of the genera 
<taxonomicName id="71D34D48D970165AFC00C388FBF5FC8C" authorityName="Weise" authorityYear="1899" box="[929,1034,765,788]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="84A7EAD9D970165AFC00C388FBF5FC8C" box="[929,1034,765,788]" italics="true" pageId="34" pageNumber="365">Toxotoma</emphasis>
</taxonomicName>
and 
<taxonomicName id="71D34D48D970165AFBE7C388FB4DFC8D" authorityName="Chevrolat" authorityYear="1836" box="[1094,1202,765,789]" class="Insecta" family="Coccinellidae" genus="Epilachna" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="84A7EAD9D970165AFBE7C388FB4DFC8D" box="[1094,1202,765,789]" italics="true" pageId="34" pageNumber="365">Epilachna</emphasis>
</taxonomicName>
, see discussion under 
<taxonomicName id="71D34D48D970165AFE3FC268FD7BFCAD" authorityName="González &amp; Větrovec" authorityYear="2021" box="[414,644,797,821]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="venezuelae" status="sp. nov.">
<emphasis id="84A7EAD9D970165AFE3FC268FD7BFCAD" box="[414,644,797,821]" italics="true" pageId="34" pageNumber="365">Toxotoma venezuelae</emphasis>
</taxonomicName>
<emphasis id="84A7EAD9D970165AFD2AC268FD31FCAD" bold="true" box="[651,718,797,821]" pageId="34" pageNumber="365">
<taxonomicNameLabel id="9F9457A2D970165AFD2AC268FD31FCAD" box="[651,718,797,821]" pageId="34" pageNumber="365" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
The new species presents the characters indicated by 
<bibRefCitation id="D2424B3AD970165AFE9BC248FD2CFCCD" author="Tomaszewska, W. &amp; Szawaryn, K." box="[314,723,829,853]" pageId="34" pageNumber="365" pagination="1 - 91" refId="ref28231" refString="Tomaszewska, W. &amp; Szawaryn, K. (2016) Epilachnini (Coleoptera: Coccinellidae) - a revision of the world genera. Journal of Insect Science, 16 (1): 1 - 91. https: // doi. org / 10.1093 / jisesa / iew 082" type="journal article" year="2016">Tomaszewska &amp; Szawaryn (2016)</bibRefCitation>
for the genus, although the abdominal postcoxal lines are clearly visible and well formed, as in many 
<taxonomicName id="71D34D48D970165AFC68C228FB64FCED" box="[969,1179,861,885]" class="Insecta" family="Coccinellidae" genus="Toxotoma" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="84A7EAD9D970165AFC68C228FBCDFCEC" box="[969,1074,861,884]" italics="true" pageId="34" pageNumber="365">Toxotoma</emphasis>
species.
</taxonomicName>
It should be noted that 
<taxonomicName id="71D34D48D970165AFE34C208FCBEFC0D" authority="(Gordon, 1975)" baseAuthorityName="Gordon" baseAuthorityYear="1975" box="[405,833,893,917]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="monovittata">
<emphasis id="84A7EAD9D970165AFE34C208FD77FC0C" box="[405,648,893,916]" italics="true" pageId="34" pageNumber="365">Toxotoma monovittata</emphasis>
(
<bibRefCitation id="D2424B3AD970165AFD36C208FCC6FC0D" author="Gordon, R. D." box="[663,825,893,917]" pageId="34" pageNumber="365" pagination="1 - 409" refId="ref26293" refString="Gordon, R. D. (1975) A revision of Epilachninae of the Western Hemisphere (Coleoptera: Coccinellidae). US Department of Agriculture Technical Bulletin, 1493: 1 - 409." type="journal article" year="1975">Gordon, 1975</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="71D34D48D970165AFCDDC208FB4AFC0D" authority="(Crotch, 1874)" baseAuthorityName="Crotch" baseAuthorityYear="1874" box="[892,1205,893,917]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="univittata">
<emphasis id="84A7EAD9D970165AFCDDC208FBF9FC0C" box="[892,1030,893,916]" italics="true" pageId="34" pageNumber="365">T. univittata</emphasis>
(
<bibRefCitation id="D2424B3AD970165AFBB4C208FB52FC0D" author="Crotch, G. R." box="[1045,1197,893,917]" pageId="34" pageNumber="365" refId="ref25482" refString="Crotch, G. R. (1874) A revision of the Coleopterous Family Coccinellidae. University Press, London, 311 pp." type="book" year="1874">Crotch, 1874</bibRefCitation>
)
</taxonomicName>
, assigned by 
<bibRefCitation id="D2424B3AD970165AFE8AC2E8FE2AFC2D" author="Gordon, R. D." box="[299,469,925,949]" pageId="34" pageNumber="365" pagination="1 - 409" refId="ref26293" refString="Gordon, R. D. (1975) A revision of Epilachninae of the Western Hemisphere (Coleoptera: Coccinellidae). US Department of Agriculture Technical Bulletin, 1493: 1 - 409." type="journal article" year="1975">Gordon (1975)</bibRefCitation>
in the same species-group as 
<taxonomicName id="71D34D48D970165AFC97C2E8FB4AFC2D" authority="(Crotch, 1874)" baseAuthorityName="Crotch" baseAuthorityYear="1874" box="[822,1205,925,949]" class="Insecta" family="Coccinellidae" genus="Epilachna" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="species" species="vittigera">
<emphasis id="84A7EAD9D970165AFC97C2E8FBF9FC2C" box="[822,1030,925,949]" italics="true" pageId="34" pageNumber="365">Epilachna vittigera</emphasis>
(
<bibRefCitation id="D2424B3AD970165AFBB4C2E8FB52FC2D" author="Crotch, G. R." box="[1045,1197,925,949]" pageId="34" pageNumber="365" refId="ref25482" refString="Crotch, G. R. (1874) A revision of the Coleopterous Family Coccinellidae. University Press, London, 311 pp." type="book" year="1874">Crotch, 1874</bibRefCitation>
)
</taxonomicName>
, had already been transferred to 
<taxonomicName id="71D34D48D970165AFDB5C2C8FD82FC4C" authorityName="Weise" authorityYear="1899" box="[532,637,957,980]" class="Insecta" family="Coccinellidae" genus="Toxotoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="365" phylum="Arthropoda" rank="genus">
<emphasis id="84A7EAD9D970165AFDB5C2C8FD82FC4C" box="[532,637,957,980]" italics="true" pageId="34" pageNumber="365">Toxotoma</emphasis>
</taxonomicName>
by 
<bibRefCitation id="D2424B3AD970165AFD09C2C8FBC6FC4D" author="Tomaszewska, W. &amp; Szawaryn, K." box="[680,1081,957,981]" pageId="34" pageNumber="365" pagination="1 - 91" refId="ref28231" refString="Tomaszewska, W. &amp; Szawaryn, K. (2016) Epilachnini (Coleoptera: Coccinellidae) - a revision of the world genera. Journal of Insect Science, 16 (1): 1 - 91. https: // doi. org / 10.1093 / jisesa / iew 082" type="journal article" year="2016">Tomaszewska &amp; Szawaryn (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FEC96540D970165AFF36C288FE1DFBAC" pageId="34" pageNumber="365" type="etymology">
<paragraph id="B66C36CBD970165AFF36C288FE1DFBAC" blockId="34.[151,1209,1021,1077]" pageId="34" pageNumber="365">
<emphasis id="84A7EAD9D970165AFF36C288FEDEFB8D" bold="true" box="[151,289,1021,1045]" pageId="34" pageNumber="365">Etymology.</emphasis>
The name is named after the collection locality, Aguas Calientes, in the Department of 
<collectingRegion id="7417F829D970165AFEEDC568FE67FBAD" box="[332,408,1053,1077]" country="Peru" name="Cusco" pageId="34" pageNumber="365">Cusco</collectingRegion>
, 
<collectingCountry id="CEC4765BD970165AFE02C568FE21FBAC" box="[419,478,1053,1076]" name="Peru" pageId="34" pageNumber="365">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>