treatments-xml/data/11/EC/A4/11ECA462290D5362BC20BAE4DB0E3ADE.xml

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<mods:title id="0F5788C381234ED4FDB603CA81B62348">Revision of the subgenus Mesopraocis Flores &amp; Pizarro-Araya of the Neotropical genus Praocis Eschscholtz (Coleoptera, Tenebrionidae, Pimeliinae)</mods:title>
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<mods:namePart id="97C10A0DCD7303782A064F7D0168A6B3">Flores, Gustavo E.</mods:namePart>
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<mods:affiliation id="C1145D2256ACC174851802288446CF12">Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA), Laboratorio de Entomologia, Centro Cientifico Tecnologico CONICET, Av. Ruiz Leal s / n, Parque General San Martin, 5500 Mendoza, Argentina</mods:affiliation>
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<mods:namePart id="239240ED32E55AEC0F0D8BD6B9FF7478">Pizarro-Araya, Jaime</mods:namePart>
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<mods:affiliation id="CD6BCE590567C01704BA423FADD4134C">Laboratorio de Entomologia Ecologica (LEULS), Departamento de Biologia, Facultad de Ciencias, Universidad de La Serena, Casilla 554, La Serena, Chile &amp; Grupo de Artropodos, Sistema Integrado de Monitoreo y Evaluacion de Ecosistemas Forestales Nativos (SIMEF), Chile &amp; Instituto de Ecologia y Biodiversidad (IEB), Santiago, Chile</mods:affiliation>
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<treatment id="11ECA462290D5362BC20BAE4DB0E3ADE" ID-GBIF-Taxon="195304379" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE" httpUri="http://treatment.plazi.org/id/11ECA462290D5362BC20BAE4DB0E3ADE" lastPageNumber="29" pageId="0" pageNumber="29" scope_family="Tenebrionidae" scope_order="Coleoptera" scope_subGenus="Praocis">
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<paragraph id="D4F39EAB69981399D012DD6DD61573B2" pageId="0" pageNumber="29">
<taxonomicName id="E193E159F1E36A7E06BFC2861631A87A" LSID="11ECA462-290D-5362-BC20-BAE4DB0E3ADE" authority="Kulzer, 1959" authorityName="Kulzer" authorityYear="1959" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis (Mesopraocis) nitens" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="nitens" subGenus="Mesopraocis">Praocis (Mesopraocis) nitens Kulzer, 1959</taxonomicName>
</paragraph>
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<subSubSection id="18AC703623C14503EFA856A1864E2C90" pageId="0" pageNumber="29" type="description">
<paragraph id="5F90AB698A09F51A52706D8B86F8458D" pageId="0" pageNumber="29">
<figureCitation id="5B26663C6C226F06606D48C9B8E3843B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genitalia of Praocis (Mesopraocis) species A-D male A, B Praocis (Mesopraocis) calderana, dorsal and ventral views C, D Praocis (Mesopraocis) nitens, dorsal and ventral views. Abbreviations: bl, basal lamina of tegmen, ls, lateral styles of tegmen, ml, median lobe E ovipositor (ventral view), spiculum ventrale and internal female reproductive tract of Praocis (Mesopraocis) pilula. Abbreviations: bc, baculi of coxite, bp, baculi of paraproct, c, coxite, g, gonostyli, m, midventral sclerite, o, oviduct, p, paraproct, r, rectum, s, spermatheca, sag, spermathecal accessory gland, sp, spiculum ventrale, v, vagina, 1, 2, 3, 4, coxite lobes. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure3" httpUri="https://binary.pensoft.net/fig/683155" pageId="0" pageNumber="29">Figs 3C, D</figureCitation>
<figureCitation id="E155B433DF16AE69EB4F03192CB77DBD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus in dorsal view A Praocis (Mesopraocis) pilula B Praocis (Mesopraocis) calderana C Praocis (Mesopraocis) nitens D Praocis (Mesopraocis) arenicola sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure4" httpUri="https://binary.pensoft.net/fig/683156" pageId="0" pageNumber="29">, 4C</figureCitation>
<figureCitation id="4328DDB151EC89ECED2FD4327E7592BD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Geographical distribution of Praocis (Mesopraocis) calderana and Praocis (Mesopraocis) nitens." figureDoi="10.3897/zookeys.1100.78769.figure5" httpUri="https://binary.pensoft.net/fig/683157" pageId="0" pageNumber="29">, 5</figureCitation>
</paragraph>
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<paragraph id="E6449AA41043203AB276830D3CE9C8D5" pageId="0" pageNumber="29">
<taxonomicName id="B155383AF54DAE8C95D87236AE340FEB" authorityName="Kulzer" authorityYear="1959" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis nitens" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="nitens">Praocis nitens</taxonomicName>
Kulzer, 1959: 561; 
<bibRefCitation id="973C600D7EE8D47BEFE171FDE743DC2A" author="Pena, LE" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="0" pageNumber="29" pagination="397 - 453" refId="B34" refString="Pena, LE, 1966. Catalogo de los Tenebrionidae (Coleoptera) de Chile. Entomologische Arbeiten aus dem Museum George Frey 17: 397 - 453" title="Catalogo de los Tenebrionidae (Coleoptera) de Chile." volume="17" year="1966">
<normalizedToken id="BD18051419B83E5F19D735B06F401D79" originalValue="Peña">Pena</normalizedToken>
1966
</bibRefCitation>
: 431 (cat.); 
<bibRefCitation id="FE25EEEB1349C3C50E98FE72BEA70ACC" author="Vidal, P" journalOrPublisher="International Journal of Insect Morphology &amp; Embryology" pageId="0" pageNumber="29" refId="B45" refString="Vidal, P, Guerrero, M, 2007. Los Tenebrionidos de Chile. Ediciones Universidad Catolica de Chile. Santiago, 478 pp." title="Los Tenebrionidos de Chile. Ediciones Universidad Catolica de Chile. Santiago, 478 pp." year="2007">Vidal and Guerrero 2007</bibRefCitation>
: 73, 218; 
<bibRefCitation id="BB91EEE3598BE29F34F7DA9BE6ED2551" author="Pizarro-Araya, J" editor="Squeo, FA" journalOrPublisher="Ediciones Universidad de La Serena, La Serena" pageId="0" pageNumber="29" pagination="257 - 274" refId="B36" refString="Pizarro-Araya, J, Cepeda-Pizarro, J, Flores, GE, 2008. Diversidad taxonomica de los artropodos epigeos de la Region de Atacama (Chile): estado del conocimiento. In: Squeo, FA, Arancio, G, Gutierrez, JR, Eds., Libro Rojo de la Flora Nativa y de los Sitios Prioritarios para su Conservacion: Region de Atacama. Ediciones Universidad de La Serena, La Serena: 257 - 274" title="Diversidad taxonomica de los artropodos epigeos de la Region de Atacama (Chile): estado del conocimiento." volumeTitle="Libro Rojo de la Flora Nativa y de los Sitios Prioritarios para su Conservacion: Region de Atacama." year="2008">Pizarro-Araya et al. 2008</bibRefCitation>
: 273 (list.); 
<bibRefCitation id="B5557EA6B1FACF98876DCF2915FBF9CD" DOI="https://doi.org/10.3897/zookeys.415.6656" author="Flores, GE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="29" pagination="53 - 80" refId="B16" refString="Flores, GE, Pizarro-Araya, J, 2014. Towards a revision of the South American genus Praocis Eschscholtz (Coleoptera: Tenebrionidae), with estimation of the diversity of each subgenus. ZooKeys 415: 53 - 80, DOI: https://doi.org/10.3897/zookeys.415.6656" title="Towards a revision of the South American genus Praocis Eschscholtz (Coleoptera: Tenebrionidae), with estimation of the diversity of each subgenus." url="https://doi.org/10.3897/zookeys.415.6656" volume="415" year="2014">Flores and Pizarro-Araya 2014</bibRefCitation>
: 60 (list).
</paragraph>
</subSubSection>
<subSubSection id="ED1661F31B4C00FD69DF49D327F73101" pageId="0" pageNumber="29" type="redescription">
<paragraph id="CC71CB843FF8422516FFD8219FF90C25" pageId="0" pageNumber="29">Redescription.</paragraph>
<paragraph id="838A2EC1F2090B6822A6BFEA0E8090D9" pageId="0" pageNumber="29">Length 7.87-10.62 mm. Body, antennae, legs black to dark brown, body widest at prothorax. Head. Clypeus with punctures bearing short setae on all surface, width of anterior margin equal to half the interocular width; frons with punctures bearing short setae on all surface, border of punctures on frons and epicanthus elevated; frontoclypeal suture with short or long setae; eyes with supraocular groove; antennae reaching anterior third of lateral margin of pronotum; antennomere 9 longer than wide, 11 wider than long, antennomere 9 longer than 10 and 11, antennomere 9 narrower than 10 and wider than 11, antennomere 10 wider than 11.</paragraph>
<paragraph id="6CDD8F6B880E7682E64915730988F7A1" pageId="0" pageNumber="29">
<emphasis id="237A92C5722C4072F4B110314A409493" bold="true" pageId="0" pageNumber="29">
<emphasis id="BB08CAEF8A7EA1185893A89734C4FD2E" italics="true" pageId="0" pageNumber="29">Thorax</emphasis>
.
</emphasis>
Pronotum with anterior angles rounded, not produced, remote from eyes and epicanthus, lateral margins declivous, disc convex reaching lateral margins (Fig. 
<figureCitation id="533352691827C0228FAEBA81866BE16C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus in dorsal view A Praocis (Mesopraocis) pilula B Praocis (Mesopraocis) calderana C Praocis (Mesopraocis) nitens D Praocis (Mesopraocis) arenicola sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure4" httpUri="https://binary.pensoft.net/fig/683156" pageId="0" pageNumber="29">4C</figureCitation>
); posterior margin bisinuate, central area not extending beyond lateral extents of posterior margin (Fig. 
<figureCitation id="126CD30338E6360FEA92011F2766E998" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus in dorsal view A Praocis (Mesopraocis) pilula B Praocis (Mesopraocis) calderana C Praocis (Mesopraocis) nitens D Praocis (Mesopraocis) arenicola sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure4" httpUri="https://binary.pensoft.net/fig/683156" pageId="0" pageNumber="29">4C</figureCitation>
), posterior angles right with apex rounded, produced. Hypomeron with a fringe of long golden setae below lateral margin of pronotum. Metaventrite with punctures on lateral thirds separated by two to four puncture diameters, transverse grooves parallel to metacoxae formed by the fusion of a row of punctures.
</paragraph>
<paragraph id="6E1480A468F85754372A427724E3ED78" pageId="0" pageNumber="29">
<emphasis id="887A65B82DA2FEEB62B5E9C9A8EF04BE" bold="true" pageId="0" pageNumber="29">
<emphasis id="574C9C267AB12CD60DCC21915DA743F4" italics="true" pageId="0" pageNumber="29">Elytron</emphasis>
.
</emphasis>
Pseudopleuron with sparse punctures of which emerge long, erect setae, becoming long, dense on anterior half of upper surface, punctures separated by three to six puncture diameters on lower surface, separated by one to two puncture diameters on anterior half of upper surface; epipleuron with a fringe of long golden setae on anterior half of edge, shiny, almost smooth, with sparse punctures and setae on anterior quarter, and with a row of punctures with same diameter and depth as on pseudopleuron located on posterior three fourths.
</paragraph>
<paragraph id="3761B31947B91461E38E36DC09A17438" pageId="0" pageNumber="29">
<emphasis id="8CF38F16EE022530E330B11D5B84B788" bold="true" pageId="0" pageNumber="29">
<emphasis id="11BE870C5446315AB16312E4CB33D675" italics="true" pageId="0" pageNumber="29">Legs</emphasis>
.
</emphasis>
Ventral surface of protrochanters with hair brush. Apical process of distal margin of protibiae equal to length of protarsomeres 1-2 combined; protibial length 2.5 times the width of distal margin. First metatarsomere 1.4-1.6 times longer than metatarsomeres 2-3 combined (Fig. 
<figureCitation id="7C8E5E68A1CB1227756EA8F42F13E778" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitus in dorsal view A Praocis (Mesopraocis) pilula B Praocis (Mesopraocis) calderana C Praocis (Mesopraocis) nitens D Praocis (Mesopraocis) arenicola sp. nov. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure4" httpUri="https://binary.pensoft.net/fig/683156" pageId="0" pageNumber="29">4C</figureCitation>
).
</paragraph>
<paragraph id="403201582E6F142FBBE126BA24491B45" pageId="0" pageNumber="29">
<emphasis id="39F095C4A9B102EF573C1013B0F2BCD0" bold="true" pageId="0" pageNumber="29">
<emphasis id="49D588B0FED6E3173CF719E597B7DEE3" italics="true" pageId="0" pageNumber="29">Abdomen</emphasis>
.
</emphasis>
Ventrites 1-2 with sparse punctures separated by two to four puncture diameters; ventrites 3-4 smooth on central area, with punctures on lateral thirds separated by two to four puncture diameters; ventrite 5 with punctures on central area separated by two to four puncture diameters, on lateral thirds separated by one to two puncture diameters.
</paragraph>
<paragraph id="F989BD0050AE4A17EBBCBC03AC45C481" pageId="0" pageNumber="29">
<emphasis id="8890E5DE8394C8C9FDCE284B1880CC68" bold="true" pageId="0" pageNumber="29">
<emphasis id="EF0C643430B50E2228E3BC13EA7D8B73" italics="true" pageId="0" pageNumber="29">Male genitalia</emphasis>
(Fig. 
<figureCitation id="AA64B3FB89B83E5412ECCA38739FCCB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Male and female genitalia of Praocis (Mesopraocis) species A-D male A, B Praocis (Mesopraocis) calderana, dorsal and ventral views C, D Praocis (Mesopraocis) nitens, dorsal and ventral views. Abbreviations: bl, basal lamina of tegmen, ls, lateral styles of tegmen, ml, median lobe E ovipositor (ventral view), spiculum ventrale and internal female reproductive tract of Praocis (Mesopraocis) pilula. Abbreviations: bc, baculi of coxite, bp, baculi of paraproct, c, coxite, g, gonostyli, m, midventral sclerite, o, oviduct, p, paraproct, r, rectum, s, spermatheca, sag, spermathecal accessory gland, sp, spiculum ventrale, v, vagina, 1, 2, 3, 4, coxite lobes. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1100.78769.figure3" httpUri="https://binary.pensoft.net/fig/683155" pageId="0" pageNumber="29">3C, D</figureCitation>
).
</emphasis>
Basal lamina of tegmen equal width throughout, with base concave. Lateral styles of tegmen with proximal margin bisinuate, with setae on distal 1/6 of ventral surface. Median lobe distally widened, half the width of tegmen, with apical aperture large.
</paragraph>
<caption id="408592BB0148B7272077ABB9F88E06DD" doi="10.3897/zookeys.1100.78769.figure5" httpUri="https://binary.pensoft.net/fig/683157" pageId="0" pageNumber="29" start="Figure 5" startId="F5">
<paragraph id="71869358A84E79D711944D70F95D6189" pageId="0" pageNumber="29">
<emphasis id="85D1AA813738E14718503500C448289B" bold="true" pageId="0" pageNumber="29">Figure 5.</emphasis>
Geographical distribution of 
<taxonomicName id="9D62C7D254C3A44F779E704D999BF6E4" authorityName="Kulzer" authorityYear="1958" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis (Mesopraocis) calderana" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="calderana" subGenus="Mesopraocis">Praocis (Mesopraocis) calderana</taxonomicName>
and 
<taxonomicName id="5FAF03A00BADF1DC110F5A34377E1FCD" authorityName="Kulzer" authorityYear="1959" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis (Mesopraocis) nitens" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="nitens" subGenus="Mesopraocis">Praocis (Mesopraocis) nitens</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DCBC37530F7ED0EC9AA4AD39BCCC1572" pageId="0" pageNumber="29" type="type specimens">
<paragraph id="13E527B96027B4DC800B3A2443761305" pageId="0" pageNumber="29">Type specimens.</paragraph>
<paragraph id="B76219FDF6999907A7DA66F5693FDAEB" pageId="0" pageNumber="29">
Holotype male: [Huasco/ Atacama/ 20-22.x.1957/ Coll: L.E. 
<normalizedToken id="546AAA3967376C872B5C6B82C9FF2720" originalValue="Peña">Pena</normalizedToken>
] [Holotypus/ 
<taxonomicName id="032D58508103971AA1EB107ADA4EAE94" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="7BC679C778996F970F9750E0C9815D26" italics="true" pageId="0" pageNumber="29">Praocis</emphasis>
</taxonomicName>
/ (
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<emphasis id="58CD45354E8C9D70FE708E038CDD568F" italics="true" pageId="0" pageNumber="29">Mesopraocis</emphasis>
</taxonomicName>
)/ 
<taxonomicName id="9CF52FC90085EF87F0A84D47CD62B96D" authorityName="Kulzer" authorityYear="1959" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis nitens" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="39E3A52FCB927C5DAFD204BC8E451AA2" italics="true" pageId="0" pageNumber="29">Praocis nitens</emphasis>
</taxonomicName>
m./ det. H. 
<bibRefCitation id="88F72FC35DA446B7B4878A39AC9703F6" author="Kulzer, H" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="0" pageNumber="29" pagination="1 - 105" refId="B26" refString="Kulzer, H, 1958. Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 9: 1 - 105" title="Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden)." volume="9" year="1958">Kulzer 1958</bibRefCitation>
] (FMNH); allotype female (FMNH) and four paratypes with the same data as holotype plus [Paratypus/ 
<taxonomicName id="B0F392BDEDF8A45D592A379BD5C5B156" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="49CD186A65FF8BFAD15347A63171851A" italics="true" pageId="0" pageNumber="29">Praocis</emphasis>
</taxonomicName>
/ 
<taxonomicName id="4194BC0DC8A67461F8F3DC5069510E62" authorityName="Kulzer" authorityYear="1959" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Praocis nitens" order="Coleoptera" pageId="0" pageNumber="29" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="22992BC774AF062F65B22C6FB189DD7D" italics="true" pageId="0" pageNumber="29">Praocis nitens</emphasis>
</taxonomicName>
m./ H. 
<bibRefCitation id="AFAAE99778B7D1B7163D2B8C370CD40E" author="Kulzer, H" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="0" pageNumber="29" pagination="1 - 105" refId="B26" refString="Kulzer, H, 1958. Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum George Frey 9: 1 - 105" title="Monographie der suedamerikanischen Tribus Praocini (Col.) (16 Beitrag zur Kenntnis der Tenebrioniden)." volume="9" year="1958">Kulzer 1958</bibRefCitation>
] 2 (FMNH), 2 (OSUC).
</paragraph>
</subSubSection>
<subSubSection id="27EE3169714E784C5AFCFFE09AFE8CF3" pageId="0" pageNumber="29" type="materials_examined">
<paragraph id="C607CA7C7657DBADF00ADB5C47E959FC" pageId="0" pageNumber="29">Other material examined.</paragraph>
<paragraph id="19CA1E6347E7D7093E9CFF4358F6E77B" pageId="0" pageNumber="29">
<materialsCitation id="8990CC85D03C0B626E80BFB35C57B710" collectingDate="1991-11" collectorName="H. Vasquez, Algarrobal" country="Chile" county="Copiapo Province" location="Caldera" municipality="Rodillo" specimenCount="1" stateProvince="Atacama Region">
<collectingCountry id="E67E3375C23286FFA41B05E493DBD8F0" name="Chile">Chile</collectingCountry>
. 
<collectingRegion id="898E18BD2177B7B78206BCE95D260511" country="Chile" name="Atacama">Atacama Region</collectingRegion>
: 
<collectingCounty id="CAB6C354D70F85FAAB2EFD66FF91B1FE">Copiapo Province</collectingCounty>
: 
<collectingMunicipality id="1B74E42083D44510852172B316FB231D">Rodillo</collectingMunicipality>
, 
<location id="EF9349EE4CEA1514064C03F9768AA3DD" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:EF9349EE4CEA1514064C03F9768AA3DD" country="Chile" county="Copiapo Province" municipality="Rodillo" name="Caldera" stateProvince="Atacama Region">Caldera</location>
, 
<collectingDate id="494CA6D021E50FD0FBE83817EA4D5B26" value="1991-11">xi.1991</collectingDate>
, 
<collectorName id="89FF7053CE311E3652CFE665D956E4AA">
H. 
<normalizedToken id="7DC3F5B62A8BAC4C3064184C8A4BF90C" originalValue="Vásquez">Vasquez</normalizedToken>
</collectorName>
, 10 (LEULS). 
<location id="42171AE6D4BCD8A6CCF934DB92407327" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:42171AE6D4BCD8A6CCF934DB92407327" country="Chile" county="Copiapo Province" municipality="Rodillo" name="Huasco Province" stateProvince="Atacama Region">Huasco Province</location>
: 
<collectorName id="D52A4F8A387F39E093E5F09CC48FE78A">Algarrobal</collectorName>
</materialsCitation>
, 
<materialsCitation id="0EBCBCD9199BA11C980276827D6A83BD" collectingDate="1957-10-20" collectingDateMax="2017-09-03" collectingDateMin="1957-10-20" collectorName="J. Pizarro-Araya, N. Huasco, L. E. Pena, F. M. Alfaro, M. Elgueta, Leg. G. Castillo, Leg. F. M. Alfaro" country="Chile" county="Huasco" latitude="-28.423916" location="Parque Nacional Llanos de Challe" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" specimenCount="53" stateProvince="Atacama" typeStatus="paratypes">
<collectingRegion id="F98257104B91139DB9FB85C78AFAFEF4" country="Chile" name="Atacama">Atacama</collectingRegion>
, 
<collectingDate id="2D38A7242F4D654D8FF1600416DACCD2" value="2002-09-15">15.ix.2002</collectingDate>
, 
<collectorName id="EF5AD5C299D78E528282DF580DF55AC6">J. Pizarro-Araya</collectorName>
, 1 (LEULS), Quebrada Carrizalillo, 
<collectorName id="0AF1F1EA276B02AAC0D559742A9D59E7">
N. 
<collectingCounty id="5BED9FD8C193C957B6DAC79C778E7C90">Huasco</collectingCounty>
</collectorName>
, 
<collectingDate id="A40ABC463FE523A69870EA902C70973F" value="1980-10-12">12.x.1980</collectingDate>
, 
<collectorName id="86E11BD24FEDBAF2AB8B0599100BB211">
L.E. 
<normalizedToken id="E73067F3D11D89B24616F6040939D292" originalValue="Peña">Pena</normalizedToken>
</collectorName>
, 1 (FMNH), 
<collectingMunicipality id="2A5FEC82BB325F1611829E267F3D0A37">Parque Nacional Llanos de Challe</collectingMunicipality>
, 
<geoCoordinate id="8DF2B9805BD568E3416F351829C9E347" degrees="28" direction="south" minutes="11" orientation="latitude" precision="1" seconds="17.36" value="-28.188154">28°11'17.36&quot;S</geoCoordinate>
, 
<geoCoordinate id="66700A8774D9A7C12A294A8E03D0B238" degrees="71" direction="west" minutes="9" orientation="longitude" precision="1" seconds="13.63" value="-71.153786">71°9'13.63&quot;W</geoCoordinate>
, 
<collectingDate id="7701957854B5D4C59387D6ED7E9E34F4" value="2016-12-20" valueMax="2016-12-22" valueMin="2016-12-20">20-22.xii.2016</collectingDate>
, 
<collectorName id="3DE3D0568EE63C676BE61CCF105E6A3D">J. Pizarro-Araya</collectorName>
&amp; 
<collectorName id="FC4408856EEB2BAC113247F7C1D5A7EC">F.M. Alfaro</collectorName>
, 9 (LEULS), 
<collectingDate id="F90D9F0C152A481B4300284F2B2227D9" value="1997-10-15">15.x.1997</collectingDate>
, Traps barber, 17 (LEULS), 
<location id="019A9684231F7E943321078542DDBEA2" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:019A9684231F7E943321078542DDBEA2" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Parque Nacional Llanos de Challe" stateProvince="Atacama">Parque Nacional Llanos de Challe</location>
, UTM 19 J 311128 W, 6883333 N, 269 msl, 
<collectingDate id="0898C47D9FCE674116DCF70BF9D30DD9" value="2017-08-31" valueMax="2017-09-03" valueMin="2017-08-31">31.viii-03.ix.2017</collectingDate>
, 
<collectorName id="0ED12C43BA14A1E3B4E22BE03B1032EF">J. Pizarro-Araya</collectorName>
&amp; 
<collectorName id="529A18B19B311EB7F2EA62C9102D7378">F.M. Alfaro</collectorName>
(SIMEF-Project), 3 (LEULS), 
<location id="A012C12671BBD790EF74DCE290D7E0AC" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:A012C12671BBD790EF74DCE290D7E0AC" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Agua de Luna" stateProvince="Atacama">Agua de Luna</location>
, 
<locationDeviation id="F247CC2F9A54A0A06BBCB5ADB5B51A88" location="Huasco">
<quantity id="CB7795C5183174E94A7D91FF8651E3FC" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
N Huasco
</locationDeviation>
, sand dunes, 
<geoCoordinate id="AEDE9185746EB38066E375B4E1C0D2F5" degrees="28" direction="south" minutes="20" orientation="latitude" precision="15" seconds="09" value="-28.335835">28°20'09&quot;S</geoCoordinate>
,71°09'&quot;W, 
<collectingDate id="07512544F31B508F250A13C9FBB43F03" value="2004-01">i.2004</collectingDate>
, 
<collectorName id="99E4B4059FF848C22ED1FEB891FFA64A">F.M. Alfaro</collectorName>
, 1 (IADIZA), 
<collectingDate id="AD75F2FE20A3EAC9073962D3404E2951" value="2010-10-27">27.x.2010</collectingDate>
, 
<collectorName id="A4EB45DFA095CCBBEC3D0137E905D63D">M. Elgueta</collectorName>
, 4 (MNNC), 
<geoCoordinate id="139255347904DE0391AC0E5B8F01F72A" degrees="28" direction="south" minutes="20" orientation="latitude" precision="1" seconds="5.2" value="-28.334778">28°20'5.2&quot;S</geoCoordinate>
, 71°09'37.&quot;W, 
<collectingDate id="138C47FD63602024B506DC31BC01285B" value="2012-10-29">29.x.2012</collectingDate>
, 
<collectorName id="5F5FF04BED52734AD8096E5585E22581">M. Elgueta</collectorName>
, 2 (IADIZA), 
<location id="0EE3BE2041533077BDB1BBCF405EB4D7" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:0EE3BE2041533077BDB1BBCF405EB4D7" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
bajo, 
<collectingDate id="2A7F1A66FAF857BD85D92993F45C3BB2" value="1976-08-08">8.viii.1976</collectingDate>
, 1 (UCCC), 
<location id="A105485CF2F5C78109D9D18DDF16049F" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:A105485CF2F5C78109D9D18DDF16049F" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="4AA65E38503AB163793394FB4AF700C6" value="1957-10-20" valueMax="1957-10-22" valueMin="1957-10-20">20-22.x.1957</collectingDate>
, 
<collectorName id="8DB132E6C74EF482FD0AC20509791444">
L.E. 
<normalizedToken id="D81B9DD7AC764815A741EA55D562978E" originalValue="Peña">Pena</normalizedToken>
</collectorName>
, 2 (OSUC), 29 (FMNH). 
<emphasis id="428462828F77EE7D90F383A4D5050516" bold="true" pageId="0" pageNumber="29">Note</emphasis>
: these 
<specimenCount id="02E26817092DBD597B8E4E8F4C9EBDCD" type="generic">29 specimens</specimenCount>
were later labeled as 
<typeStatus id="688D83A5D70DFA8DBBD28D320C289247">paratypes</typeStatus>
by 
<normalizedToken id="0CE1A388D059E45B471AD4CCD6C8B34D" originalValue="Peña">Pena</normalizedToken>
but these were not originally designated by 
<bibRefCitation id="1EE9D776239121EA4114A632C2E71B5A" author="Kulzer, H" journalOrPublisher="Entomologische Arbeiten aus dem Museum George Frey" pageId="0" pageNumber="29" pagination="523 - 567" refId="B27" refString="Kulzer, H, 1959. Neue Tenebrioniden aus Suedamerika (Col). (18. Beitrag zur Kenntnis der Tenebrioniden). 3. Verschiedene Neue Arten. Entomologische Arbeiten aus dem Museum George Frey 10: 523 - 567" title="Neue Tenebrioniden aus Suedamerika (Col). (18. Beitrag zur Kenntnis der Tenebrioniden). 3. Verschiedene Neue Arten." volume="10" year="1959">Kulzer (1959)</bibRefCitation>
, who stated that he designated 
<specimenCount id="3854E1E64D68EF1240E04B26743BE4CC" type="generic" typeStatus="paratypes">24 paratypes</specimenCount>
with the same data as holotype and 
<normalizedToken id="072E3EEFF595382AF430EE85878E39BE" originalValue="Peña">Pena</normalizedToken>
kept 88 more specimens; 
<collectingDate id="2161F0D6CA140D83B1D04A198BF99E58" value="1974-10-17" valueMax="1974-10-21" valueMin="1974-10-17">17-21.x.1974</collectingDate>
, 
<collectorName id="97A556EC1569274B0625983536EDBDA7">
L.E. 
<normalizedToken id="278E2BB5D1D3EF65FC25569742D282C3" originalValue="Peña">Pena</normalizedToken>
</collectorName>
, 68 (FMNH), 
<location id="33EE3011473565A8CA8C7877EB616112" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:33EE3011473565A8CA8C7877EB616112" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, sand dunes, 
<collectingDate id="226BE5FC72E94E666C7CC6DB7509DD83" value="1981-09-28" valueMax="1981-09-29" valueMin="1981-09-28">28-29.ix.1981</collectingDate>
, 
<collectorName id="A7B500207AE3EA535D654934BA287731">M. Elgueta</collectorName>
, 3 (MNNC), 
<geoCoordinate id="7654A440712BB188C42158E8994919FB" degrees="28" direction="south" minutes="27" orientation="latitude" precision="1" seconds="19.5" value="-28.455418">28°27'19.5&quot;S</geoCoordinate>
, 71°12'6.&quot;W, 
<collectingDate id="FDE7972F8D366DA9951238DBE0AFF7E3" value="2012-10-28">28.x.2012</collectingDate>
, 
<collectorName id="E934E3C5715C8B376E1315468DADDA20">M. Elgueta</collectorName>
, 1 (IADIZA), 
<locationDeviation id="E2AC8736478F07CC941AD61E996DCD9E" location="Huasco">
<quantity id="99A53F262EC6E2C8DCAA9ACC01EF2CFE" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
N Huasco
</locationDeviation>
, 
<collectingDate id="D2926A682B62F3ED7F535974E8AABD6A" value="1996-09-13">13.ix.1996</collectingDate>
, 
<collectorName id="4957D433CC20800B4E203B7A92D0014C">Leg. G. Castillo</collectorName>
, 1 (LEULS), 
<location id="6E37BB7173A9773A83BF193422EC39BA" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:6E37BB7173A9773A83BF193422EC39BA" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Agua de Luna" stateProvince="Atacama">Agua de Luna</location>
, 
<location id="FB2EB475EA2E54FD052D8AA79D7EEC36" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:FB2EB475EA2E54FD052D8AA79D7EEC36" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="58FDC58F679CC94E6F1B424BBE7D8E22" value="2008-11-12" valueMax="2008-11-16" valueMin="2008-11-12">12-16.xi.2008</collectingDate>
, 
<collectorName id="C176E4A816FDFE72995001227FA5B182">Leg. F. M. Alfaro</collectorName>
, 5 (LEULS), 
<collectingDate id="B08CE563D28386C47E02E61C6A2BC54A" value="2004-01">i.2004</collectingDate>
, 
<collectorName id="DB36666C970C5AE256F640859E30357D">F.M. Alfaro</collectorName>
4 (LEULS), Caleta Angosta, 
<location id="FC6769956B431814D16B0CB0D428D412" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:FC6769956B431814D16B0CB0D428D412" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="43499E1AC6F4E8646686071B35040523" value="2009-08-18">18.viii.2009</collectingDate>
, 
<collectorName id="D17CE565502421D359418EA1F8B4F1EF">F. M. Alfaro</collectorName>
, 1 (LEULS), Tres Playitas, 
<location id="1CACCC61190ADA2330EBF32F554C1DBB" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:1CACCC61190ADA2330EBF32F554C1DBB" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="E64BF8278AFC5969DBB15654BCFBF951" value="2015-11">xi.2015</collectingDate>
, 
<collectorName id="6B3E17AF55E58BE9A355691D772A193C">F.M. Alfaro</collectorName>
, 1 (IADIZA), 
<collectingDate id="0DC6CA7A7BE26735A08B650188DBBEB1" value="2014-11-16" valueMax="2014-11-21" valueMin="2014-11-16">16-21.xi.2014</collectingDate>
, 
<geoCoordinate id="081AFB99E1D9DE47C90E4B9E7BF0B2E3" degrees="28" direction="south" minutes="25" orientation="latitude" precision="1" seconds="26.1" value="-28.423916">28°25' 26.1&quot;S</geoCoordinate>
, 
<geoCoordinate id="A36F087343AA1FD92C07294BBC77537D" degrees="71" direction="west" minutes="11" orientation="longitude" precision="1" seconds="19.5" value="-71.18875">71°11'19.5&quot;W</geoCoordinate>
, 
<collectorName id="6FA4E8E1DB91455E09B726C1223771DC">J. Pizarro-Araya</collectorName>
&amp; 
<collectorName id="5D1666FBEA20803A6ED19611382FD57B">F.M. Alfaro</collectorName>
, 10 (LEULS), Los Bronces, 
<location id="D9F1565014B7225B826827EDBE2AB298" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:D9F1565014B7225B826827EDBE2AB298" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="3157A9604F255D696E1ACF39DDACC9E5" value="2006-10-15">15.x.2006</collectingDate>
, 
<collectorName id="DF145C456049F027B38D4EE200F9EA58">F.M. Alfaro</collectorName>
, 3 (IADIZA), 
<collectingDate id="26C4FAA5A89DA51A026954DF1BC938A7" value="2006-10-15">15.x.2006</collectingDate>
, 6 (LEULS), Playa Tontado, 
<location id="BE247141BC8377796077D351A2DAFFB6" LSID="urn:lsid:plazi:treatment:11ECA462290D5362BC20BAE4DB0E3ADE:BE247141BC8377796077D351A2DAFFB6" country="Chile" county="Huasco" latitude="-28.423916" longLatPrecision="1" longitude="-71.18875" municipality="Parque Nacional Llanos de Challe" name="Huasco" stateProvince="Atacama">Huasco</location>
, 
<collectingDate id="C972758013AF0D21B2B991FEF50866C3" value="2008-11-28">28.xi.2008</collectingDate>
, 
<collectorName id="58FBCE81E46B9DDCB23FBD0055E89B7B">Leg. F.M. Alfaro</collectorName>
, 5 (LEULS)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>