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<document id="AEE5B6DFCDB91CDA1406F058E918D1B6" ID-CLB-Dataset="24722" ID-DOI="10.11646/zootaxa.4705.1.1" ID-GBIF-Dataset="3e84f284-4d30-4c6e-a801-f9822d49edfc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3563677" ID-ZooBank="F534C43A-AB09-4CB3-9B08-FD5BDFD90298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575538349781" checkinUser="plazi" docAuthor="Weissman, David B. &amp; Gray, David A." docDate="2019" docId="182387A80948FFBF51F6FBA103CDFF61" docLanguage="en" docName="zootaxa.4705.1.1.pdf" docOrigin="Zootaxa 4705 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Gryllus thinos Weissman &amp; Gray 2019, n. sp." docType="treatment" docVersion="10" lastPageNumber="46" masterDocId="E41AFFD00961FF925161FFD20157FF9A" masterDocTitle="Crickets of the genus Gryllus in the United States (Orthoptera: Gryllidae: Gryllinae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="42" updateTime="1698772641233" updateUser="ExternalLinkService">
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<mods:title id="8BBC7AF1B9AC1200DD84D196D4D02351">Crickets of the genus Gryllus in the United States (Orthoptera: Gryllidae: Gryllinae)</mods:title>
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<mods:namePart id="F233B264F6AEE18936F2598ACE34B760">Weissman, David B.</mods:namePart>
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<mods:namePart id="A607C1E54A7C08EDF94922FE825F335A">Gray, David A.</mods:namePart>
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<mods:number id="832A70EBD772DB03CA271FD180E64FFA">2019-12-05</mods:number>
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<treatment id="182387A80948FFBF51F6FBA103CDFF61" ID-GBIF-Taxon="160630886" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61" httpUri="http://treatment.plazi.org/id/182387A80948FFBF51F6FBA103CDFF61" lastPageId="45" lastPageNumber="46" pageId="41" pageNumber="42">
<subSubSection id="D89065350948FFBB51F6FBA100E8FB2B" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="903536BE0948FFBB51F6FBA103D6FB14" blockId="41.[151,641,1139,1237]" box="[151,641,1139,1166]" pageId="41" pageNumber="42">
<heading id="CB7D81D20948FFBB51F6FBA103D6FB14" bold="true" box="[151,641,1139,1166]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<emphasis id="A2FEEAAC0948FFBB51F6FBA103D6FB14" bold="true" box="[151,641,1139,1166]" pageId="41" pageNumber="42">
<taxonomicName id="578A4D3D0948FFBB51F6FBA10366FB17" authority="Weissman &amp; Gray" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[151,561,1139,1166]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="thinos" status="sp. nov.">
<emphasis id="A2FEEAAC0948FFBB51F6FBA10014FB14" bold="true" box="[151,323,1139,1166]" italics="true" pageId="41" pageNumber="42">Gryllus thinos</emphasis>
Weissman &amp; Gray
</taxonomicName>
,
<taxonomicNameLabel id="B9CD57D70948FFBB535FFBA603D6FB14" box="[574,641,1140,1166]" pageId="41" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="903536BE0948FFBB51F6FB4A00E8FB2B" blockId="41.[151,641,1139,1237]" box="[151,447,1175,1202]" pageId="41" pageNumber="42">
<materialsCitation id="20E23CE30948FFBB51F6FB4A00E8FB2B" ID-GBIF-Occurrence="2464588165" box="[151,447,1175,1202]" location="Beach Field Cricket" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Texas">
<collectingRegion id="524EF85C0948FFBB51F6FB4A018FFB28" box="[151,216,1176,1202]" country="United States of America" name="Texas" pageId="41" pageNumber="42">Texas</collectingRegion>
<location id="955560650948FFBB51BEFB4500E8FB2B" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB51BEFB4500E8FB2B" box="[223,447,1175,1201]" name="Beach Field Cricket" pageId="41" pageNumber="42" stateProvince="Texas">Beach Field Cricket</location>
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</paragraph>
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<subSubSection id="D89065350948FFBF51F6FB6903CDFF61" lastPageId="45" lastPageNumber="46" pageId="41" pageNumber="42" type="description">
<paragraph id="903536BE0948FFBB51F6FB690022FB4F" blockId="41.[151,641,1139,1237]" box="[151,373,1211,1237]" pageId="41" pageNumber="42">
<figureCitation id="08B12A3B0948FFBB51F6FB690040FB4F" box="[151,279,1211,1237]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStart-3="FIGURE 27" captionStartId-0="42.[151,250,1147,1172]" captionStartId-1="43.[151,250,1827,1852]" captionStartId-2="44.[151,250,1282,1307]" captionStartId-3="44.[151,250,1738,1763]" captionTargetBox-0="[185,1420,271,1104]" captionTargetBox-1="[156,1402,642,1774]" captionTargetBox-2="[151,1058,174,1214]" captionTargetBox-3="[151,1388,1322,1703]" captionTargetId-0="figure@42.[151,1435,265,1128]" captionTargetId-1="figure@43.[151,1435,625,1807]" captionTargetId-2="figure@44.[151,1058,174,1263]" captionTargetId-3="figure@44.[151,1435,1322,1719]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionTargetPageId-2="44" captionTargetPageId-3="44" captionText-0="FIGURE 24. Holotype male (left), G. thinos. Female (right) from type locality showing typical, linear head stripes." captionText-1="FIGURE 25. Calling song of G. thinos (R07-125) Cameron Co., TX (S07-25), at 25°C: (A) five second waveform, (B) one second spectrogram." captionText-2="FIGURE 26. Known distribution of G. thinos." captionText-3="FIGURE 27. ITS2 gene tree showing some separation between Padre Island and Boca Chica samples of G. thinos." figureDoi-0="http://doi.org/10.5281/zenodo.3563725" figureDoi-1="http://doi.org/10.5281/zenodo.3563727" figureDoi-2="http://doi.org/10.5281/zenodo.3563729" figureDoi-3="http://doi.org/10.5281/zenodo.3563731" httpUri-0="https://zenodo.org/record/3563725/files/figure.png" httpUri-1="https://zenodo.org/record/3563727/files/figure.png" httpUri-2="https://zenodo.org/record/3563729/files/figure.png" httpUri-3="https://zenodo.org/record/3563731/files/figure.png" pageId="41" pageNumber="42">Figs 24–27</figureCitation>
,
<tableCitation id="DD0803050948FFBB5043FB690022FB4F" box="[290,373,1211,1237]" captionStart="TABLE 1" captionStartId="17.[150,238,466,492]" captionTargetPageId="17" captionText="TABLE 1. Summary measurements for morphological and song characters for Gryllus species discussed in this paper." httpUri="http://table.plazi.org/id/C4F566360970FF8351F7FE0004D1FE76" pageId="41" pageNumber="42" tableUuid="C4F566360970FF8351F7FE0004D1FE76">Table 1</tableCitation>
</paragraph>
<paragraph id="903536BE0948FFBB51F6FB2D03DEFA82" blockId="41.[151,649,1279,1304]" box="[151,649,1279,1304]" pageId="41" pageNumber="42">
‘
<taxonomicName id="578A4D3D0948FFBB51FAFB2D01BCFA82" authorityName="Linnaeus" authorityYear="1758" box="[155,235,1279,1304]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="genus">Gryllus</taxonomicName>
#21’ and ‘near #21’ of DBW notebooks.
</paragraph>
<paragraph id="903536BE0948FFBB51F6FA940281FAFA" blockId="41.[151,1437,1350,2024]" box="[151,982,1350,1376]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB51F6FA94007FFAFA" box="[151,296,1350,1376]" italics="true" pageId="41" pageNumber="42">Distribution.</emphasis>
Texas—restricted to
<collectingRegion id="524EF85C0948FFBB5377FA940300FAFA" box="[534,599,1350,1376]" country="United States of America" name="Texas" pageId="41" pageNumber="42">Texas</collectingRegion>
Gulf coast and up to
<quantity id="57729B5B0948FFBB5229FA9402D6FAFA" box="[840,897,1350,1376]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="41" pageNumber="42" unit="km" value="8.0">8 km</quantity>
inland.
</paragraph>
<paragraph id="903536BE0948FFBB51A6FAB900D1F9C6" blockId="41.[151,1437,1350,2024]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB51A6FAB9036AFA1E" box="[199,573,1386,1412]" italics="true" pageId="41" pageNumber="42">Recognition characters and song</emphasis>
. Small to medium sized (
<figureCitation id="08B12A3B0948FFBB5201FAB802E5FA1E" box="[864,946,1386,1412]" captionStart="FIGURE 24" captionStartId="42.[151,250,1147,1172]" captionTargetBox="[185,1420,271,1104]" captionTargetId="figure@42.[151,1435,265,1128]" captionTargetPageId="42" captionText="FIGURE 24. Holotype male (left), G. thinos. Female (right) from type locality showing typical, linear head stripes." figureDoi="http://doi.org/10.5281/zenodo.3563725" httpUri="https://zenodo.org/record/3563725/files/figure.png" pageId="41" pageNumber="42">Fig. 24</figureCitation>
), always short hind wings, medium length cerci, almost always in sandy habitats, males frequently sing from up in vegetation.
<emphasis id="A2FEEAAC0948FFBB555FFA5C0521FA32" box="[1086,1142,1422,1448]" italics="true" pageId="41" pageNumber="42">Song</emphasis>
(
<figureCitation id="08B12A3B0948FFBB55E5FA5C0580FA32" box="[1156,1239,1421,1448]" captionStart="FIGURE 25" captionStartId="43.[151,250,1827,1852]" captionTargetBox="[156,1402,642,1774]" captionTargetId="figure@43.[151,1435,625,1807]" captionTargetPageId="43" captionText="FIGURE 25. Calling song of G. thinos (R07-125) Cameron Co., TX (S07-25), at 25°C: (A) five second waveform, (B) one second spectrogram." figureDoi="http://doi.org/10.5281/zenodo.3563727" httpUri="https://zenodo.org/record/3563727/files/figure.png" pageId="41" pageNumber="42">Fig. 25</figureCitation>
; R07-125) a slow chirp, CR usually &lt;90, 4–7 p/c,
<collectingRegion id="524EF85C0948FFBB5372FA600362FA56" box="[531,565,1458,1484]" country="Brazil" name="Para" pageId="41" pageNumber="42">PR</collectingRegion>
20–35. Can be distinguished from only other
<collectingRegion id="524EF85C0948FFBB5529FA6005DEFA56" box="[1096,1161,1458,1484]" country="United States of America" name="Texas" pageId="41" pageNumber="42">Texas</collectingRegion>
Gulf coast, sandy-environment field cricket,
<taxonomicName id="578A4D3D0948FFBB50F2FA04030BFA6A" authority="(Texas)" baseAuthorityName="Texas" box="[403,604,1494,1520]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC0948FFBB50F2FA0400A9FA6A" box="[403,510,1494,1520]" italics="true" pageId="41" pageNumber="42">G. firmus</emphasis>
(
<collectingRegion id="524EF85C0948FFBB536FFA040303FA6A" box="[526,596,1494,1520]" country="United States of America" name="Texas" pageId="41" pageNumber="42">Texas</collectingRegion>
)
</taxonomicName>
, by DNA, non-overlapping file teeth, teeth/mm and a generally lower
<collectingRegion id="524EF85C0948FFBB541BFA0404CBFA6A" box="[1402,1436,1494,1520]" country="Brazil" name="Para" pageId="41" pageNumber="42">PR</collectingRegion>
(
<tableCitation id="DD0803050948FFBB51FEFA2801A5F98E" box="[159,242,1530,1556]" captionStart="TABLE 1" captionStartId="17.[150,238,466,492]" captionTargetPageId="17" captionText="TABLE 1. Summary measurements for morphological and song characters for Gryllus species discussed in this paper." httpUri="http://table.plazi.org/id/C4F566360970FF8351F7FE0004D1FE76" pageId="41" pageNumber="42" tableUuid="C4F566360970FF8351F7FE0004D1FE76">Table 1</tableCitation>
, p. 18). Singing males of
<taxonomicName id="578A4D3D0948FFBB536EFA280323F98E" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[527,628,1530,1556]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC0948FFBB536EFA280323F98E" box="[527,628,1530,1556]" italics="true" pageId="41" pageNumber="42">G. thinos</emphasis>
</taxonomicName>
also easier to approach than singing, microsympatric
<taxonomicName id="578A4D3D0948FFBB55A8FA280465F98E" box="[1225,1330,1530,1556]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC0948FFBB55A8FA280465F98E" box="[1225,1330,1530,1556]" italics="true" pageId="41" pageNumber="42">G. firmus</emphasis>
</taxonomicName>
. Females of the two taxa can be difficult to separate although
<taxonomicName id="578A4D3D0948FFBB5391F9CC020BF9A2" box="[752,860,1566,1592]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC0948FFBB5391F9CC020BF9A2" box="[752,860,1566,1592]" italics="true" pageId="41" pageNumber="42">G. firmus</emphasis>
</taxonomicName>
ovipositor length is generally longer in that larger species (see
<tableCitation id="DD0803050948FFBB5042F9900023F9C6" box="[291,372,1602,1628]" captionStart="TABLE 1" captionStartId="17.[150,238,466,492]" captionTargetPageId="17" captionText="TABLE 1. Summary measurements for morphological and song characters for Gryllus species discussed in this paper." httpUri="http://table.plazi.org/id/C4F566360970FF8351F7FE0004D1FE76" pageId="41" pageNumber="42" tableUuid="C4F566360970FF8351F7FE0004D1FE76">Table 1</tableCitation>
).
</paragraph>
<paragraph id="903536BE0948FFBB51A6F9B402C9F976" blockId="41.[151,1437,1350,2024]" pageId="41" pageNumber="42">
<materialsCitation id="20E23CE30948FFBB51A6F9B402C9F976" ID-GBIF-Occurrence="2464588162" collectingDate="2011-06-11" collectionCode="MK, CAS" collectorName="D. B. Weissman. &amp; S GenBank &amp; Right &amp; Type &amp; Entomology Type" country="United States of America" county="Kleberg Co." location="Padre Island National Seashore" municipality="Male" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Texas" typeStatus="holotype">
<emphasis id="A2FEEAAC0948FFBB51A6F9B40079F91A" box="[199,302,1638,1664]" italics="true" pageId="41" pageNumber="42">
<typeStatus id="4F31881C0948FFBB51A6F9B40079F91A" box="[199,302,1638,1664]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="7051ACC40948FFBB505DF9B40021F91A" box="[316,374,1638,1664]" pageId="41" pageNumber="42">Male</collectingMunicipality>
(
<figureCitation id="08B12A3B0948FFBB50E7F9B4008DF91A" box="[390,474,1638,1664]" captionStart="FIGURE 24" captionStartId="42.[151,250,1147,1172]" captionTargetBox="[185,1420,271,1104]" captionTargetId="figure@42.[151,1435,265,1128]" captionTargetPageId="42" captionText="FIGURE 24. Holotype male (left), G. thinos. Female (right) from type locality showing typical, linear head stripes." figureDoi="http://doi.org/10.5281/zenodo.3563725" httpUri="https://zenodo.org/record/3563725/files/figure.png" pageId="41" pageNumber="42">Fig. 24</figureCitation>
).
<collectingCountry id="E89D762E0948FFBB508FF9B4037EF91A" box="[494,553,1638,1664]" name="United States of America" pageId="41" pageNumber="42">USA</collectingCountry>
,
<collectingRegion id="524EF85C0948FFBB5355F9B4032FF91A" box="[564,632,1638,1664]" country="United States of America" name="Texas" pageId="41" pageNumber="42">Texas</collectingRegion>
,
<collectingCounty id="79544E320948FFBB53E5F9B40258F91A" box="[644,783,1638,1664]" pageId="41" pageNumber="42">Kleberg Co.</collectingCounty>
,
<location id="955560650948FFBB527AF9B4052CF91A" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB527AF9B4052CF91A" box="[795,1147,1638,1664]" country="United States of America" county="Kleberg Co." municipality="Male" name="Padre Island National Seashore" pageId="41" pageNumber="42" stateProvince="Texas">
<location id="955560650948FFBB527AF9B402F1F91A" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB527AF9B402F1F91A" box="[795,934,1638,1664]" country="United States of America" county="Kleberg Co." municipality="Male" name="Padre Island" pageId="41" pageNumber="42" stateProvince="Texas">Padre Island</location>
National Seashore
</location>
around
<location id="955560650948FFBB55B9F9B401B4F93E" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB55B9F9B401B4F93E" country="United States of America" county="Kleberg Co." municipality="Male" name="Malaquite Visitor Center" pageId="41" pageNumber="42" stateProvince="Texas">Malaquite Visitor Center</location>
,
<date id="E434107E0948FFBB5190F958003AF93E" box="[241,365,1674,1700]" pageId="41" pageNumber="42" value="2011-06-11">
<collectingDate id="F470E9960948FFBB5190F958003AF93E" box="[241,365,1674,1700]" pageId="41" pageNumber="42" value="2011-06-11">11-vi-2011</collectingDate>
</date>
, 15’. 27° 25’ 23.2” -97° 18’ 07.7”.
<collectorName id="3D7F53680948FFBB5264F95802EBF93E" box="[773,956,1674,1700]" pageId="41" pageNumber="42">D.B. Weissman.</collectorName>
S11-35, R11-8, DNA sample G2018. 16
<collectorName id="3D7F53680948FFBB54ECF9580056F952" pageId="41" pageNumber="42">S GenBank</collectorName>
accession #
<accessionNumber id="8FD9AB5D0948FFBB50E9F97C035AF952" box="[392,525,1710,1736]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MK446632" pageId="41" pageNumber="42">
<collectionCode id="F69BAE7B0948FFBB50E9F97C00E2F952" box="[392,437,1710,1736]" country="Kenya" name="National Museum of Kenya" pageId="41" pageNumber="42">MK</collectionCode>
446632
</accessionNumber>
. BL 17.28, HF11.53, LC 10.76.
<collectorName id="3D7F53680948FFBB5214F97C02E4F952" box="[885,947,1710,1736]" pageId="41" pageNumber="42">Right</collectorName>
tegmen removed: 119 teeth, file length 3.1, TL 10.0, TW 4.6.
<collectorName id="3D7F53680948FFBB5001F90000CFF976" box="[352,408,1746,1772]" pageId="41" pageNumber="42">Type</collectorName>
deposited in
<collectionCode id="F69BAE7B0948FFBB534FF9000330F976" box="[558,615,1746,1772]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="41" pageNumber="42" type="Museum">CAS</collectionCode>
,
<collectorName id="3D7F53680948FFBB5312F900026CF976" box="[627,827,1746,1772]" pageId="41" pageNumber="42">Entomology Type</collectorName>
#19274.
</materialsCitation>
</paragraph>
<paragraph id="903536BE0948FFB851A6F925052AFF61" blockId="41.[151,1437,1350,2024]" lastBlockId="42.[151,1436,152,251]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<materialsCitation id="20E23CE30948FFBB51A6F9250085F8AE" ID-GBIF-Occurrence="2464588150" collectingDate="2013-07-12" county="Texas. Aransas Co." location="Total" municipality="Rockport" pageId="41" pageNumber="42" specimenCount="82" specimenCount-female="14" specimenCount-male="68" typeStatus="paratype">
<emphasis id="A2FEEAAC0948FFBB51A6F925006EF88A" box="[199,313,1783,1808]" italics="true" pageId="41" pageNumber="42">
<typeStatus id="4F31881C0948FFBB51A6F925006EF88A" box="[199,313,1783,1808]" pageId="41" pageNumber="42" type="paratype">Paratypes</typeStatus>
</emphasis>
. (
<location id="955560650948FFBB502EF92400D9F88A" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB502EF92400D9F88A" box="[335,398,1782,1808]" county="Texas. Aransas Co." municipality="Rockport" name="Total" pageId="41" pageNumber="42">Total</location>
:
<specimenCount id="868CFD370948FFBB50F8F924009CF88A" box="[409,459,1782,1808]" pageId="41" pageNumber="42" type="male">67♂</specimenCount>
<specimenCount id="868CFD370948FFBB50B3F9240354F88A" box="[466,515,1782,1808]" pageId="41" pageNumber="42" type="female">14♀</specimenCount>
)
<collectingCounty id="79544E320948FFBB5375F92403A7F88A" box="[532,752,1782,1808]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB5375F9240300F88A" bold="true" box="[532,599,1782,1808]" pageId="41" pageNumber="42">Texas</emphasis>
.
<emphasis id="A2FEEAAC0948FFBB5307F92403BCF88A" box="[614,747,1782,1808]" italics="true" pageId="41" pageNumber="42">Aransas Co</emphasis>
.
</collectingCounty>
,
<location id="955560650948FFBB5261F9240233F88A" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB5261F9240233F88A" box="[768,868,1782,1808]" county="Texas. Aransas Co." municipality="Rockport" name="Business" pageId="41" pageNumber="42">Business</location>
35 near
<collectingMunicipality id="7051ACC40948FFBB52A6F9240565F88A" box="[967,1074,1782,1808]" pageId="41" pageNumber="42">Rockport</collectingMunicipality>
, 10’,
<date id="E434107E0948FFBB5513F92405AFF88A" box="[1138,1272,1782,1808]" pageId="41" pageNumber="42" value="2013-07-12">
<collectingDate id="F470E9960948FFBB5513F92405AFF88A" box="[1138,1272,1782,1808]" pageId="41" pageNumber="42" value="2013-07-12">12-vii-2013</collectingDate>
</date>
, 27° 57’ 32.7” -97° 06’ 14.4” (S13-55)
<specimenCount id="868CFD370948FFBB50CDF8C80085F8AE" box="[428,466,1818,1844]" pageId="41" pageNumber="42" type="male">1♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE30948FFBB50BFF8C80446F8C2" ID-GBIF-Occurrence="2464588051" collectingDate="1991-06-03" collectingDateMax="2013-07-12" collectingDateMin="1991-06-03" county="Cameron Co." location="Texas" pageId="41" pageNumber="42" specimenCount="8" specimenCount-female="1" specimenCount-male="7" typeStatus="paratype">
<location id="955560650948FFBB50BFF8C80348F8AE" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB50BFF8C80348F8AE" box="[478,543,1818,1844]" county="Cameron Co." name="Texas" pageId="41" pageNumber="42">Texas</location>
188
<locationDeviation id="81872F9C0948FFBB5338F8C80257F8AE" box="[601,768,1818,1844]" location="Hwy" pageId="41" pageNumber="42">
<quantity id="57729B5B0948FFBB5338F8C803CDF8AE" box="[601,666,1818,1844]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="41" pageNumber="42" unit="m" value="0.2">0.2 m</quantity>
SE Hwy
</locationDeviation>
35, 19’,
<date id="E434107E0948FFBB5209F8C802B8F8AE" box="[872,1007,1818,1844]" pageId="41" pageNumber="42" value="2013-07-12">
<collectingDate id="F470E9960948FFBB5209F8C802B8F8AE" box="[872,1007,1818,1844]" pageId="41" pageNumber="42" value="2013-07-12">12-vii-2013</collectingDate>
</date>
, 27° 57’ 56.1 -97° 07’ 13.7” (S13-54)
<specimenCount id="868CFD370948FFBB51F6F8EC01EAF8C2" box="[151,189,1854,1880]" pageId="41" pageNumber="42" type="male">2♂</specimenCount>
.
<collectingCounty id="79544E320948FFBB51A8F8EC0037F8C2" box="[201,352,1854,1880]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB51A8F8EC000DF8C2" box="[201,346,1854,1880]" italics="true" pageId="41" pageNumber="42">Cameron Co</emphasis>
.
</collectingCounty>
, Boca Chica State Park, 10’, 25° 59.827” -97° 09.146”
<date id="E434107E0948FFBB52BEF8EC051AF8C2" box="[991,1101,1854,1880]" pageId="41" pageNumber="42" value="1991-06-03">
<collectingDate id="F470E9960948FFBB52BEF8EC051AF8C2" box="[991,1101,1854,1880]" pageId="41" pageNumber="42" value="1991-06-03">3-vi-1991</collectingDate>
</date>
(S91-36)
<specimenCount id="868CFD370948FFBB55A0F8EF05B3F8C2" box="[1217,1252,1853,1880]" pageId="41" pageNumber="42" type="male">5♂</specimenCount>
<specimenCount id="868CFD370948FFBB558CF8EC0446F8C2" box="[1261,1297,1854,1880]" pageId="41" pageNumber="42" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="20E23CE30948FFBB5441F8EC0004F8E6" ID-GBIF-Occurrence="2464588046" collectingDate="2007-06-10" collectingDateMax="2013-07-12" collectingDateMin="1991-06-03" county="Cameron Co." isTailEnumeration="true" location="Texas" pageId="41" pageNumber="42" specimenCount="10" specimenCount-female="4" specimenCount-male="6" typeStatus="paratype">
<date id="E434107E0948FFBB5441F8EC04CBF8C2" box="[1312,1436,1854,1880]" pageId="41" pageNumber="42" value="2007-06-10">
<collectingDate id="F470E9960948FFBB5441F8EC04CBF8C2" box="[1312,1436,1854,1880]" pageId="41" pageNumber="42" value="2007-06-10">10-vi-2007</collectingDate>
</date>
(S07-25)
<specimenCount id="868CFD370948FFBB5062F8B00071F8E6" box="[259,294,1890,1916]" pageId="41" pageNumber="42" type="male">6♂</specimenCount>
<specimenCount id="868CFD370948FFBB504FF8B00004F8E6" box="[302,339,1890,1916]" pageId="41" pageNumber="42" type="female">4♀</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE30948FFBB503EF8B004CFF8E6" ID-GBIF-Occurrence="2464588169" box="[351,1432,1889,1916]" collectingDate="2013-07-10" country="Iceland" location="South Padre Island" municipality="Blanca Park" pageId="41" pageNumber="42" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<location id="955560650948FFBB503EF8B00360F8E6" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB503EF8B00360F8E6" box="[351,567,1890,1916]" country="Iceland" municipality="Blanca Park" name="South Padre Island" pageId="41" pageNumber="42">South Padre Island</location>
,
<collectingCountry id="E89D762E0948FFBB5325F8B0033BF8E6" box="[580,620,1890,1916]" name="Iceland" pageId="41" pageNumber="42">Isla</collectingCountry>
<collectingMunicipality id="7051ACC40948FFBB5315F8B00257F8E6" box="[628,768,1890,1916]" pageId="41" pageNumber="42">Blanca Park</collectingMunicipality>
, 5’,
<date id="E434107E0948FFBB5253F8B002EEF8E6" box="[818,953,1890,1916]" pageId="41" pageNumber="42" value="2013-07-10">
<collectingDate id="F470E9960948FFBB5253F8B002EEF8E6" box="[818,953,1890,1916]" pageId="41" pageNumber="42" value="2013-07-10">10-vii-2013</collectingDate>
</date>
, 26° 04” 09.3’ -97° 09” 41.7 (S13-41)
<specimenCount id="868CFD370948FFBB5413F8B004CFF8E6" box="[1394,1432,1890,1916]" pageId="41" pageNumber="42" type="male">3♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE30948FFBB51F6F85401EAF85E" ID-GBIF-Occurrence="2464588180" collectingDate="2011-06-11" country="Iceland" county="Kleberg Co." location="Padre Island National Seashore" municipality="Park" pageId="41" pageNumber="42" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCounty id="79544E320948FFBB51F6F854004AF83A" box="[151,285,1926,1952]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB51F6F854004FF83A" box="[151,280,1926,1952]" italics="true" pageId="41" pageNumber="42">Kleberg Co</emphasis>
.
</collectingCounty>
,
<location id="955560650948FFBB504DF85403DBF83A" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB504DF85403DBF83A" box="[300,652,1926,1952]" country="Iceland" county="Kleberg Co." municipality="Park" name="Padre Island National Seashore" pageId="41" pageNumber="42">Padre Island National Seashore</location>
, near
<collectingMunicipality id="7051ACC40948FFBB53ACF8540257F83A" box="[717,768,1926,1952]" pageId="41" pageNumber="42">Park</collectingMunicipality>
entrance,
<date id="E434107E0948FFBB5212F85402B8F83A" box="[883,1007,1926,1952]" pageId="41" pageNumber="42" value="2011-06-11">
<collectingDate id="F470E9960948FFBB5212F85402B8F83A" box="[883,1007,1926,1952]" pageId="41" pageNumber="42" value="2011-06-11">11-vi-2011</collectingDate>
</date>
, 27° 28” 39.9’ -97° 16” 28.7’ (S11-34)
<specimenCount id="868CFD370948FFBB51F6F87801EAF85E" box="[151,189,1962,1988]" pageId="41" pageNumber="42" type="male">1♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE30948FFBB51ABF8780263F85E" ID-GBIF-Occurrence="2464588039" box="[202,820,1961,1988]" collectingDate="2011-06-11" country="Iceland" location="Malaquite Visitor Center" pageId="41" pageNumber="42" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paratype">
<location id="955560650948FFBB51ABF87800B1F85E" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB51ABF87800B1F85E" box="[202,486,1962,1988]" country="Iceland" name="Malaquite Visitor Center" pageId="41" pageNumber="42">Malaquite Visitor Center</location>
,
<date id="E434107E0948FFBB5095F8780339F85E" box="[500,622,1962,1988]" pageId="41" pageNumber="42" value="2011-06-11">
<collectingDate id="F470E9960948FFBB5095F8780339F85E" box="[500,622,1962,1988]" pageId="41" pageNumber="42" value="2011-06-11">11-vi-2011</collectingDate>
</date>
(S11-35)
<specimenCount id="868CFD370948FFBB5382F8780251F85E" box="[739,774,1962,1988]" pageId="41" pageNumber="42" type="male">2♂</specimenCount>
<specimenCount id="868CFD370948FFBB5271F8780263F85E" box="[784,820,1962,1988]" pageId="41" pageNumber="42" type="female">3♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="20E23CE30948FFBB5225F87801A3F872" ID-GBIF-Occurrence="2464588075" collectingDate="1991-06-02" country="Iceland" elevation="3" location="Malaquite Visitor Center" pageId="41" pageNumber="42" specimenCount="16" specimenCount-female="5" specimenCount-male="11" typeStatus="paratype">
<quantity id="57729B5B0948FFBB5225F8780226F85E" box="[836,881,1962,1988]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="41" pageNumber="42" unit="m" value="3.0">
<elevation id="1BA7D18D0948FFBB5225F8780226F85E" box="[836,881,1962,1988]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="41" pageNumber="42" unit="m" value="3.0">3 m</elevation>
</quantity>
N
<location id="955560650948FFBB52F6F87805E4F85E" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB52F6F87805E4F85E" box="[919,1203,1962,1988]" country="Iceland" name="Malaquite Visitor Center" pageId="41" pageNumber="42">Malaquite Visitor Center</location>
,
<date id="E434107E0948FFBB55A0F8780478F85E" box="[1217,1327,1962,1988]" pageId="41" pageNumber="42" value="1991-06-02">
<collectingDate id="F470E9960948FFBB55A0F8780478F85E" box="[1217,1327,1962,1988]" pageId="41" pageNumber="42" value="1991-06-02">2-vi-1991</collectingDate>
</date>
(S91-33)
<specimenCount id="868CFD370948FFBB51F6F81C0190F872" box="[151,199,1998,2024]" pageId="41" pageNumber="42" type="male">11♂</specimenCount>
<specimenCount id="868CFD370948FFBB51B1F81F01A3F872" box="[208,244,1997,2024]" pageId="41" pageNumber="42" type="female">5♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="20E23CE30948FFBB5062F81C03FCF872" ID-GBIF-Occurrence="2464588053" box="[259,683,1998,2024]" collectingDate="1991-06-02" country="Iceland" location="South Beach" pageId="41" pageNumber="42" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<location id="955560650948FFBB5062F81C00C1F872" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB5062F81C00C1F872" box="[259,406,1998,2024]" country="Iceland" name="South Beach" pageId="41" pageNumber="42">South Beach</location>
,
<date id="E434107E0948FFBB50C3F81C0346F872" box="[418,529,1998,2024]" pageId="41" pageNumber="42" value="1991-06-02">
<collectingDate id="F470E9960948FFBB50C3F81C0346F872" box="[418,529,1998,2024]" pageId="41" pageNumber="42" value="1991-06-02">2-vi-1991</collectingDate>
</date>
(S91-34)
<specimenCount id="868CFD370948FFBB53E4F81C03FCF872" box="[645,683,1998,2024]" pageId="41" pageNumber="42" type="male">2♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE30948FFB853D9F81C00B5FF4D" ID-GBIF-Occurrence="2464588033" collectingDate="1986-06-29" collectingDateMax="2013-07-13" collectingDateMin="1986-06-29" collectorName="S. Padre Island Dr." country="Iceland" county="Matagorda Co." lastPageId="42" lastPageNumber="43" location="Hog Island" pageId="41" pageNumber="42" specimenCount="25" specimenCount-female="1" specimenCount-male="24" typeStatus="paratype">
<collectingCounty id="79544E320948FFBB53D9F81C0233F872" box="[696,868,1998,2024]" pageId="41" pageNumber="42">
<emphasis id="A2FEEAAC0948FFBB53D9F81C0208F872" box="[696,863,1998,2024]" italics="true" pageId="41" pageNumber="42">Matagorda Co</emphasis>
.
</collectingCounty>
,
<location id="955560650948FFBB5214F81C02A2F872" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:955560650948FFBB5214F81C02A2F872" box="[885,1013,1998,2024]" country="Iceland" county="Matagorda Co." name="Hog Island" pageId="41" pageNumber="42">Hog Island</location>
, 21’,
<date id="E434107E0948FFBB5557F81C05EAF872" box="[1078,1213,1998,2024]" pageId="41" pageNumber="42" value="2013-07-13">
<collectingDate id="F470E9960948FFBB5557F81C05EAF872" box="[1078,1213,1998,2024]" pageId="41" pageNumber="42" value="2013-07-13">13-vii-2013</collectingDate>
</date>
, 28° 36’ 17.4” -95° 57’ 34.8” (S13-60)
<specimenCount id="868CFD37094BFFB85014FF4B00CCFF29" box="[373,411,153,179]" pageId="42" pageNumber="43" type="male">4♂</specimenCount>
.
<emphasis id="A2FEEAAC094BFFB850C6FF480376FF29" box="[423,545,153,179]" italics="true" pageId="42" pageNumber="43">Nueces Co</emphasis>
., Corpus Christi near 10421
<collectorName id="3D7F5368094BFFB8520DFF4B0511FF29" box="[876,1094,153,179]" pageId="42" pageNumber="43">S. Padre Island Dr.</collectorName>
, 27° 40’ 02.5” -97° 16’ 54.4”,
<date id="E434107E094BFFB851F6FF6F0043FF4D" box="[151,276,189,215]" pageId="42" pageNumber="43" value="1986-06-29">
<collectingDate id="F470E996094BFFB851F6FF6F0043FF4D" box="[151,276,189,215]" pageId="42" pageNumber="43" value="1986-06-29">29-vi-1986</collectingDate>
</date>
(S86-57)
<specimenCount id="868CFD37094BFFB850E7FF6F00E1FF4D" box="[390,438,189,215]" pageId="42" pageNumber="43" type="male">20♂</specimenCount>
<specimenCount id="868CFD37094BFFB850DFFF6F00B5FF4D" box="[446,482,189,215]" pageId="42" pageNumber="43" type="female">1♀</specimenCount>
</materialsCitation>
;
<materialsCitation id="20E23CE3094BFFB85091FF6F025BFF4D" ID-GBIF-Occurrence="2464588042" box="[496,780,188,215]" collectingDate="2013-07-12" collectingDateMax="2013-07-13" collectingDateMin="1986-06-29" collectorName="S. Padre Island Dr." country="Iceland" county="Matagorda Co." isTailEnumeration="true" location="Hog Island" pageId="42" pageNumber="43" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<date id="E434107E094BFFB85091FF6F0323FF4D" box="[496,628,189,215]" pageId="42" pageNumber="43" value="2013-07-12">
<collectingDate id="F470E996094BFFB85091FF6F0323FF4D" box="[496,628,189,215]" pageId="42" pageNumber="43" value="2013-07-12">12-vii-2013</collectingDate>
</date>
(S13-53)
<specimenCount id="868CFD37094BFFB85387FF6F025BFF4D" box="[742,780,189,215]" pageId="42" pageNumber="43" type="male">1♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE3094BFFB85279FF6F009AFF61" ID-GBIF-Occurrence="2464588173" collectingDate="2013-07-12" country="Iceland" location="Mustang Island State Park" pageId="42" pageNumber="43" specimenCount="7" specimenCount-male="7" typeStatus="paratype">
<location id="95556065094BFFB85279FF6F0515FF4D" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:95556065094BFFB85279FF6F0515FF4D" box="[792,1090,189,215]" country="Iceland" name="Mustang Island State Park" pageId="42" pageNumber="43">Mustang Island State Park</location>
, 5’,
<date id="E434107E094BFFB85512FF6F05ADFF4D" box="[1139,1274,189,215]" pageId="42" pageNumber="43" value="2013-07-12">
<collectingDate id="F470E996094BFFB85512FF6F05ADFF4D" box="[1139,1274,189,215]" pageId="42" pageNumber="43" value="2013-07-12">12-vii-2013</collectingDate>
</date>
, 27° 40’ 25.9” -97° 10’ 30.1” (S13-51)
<specimenCount id="868CFD37094BFFB850C9FF33009AFF61" box="[424,461,225,251]" pageId="42" pageNumber="43" type="male">7♂</specimenCount>
</materialsCitation>
.
<materialsCitation id="20E23CE3094BFFB850B8FF33052EFF61" ID-GBIF-Occurrence="2464588058" box="[473,1145,224,251]" collectingDate="2013-07-12" country="Iceland" elevation="2" location="Hwy" municipality="Port Aransas" pageId="42" pageNumber="43" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<location id="95556065094BFFB850B8FF330358FF61" LSID="urn:lsid:plazi:treatment:182387A80948FFBF51F6FBA103CDFF61:95556065094BFFB850B8FF330358FF61" box="[473,527,225,251]" country="Iceland" municipality="Port Aransas" name="Hwy" pageId="42" pageNumber="43">Hwy</location>
361
<quantity id="57729B5B094BFFB85326FF330325FF61" box="[583,626,225,251]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2.0">
<elevation id="1BA7D18D094BFFB85326FF330325FF61" box="[583,626,225,251]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="42" pageNumber="43" unit="m" value="2.0">2 m</elevation>
</quantity>
S
<collectingMunicipality id="7051ACC4094BFFB853F1FF330276FF61" box="[656,801,225,251]" pageId="42" pageNumber="43">Port Aransas</collectingMunicipality>
,
<date id="E434107E094BFFB8524DFF3302E5FF61" box="[812,946,225,251]" pageId="42" pageNumber="43" value="2013-07-12">
<collectingDate id="F470E996094BFFB8524DFF3302E5FF61" box="[812,946,225,251]" pageId="42" pageNumber="43" value="2013-07-12">12-vii-2013</collectingDate>
</date>
, 15’ (S13-52)
<specimenCount id="868CFD37094BFFB85532FF33052EFF61" box="[1107,1145,225,251]" pageId="42" pageNumber="43" type="male">2♂</specimenCount>
</materialsCitation>
.
</paragraph>
<caption id="C4F56636094BFFB851F6FBA9047BFB0E" ID-DOI="http://doi.org/10.5281/zenodo.3563725" ID-Zenodo-Dep="3563725" box="[151,1324,1147,1172]" httpUri="https://zenodo.org/record/3563725/files/figure.png" pageId="42" pageNumber="43" startId="42.[151,250,1147,1172]" targetBox="[185,1420,271,1104]" targetPageId="42">
<paragraph id="903536BE094BFFB851F6FBA9047BFB0E" blockId="42.[151,1324,1147,1172]" box="[151,1324,1147,1172]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851F6FBA90077FB0E" bold="true" box="[151,288,1147,1172]" pageId="42" pageNumber="43">FIGURE 24.</emphasis>
Holotype male (left),
<emphasis id="A2FEEAAC094BFFB85363FBAE0331FB0E" box="[514,614,1148,1172]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="578A4D3D094BFFB85363FBAE0336FB0E" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[514,609,1148,1172]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="thinos">G. thinos</taxonomicName>
.
</emphasis>
Female (right) from type locality showing typical, linear head stripes.
</paragraph>
</caption>
<paragraph id="903536BE094BFFB851A6FB1104D3FB47" blockId="42.[151,1437,1218,2037]" box="[199,1412,1218,1245]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6FB1100FEFB46" box="[199,425,1218,1244]" italics="true" pageId="42" pageNumber="43">Derivation of name.</emphasis>
“thinos” is Greek for beach, shore, or strand in recognition of where this cricket occurs.
</paragraph>
<paragraph id="903536BE094BFFB851A6FB340475FAD2" blockId="42.[151,1437,1218,2037]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6FB3400FDFA9A" box="[199,426,1254,1280]" italics="true" pageId="42" pageNumber="43">Geographical range</emphasis>
.
<figureCitation id="08B12A3B094BFFB850D6FB34035EFA9A" box="[439,521,1254,1280]" captionStart="FIGURE 26" captionStartId="44.[151,250,1282,1307]" captionTargetBox="[151,1058,174,1214]" captionTargetId="figure@44.[151,1058,174,1263]" captionTargetPageId="44" captionText="FIGURE 26. Known distribution of G. thinos." figureDoi="http://doi.org/10.5281/zenodo.3563729" httpUri="https://zenodo.org/record/3563729/files/figure.png" pageId="42" pageNumber="43">Fig. 26</figureCitation>
. Known only from coastal and barrier island areas of Texas and inland for no more than
<quantity id="57729B5B094BFFB851AEFAD8005CFABE" box="[207,267,1290,1316]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="42" pageNumber="43" unit="km" value="8.0">8 km</quantity>
. May get into adjacent coastal
<collectingCountry id="E89D762E094BFFB85309FAD803E9FABE" box="[616,702,1290,1316]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
but areas not checked due to limited access. We did find suitable sandy areas just south of Tampico,
<collectingCountry id="E89D762E094BFFB85340FAFC032EFAD2" box="[545,633,1326,1352]" name="Mexico" pageId="42" pageNumber="43">Mexico</collectingCountry>
, and around Galveston, Texas, but no
<taxonomicName id="578A4D3D094BFFB85545FAFD05DDFAD2" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[1060,1162,1326,1353]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094BFFB85545FAFD05DDFAD2" box="[1060,1162,1326,1353]" italics="true" pageId="42" pageNumber="43">G. thinos</emphasis>
</taxonomicName>
at either site.
</paragraph>
<paragraph id="903536BE094BFFB851A6FA800043F94F" blockId="42.[151,1437,1218,2037]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6FA80004AFAF6" box="[199,285,1362,1388]" italics="true" pageId="42" pageNumber="43">Habitat</emphasis>
. Almost always on sandy substrate in open, low rolling coastal dunes. In Corpus Christi (S86-57 and S13-53), somewhat sandy-substrate vacant field with grasses, forbs, and small shrubs (all three vegetation
<typeStatus id="4F31881C094BFFB85400FAA504CBFA0B" box="[1377,1436,1399,1425]" pageId="42" pageNumber="43">types</typeStatus>
from
<quantity id="57729B5B094BFFB851B5FA480000FA2F" box="[212,343,1434,1461]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="42" pageNumber="43" unit="cm" value="25.0" valueMax="40.0" valueMin="10.0">10 to 40 cm</quantity>
tall), males almost always singing from vegetation (but never at top of plants) and initially visible on triangulating but quickly climbed (not jumped) down. Also heard throughout town of Corpus Christi in
<quantity id="57729B5B094BFFB85426FA6C04CBFA42" box="[1351,1436,1470,1496]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="42" pageNumber="43" unit="in" value="1986.0">1986 in</quantity>
road median areas, mowed lawns, etc. At Padre Island National Seashore (S91-33), common, with some in plants, but many just along mowed edges of road in vegetation
<quantity id="57729B5B094BFFB8526BF9D70216F9BB" box="[778,833,1541,1569]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="42" pageNumber="43" unit="cm" value="5.0">5 cm</quantity>
tall and singing with females also walking around. At Padre Island National Seashore (S11-34), nymphs and one adult male collected under boards just above high tide line. At Mustang Island State Park (S13-51), several reddish/pale mid instar nymphs under boards on low rolling dunes during daytime. At night at Mustang Island, all 7 singing males
<quantity id="57729B5B094BFFB852FDF9A30549F917" box="[924,1054,1649,1677]" metricMagnitude="-1" metricUnit="m" metricValue="3.05" metricValueMax="4.6" metricValueMin="1.5" pageId="42" pageNumber="43" unit="cm" value="30.5" valueMax="46.0" valueMin="15.0">15 to 46 cm</quantity>
up in grass, dead twigs, and
<taxonomicName id="578A4D3D094BFFB8543BF9A104CBF916" box="[1370,1436,1651,1676]" class="Liliopsida" family="Asparagaceae" genus="Yucca" kingdom="Plantae" order="Asparagales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="A2FEEAAC094BFFB8543BF9A104CBF916" box="[1370,1436,1651,1676]" italics="true" pageId="42" pageNumber="43">Yucca</emphasis>
</taxonomicName>
spp. Two of those males quickly climbed down and retreated into nearby holes in the sand and were easily flushed with water.
</paragraph>
<paragraph id="903536BE094BFFB851A6F90C0523F8FF" blockId="42.[151,1437,1218,2037]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6F90C0302F962" box="[199,597,1758,1784]" italics="true" pageId="42" pageNumber="43">Life cycle and seasonal occurrence</emphasis>
. No egg diapause: Corpus Christi (S86-57). Probably one generation/year but many late spring and early summer mid-instar nymphs, at Padre Island National Seashore (S91-33) and Mustang Island State Park (S13-51), either represent a second generation or, more likely, show that maturation occurs over several months. Mid instar nymphs from those two localities molted to adults in August.
</paragraph>
<paragraph id="903536BE094BFFB851A6F8BD0273F84A" blockId="42.[151,1437,1218,2037]" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6F8BD0060F812" box="[199,311,1903,1928]" italics="true" pageId="42" pageNumber="43">Variation.</emphasis>
<emphasis id="A2FEEAAC094BFFB8505FF8BF00DBF812" bold="true" box="[318,396,1901,1928]" pageId="42" pageNumber="43">Color:</emphasis>
tegmina almost equally divided between brown and black. Rear leg femur and tibia usually tan/orange but also black.
<emphasis id="A2FEEAAC094BFFB850DDF84100ABF837" bold="true" box="[444,508,1939,1965]" pageId="42" pageNumber="43">Head</emphasis>
:
<specimenCount id="868CFD37094BFFB8536BF8400332F836" box="[522,613,1938,1964]" pageId="42" pageNumber="43" type="adult">13 adult</specimenCount>
males (out of 67) and
<specimenCount id="868CFD37094BFFB85201F84002FBF836" box="[864,940,1938,1964]" pageId="42" pageNumber="43" type="adult">3 adult</specimenCount>
females (out of 14) with defined linear head stripes (
<figureCitation id="08B12A3B094BFFB85191F8640014F84A" box="[240,323,1974,2000]" captionStart="FIGURE 24" captionStartId="42.[151,250,1147,1172]" captionTargetBox="[185,1420,271,1104]" captionTargetId="figure@42.[151,1435,265,1128]" captionTargetPageId="42" captionText="FIGURE 24. Holotype male (left), G. thinos. Female (right) from type locality showing typical, linear head stripes." figureDoi="http://doi.org/10.5281/zenodo.3563725" httpUri="https://zenodo.org/record/3563725/files/figure.png" pageId="42" pageNumber="43">Fig. 24</figureCitation>
), a character also seen in some late instars.
</paragraph>
<paragraph id="903536BE094BFFB951A6F80900A1FF4D" blockId="42.[151,1437,1218,2037]" lastBlockId="43.[151,1436,152,575]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<emphasis id="A2FEEAAC094BFFB851A6F80901A8F86F" box="[199,255,2011,2037]" italics="true" pageId="42" pageNumber="43">DNA</emphasis>
. Multilocus G1209 (Boca Chica State Park, TX, S07-25) maps (Gray
<emphasis id="A2FEEAAC094BFFB85571F809051FF86E" box="[1040,1096,2010,2036]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2019) with
<specimenCount id="868CFD37094BFFB855AEF8080472F86E" box="[1231,1317,2010,2036]" pageId="42" pageNumber="43" type="male">2 males</specimenCount>
from Cor- pus Christi, TX (G2018, G2022 from S11-35). Surprisingly, the single, fall generation, egg diapausing
<taxonomicName id="578A4D3D094AFFB9547FFF4B04CBFF29" box="[1310,1436,153,179]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="ovisopis">
<emphasis id="A2FEEAAC094AFFB9547FFF4B04CBFF29" box="[1310,1436,153,179]" italics="true" pageId="43" pageNumber="44">G. ovisopis</emphasis>
</taxonomicName>
appears to be the sister species.
</paragraph>
<paragraph id="903536BE094AFFB951A6FF33051DFDA5" blockId="43.[151,1436,152,575]" pageId="43" pageNumber="44">
Much DNA variation between habitats although no morphologically corresponding characters. When South Padre Island specimens (S11-35) were genetically compared with those from Boca Chica (S07-25), less than
<quantity id="57729B5B094AFFB95413FED701ECFED9" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="43" pageNumber="44" unit="km" value="200.0">200 km</quantity>
south and in a similar habitat, we found, for CO1, 17 fixed nucleotide differences in 700+ base pairs; and for 16S, 6 fixed nucleotide differences in 500+ base pairs. We are concerned, however, that the COI variation might be an artifact of pseudogenes (see general DNA discussion, pages 16 &amp; 17). ITS2 also shows very slight separation within
<taxonomicName id="578A4D3D094AFFB95186FE470019FE35" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[231,334,405,431]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094AFFB95186FE470019FE35" box="[231,334,405,431]" italics="true" pageId="43" pageNumber="44">G. thinos</emphasis>
</taxonomicName>
from these localities (
<figureCitation id="08B12A3B094AFFB9532AFE4703F6FE35" box="[587,673,405,431]" captionStart="FIGURE 27" captionStartId="44.[151,250,1738,1763]" captionTargetBox="[151,1388,1322,1703]" captionTargetId="figure@44.[151,1435,1322,1719]" captionTargetPageId="44" captionText="FIGURE 27. ITS2 gene tree showing some separation between Padre Island and Boca Chica samples of G. thinos." figureDoi="http://doi.org/10.5281/zenodo.3563731" httpUri="https://zenodo.org/record/3563731/files/figure.png" pageId="43" pageNumber="44">Fig. 27</figureCitation>
). Further analysis might support species status for these genetically very different populations but in a comparison of their morphological characters (see
<tableCitation id="DD080305094AFFB9552CFE6B05C8FE49" box="[1101,1183,441,467]" captionStart="TABLE 1" captionStartId="17.[150,238,466,492]" captionTargetPageId="17" captionText="TABLE 1. Summary measurements for morphological and song characters for Gryllus species discussed in this paper." httpUri="http://table.plazi.org/id/C4F566360970FF8351F7FE0004D1FE76" pageId="43" pageNumber="44" tableUuid="C4F566360970FF8351F7FE0004D1FE76">Table 1</tableCitation>
, p. 18), nothing stands out. Plus, all 3 individuals are each other’s nearest neighbor in multilocus tree. The Boca Chica habitat, while appearing very similar to Padre Island, is somehow ecologically different in that the former locality is also inhabited by microsympatric
<taxonomicName id="578A4D3D094AFFB95010FDF7008CFDA5" box="[369,475,549,575]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC094AFFB95010FDF7008CFDA5" box="[369,475,549,575]" italics="true" pageId="43" pageNumber="44">G. firmus</emphasis>
</taxonomicName>
, while no other
<taxonomicName id="578A4D3D094AFFB953F0FDF703B2FDA5" authorityName="Linnaeus" authorityYear="1758" box="[657,741,549,575]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="A2FEEAAC094AFFB953F0FDF703B2FDA5" box="[657,741,549,575]" italics="true" pageId="43" pageNumber="44">Gryllus</emphasis>
</taxonomicName>
species occurs on Padre Island.
</paragraph>
<caption id="C4F56636094AFFB951F6F8F1003FF8FA" ID-DOI="http://doi.org/10.5281/zenodo.3563727" ID-Zenodo-Dep="3563727" httpUri="https://zenodo.org/record/3563727/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,250,1827,1852]" targetBox="[156,1402,642,1774]" targetPageId="43">
<paragraph id="903536BE094AFFB951F6F8F1003FF8FA" blockId="43.[151,1437,1827,1888]" pageId="43" pageNumber="44">
<emphasis id="A2FEEAAC094AFFB951F6F8F10075F8A6" bold="true" box="[151,290,1827,1852]" pageId="43" pageNumber="44">FIGURE 25.</emphasis>
Calling song of
<taxonomicName id="578A4D3D094AFFB950B0F8F60367F8A1" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[465,560,1827,1851]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094AFFB950B0F8F60367F8A1" box="[465,560,1827,1851]" italics="true" pageId="43" pageNumber="44">G. thinos</emphasis>
</taxonomicName>
(R07-125) Cameron Co., TX (S07-25), at 25°C: (A) five second waveform, (B) one second spectrogram.
</paragraph>
</caption>
<caption id="C4F56636094DFFBE51F6FAD00327FA81" ID-DOI="http://doi.org/10.5281/zenodo.3563729" ID-Zenodo-Dep="3563729" box="[151,624,1282,1307]" httpUri="https://zenodo.org/record/3563729/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1282,1307]" targetBox="[151,1058,174,1214]" targetPageId="44">
<paragraph id="903536BE094DFFBE51F6FAD00327FA81" blockId="44.[151,624,1282,1307]" box="[151,624,1282,1307]" pageId="44" pageNumber="45">
<emphasis id="A2FEEAAC094DFFBE51F6FAD00077FA81" bold="true" box="[151,288,1282,1307]" pageId="44" pageNumber="45">FIGURE 26.</emphasis>
Known distribution of
<taxonomicName id="578A4D3D094DFFBE536FFAD1033CFA81" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[526,619,1283,1307]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094DFFBE536FFAD1033CFA81" box="[526,619,1283,1307]" italics="true" pageId="44" pageNumber="45">G. thinos</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="C4F56636094DFFBE51F6F9180473F978" ID-DOI="http://doi.org/10.5281/zenodo.3563731" ID-Zenodo-Dep="3563731" box="[151,1316,1738,1763]" httpUri="https://zenodo.org/record/3563731/files/figure.png" pageId="44" pageNumber="45" startId="44.[151,250,1738,1763]" targetBox="[151,1388,1322,1703]" targetPageId="44">
<paragraph id="903536BE094DFFBE51F6F9180473F978" blockId="44.[151,1316,1738,1763]" box="[151,1316,1738,1763]" pageId="44" pageNumber="45">
<emphasis id="A2FEEAAC094DFFBE51F6F9180077F979" bold="true" box="[151,288,1738,1763]" pageId="44" pageNumber="45">FIGURE 27.</emphasis>
ITS2 gene tree showing some separation between Padre Island and Boca Chica samples of
<taxonomicName id="578A4D3D094DFFBE55A0F9190449F979" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[1217,1310,1739,1763]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094DFFBE55A0F9190449F979" box="[1217,1310,1739,1763]" italics="true" pageId="44" pageNumber="45">G. thinos</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="903536BE094DFFBE51A6F8C30201F80D" blockId="44.[151,1437,1809,2015]" pageId="44" pageNumber="45">
The
<collectingRegion id="524EF85C094DFFBE519BF8C3006CF8B1" box="[250,315,1809,1835]" country="United States of America" name="Texas" pageId="44" pageNumber="45">Texas</collectingRegion>
beach inhabiting crickets,
<taxonomicName id="578A4D3D094DFFBE530BF8C30386F8B1" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[618,721,1809,1835]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094DFFBE530BF8C30386F8B1" box="[618,721,1809,1835]" italics="true" pageId="44" pageNumber="45">G. thinos</emphasis>
</taxonomicName>
and
<taxonomicName id="578A4D3D094DFFBE5268F8C30286F8B1" authority="(Texas)" baseAuthorityName="Texas" box="[777,977,1809,1835]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC094DFFBE5268F8C30224F8B1" box="[777,883,1809,1835]" italics="true" pageId="44" pageNumber="45">G. firmus</emphasis>
(
<collectingRegion id="524EF85C094DFFBE52E3F8C3029FF8B1" box="[898,968,1809,1835]" country="United States of America" name="Texas" pageId="44" pageNumber="45">Texas</collectingRegion>
)
</taxonomicName>
, are clearly genetically distinct: Tamura- Nei genetic distance within
<taxonomicName id="578A4D3D094DFFBE50ADF8E70366F8D5" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[460,561,1845,1871]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094DFFBE50ADF8E70366F8D5" box="[460,561,1845,1871]" italics="true" pageId="44" pageNumber="45">G. thinos</emphasis>
</taxonomicName>
ITS2 samples is low (N = 6, mean ±
<collectingRegion id="524EF85C094DFFBE52ADF8E702A5F8D5" box="[972,1010,1845,1871]" country="Serbia" name="Sid" pageId="44" pageNumber="45">SD</collectingRegion>
, 0.0044 ± 0.0037), similar to variation within
<taxonomicName id="578A4D3D094DFFBE5184F88B00FFF8E9" authority="(Texas)" baseAuthorityName="Texas" box="[229,424,1881,1907]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC094DFFBE5184F88B0019F8E9" box="[229,334,1881,1907]" italics="true" pageId="44" pageNumber="45">G. firmus</emphasis>
(
<collectingRegion id="524EF85C094DFFBE503DF88B00F7F8E9" box="[348,416,1881,1907]" country="United States of America" name="Texas" pageId="44" pageNumber="45">Texas</collectingRegion>
)
</taxonomicName>
samples (N = 12, mean ±
<collectingRegion id="524EF85C094DFFBE53A8F88B03B8F8E9" box="[713,751,1881,1907]" country="Serbia" name="Sid" pageId="44" pageNumber="45">SD</collectingRegion>
, 0.0034 ± 0.0040), but an order of magnitude greater between
<taxonomicName id="578A4D3D094DFFBE51F6F8AF01AAF80D" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[151,253,1917,1943]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094DFFBE51F6F8AF01AAF80D" box="[151,253,1917,1943]" italics="true" pageId="44" pageNumber="45">G. thinos</emphasis>
</taxonomicName>
and
<taxonomicName id="578A4D3D094DFFBE5055F8AF00AEF80D" authority="(Texas)" baseAuthorityName="Texas" box="[308,505,1917,1943]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC094DFFBE5055F8AF00C8F80D" box="[308,415,1917,1943]" italics="true" pageId="44" pageNumber="45">G. firmus</emphasis>
(
<collectingRegion id="524EF85C094DFFBE50CCF8AF00A6F80D" box="[429,497,1917,1943]" country="United States of America" name="Texas" pageId="44" pageNumber="45">Texas</collectingRegion>
)
</taxonomicName>
(0.0240 ± 0.0027 mean ±
<collectingRegion id="524EF85C094DFFBE5240F8AF021EF80D" box="[801,841,1917,1943]" country="Serbia" name="Sid" pageId="44" pageNumber="45">SD</collectingRegion>
).
</paragraph>
<paragraph id="903536BE094DFFBE51A6F8700400F845" blockId="44.[151,1437,1809,2015]" pageId="44" pageNumber="45">
<emphasis id="A2FEEAAC094DFFBE51A6F870001DF821" box="[199,330,1954,1979]" italics="true" pageId="44" pageNumber="45">Discussion.</emphasis>
Singing males easy to approach. On
<date id="E434107E094DFFBE5391F8730238F821" box="[752,879,1953,1979]" pageId="44" pageNumber="45" value="2007-06-10">10-vi-2007</date>
, males not singing well until 21:45, considerably after sunset, and without a significant moon. Oatmeal trails in dunes helped to attract wandering individuals.
</paragraph>
<paragraph id="903536BE094CFFBF51A6FF4B03CDFF61" blockId="45.[151,1437,153,251]" pageId="45" pageNumber="46">
Besides finding
<taxonomicName id="578A4D3D094CFFBF501CFF4B00B5FF29" authorityName="Weissman &amp; Gray" authorityYear="2019" box="[381,482,153,179]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="thinos">
<emphasis id="A2FEEAAC094CFFBF501CFF4B00B5FF29" box="[381,482,153,179]" italics="true" pageId="45" pageNumber="46">G. thinos</emphasis>
</taxonomicName>
microsympatric with
<taxonomicName id="578A4D3D094CFFBF53B9FF4B0215FF29" box="[728,834,153,179]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="firmus">
<emphasis id="A2FEEAAC094CFFBF53B9FF4B0215FF29" box="[728,834,153,179]" italics="true" pageId="45" pageNumber="46">G. firmus</emphasis>
</taxonomicName>
at Boca Chica, we also found them together near Port Aransas (S13-52), and Mustang Island State Park (S13-51). Singing males of the two species are easily separated in the field by the slower pulse rate in
<emphasis id="A2FEEAAC094CFFBF5349FF3303CDFF61" box="[552,666,225,251]" italics="true" pageId="45" pageNumber="46">
<taxonomicName id="578A4D3D094CFFBF5349FF3303C1FF61" box="[552,662,225,251]" class="Insecta" family="Haglotettigoniidae" genus="Gryllus" kingdom="Animalia" order="Orthoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="firmus">G. firmus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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