treatments-xml/data/BF/3A/87/BF3A87E0FFF1FFCDFF5EFE7819BCFAE4.xml

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<mods:title id="95B6916E0D1CFB6BCE744C594C8ACE65">Taxonomy and distribution of Cylindera germanica (Linnaeus, 1758) in the Middle East with the description of two new subspecies (Coleoptera, Carabidae Cicindelinae)</mods:title>
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<mods:namePart id="A6DA36A740BB0D61F089443EA67D0CEC">Boetzl, Fabian A.</mods:namePart>
<mods:affiliation id="D335046C80BF751D4A0C232405309ECB">Department of Animal Ecology and Tropical Biology, Biocenter, University of Würzburg, Am Hubland, 97074 Würzburg, Germany fabian. boetzl @ uni-wuerzburg. de; https: // orcid. org / 0000 - 0001 - 5121 - 3370</mods:affiliation>
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<subSubSection id="7F89657DFFF1FFCFFF5EFE781E09FE79" box="[151,1079,405,432]" pageId="9" pageNumber="104" type="nomenclature">
<paragraph id="372C36F6FFF1FFCFFF5EFE781E09FE79" blockId="9.[151,1079,405,432]" box="[151,1079,405,432]" pageId="9" pageNumber="104">
<heading id="6C64819AFFF1FFCFFF5EFE781E09FE79" bold="true" box="[151,1079,405,432]" fontSize="11" level="1" pageId="9" pageNumber="104" reason="1">
<emphasis id="05E7EAE4FFF1FFCFFF5EFE781E09FE79" bold="true" box="[151,1079,405,432]" pageId="9" pageNumber="104">
<taxonomicName id="F0934D75FFF1FFCFFF5EFE7818A9FE79" ID-CoL="8MMX9" authority="Boetzl &amp; Franzen, 2020" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[151,663,405,432]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="subSpecies" species="germanica" status="subsp. nov." subGenus="Cylindera" subSpecies="persiana">
<emphasis id="05E7EAE4FFF1FFCFFF5EFE781B30FE66" bold="true" box="[151,270,405,432]" italics="true" pageId="9" pageNumber="104">Cylindera</emphasis>
(
<emphasis id="05E7EAE4FFF1FFCFFED6FE781BA8FE66" bold="true" box="[287,406,405,432]" italics="true" pageId="9" pageNumber="104">Cylindera</emphasis>
) 
<emphasis id="05E7EAE4FFF1FFCFFE6EFE7818A9FE79" bold="true" box="[423,663,405,431]" italics="true" pageId="9" pageNumber="104">germanica persiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED4579FFFF1FFCFFD56FE781936FE79" box="[671,776,405,431]" pageId="9" pageNumber="104" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
Boetzl &amp; Franzen, 2020
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7F89657DFFF1FFCFFF5EFE3018CBFBE1" pageId="9" pageNumber="104" type="materials_examined">
<paragraph id="372C36F6FFF1FFCFFF5EFE301968FE21" blockId="9.[151,1437,476,2015]" box="[151,854,476,503]" pageId="9" pageNumber="104">
<emphasis id="05E7EAE4FFF1FFCFFF5EFE301B07FE21" bold="true" box="[151,313,476,503]" pageId="9" pageNumber="104">Type locality:</emphasis>
<collectingCountry id="4F847666FFF1FFCFFE89FE301B4EFE21" box="[320,368,477,503]" name="Iran" pageId="9" pageNumber="104">Iran</collectingCountry>
, 
<collectingRegion id="F557F814FFF1FFCFFEB2FE30187FFE21" box="[379,577,477,503]" country="Iran" name="Kerman" pageId="9" pageNumber="104">Kerman province</collectingRegion>
, Dschiroft, Anbar-Abad.
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EFDED1B44FD25" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
<emphasis id="05E7EAE4FFF1FFCFFF0EFDED1B96FDCD" bold="true" box="[199,424,512,539]" pageId="9" pageNumber="104">Material examined</emphasis>
: 
<emphasis id="05E7EAE4FFF1FFCFFE7CFDEC181CFDCD" bold="true" box="[437,546,513,539]" pageId="9" pageNumber="104">
<typeStatus id="E8288854FFF1FFCFFE7CFDEC181CFDCD" box="[437,546,513,539]" pageId="9" pageNumber="104" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<specimenCount id="2195FD7FFFF1FFCFFDF9FDEC1879FDCD" box="[560,583,513,539]" pageId="9" pageNumber="104" type="male">♂</specimenCount>
, ‘S.O. 
<collectingCountry id="4F847666FFF1FFCFFD5BFDEC18E7FDCD" box="[658,729,513,539]" name="Iran" pageId="9" pageNumber="104">IRAN</collectingCountry>
, [Djiroft], Anbar-Abad, 
<date id="432D1036FFF1FFCFFC2DFDEC1EA5FDCD" box="[996,1179,512,539]" pageId="9" pageNumber="104" value="1956-04-21" valueMax="1956-04-30" valueMin="1956-04-21">21. - 30.IV.1956</date>
, W. Richter leg.’ [black print on greyish-green cardboard label]; ‘GK 13’ [black ink written on mounting board]; ‘Cic. 
<taxonomicName id="F0934D75FFF1FFCFFB73FDCB1F0CFDE9" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1210,1330,550,575]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF1FFCFFB73FDCB1F0CFDE9" box="[1210,1330,550,575]" italics="true" pageId="9" pageNumber="104">germanica</emphasis>
</taxonomicName>
<taxonomicName id="F0934D75FFF1FFCFFAF0FDC81B37FDB5" authority="Fisch." authorityName="Fisch." class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF1FFCFFAF0FDC81A82FDB5" italics="true" pageId="9" pageNumber="104">descendens</emphasis>
Fisch.
</taxonomicName>
, det. Ing. K. 
<bibRefCitation id="53024B07FFF1FFCFFE5FFDA4181EFDB5" author="Mandl, K." box="[406,544,584,611]" pageId="9" pageNumber="104" pagination="1 - 3" refId="ref10348" refString="Mandl, K. (1959) Eine Ausbeute an Cicindeliden aus Iran. Ergebnisse der Entomologischen Reisen Willi Richter, Stuttgart, im Iran 1954 und 1956. Nr. 19. Stuttgarter Beitrage zur Naturkunde, 18, 1 - 3." type="journal article" year="1959">Mandl 1959</bibRefCitation>
’ [black print on yellowish cardboard label, ‘ 
<taxonomicName id="F0934D75FFF1FFCFFBC1FDA71EBEFDB5" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1032,1152,586,611]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF1FFCFFBC1FDA71EBEFDB5" box="[1032,1152,586,611]" italics="true" pageId="9" pageNumber="104">germanica</emphasis>
</taxonomicName>
<taxonomicName id="F0934D75FFF1FFCFFB4FFDA41F6DFDB5" authority="Fisch." authorityName="Fisch." box="[1158,1363,585,611]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF1FFCFFB4FFDA41F38FDB5" box="[1158,1286,585,611]" italics="true" pageId="9" pageNumber="104">descendens</emphasis>
Fisch.
</taxonomicName>
’ hand- written in black ink]; ‘MUSEUM STUTTGART’ [black print on white label]; ‘ 
<taxonomicName id="F0934D75FFF1FFCFFBE9FD801EAAFD51" box="[1056,1172,621,647]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="genus">Cylindera</taxonomicName>
(s.str.), obliquefasciata oblique-, 
<taxonomicName id="F0934D75FFF1FFCFFECAFD7C1835FD7D" authority="(Adams, 1817)" baseAuthorityName="Adams" baseAuthorityYear="1817" box="[259,523,657,683]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="fasciata">fasciata (Adams, 1817)</taxonomicName>
, det. J. 
<bibRefCitation id="53024B07FFF1FFCFFDABFD7C18CFFD7D" author="Gebert, J." box="[610,753,657,683]" pageId="9" pageNumber="104" pagination="217 - 222" refId="ref9936" refString="Gebert, J. (1992) Zur Taxonomie, Systematik und Verbreitung der Cylindera &quot; obliquefasciata &quot; - Gruppe (Col., Cicindelidae). Entomologische Nachrichten und Berichte, 39, 217 - 222." type="journal article" year="1992">Gebert 1992</bibRefCitation>
’ [black print on yellowish cardboard label]; ‘ 
<typeStatus id="E8288854FFF1FFCFFB23FD7C1FA6FD7D" box="[1258,1432,657,683]" pageId="9" pageNumber="104" type="holotype">HOLOTYPUS</typeStatus>
, 
<taxonomicName id="F0934D75FFF1FFCFFF5EFD581845FD19" authority="Boetzl &amp; Franzen, 2020" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[151,635,693,719]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="subSpecies" species="germanica" status="subsp. nov." subGenus="Cylindera" subSpecies="persiana">
<emphasis id="05E7EAE4FFF1FFCFFF5EFD581845FD19" box="[151,635,693,719]" italics="true" pageId="9" pageNumber="104">Cylindera (Cylindera) germanica persiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED4579FFFF1FFCFFD4AFD5818D9FD19" box="[643,743,693,719]" pageId="9" pageNumber="104" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
, des. F. Boetzl &amp; M. Franzen 2020’ (black print on red label) deposited in SMNS;
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EFD1018CBFBE1" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
<emphasis id="05E7EAE4FFF1FFCFFF0EFD101B01FCC1" bold="true" box="[199,319,765,791]" pageId="9" pageNumber="104">
<typeStatus id="E8288854FFF1FFCFFF0EFD101B01FCC1" box="[199,319,765,791]" pageId="9" pageNumber="104" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="2195FD7FFFF1FFCFFE86FD101B4AFCC1" box="[335,372,765,791]" pageId="9" pageNumber="104" type="female">2♀</specimenCount>
, ‘ 
<collectingCountry id="4F847666FFF1FFCFFE4FFD101BECFCC1" box="[390,466,765,791]" name="Iran" pageId="9" pageNumber="104">IRAN</collectingCountry>
, Khuzistan, Shadegan, 
<date id="432D1036FFF1FFCFFD14FD1019A4FCC1" box="[733,922,764,791]" pageId="9" pageNumber="104" value="1956-03-26" valueMax="1956-03-31" valueMin="1956-03-26">26. - 31.III.1956</date>
, Richter u. Schäuffele’, [black print on grey- ish-green cardboard label]; ‘Cic. 
<taxonomicName id="F0934D75FFF1FFCFFDC5FCCF18BAFCED" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[524,644,802,827]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF1FFCFFDC5FCCF18BAFCED" box="[524,644,802,827]" italics="true" pageId="9" pageNumber="104">germanica</emphasis>
</taxonomicName>
<taxonomicName id="F0934D75FFF1FFCFFD42FCCC1967FCED" authority="Fisch." authorityName="Fisch." box="[651,857,801,827]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF1FFCFFD42FCCC1935FCED" box="[651,779,801,827]" italics="true" pageId="9" pageNumber="104">descendens</emphasis>
Fisch.
</taxonomicName>
, det. Ing. K. 
<bibRefCitation id="53024B07FFF1FFCFFC22FCCC1E48FCED" author="Mandl, K." box="[1003,1142,800,827]" pageId="9" pageNumber="104" pagination="1 - 3" refId="ref10348" refString="Mandl, K. (1959) Eine Ausbeute an Cicindeliden aus Iran. Ergebnisse der Entomologischen Reisen Willi Richter, Stuttgart, im Iran 1954 und 1956. Nr. 19. Stuttgarter Beitrage zur Naturkunde, 18, 1 - 3." type="journal article" year="1959">Mandl 1959</bibRefCitation>
’ [black print on yellowish cardboard label, ‘ 
<taxonomicName id="F0934D75FFF1FFCFFE95FCAB1BEAFC89" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[348,468,838,863]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF1FFCFFE95FCAB1BEAFC89" box="[348,468,838,863]" italics="true" pageId="9" pageNumber="104">germanica</emphasis>
</taxonomicName>
<taxonomicName id="F0934D75FFF1FFCFFE12FCA81897FC89" authority="Fisch." authorityName="Fisch." box="[475,681,837,863]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF1FFCFFE12FCA81865FC89" box="[475,603,837,863]" italics="true" pageId="9" pageNumber="104">descendens</emphasis>
Fisch.
</taxonomicName>
’ handwritten in black ink]; ‘MUSEUM STUTTGART’ [black print on white label]; ‘ 
<taxonomicName id="F0934D75FFF1FFCFFE93FC841BF0FC55" box="[346,462,873,899]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="genus">Cylindera</taxonomicName>
(s.str.), obliquefasciata oblique-, 
<taxonomicName id="F0934D75FFF1FFCFFC86FC841E65FC55" authority="(Adams, 1817)" baseAuthorityName="Adams" baseAuthorityYear="1817" box="[847,1115,873,899]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="species" species="fasciata">fasciata (Adams, 1817)</taxonomicName>
, det. J. 
<bibRefCitation id="53024B07FFF1FFCFFB71FC841F76FC55" author="Gebert, J." box="[1208,1352,873,899]" pageId="9" pageNumber="104" pagination="217 - 222" refId="ref9936" refString="Gebert, J. (1992) Zur Taxonomie, Systematik und Verbreitung der Cylindera &quot; obliquefasciata &quot; - Gruppe (Col., Cicindelidae). Entomologische Nachrichten und Berichte, 39, 217 - 222." type="journal article" year="1992">Gebert 1992</bibRefCitation>
’ [black print on yellowish cardboard label]; ‘ 
<typeStatus id="E8288854FFF1FFCFFDF4FC6018DDFC71" box="[573,739,909,935]" pageId="9" pageNumber="104" type="paratype">PARATYPUS</typeStatus>
, 
<taxonomicName id="F0934D75FFF1FFCFFD39FC601EE9FC71" authority="Boetzl &amp; Franzen, 2020" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[752,1239,909,935]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="subSpecies" species="germanica" status="subsp. nov." subGenus="Cylindera" subSpecies="persiana">
<emphasis id="05E7EAE4FFF1FFCFFD39FC601EE9FC71" box="[752,1239,909,935]" italics="true" pageId="9" pageNumber="104">Cylindera (Cylindera) germanica persiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED4579FFFF1FFCFFB29FC601F78FC71" box="[1248,1350,909,935]" pageId="9" pageNumber="104" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
, des. F. Boetzl &amp; M. Franzen 2020’ (black print on red label), [
<collectionCode id="5182AE33FFF1FFCFFCC7FC5D195CFC1D" box="[782,866,944,971]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="9" pageNumber="104">SMNS</collectionCode>
]; 
<specimenCount id="2195FD7FFFF1FFCFFCBEFC5C19A5FC1D" box="[887,923,945,971]" pageId="9" pageNumber="104" type="female">2♀</specimenCount>
, ‘ 
<collectingCountry id="4F847666FFF1FFCFFC67FC5C19C4FC1D" box="[942,1018,945,971]" name="Iran" pageId="9" pageNumber="104">IRAN</collectingCountry>
, Busher prov., 
<quantity id="F06B9B13FFF1FFCFFB6EFC5D1ECEFC1D" box="[1191,1264,944,971]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="104" unit="km" value="50.0">50 km</quantity>
N Busher city, Borazjan, Khoshmakan, river side, 
<quantity id="F06B9B13FFF1FFCFFDE2FC391856FC39" box="[555,616,980,1007]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="104" unit="m" value="35.0">35 m</quantity>
, leg. S. Azadbakhsh, 
<date id="432D1036FFF1FFCFFCA9FC3819DFFC39" box="[864,993,981,1007]" pageId="9" pageNumber="104" value="2018-03-21">21.III.2018</date>
’ [black print on white label, collection numbers CI 01643 &amp; CI 01644], ‘ 
<typeStatus id="E8288854FFF1FFCFFDC1FC141890FBC5" box="[520,686,1017,1043]" pageId="9" pageNumber="104" type="paratype">PARATYPUS</typeStatus>
, 
<taxonomicName id="F0934D75FFF1FFCFFD71FC141EABFBC5" authority="Boetzl &amp; Franzen, 2020" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[696,1173,1017,1043]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="subSpecies" species="germanica" status="subsp. nov." subGenus="Cylindera" subSpecies="persiana">
<emphasis id="05E7EAE4FFF1FFCFFD71FC141EABFBC5" box="[696,1173,1017,1043]" italics="true" pageId="9" pageNumber="104">Cylindera (Cylindera) germanica persiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED4579FFFF1FFCFFB52FC141EC3FBC5" box="[1179,1277,1017,1043]" pageId="9" pageNumber="104" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
, des. F. Boetzl &amp; M. Franzen 2020’ (black print on red label), [
<collectionCode id="5182AE33FFF1FFCFFD67FBF118D5FBE1" box="[686,747,1052,1079]" country="Canada" lsid="urn:lsid:biocol.org:col:13058" name="Northern Forestry Centre, Canadian Forest Service" pageId="9" pageNumber="104" type="Herbarium">CFB</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="7F89657DFFF1FFCCFF0EFBAD1A82FC39" lastPageId="10" lastPageNumber="105" pageId="9" pageNumber="104" type="description">
<paragraph id="372C36F6FFF1FFCFFF0EFBAD1AEAFBA9" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
<emphasis id="05E7EAE4FFF1FFCFFF0EFBAD1B6FFB8D" bold="true" box="[199,337,1088,1115]" pageId="9" pageNumber="104">Description</emphasis>
: Total length of the 
<typeStatus id="E8288854FFF1FFCFFDFEFBAC18A7FB8D" box="[567,665,1089,1115]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
male is 
<quantity id="F06B9B13FFF1FFCFFD31FBAC1962FB8D" box="[760,860,1089,1115]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="104" unit="mm" value="10.6">10.6 mm</quantity>
(type series: 
<specimenCount id="2195FD7FFFF1FFCFFC3AFBAC1E23FB8D" box="[1011,1053,1089,1115]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFBEDFBAC1EFAFB8D" box="[1060,1220,1089,1115]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="9.8" metricValueMin="9.2" pageId="9" pageNumber="104" unit="mm" value="9.5" valueMax="9.8" valueMin="9.2">9.2 to 9.8 mm</quantity>
, mean: 9.58 ± 
<quantity id="F06B9B13FFF1FFCFFAA2FBAC1AF9FBA9" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="104" unit="mm" value="0.13">0.13 mm</quantity>
).
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EFB6418D2FAE5" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
Labrum unidentate, yellowish brown except tip of spine black, with 7 submarginal setae (
<typeStatus id="E8288854FFF1FFCFFB0EFB641EC7FB75" box="[1223,1273,1161,1187]" pageId="9" pageNumber="104">type</typeStatus>
series: 7 to 9, mean: 8 ± 0.41), transverse (LL / LW: 
<specimenCount id="2195FD7FFFF1FFCFFD89FB401866FB11" box="[576,600,1197,1223]" pageId="9" pageNumber="104" type="male">♂</specimenCount>
: 0.59; 
<specimenCount id="2195FD7FFFF1FFCFFD68FB4018F5FB11" box="[673,715,1197,1223]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
0.54 to 0.60, mean: 0.55 ± 0.02), 
<quantity id="F06B9B13FFF1FFCFFB8BFB401EA9FB11" box="[1090,1175,1197,1223]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="104" unit="mm" value="0.8">0.8 mm</quantity>
long (
<quantity id="F06B9B13FFF1FFCFFB15FB411F7FFB11" box="[1244,1345,1196,1223]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="104" unit="mm" value="0.75">0.75 mm</quantity>
without spine; 
<typeStatus id="E8288854FFF1FFCFFF28FB3C1B2FFB3D" box="[225,273,1233,1259]" pageId="9" pageNumber="104">type</typeStatus>
series: 
<specimenCount id="2195FD7FFFF1FFCFFEA1FB3C1BACFB3D" box="[360,402,1233,1259]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFE50FB3D186BFB3D" box="[409,597,1232,1259]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="7.5" metricValueMin="6.5" pageId="9" pageNumber="104" unit="mm" value="0.7" valueMax="0.75" valueMin="0.65">0.65 to 0.75 mm</quantity>
, mean: 0.70 ± 
<quantity id="F06B9B13FFF1FFCFFD35FB3C195DFB3D" box="[764,867,1233,1259]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
; without spine: 
<quantity id="F06B9B13FFF1FFCFFBDFFB3D1EFAFB3D" box="[1046,1220,1232,1259]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="9" pageNumber="104" unit="mm" value="0.575" valueMax="0.6" valueMin="0.55">0.55 to 0.6 mm</quantity>
, mean: 0.59 ± 
<quantity id="F06B9B13FFF1FFCFFAA2FB3C1AFAFAD9" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="104" unit="mm" value="0.01">0.01 mm</quantity>
) and 
<quantity id="F06B9B13FFF1FFCFFEC8FB191B5BFAD9" box="[257,357,1268,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="9" pageNumber="104" unit="mm" value="1.35">1.35 mm</quantity>
wide (
<typeStatus id="E8288854FFF1FFCFFE66FB181BDFFAD9" box="[431,481,1269,1295]" pageId="9" pageNumber="104">type</typeStatus>
series: 
<specimenCount id="2195FD7FFFF1FFCFFDFFFB18185EFAD9" box="[566,608,1269,1295]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFDAFFB18193DFAD9" box="[614,771,1269,1295]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="9" pageNumber="104" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.2 to 1.3 mm</quantity>
, mean: 1.26 ± 
<quantity id="F06B9B13FFF1FFCFFC6FFB181E30FAD9" box="[934,1038,1269,1295]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
). Maxillary and labial palpi yellow- ish brown with the last segment noticeably darkened.
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EFAD018AEF9F9" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
Head wider than pronotum (
<typeStatus id="E8288854FFF1FFCFFDC1FAD0184EFA81" box="[520,624,1341,1367]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
: 
<quantity id="F06B9B13FFF1FFCFFDB2FAD018EDFA81" box="[635,723,1341,1367]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="9" pageNumber="104" unit="mm" value="2.9">2.9 mm</quantity>
wide; type series: 
<specimenCount id="2195FD7FFFF1FFCFFC60FAD019EDFA81" box="[937,979,1341,1367]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFC12FAD01E40FA81" box="[987,1150,1341,1367]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="9" pageNumber="104" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2 to 2.7 mm</quantity>
, mean: 2.53 ± 
<quantity id="F06B9B13FFF1FFCFFAE1FAD01FAEFA81" box="[1320,1424,1341,1367]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="9" pageNumber="104" unit="mm" value="0.11">0.11 mm</quantity>
), glabrous, metallic green sometimes with coppery sheen, longitudinally wrinkled with circular wrinkles on the frons typical for 
<taxonomicName id="F0934D75FFF1FFCFFEDAFA681BBDFA49" box="[275,387,1413,1439]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="05E7EAE4FFF1FFCFFEDAFA681BBDFA49" box="[275,387,1413,1439]" italics="true" pageId="9" pageNumber="104">Cylindera</emphasis>
</taxonomicName>
. Genae glabrous, metallic dark blue. Two supraorbital setae, one at the anterior margin of the eye, one at the widest point of the head between the eyes. Antennae reaching approximately to the end of the first third of the elytra, segments 1 to 4 glabrous, metallic green with coppery reflections, segments 5 to 11 light brown and densely setose with short, thin setae. Scapus with one long apical seta, antennal segment 3 with 6 shorter setae (
<typeStatus id="E8288854FFF1FFCFFF56F9F81B3AF9E6" box="[159,260,1557,1584]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
) apart from the longer apical setae.
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EF9D4194FF8FD" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">
Pronotum cylindrical, with 2.0 mm wider (type series: 
<specimenCount id="2195FD7FFFF1FFCFFCF9F9D41964F985" box="[816,858,1593,1619]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFCA8F9D51E24F985" box="[865,1050,1592,1619]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="9" pageNumber="104" unit="mm" value="1.9" valueMax="1.95" valueMin="1.85">1.85 to 1.95 mm</quantity>
- mean: 1.9 ± 
<quantity id="F06B9B13FFF1FFCFFB77F9D41F1AF985" box="[1214,1316,1593,1619]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="104" unit="mm" value="0.02">0.02 mm</quantity>
) than long (
<quantity id="F06B9B13FFF1FFCFFF56F9B11B39F9A1" box="[159,263,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="9" pageNumber="104" unit="mm" value="1.95">1.95 mm</quantity>
; type series: 
<specimenCount id="2195FD7FFFF1FFCFFE5EF9B01BFFF9A1" box="[407,449,1629,1655]" pageId="9" pageNumber="104" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF1FFCFFE01F9B018BFF9A1" box="[456,641,1628,1655]" metricMagnitude="-3" metricUnit="m" metricValue="1.775" metricValueMax="1.85" metricValueMin="1.7" pageId="9" pageNumber="104" unit="mm" value="1.775" valueMax="1.85" valueMin="1.7">1.70 to 1.85 mm</quantity>
, mean: 1.75 ± 
<quantity id="F06B9B13FFF1FFCFFCECF9B019B3F9A1" box="[805,909,1629,1655]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="104" unit="mm" value="0.04">0.04 mm</quantity>
), lateral edges setose, its sides glabrous. Dorsal colouration in 
<typeStatus id="E8288854FFF1FFCFFEF1F96C1BA4F94D" box="[312,410,1665,1691]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
male dark coppery with dull green sheen, in the 
<typeStatus id="E8288854FFF1FFCFFC78F96C1E2FF94D" box="[945,1041,1665,1691]" pageId="9" pageNumber="104" type="paratype">paratype</typeStatus>
series typically azure blueish green metallic. Pronotum disc convex, with clear midline and deep impressions forming a basal furrow near the posterior margin, wrinkled, especially near the anterior and posterior margins as well as around the midline. Sparsely setose along the basal furrow (
<figureCitation id="AFA82A73FFF1FFCFFE68F900182CF8D1" box="[417,530,1773,1799]" captionStart="FIGURES 20-23" captionStartId="10.[151,264,1953,1978]" captionTargetBox="[156,1432,1041,1923]" captionTargetId="figure@10.[151,1436,1029,1929]" captionTargetPageId="10" captionText="FIGURES 20-23: Cylindera germanica persiana ssp. nov. – (20) holotype (male), (21) paratype (female), (22) pronotum of female paratype, (23) adeagus of holotype male (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3933986" httpUri="https://zenodo.org/record/3933986/files/figure.png" pageId="9" pageNumber="104">Figure 22</figureCitation>
). Thoracic segments dark metallic blue, all segments glabrous except for the metathorax which is covered with a brush of thick short white setae.
</paragraph>
<paragraph id="372C36F6FFF1FFCFFF0EF8D81E0FF86D" blockId="9.[151,1437,476,2015]" pageId="9" pageNumber="104">Legs metallic green with dark coppery reflections except for the dark brown tibiae and trochanters, fore and middle femora with sparse white setae along the anterior and posterior margins and on both sides in between, hind femora without setae between he margins on the side that faces the abdomen. Fore and middle trochanters with a single, long subapical seta. Basal three tarsomeres of forelegs in males broadened.</paragraph>
<paragraph id="372C36F6FFF1FFCCFF0EF8281B13FD7D" blockId="9.[151,1437,476,2015]" lastBlockId="10.[151,1437,152,1007]" lastPageId="10" lastPageNumber="105" pageId="9" pageNumber="104">
Elytra elliptical, slightly dilated in apical third in females, more parallel in the 
<typeStatus id="E8288854FFF1FFCFFBF9F8281EACF809" box="[1072,1170,1989,2015]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
male (length: 
<typeStatus id="E8288854FFF1FFCFFAFBF8281FA6F809" box="[1330,1432,1989,2015]" pageId="9" pageNumber="104" type="holotype">holotype</typeStatus>
: 
<quantity id="F06B9B13FFF2FFCCFF5EFF741ACFFF65" box="[151,241,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="105" unit="mm" value="6.2">6.2 mm</quantity>
; type series: 
<specimenCount id="2195FD7FFFF2FFCCFE4FFF741B8EFF65" box="[390,432,153,179]" pageId="10" pageNumber="105" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF2FFCCFE71FF751865FF65" box="[440,603,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" metricValueMax="5.9" metricValueMin="5.5" pageId="10" pageNumber="105" unit="mm" value="5.7" valueMax="5.9" valueMin="5.5">5.5 to 5.9 mm</quantity>
, mean: 5.7 ± 
<quantity id="F06B9B13FFF2FFCCFD3EFF741961FF65" box="[759,863,153,179]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="105" unit="mm" value="0.08">0.08 mm</quantity>
; width (at broadest point): 
<typeStatus id="E8288854FFF2FFCCFB5AFF741EC7FF65" box="[1171,1273,153,179]" pageId="10" pageNumber="105" type="holotype">holotype</typeStatus>
: 
<quantity id="F06B9B13FFF2FFCCFACCFF741F61FF65" box="[1285,1375,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="10" pageNumber="105" unit="mm" value="3.7">3.7 mm</quantity>
; type series: 
<specimenCount id="2195FD7FFFF2FFCCFF2CFF501B31FF01" box="[229,271,189,215]" pageId="10" pageNumber="105" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF2FFCCFEDFFF501B8DFF01" box="[278,435,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="10" pageNumber="105" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.3 to 3.5 mm</quantity>
, mean: 3.44 ± 
<quantity id="F06B9B13FFF2FFCCFD9EFF511883FF01" box="[599,701,188,215]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="105" unit="mm" value="0.05">0.05 mm</quantity>
; width (at the tip of the scutellum): 
<typeStatus id="E8288854FFF2FFCCFB83FF501E8EFF01" box="[1098,1200,189,215]" pageId="10" pageNumber="105" type="holotype">holotype</typeStatus>
: 
<quantity id="F06B9B13FFF2FFCCFB72FF501F2DFF01" box="[1211,1299,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="105" unit="mm" value="3.1">3.1 mm</quantity>
; type series: 
<specimenCount id="2195FD7FFFF2FFCCFF5EFF0C1AFFFF2D" box="[151,193,225,251]" pageId="10" pageNumber="105" type="female">♀♀</specimenCount>
<quantity id="F06B9B13FFF2FFCCFF01FF0D1B4AFF2D" box="[200,372,224,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.825" metricValueMax="2.9" metricValueMin="2.75" pageId="10" pageNumber="105" unit="mm" value="2.825" valueMax="2.9" valueMin="2.75">2.75 to 2.9 mm</quantity>
, mean: 2.81 ± 
<quantity id="F06B9B13FFF2FFCCFDD0FF0C18BFFF2D" box="[537,641,225,251]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="105" unit="mm" value="0.03">0.03 mm</quantity>
). Scutellum dark coppery metallic. Elytral disc dark blackish grey with dark coppery to anthracite metallic sheen, with a faint black band on each elytron reaching curved from the humeral (shoulder) spot over the basal spot (endpoint of the not present apical lunule) and the end tip of the middle band towards the apical lunule (recognizable as area without metallic sheen), the rest of the elytra covered more or less with small bright turquoise to blue metallic dots, with a line of large bright turquoise blue dots along the suture and a group of the same dots at the shoulder. Shoulders prominent with small round maculation. Basal maculation (endpoint of the not present apical lunule) centrally on each elytron at the end of the first quarter variable in size but always present, the middle band reaching straight from a very large marginal spot that is slightly distanced from the lateral margin of the elytron towards the suture near the elytral apex, thinning out towards the middle of the elytron but ending in a clear hook projecting towards the elytral base. The apical lunule is starting at the elytral apex, running along the margin to a spot shortly before the broadest point of the elytron and then projecting in a hook towards the central point of the middle band. Epipleura mid brown. Elytral apex with a clear but short, coppery metallic sutural spine.
</paragraph>
<paragraph id="372C36F6FFF2FFCCFF0EFD581B57FD25" blockId="10.[151,1437,152,1007]" pageId="10" pageNumber="105">
Abdominal sternites glabrous, black, 1 
<superScript id="C0E69BBEFFF2FFCCFDBEFD5918BCFD14" attach="left" box="[631,642,692,706]" fontSize="6" pageId="10" pageNumber="105">st</superScript>
to 5 
<superScript id="C0E69BBEFFF2FFCCFD7AFD5918FEFD14" attach="left" box="[691,704,692,706]" fontSize="6" pageId="10" pageNumber="105">th</superScript>
segment with blue to violet sheen, 6 
<superScript id="C0E69BBEFFF2FFCCFB93FD591E59FD14" attach="left" box="[1114,1127,692,706]" fontSize="6" pageId="10" pageNumber="105">th</superScript>
segment more or less without metallic lustre.
</paragraph>
<paragraph id="372C36F6FFF2FFCCFF0EFD1018EEFC55" blockId="10.[151,1437,152,1007]" pageId="10" pageNumber="105">
The aedeagus generally resembles that of 
<taxonomicName id="F0934D75FFF2FFCCFD6AFD101983FCC1" authority="germanica" authorityName="germanica (Linnaeus" authorityYear="1758" box="[675,957,765,791]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="105" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="germanica">
<emphasis id="05E7EAE4FFF2FFCCFD6AFD101983FCC1" box="[675,957,765,791]" italics="true" pageId="10" pageNumber="105">C. germanica germanica</emphasis>
</taxonomicName>
in shape (
<figureCitation id="AFA82A73FFF2FFCCFBFBFD101E9AFCC1" box="[1074,1188,765,791]" captionStart="FIGURES 20-23" captionStartId="10.[151,264,1953,1978]" captionTargetBox="[156,1432,1041,1923]" captionTargetId="figure@10.[151,1436,1029,1929]" captionTargetPageId="10" captionText="FIGURES 20-23: Cylindera germanica persiana ssp. nov. – (20) holotype (male), (21) paratype (female), (22) pronotum of female paratype, (23) adeagus of holotype male (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3933986" httpUri="https://zenodo.org/record/3933986/files/figure.png" pageId="10" pageNumber="105">Figure 23</figureCitation>
). The aedeagus of the 
<typeStatus id="E8288854FFF2FFCCFF5EFCCC1AC7FCED" box="[151,249,801,827]" pageId="10" pageNumber="105" type="holotype">holotype</typeStatus>
is 
<quantity id="F06B9B13FFF2FFCCFED6FCCC1B49FCED" box="[287,375,801,827]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="10" pageNumber="105" unit="mm" value="3.3">3.3 mm</quantity>
long and embedded in Xylol and therefore bleached. Due to the previous embedding of the genital of the only accessible male, we refrained from extracting the genital and from inflating the internal sack for the sake of conservation of the 
<typeStatus id="E8288854FFF2FFCCFE3CFC841869FC55" box="[501,599,873,899]" pageId="10" pageNumber="105" type="holotype">holotype</typeStatus>
specimen.
</paragraph>
<paragraph id="372C36F6FFF2FFCCFF0EFC601A82FC39" blockId="10.[151,1437,152,1007]" pageId="10" pageNumber="105">
<typeStatus id="E8288854FFF2FFCCFF0EFC601B11FC71" box="[199,303,909,935]" pageId="10" pageNumber="105" type="holotype">Holotype</typeStatus>
glued onto yellowish cardboard mounting board, pinhole from previous mounting on right elytron, centrally at the end of the first fifth of the elytron, left foreleg lacking all tarsomeres, aedeagus embedded in Xy- lol.
</paragraph>
</subSubSection>
<caption id="63EC667EFFF2FFCCFF5EF84C1906F808" ID-DOI="http://doi.org/10.5281/zenodo.3933986" ID-Zenodo-Dep="3933986" httpUri="https://zenodo.org/record/3933986/files/figure.png" pageId="10" pageNumber="105" startId="10.[151,264,1953,1978]" targetBox="[156,1432,1041,1923]" targetPageId="10">
<paragraph id="372C36F6FFF2FFCCFF5EF84C1906F808" blockId="10.[151,1436,1953,2014]" pageId="10" pageNumber="105">
<emphasis id="05E7EAE4FFF2FFCCFF5EF84C1B6DF86C" bold="true" box="[151,339,1953,1978]" pageId="10" pageNumber="105">FIGURES 20-23:</emphasis>
<taxonomicName id="F0934D75FFF2FFCCFE92F84F18ABF86C" authority="Boetzl &amp; Franzen, 2020" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[347,661,1954,1978]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="105" phylum="Arthropoda" rank="subSpecies" species="germanica" status="subsp. nov." subSpecies="persiana">
<emphasis id="05E7EAE4FFF2FFCCFE92F84F18ABF86C" box="[347,661,1954,1978]" italics="true" pageId="10" pageNumber="105">Cylindera germanica persiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1ED4579FFFF2FFCCFD54F84F18C9F86F" box="[669,759,1954,1977]" pageId="10" pageNumber="105" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
– (20) holotype (male), (21) paratype (female), (22) pronotum of female paratype, (23) adeagus of holotype male (left lateral view).
</paragraph>
</caption>
<subSubSection id="7F89657DFFF3FFCDFF0EFF741B8AFF00" pageId="11" pageNumber="106" type="etymology">
<paragraph id="372C36F6FFF3FFCDFF0EFF741B8AFF00" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
<emphasis id="05E7EAE4FFF3FFCDFF0EFF741B76FF65" bold="true" box="[199,328,153,179]" pageId="11" pageNumber="106">Etymology</emphasis>
: The given name originates form the range of this new subspecies, the lowlands along the north coast of the Persian Gulf.
</paragraph>
</subSubSection>
<subSubSection id="7F89657DFFF3FFCDFF0EFF0D19BCFAE4" pageId="11" pageNumber="106" type="diagnosis">
<paragraph id="372C36F6FFF3FFCDFF0EFF0D19B0FEB1" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
<emphasis id="05E7EAE4FFF3FFCDFF0EFF0D1BF9FF2D" bold="true" box="[199,455,224,251]" pageId="11" pageNumber="106">Differential diagnosis</emphasis>
: This new subspecies resembles 
<taxonomicName id="F0934D75FFF3FFCDFCF4FF0C19E0FF2C" box="[829,990,224,251]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF3FFCDFCF4FF0C19E0FF2C" box="[829,990,224,251]" italics="true" pageId="11" pageNumber="106">C. descendens</emphasis>
</taxonomicName>
in elytral pattern and colouration but is easily differentiated by the lack of setae on the abdominal sternites and the sparsely setose front and hind edges of the pronotum (glabrous in 
<taxonomicName id="F0934D75FFF3FFCDFE0AFEC4185AFE94" box="[451,612,296,323]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF3FFCDFE0AFEC4185AFE94" box="[451,612,296,323]" italics="true" pageId="11" pageNumber="106">C. descendens</emphasis>
</taxonomicName>
). Due to these differences and an apparent co-existence of 
<taxonomicName id="F0934D75FFF3FFCDFB35FEC41FA3FE94" box="[1276,1437,296,323]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF3FFCDFB35FEC41FA3FE94" box="[1276,1437,296,323]" italics="true" pageId="11" pageNumber="106">C. descendens</emphasis>
</taxonomicName>
in the same region, the new subspecies is assigned to 
<taxonomicName id="F0934D75FFF3FFCDFD26FEA019B9FEB0" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[751,903,333,359]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF3FFCDFD26FEA019B9FEB0" box="[751,903,333,359]" italics="true" pageId="11" pageNumber="106">C. germanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="372C36F6FFF3FFCDFF0EFE9C19A8FE05" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
From 
<emphasis id="05E7EAE4FFF3FFCDFEC5FE9C1813FE5C" box="[268,557,369,395]" italics="true" pageId="11" pageNumber="106">
<taxonomicName id="F0934D75FFF3FFCDFEC5FE9C1817FE5C" authority="germanica" authorityName="germanica (Linnaeus" authorityYear="1758" box="[268,553,369,395]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="germanica">C. germanica germanica</taxonomicName>
,
</emphasis>
occurring in northern 
<collectingCountry id="4F847666FFF3FFCDFCE7FE9C1960FE5D" box="[814,862,369,395]" name="Iran" pageId="11" pageNumber="106">Iran</collectingCountry>
, this new subspecies is easily differentiated by the dark colouration, the broadly extended, elongated and hooked elytral maculations and the setose posterior edge of the pronotum (glabrous in 
<emphasis id="05E7EAE4FFF3FFCDFE0BFE5418DCFE04" box="[450,738,441,467]" italics="true" pageId="11" pageNumber="106">
<taxonomicName id="F0934D75FFF3FFCDFE0BFE5418E5FE04" authority="germanica" authorityName="germanica (Linnaeus" authorityYear="1758" box="[450,731,441,467]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="germanica">C. germanica germanica</taxonomicName>
;
</emphasis>
<figureCitation id="AFA82A73FFF3FFCDFD20FE5519B4FE05" box="[745,906,440,467]" captionStart="FIGURES 20-23" captionStartId="10.[151,264,1953,1978]" captionTargetBox="[156,1432,1041,1923]" captionTargetId="figure@10.[151,1436,1029,1929]" captionTargetPageId="10" captionText="FIGURES 20-23: Cylindera germanica persiana ssp. nov. – (20) holotype (male), (21) paratype (female), (22) pronotum of female paratype, (23) adeagus of holotype male (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3933986" httpUri="https://zenodo.org/record/3933986/files/figure.png" pageId="11" pageNumber="106">Figures 20-22</figureCitation>
).
</paragraph>
<paragraph id="372C36F6FFF3FFCDFF0EFE311F65FD51" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
<emphasis id="05E7EAE4FFF3FFCDFF0EFE311B79FE21" bold="true" box="[199,327,476,503]" pageId="11" pageNumber="106">Variability</emphasis>
: In the small series of specimens available for study, we detected little variation despite the large geographical range the specimens cover. The elytral maculation is rather conserved and all specimens show a very similar colouration. In some individuals, head and pronotum are more blueish-green metallic, while in others, they have a predominant coppery sheen. Also, while in all individuals the posterior edge of the pronotum is sparsely setose, only the 
<typeStatus id="E8288854FFF3FFCDFE85FD811B90FD50" box="[332,430,620,646]" pageId="11" pageNumber="106" type="holotype">holotype</typeStatus>
male also shows a sparsely setose anterior edge of the pronotum (
<figureCitation id="AFA82A73FFF3FFCDFB5BFD811F73FD51" box="[1170,1357,620,647]" captionStart="FIGURES 20-23" captionStartId="10.[151,264,1953,1978]" captionTargetBox="[156,1432,1041,1923]" captionTargetId="figure@10.[151,1436,1029,1929]" captionTargetPageId="10" captionText="FIGURES 20-23: Cylindera germanica persiana ssp. nov. – (20) holotype (male), (21) paratype (female), (22) pronotum of female paratype, (23) adeagus of holotype male (left lateral view)." figureDoi="http://doi.org/10.5281/zenodo.3933986" httpUri="https://zenodo.org/record/3933986/files/figure.png" pageId="11" pageNumber="106">Figures 20 &amp; 21</figureCitation>
).
</paragraph>
<paragraph id="372C36F6FFF3FFCDFF0EFD7D18F2FCC0" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
<emphasis id="05E7EAE4FFF3FFCDFF0EFD7D1BD3FD7D" bold="true" box="[199,493,656,683]" pageId="11" pageNumber="106">Habitat and distribution</emphasis>
: Although the 
<typeStatus id="E8288854FFF3FFCDFD5CFD7C18FBFD7D" box="[661,709,657,683]" pageId="11" pageNumber="106">type</typeStatus>
series is small, the few available records of 
<taxonomicName id="F0934D75FFF3FFCDFB73FD7C1AFFFD18" authorityName="Boetzl &amp; Franzen" authorityYear="2020" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="persiana">
<emphasis id="05E7EAE4FFF3FFCDFB73FD7C1AFFFD18" italics="true" pageId="11" pageNumber="106">C. germanica persiana</emphasis>
</taxonomicName>
ssp. nov. show an activity period early in the year in March and April, likely driven by the extremely hot and dry climate during the summer months. Populations of 
<taxonomicName id="F0934D75FFF3FFCDFD14FD3419C8FD24" authority="germanica" authorityName="germanica (Linnaeus" authorityYear="1758" box="[733,1014,729,755]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="germanica">
<emphasis id="05E7EAE4FFF3FFCDFD14FD3419C8FD24" box="[733,1014,729,755]" italics="true" pageId="11" pageNumber="106">C. germanica germanica</emphasis>
</taxonomicName>
in northern 
<collectingCountry id="4F847666FFF3FFCDFB4AFD341E8EFD25" box="[1155,1200,729,755]" name="Iran" pageId="11" pageNumber="106">Iran</collectingCountry>
as well as in 
<collectingCountry id="4F847666FFF3FFCDFA85FD351FA2FD24" box="[1356,1436,728,754]" name="Turkey" pageId="11" pageNumber="106">Turkey</collectingCountry>
however are active later, mostly from May to July.
</paragraph>
<paragraph id="372C36F6FFF3FFCDFF0EFCCC19BAFC54" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
<taxonomicName id="F0934D75FFF3FFCDFF0EFCCC1BF7FCEC" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[199,457,801,827]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="persiana">
<emphasis id="05E7EAE4FFF3FFCDFF0EFCCC1BF7FCEC" box="[199,457,801,827]" italics="true" pageId="11" pageNumber="106">C. germanica persiana</emphasis>
</taxonomicName>
ssp. nov. seems to be distributed south of the main ridge of the Zargos mountain range in the lowlands along the north coast of the Persian Gulf in southern 
<collectingCountry id="4F847666FFF3FFCDFC65FCA819E7FC89" box="[940,985,837,863]" name="Iran" pageId="11" pageNumber="106">Iran</collectingCountry>
and probably also in south-eastern 
<collectingCountry id="4F847666FFF3FFCDFAA6FCA81FA2FC89" box="[1391,1436,837,863]" name="Iraq" pageId="11" pageNumber="106">Iraq</collectingCountry>
(
<figureCitation id="AFA82A73FFF3FFCDFF56FC851AC0FC55" box="[159,254,872,899]" captionStart="FIGURE 1" captionStartId="3.[151,250,1002,1027]" captionTargetBox="[156,1430,187,965]" captionTargetId="figure@3.[151,1436,179,975]" captionTargetPageId="3" captionText="FIGURE 1: Distribution of Cylindera germanica (Linnaeus, 1758) in the Middle East." figureDoi="http://doi.org/10.5281/zenodo.3933977" httpUri="https://zenodo.org/record/3933977/files/figure.png" pageId="11" pageNumber="106">Figure 1</figureCitation>
). All specimens investigated originate from river valleys.
</paragraph>
<paragraph id="372C36F6FFF3FFCDFF0EFC6019BCFAE4" blockId="11.[151,1437,152,1331]" pageId="11" pageNumber="106">
Most of the specimens investigated by us were previously studied but assigned to 
<taxonomicName id="F0934D75FFF3FFCDFBA2FC611F5DFC70" box="[1131,1379,908,934]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF3FFCDFBA2FC611F5DFC70" box="[1131,1379,908,934]" italics="true" pageId="11" pageNumber="106">Cylindera descendens</emphasis>
</taxonomicName>
(e.g. 
<bibRefCitation id="53024B07FFF3FFCDFF5EFC5D1B08FC1D" author="Mandl, K." box="[151,310,944,971]" pageId="11" pageNumber="106" pagination="1 - 3" refId="ref10348" refString="Mandl, K. (1959) Eine Ausbeute an Cicindeliden aus Iran. Ergebnisse der Entomologischen Reisen Willi Richter, Stuttgart, im Iran 1954 und 1956. Nr. 19. Stuttgarter Beitrage zur Naturkunde, 18, 1 - 3." type="journal article" year="1959">Mandl (1959)</bibRefCitation>
, 
<bibRefCitation id="53024B07FFF3FFCDFE8DFC5D1BCEFC1D" author="Gebert, J." box="[324,496,944,971]" pageId="11" pageNumber="106" pagination="217 - 222" refId="ref9936" refString="Gebert, J. (1992) Zur Taxonomie, Systematik und Verbreitung der Cylindera &quot; obliquefasciata &quot; - Gruppe (Col., Cicindelidae). Entomologische Nachrichten und Berichte, 39, 217 - 222." type="journal article" year="1992">Gebert (1992))</bibRefCitation>
, which is clearly a misidentification due to their glabrous abdominal sternites. By contrast, specimens of 
<taxonomicName id="F0934D75FFF3FFCDFE51FC38180FFC38" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[408,561,981,1007]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="germanica">
<emphasis id="05E7EAE4FFF3FFCDFE51FC38180FFC38" box="[408,561,981,1007]" italics="true" pageId="11" pageNumber="106">C. germanica</emphasis>
</taxonomicName>
from Amara / 
<collectingCountry id="4F847666FFF3FFCDFD1CFC38193CFC39" box="[725,770,981,1007]" name="Iraq" pageId="11" pageNumber="106">Iraq</collectingCountry>
mentioned in 
<bibRefCitation id="53024B07FFF3FFCDFC6AFC381E11FC39" author="Horn, W." box="[931,1071,981,1007]" pageId="11" pageNumber="106" pagination="53 - 60" refId="ref10042" refString="Horn, W. (1927) Zur Erforschung des Persischen Golfes. (Beitrag Nr. 1). Einleitung; und Cicindelinae (Col). Entomologische Mitteilungen, XVI, 53 - 60." type="journal article" year="1927">Horn (1927)</bibRefCitation>
were identified as 
<taxonomicName id="F0934D75FFF3FFCDFACDFC381B8BFBC5" authority="(Kraatz, 1890)" baseAuthorityName="Kraatz" baseAuthorityYear="1890" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="martorelli">
<emphasis id="05E7EAE4FFF3FFCDFACDFC381B36FBC4" italics="true" pageId="11" pageNumber="106">C. germanica martorelli</emphasis>
(
<bibRefCitation id="53024B07FFF3FFCDFED1FC141B93FBC5" author="Kraatz, G." box="[280,429,1017,1043]" pageId="11" pageNumber="106" pagination="135 - 137" refId="ref10194" refString="Kraatz, G. (1890) Uber Cicindela maura Linne und andere. Entomologische Nachrichten, 9, 135 - 137. https: // doi. org / 10.1002 / mmnd. 48018900320" type="journal article" year="1890">Kraatz, 1890</bibRefCitation>
)
</taxonomicName>
(described as a colour morph of the Iberian 
<emphasis id="05E7EAE4FFF3FFCDFC7AFC141E57FBC4" box="[947,1129,1016,1043]" italics="true" pageId="11" pageNumber="106">
C. g. 
<taxonomicName id="F0934D75FFF3FFCDFC3AFC151E57FBC4" baseAuthorityName="Beuthin" baseAuthorityYear="1890" box="[1011,1129,1016,1042]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="catalonica">catalonica</taxonomicName>
</emphasis>
[Beuthin, 1890] by 
<bibRefCitation id="53024B07FFF3FFCDFA9BFC141ADCFBE1" author="Kraatz, G." pageId="11" pageNumber="106" pagination="135 - 137" refId="ref10194" refString="Kraatz, G. (1890) Uber Cicindela maura Linne und andere. Entomologische Nachrichten, 9, 135 - 137. https: // doi. org / 10.1002 / mmnd. 48018900320" type="journal article" year="1890">Kraatz (1890)</bibRefCitation>
and therefore a junior synonym of 
<taxonomicName id="F0934D75FFF3FFCDFDA4FBF0195CFBE0" baseAuthorityName="Gory" baseAuthorityYear="1833" box="[621,866,1052,1079]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="sobrina">
<emphasis id="05E7EAE4FFF3FFCDFDA4FBF0195CFBE0" box="[621,866,1052,1079]" italics="true" pageId="11" pageNumber="106">C. germanica sobrina</emphasis>
</taxonomicName>
[Gory, 1833]). The elytral pattern of 
<taxonomicName id="F0934D75FFF3FFCDFACDFBF01B36FB8C" baseAuthorityName="Kraatz" baseAuthorityYear="1890" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="martorelli">
<emphasis id="05E7EAE4FFF3FFCDFACDFBF01B36FB8C" italics="true" pageId="11" pageNumber="106">C. germanica martorelli</emphasis>
</taxonomicName>
is similar to that of 
<taxonomicName id="F0934D75FFF3FFCDFE24FBAC18D1FB8C" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[493,751,1089,1115]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="persiana">
<emphasis id="05E7EAE4FFF3FFCDFE24FBAC18D1FB8C" box="[493,751,1089,1115]" italics="true" pageId="11" pageNumber="106">C. germanica persiana</emphasis>
</taxonomicName>
ssp. nov. but specimens of all Iberian populations, including individuals of the before mentioned populations, lack the setae at the hind margin of the pronotum. We therefore expect the individuals from Amara to have been misidentified by W. Horn and actually belong to 
<taxonomicName id="F0934D75FFF3FFCDFB19FB641AEAFB10" authorityName="Boetzl &amp; Franzen" authorityYear="2020" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="persiana">
<emphasis id="05E7EAE4FFF3FFCDFB19FB641AEAFB10" italics="true" pageId="11" pageNumber="106">C. germanica persiana</emphasis>
</taxonomicName>
ssp. nov.. Another record of 
<taxonomicName id="F0934D75FFF3FFCDFDDFFB401889FB10" box="[534,695,1196,1223]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="species" species="descendens">
<emphasis id="05E7EAE4FFF3FFCDFDDFFB401889FB10" box="[534,695,1196,1223]" italics="true" pageId="11" pageNumber="106">C. descendens</emphasis>
</taxonomicName>
from Kusha / Hormozgan province listed in 
<bibRefCitation id="53024B07FFF3FFCDFB60FB411FA2FB11" author="Tschitscherine, T." box="[1193,1436,1196,1223]" pageId="11" pageNumber="106" pagination="1 - 21" refId="ref10612" refString="Tschitscherine, T. (1903) Memoire sur les Cicindelides des voyages de N. Zarudny dans la Perse orientale. Horae Societatis Entomologicae Rossicae, 36, 1 - 21." type="journal article" year="1903">Tschitschérine (1903)</bibRefCitation>
may also refer to 
<taxonomicName id="F0934D75FFF3FFCDFEAAFB3C1857FB3C" authorityName="Boetzl &amp; Franzen" authorityYear="2020" box="[355,617,1233,1259]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="106" phylum="Arthropoda" rank="subSpecies" species="germanica" subSpecies="persiana">
<emphasis id="05E7EAE4FFF3FFCDFEAAFB3C1857FB3C" box="[355,617,1233,1259]" italics="true" pageId="11" pageNumber="106">C. germanica persiana</emphasis>
</taxonomicName>
ssp. nov.. These populations south of the alpidic mountain ranges have clearly been overlooked so far. As there is a clear lack in collections from 
<collectingCountry id="4F847666FFF3FFCDFC1FFB191E2AFAD8" box="[982,1044,1268,1294]" name="Syria" pageId="11" pageNumber="106">Syria</collectingCountry>
, 
<collectingCountry id="4F847666FFF3FFCDFBE9FB181E73FAD9" box="[1056,1101,1269,1295]" name="Iraq" pageId="11" pageNumber="106">Iraq</collectingCountry>
and south-eastern 
<collectingCountry id="4F847666FFF3FFCDFAD6FB191F4EFAD8" box="[1311,1392,1268,1294]" name="Turkey" pageId="11" pageNumber="106">Turkey</collectingCountry>
, we cannot exclude the possibility of further populations in this region.
</paragraph>
</subSubSection>
</treatment>
</document>