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<document id="E7FF56FB36D48F917AC571F82E3749E8" ID-CLB-Dataset="36851" ID-DOI="10.11646/zootaxa.3799.1.1" ID-GBIF-Dataset="d9f29df5-52ff-451f-a367-f9fdb29b180f" ID-ISSN="1175-5326" ID-Zenodo-Dep="286090" ID-ZooBank="56DD05E1-C61C-4D37-9454-396840EB67C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460422790352" checkinUser="plazi" docAuthor="Gibbs, David" docDate="2014" docId="A96887E8FFFCFFBCFF43FB66FBEF06C3" docLanguage="en" docName="zt03799p085.pdf" docOrigin="Zootaxa 3799 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Usia putilla Becker 1906" docType="treatment" docVersion="11" lastPageNumber="45" masterDocId="5551FF90FFD5FF90FFD4FFCAFFF60616" masterDocTitle="A world revision of the bee fly tribe Usiini (Diptera, Bombyliidae) Part 2: Usia sensu stricto" masterLastPageNumber="85" masterPageNumber="1" pageNumber="42" updateTime="1698330265731" updateUser="ExternalLinkService">
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<mods:titleInfo id="9B42D9231608AF1E3F7518BE0F1E48F9">
<mods:title id="BECA0B21AF6B04A844786C7920A31A03">A world revision of the bee fly tribe Usiini (Diptera, Bombyliidae) Part 2: Usia sensu stricto</mods:title>
</mods:titleInfo>
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<mods:namePart id="AD1C1DC4207C5A5604EDEC1154103357">Gibbs, David</mods:namePart>
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<mods:title id="6540F597966F47062A04F3560B4B96F1">Zootaxa</mods:title>
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<mods:part id="53A9C481CA65D8A6E4550CAC4B331B19">
<mods:date id="CDA0EB09C8A53BED45E7185AC2BF7DF8">2014</mods:date>
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<mods:number id="154CB35764F2EF66B71E14F862753D94">2014-05-22</mods:number>
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<mods:number id="43BE80F6E1792D134CC83AFCC852ED18">3799</mods:number>
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<mods:identifier id="460DB8D4D6765BCC367ADCF88DE8A6E2" type="DOI">10.11646/zootaxa.3799.1.1</mods:identifier>
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<treatment id="A96887E8FFFCFFBCFF43FB66FBEF06C3" ID-DOI="http://doi.org/10.5281/zenodo.6135498" ID-GBIF-Taxon="127667889" ID-Zenodo-Dep="6135498" LSID="urn:lsid:plazi:treatment:A96887E8FFFCFFBCFF43FB66FBEF06C3" httpUri="http://treatment.plazi.org/id/A96887E8FFFCFFBCFF43FB66FBEF06C3" lastPageId="44" lastPageNumber="45" pageId="41" pageNumber="42">
<subSubSection id="69DB6575FFFCFFB9FF43FB66FE7702F1" pageId="41" pageNumber="42" type="nomenclature">
<paragraph id="217E36FEFFFCFFB9FF43FB66FE3C02D0" blockId="41.[151,458,1196,1255]" box="[151,458,1196,1222]" pageId="41" pageNumber="42">
<heading id="7A368192FFFCFFB9FF43FB66FE3C02D0" bold="true" box="[151,458,1196,1222]" fontSize="11" level="1" pageId="41" pageNumber="42" reason="1">
<taxonomicName id="E6C14D7DFFFCFFB9FF43FB66FE3C02D0" ID-CoL="8FL6P" authority="Becker, 1906" authorityName="Becker" authorityYear="1906" box="[151,458,1196,1222]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FF43FB66FE3C02D0" bold="true" box="[151,458,1196,1222]" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FF43FB66FED502D0" bold="true" box="[151,291,1196,1222]" italics="true" pageId="41" pageNumber="42">Usia putilla</emphasis>
<bibRefCitation id="45504B0FFFFCFFB9FEFFFB66FE3C02D0" author="Becker, T." box="[299,458,1196,1222]" firstAuthor="Becker" journalOrPublisher="Zeitschrift fur Systematische Hymenopterologie und Dipterologie" pageId="41" pageNumber="42" pagination="1 - 16" part="6" refId="ref58366" refString="Becker, T. (1906 a) Die Ergebnisse meiner dipterologischen Fruhjahrsreise nach Algier und Tunis 1906 [part]. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 6, 1 - 16." title="Die Ergebnisse meiner dipterologischen Fruhjahrsreise nach Algier und Tunis 1906 [part]" type="journal article" year="1906">Becker, 1906</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="217E36FEFFFCFFB9FF43FB04FE7702F1" blockId="41.[151,458,1196,1255]" box="[151,385,1230,1255]" pageId="41" pageNumber="42">
(
<figureCitation id="B9FA2A7BFFFCFFB9FF4AFB04FEF902F1" box="[158,271,1230,1255]" captionStart="FIGURE 12" captionStartId="13.[151,250,1941,1963]" captionTargetId="figure@13.[358,1228,1265,1920]" captionTargetPageId="13" captionText="FIGURE 12. ♂ U. putilla wing." figureDoi="http://doi.org/10.5281/zenodo.6376821" httpUri="https://zenodo.org/record/6376821/files/figure.png" pageId="41" pageNumber="42">Figure 12</figureCitation>
; Plate IX)
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFCFFB9FF43FAD2FDB5033A" box="[151,579,1302,1325]" pageId="41" pageNumber="42" type="reference_group">
<paragraph id="217E36FEFFFCFFB9FF43FAD2FDB5033A" blockId="41.[151,579,1302,1325]" box="[151,579,1302,1325]" pageId="41" pageNumber="42">
<treatmentCitationGroup id="01D111D0FFFCFFB9FF43FAD2FDB5033A" box="[151,579,1302,1325]" pageId="41" pageNumber="42">
<taxonomicName id="E6C14D7DFFFCFFB9FF43FAD2FE2B033A" ID-CoL="8FL6P" authority="Becker, 1906b: 215" authorityName="Becker" authorityPageNumber="215" authorityYear="1906" box="[151,477,1302,1325]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla" status="stat. nov.">
<emphasis id="13B5EAECFFFCFFB9FF43FAD2FEF9033A" box="[151,271,1302,1325]" italics="true" pageId="41" pageNumber="42">Usia putilla</emphasis>
<treatmentCitation id="A06010EFFFFCFFB9FEC1FADCFE2B033A" author="Becker" box="[277,477,1302,1324]" page="215" pageId="41" pageNumber="42" year="1906">
<bibRefCitation id="45504B0FFFFCFFB9FEC1FADCFE2B033A" author="Becker, T." box="[277,477,1302,1324]" firstAuthor="Becker" journalOrPublisher="Berliner Entomologische Zeitschrift" page="215" pageId="41" pageNumber="42" pagination="193 - 228" part="50" refId="ref58401" refString="Becker, T. (1906 b) Usia Latr. Berliner Entomologische Zeitschrift, 50 (1905), 193 - 228." title="Usia Latr" type="journal article" year="1906" yearSuffix="b">Becker, 1906b: 215</bibRefCitation>
</treatmentCitation>
</taxonomicName>
,
<taxonomicNameLabel id="08865797FFFCFFB9FE3CFADCFDB5033A" box="[488,579,1302,1324]" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FE3CFADCFDB5033A" bold="true" box="[488,579,1302,1324]" pageId="41" pageNumber="42">stat. rev.</emphasis>
</taxonomicNameLabel>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFCFFB9FF43FA93FAB500A3" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="217E36FEFFFCFFB9FF43FA93FEFA0033" blockId="41.[151,1436,1369,2006]" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FF43FA93FE3E0364" bold="true" box="[151,456,1369,1394]" pageId="41" pageNumber="42">
<typeStatus id="FE7A885CFFFCFFB9FF43FA93FF240364" box="[151,210,1369,1394]" pageId="41" pageNumber="42">Type</typeStatus>
material examined.
</emphasis>
<materialsCitation id="91A93CA3FFFCFFB9FE07FA90FC460383" ID-GBIF-Occurrence="3712321446" pageId="41" pageNumber="42" typeStatus="lectotype">
<typeStatus id="FE7A885CFFFCFFB9FE07FA90FD850364" box="[467,627,1370,1394]" pageId="41" pageNumber="42" type="lectotype">LECTOTYPE</typeStatus>
<emphasis id="13B5EAECFFFCFFB9FDABFA93FCC90364" bold="true" box="[639,831,1369,1394]" pageId="41" pageNumber="42">here designated</emphasis>
;
<emphasis id="13B5EAECFFFCFFB9FC86FA93FC5A0364" bold="true" box="[850,940,1369,1394]" pageId="41" pageNumber="42">
<collectingCountry id="59D6766EFFFCFFB9FC86FA93FC5E0364" box="[850,936,1369,1394]" name="France" pageId="41" pageNumber="42">France</collectingCountry>
,
</emphasis>
<taxonomicName id="E6C14D7DFFFCFFB9FC6CFA93FAFD0364" authority="Lyon, Coll. Winthem" authorityName="Lyon, Coll. Winthem" box="[952,1291,1369,1394]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FC6CFA93FBF70364" box="[952,1025,1369,1394]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
, Lyon, Coll. Winthem
</taxonomicName>
/
<taxonomicName id="E6C14D7DFFFCFFB9FACAFA93FA910364" box="[1310,1383,1369,1394]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FACAFA93FA910364" box="[1310,1383,1369,1394]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
</taxonomicName>
det. Becker/
<typeStatus id="FE7A885CFFFCFFB9FF22FAB4FEDB0380" box="[246,301,1406,1430]" pageId="41" pageNumber="42">Type</typeStatus>
/
<taxonomicName id="E6C14D7DFFFCFFB9FEEAFAB4FDF00380" box="[318,518,1406,1430]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFFCFFB9FEEAFAB4FDF00380" box="[318,518,1406,1430]" italics="true" pageId="41" pageNumber="42">Usia angustifrons</emphasis>
</taxonomicName>
B., Dr E.O. Engel det. [♂ in
<collectionCode id="47D0AE3BFFFCFFB9FC8FFAB4FC5F0383" box="[859,937,1406,1429]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="91A93CA3FFFCFFB9FC6FFAB4FB4B03AF" ID-GBIF-Occurrence="3712321353" pageId="41" pageNumber="42" typeStatus="paralectotype">
<typeStatus id="FE7A885CFFFCFFB9FC6FFAB4FB400380" box="[955,1206,1406,1430]" pageId="41" pageNumber="42" type="paralectotype">PARALECTOTYPES</typeStatus>
;
<taxonomicName id="E6C14D7DFFFCFFB9FB17FAB7FF0803AC" authority="Lyon, Coll. Winthem" authorityName="Lyon, Coll. Winthem" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FB17FAB7FAFA0380" box="[1219,1292,1405,1430]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
, Lyon, Coll. Winthem
</taxonomicName>
/
<taxonomicName id="E6C14D7DFFFCFFB9FED9FA6BFEA003AC" box="[269,342,1441,1466]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FED9FA6BFEA003AC" box="[269,342,1441,1466]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
</taxonomicName>
det. Becker/
<typeStatus id="FE7A885CFFFCFFB9FE38FA68FDD503AC" box="[492,547,1442,1466]" pageId="41" pageNumber="42">Type</typeStatus>
/
<taxonomicName id="E6C14D7DFFFCFFB9FDE6FA68FC1903AC" authority="Beck., Dr E.O. Engel" authorityName="Beck., Dr E.O. Engel" box="[562,1007,1441,1466]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFFCFFB9FDE6FA68FD0F03AC" box="[562,761,1442,1466]" italics="true" pageId="41" pageNumber="42">Usia angustifrons</emphasis>
Beck., Dr E.O. Engel
</taxonomicName>
det. [♂ in
<collectionCode id="47D0AE3BFFFCFFB9FBBCFA68FB4003AF" box="[1128,1206,1442,1465]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="91A93CA3FFFCFFB9FB1CFA68FEFF0033" ID-GBIF-Occurrence="3712321498" pageId="41" pageNumber="42">
103 [on micropin]/
<taxonomicName id="E6C14D7DFFFCFFB9FF43FA0FFE1503C8" authority="Lyon, Coll. Winthem" authorityName="Lyon, Coll. Winthem" box="[151,483,1477,1502]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FF43FA0FFF1603C8" box="[151,224,1477,1502]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
, Lyon, Coll. Winthem
</taxonomicName>
/
<taxonomicName id="E6C14D7DFFFCFFB9FE21FA0FFDC803C8" box="[501,574,1477,1502]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FE21FA0FFDC803C8" box="[501,574,1477,1502]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
</taxonomicName>
det. Becker/
<typeStatus id="FE7A885CFFFCFFB9FD0FFA0CFCE403C8" box="[731,786,1478,1502]" pageId="41" pageNumber="42">Type</typeStatus>
/
<taxonomicName id="E6C14D7DFFFCFFB9FCF1FA0CFC1903C8" box="[805,1007,1478,1502]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFFCFFB9FCF1FA0CFC1903C8" box="[805,1007,1478,1502]" italics="true" pageId="41" pageNumber="42">Usia angustifrons</emphasis>
</taxonomicName>
B., Dr E.O. Engel det. [♀in
<collectionCode id="47D0AE3BFFFCFFB9FA97FA0CFA6703CB" box="[1347,1425,1478,1501]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
];
<taxonomicName id="E6C14D7DFFFCFFB9FF43FA23FED10014" authority="Lyon" authorityName="Lyon" box="[151,295,1513,1538]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FF43FA23FF160014" box="[151,224,1513,1538]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
, Lyon
</taxonomicName>
/
<taxonomicName id="E6C14D7DFFFCFFB9FEE2FA23FD8D0014" authority="Lyon, Coll. Winthem" authorityName="Lyon, Coll. Winthem" box="[310,635,1513,1538]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FEE2FA23FE890014" box="[310,383,1513,1538]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
, Lyon, Coll. Winthem
</taxonomicName>
/
<taxonomicName id="E6C14D7DFFFCFFB9FD5EFA23FD250014" box="[650,723,1513,1538]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFCFFB9FD5EFA23FD250014" box="[650,723,1513,1538]" italics="true" pageId="41" pageNumber="42">putilla</emphasis>
</taxonomicName>
det. Becker/
<typeStatus id="FE7A885CFFFCFFB9FCBDFA20FC560014" box="[873,928,1514,1538]" pageId="41" pageNumber="42">Type</typeStatus>
/
<taxonomicName id="E6C14D7DFFFCFFB9FC7BFA20FB810014" box="[943,1143,1514,1538]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFFCFFB9FC7BFA20FB810014" box="[943,1143,1514,1538]" italics="true" pageId="41" pageNumber="42">Usia angustifrons</emphasis>
</taxonomicName>
B., Dr E.O. Engel det. [♀ in
<collectionCode id="47D0AE3BFFFCFFB9FF60F9C4FEF40033" box="[180,258,1550,1573]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="217E36FEFFFCFFB9FF13F9FBFB530078" blockId="41.[151,1436,1369,2006]" pageId="41" pageNumber="42">
Originally described from an unspecified number of specimens of both sexes from Winthem’s collection, Vienna. Four
<typeStatus id="FE7A885CFFFCFFB9FEE4F99CFE640078" box="[304,402,1622,1646]" pageId="41" pageNumber="42" type="syntype">syntypes</typeStatus>
were found in
<collectionCode id="47D0AE3BFFFCFFB9FDEFF99CFD75007B" box="[571,643,1622,1645]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
, the male in best condition selected as
<typeStatus id="FE7A885CFFFCFFB9FBE2F99FFB570078" box="[1078,1185,1621,1646]" pageId="41" pageNumber="42" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="217E36FEFFFCFFB9FF13F9B3FAB500A3" blockId="41.[151,1436,1369,2006]" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FF13F9B3FDFF0084" bold="true" box="[199,521,1657,1682]" pageId="41" pageNumber="42">Other material examined.</emphasis>
<materialsCitation id="91A93CA3FFFCFFB9FDC3F9B3FF0E00A0" ID-GBIF-Occurrence="3712321306" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FDC3F9B3FDAD0084" bold="true" box="[535,603,1657,1682]" pageId="41" pageNumber="42">
<collectingCountry id="59D6766EFFFCFFB9FDC3F9B3FDAD0084" box="[535,603,1657,1682]" name="Spain" pageId="41" pageNumber="42">Spain</collectingCountry>
</emphasis>
, Chañe (Segovia)
<date id="557F103EFFFCFFB9FC95F9B3FC0D0084" box="[833,1019,1657,1682]" pageId="41" pageNumber="42" value="2000-05-18" valueMax="2000-07-03" valueMin="2000-05-18">18.v–3.vii 2000</date>
, T. Malaise [♂] (leg. J.F. Gómez) (
<collectionCode id="47D0AE3BFFFCFFB9FF4BF954FF0600A0" box="[159,240,1694,1718]" pageId="41" pageNumber="42">PCDG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="91A93CA3FFFCFFB9FED6F957FAB500A3" ID-GBIF-Occurrence="3712321382" box="[258,1347,1693,1718]" pageId="41" pageNumber="42">
<collectingCountry id="59D6766EFFFCFFB9FED6F957FEA000A0" box="[258,342,1693,1718]" name="France" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FED6F957FEA000A0" bold="true" box="[258,342,1693,1718]" pageId="41" pageNumber="42">France</emphasis>
</collectingCountry>
, Cavaliea [sic. = Cavalaire, Var], June [19]06, Museum Paris, ex coll. Hesse [♀
<collectionCode id="47D0AE3BFFFCFFB9FB08F954FACE00A3" LSID="urn:lsid:biocol.org:col:34988" box="[1244,1336,1694,1717]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="41" pageNumber="42">MNHN</collectionCode>
].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFCFFB9FF13F90BFE5D00E8" pageId="41" pageNumber="42" type="etymology">
<paragraph id="217E36FEFFFCFFB9FF13F90BFE5D00E8" blockId="41.[151,1436,1369,2006]" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FF13F90BFEB800CC" bold="true" box="[199,334,1729,1754]" pageId="41" pageNumber="42">Etymology.</emphasis>
from Latin
<emphasis id="13B5EAECFFFCFFB9FE37F908FDD000CF" box="[483,550,1730,1753]" italics="true" pageId="41" pageNumber="42">putus,</emphasis>
meaning pure or clear, +
<emphasis id="13B5EAECFFFCFFB9FCB0F90BFC6600CC" box="[868,912,1729,1754]" italics="true" pageId="41" pageNumber="42">illa,</emphasis>
a diminutive suffix; perhaps referring to its shininess and small size.
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFCFFBAFF13F8C3FC3906EE" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="diagnosis">
<paragraph id="217E36FEFFFCFFBAFF13F8C3FC3906EE" blockId="41.[151,1436,1369,2006]" lastBlockId="42.[151,1437,151,2013]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="13B5EAECFFFCFFB9FF13F8C3FEB70134" bold="true" box="[199,321,1801,1826]" pageId="41" pageNumber="42">Diagnosis.</emphasis>
A small to very small shining black species, known only from a few specimens from
<collectingCountry id="59D6766EFFFCFFB9FACBF8C0FA9D0137" box="[1311,1387,1802,1825]" name="France" pageId="41" pageNumber="42">France</collectingCountry>
and
<collectingCountry id="59D6766EFFFCFFB9FF43F8E7FF2C0150" box="[151,218,1837,1862]" name="Spain" pageId="41" pageNumber="42">Spain</collectingCountry>
. Entirely black, vestiture fine, relatively sparse and silvery white, hairs erect or slightly inclined. Occiput shining, male frons narrow, eyes separated by less than distance across lateral ocelli, rear half of frons parallel, silvery dusting coalescing in front of anterior ocellus or, in females, dust spots separated. Acrostichal setulae proclinate in all specimens seen. Pleurae dusted apart from small shining dorsal and posterior margin on anepisternum in some individuals. Wings faintly yellow-grey tinged, distinctly yellow basally, veins yellow, becoming darker distally, anal lobe narrow but trailing edge clearly curved, more than half width of anal cell. Genitalia typical of group, differing only subtly from close congeners, most importantly in membranous apical margin of epandrium, which extends for more than half length of sclerite.
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFFFFBAFF13FECEFAB2003B" pageId="42" pageNumber="43" type="description">
<paragraph id="217E36FEFFFFFFBAFF13FECEFB80070A" blockId="42.[151,1437,151,2013]" box="[199,1142,260,285]" pageId="42" pageNumber="43">
<emphasis id="13B5EAECFFFFFFBAFF13FECEFDDC070B" bold="true" box="[199,554,260,285]" pageId="42" pageNumber="43">Redescription. Measurements</emphasis>
.
<emphasis id="13B5EAECFFFFFFBAFDECFECEFD49070B" box="[568,703,260,285]" italics="true" pageId="42" pageNumber="43">Body length</emphasis>
.
<quantity id="E6399B1BFFFFFFBAFD18FECEFCA6070B" box="[716,848,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="42" pageNumber="43" unit="mm" value="3.95" valueMax="4.5" valueMin="3.4">3.4–4.5mm</quantity>
.
<emphasis id="13B5EAECFFFFFFBAFC8FFECFFC17070B" box="[859,993,260,285]" italics="true" pageId="42" pageNumber="43">Wing length</emphasis>
.
<quantity id="E6399B1BFFFFFFBAFC3BFECEFB84070A" box="[1007,1138,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.4" metricValueMin="2.7" pageId="42" pageNumber="43" unit="mm" value="3.05" valueMax="3.4" valueMin="2.7">2.7–3.4mm</quantity>
.
</paragraph>
<paragraph id="217E36FEFFFFFFBAFF13FEEDFF190343" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
<emphasis id="13B5EAECFFFFFFBAFF13FEEDFEF20756" bold="true" box="[199,260,295,320]" pageId="42" pageNumber="43">Male</emphasis>
.
<emphasis id="13B5EAECFFFFFFBAFEC5FEEDFEBB0756" box="[273,333,295,320]" italics="true" pageId="42" pageNumber="43">Head</emphasis>
. Mouth-margin very narrow, almost absent towards rear of oral opening, frons at narrowest point a little less than distance across lateral ocelli, equal to three times diameter of anterior ocellus, lateral ocellus separated from the eye margin by about its radius or less. Ocellar tubercle shining black, anterolateral silvery grey dust spots on frons extending back to narrowest part where they coalesce, a covering of fine pale setulae on anterior part of frons. Occiput shining black without any noticeable pollinosity except in the two vertical occipital tumescence but even here black ground colour not obscured. Hairing moderate length, on ocellar area about as long as width of vertex at narrowest point. Proboscis black, hairless, about as long as the head and thorax together; palps very short and thin, black to brown with short pale hairs apically; antennae black, third antennal segment longer than the scape and pedicel together, all segments shortly-haired above.
<emphasis id="13B5EAECFFFFFFBAFC6CFD8DFBFF0476" box="[952,1033,583,608]" italics="true" pageId="42" pageNumber="43">Thorax</emphasis>
. Mesonotum and scutellum shining black except for dorsum of post pronotal lobes, anteroventral corner of notopleuron and small areas above the wings. Shining between punctures, in some individuals these punctures rather stellate, bare paramedian lines relatively conspicuous and extending back to the mid point of the mesonotum, each a little broader than the acrostichal area which has 5–6 lines of proclinate hairs, most of hairs on disc of mesonotum also proclinate. Mesonotum covered with short white hairs, that on the disc no longer than distance between lateral ocelli, on scutellum about as long as vertex at level of lateral ocelli. Laterally hairing a little longer and denser, the hairs erect or inclined inwards. Pleurae black with a covering of grey dust sufficiently dense to obscure the black ground colour except for small shining patch along dorsal and posterior margin of anepisternum. Posterior half of anepisternum and pronotum with rather longer white hairs.
<emphasis id="13B5EAECFFFFFFBAFC8BFC47FC6105B2" box="[863,919,909,932]" italics="true" pageId="42" pageNumber="43">Wing</emphasis>
. Wing membrane clear, only faintly tinged yellowish-grey, becoming yellow basally, the veins also yellow basally becoming conspicuously darker brownish in the apical half. Anal lobe narrow, but with rounded margin, just a little narrower than anal cell (
<collectingCountry id="59D6766EFFFFFFBAFB0FFC1EFADD05FA" box="[1243,1323,980,1004]" name="France" pageId="42" pageNumber="43">France</collectingCountry>
) to a little over half width of anal cell (
<collectingCountry id="59D6766EFFFFFFBAFE35FC3DFDD30206" box="[481,549,1015,1040]" name="Spain" pageId="42" pageNumber="43">Spain</collectingCountry>
,
<figureCitation id="B9FA2A7BFFFFFFBAFDE6FC3DFD550206" box="[562,675,1015,1040]" captionStart="FIGURE 12" captionStartId="13.[151,250,1941,1963]" captionTargetId="figure@13.[358,1228,1265,1920]" captionTargetPageId="13" captionText="FIGURE 12. ♂ U. putilla wing." figureDoi="http://doi.org/10.5281/zenodo.6376821" httpUri="https://zenodo.org/record/6376821/files/figure.png" pageId="42" pageNumber="43">Figure 12</figureCitation>
), r-m a little before the middle of the discal cell.
<emphasis id="13B5EAECFFFFFFBAFB0AFC3DFAC40206" box="[1246,1330,1015,1040]" italics="true" pageId="42" pageNumber="43">Haltere</emphasis>
. Whitish yellow, slightly infuscated at base of stem.
<emphasis id="13B5EAECFFFFFFBAFD4EFBD7FD390222" box="[666,719,1053,1076]" italics="true" pageId="42" pageNumber="43">Legs</emphasis>
. Entirely black, the coxae grey dusted as in pleurae; femora ventrally with mid-length, pale hairs, up to length of width of tibia, tibia and tarsi with a covering of short, white setulae, longer on tibia.
<emphasis id="13B5EAECFFFFFFBAFE72FBAEFDE6026B" box="[422,528,1124,1149]" italics="true" pageId="42" pageNumber="43">Abdomen</emphasis>
. Shining aeneous black, the disc of the tergites covered with short, whitish hairs issuing from transverse welts. Laterally and on the down-curved margins the tergites are clothed with rather longer, whitish hairs. Middle tergites with a shining, impunctate apical margin often with longitudinal grooves. Sternites grey dusted with whitish hairs confined to the posterior margin.
<emphasis id="13B5EAECFFFFFFBAFC8BFB05FC3F02FE" box="[863,969,1231,1256]" italics="true" pageId="42" pageNumber="43">Genitalia</emphasis>
. Relatively large and conspicuous for this species group, few useful details visible in pinned specimens, shining black and covered in whitish hairs. Most useful character is the membranous apical ‘V’ notch of the epandrium which extends for more than half of the sclerite.
</paragraph>
<paragraph id="217E36FEFFFFFFBAFF13FA95FAB2003B" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
<emphasis id="13B5EAECFFFFFFBAFF13FA95FEE8036E" bold="true" box="[199,286,1375,1400]" pageId="42" pageNumber="43">Female</emphasis>
. As male except as follows: broader, though still very narrow, oral margin, and conspicuously wider frons, lateral ocellus separated the eye margin by about diameter of that ocellus. Posterior half of frons shining black, parallel sided, about four times diameter of anterior ocellus, as wide or wider than length of longest hairs on ocellar tubercle. Anterior half of frons widening, lateral margins each side covered in dense whitish dust obscuring ground colour, central two-thirds shining black or dark brown. Third antennal segment sometimes a little shorter. Anal lobe broader, margin more convex, as broad as anal cell. Abdomen conical tapering to rounded point.
</paragraph>
</subSubSection>
<subSubSection id="69DB6575FFFFFFBAFF13F9FDFABD01CA" pageId="42" pageNumber="43" type="discussion">
<paragraph id="217E36FEFFFFFFBAFF13F9FDFA6B00AB" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
<emphasis id="13B5EAECFFFFFFBAFF13F9FDFEBA0046" bold="true" box="[199,332,1591,1616]" pageId="42" pageNumber="43">Discussion.</emphasis>
This small shining black species is clearly very closely allied to
<taxonomicName id="E6C14D7DFFFFFFBAFB8BF9F3FAEF0059" authority="Loew" authorityName="Loew" box="[1119,1305,1592,1616]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="13B5EAECFFFFFFBAFB8BF9F3FB3A0046" box="[1119,1228,1593,1616]" italics="true" pageId="42" pageNumber="43">U. manca</emphasis>
Loew
</taxonomicName>
, and could possibly be the end of a cline of this species. If specimens come to light between north
<collectingCountry id="59D6766EFFFFFFBAFB4EF996FB3B0063" box="[1178,1229,1628,1653]" name="Italy" pageId="42" pageNumber="43">Italy</collectingCountry>
and central south
<collectingCountry id="59D6766EFFFFFFBAFF43F94AFF150081" box="[151,227,1664,1687]" name="France" pageId="42" pageNumber="43">France</collectingCountry>
that show intermediate characters then
<taxonomicName id="E6C14D7DFFFFFFBAFD1CF94BFC66008E" authority="Becker" authorityName="Becker" box="[712,912,1663,1688]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFFFFBAFD1CF94BFCC5008E" box="[712,819,1663,1688]" italics="true" pageId="42" pageNumber="43">U. putilla</emphasis>
Becker
</taxonomicName>
might need synonymising under
<taxonomicName id="E6C14D7DFFFFFFBAFAFCF94BFA63008E" box="[1320,1429,1665,1688]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="manca">
<emphasis id="13B5EAECFFFFFFBAFAFCF94BFA63008E" box="[1320,1429,1665,1688]" italics="true" pageId="42" pageNumber="43">U. manca</emphasis>
</taxonomicName>
. However, in the absence of such specimens the differences, though small, are sufficient to retain it as a full species.
</paragraph>
<paragraph id="217E36FEFFFFFFBAFF13F90DFABD01CA" blockId="42.[151,1437,151,2013]" pageId="42" pageNumber="43">
<bibRefCitation id="45504B0FFFFFFFBAFF13F90DFE9700F6" author="Engel, E. O." box="[199,353,1735,1760]" editor="Lindner, E." firstAuthor="Engel" journalOrPublisher="E. Schweizerbart, Stuttgart" pageId="42" pageNumber="43" pagination="49 - 96" part="4" refId="ref58498" refString="Engel, E. O. (1932) Bombyliidae. In: Lindner, E., (Ed.), Die Fliegen der palaarktischen Region. Vol. 4. pt. 3. E. Schweizerbart, Stuttgart, pp. 49 - 96." title="Bombyliidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region" year="1932">Engel (1932)</bibRefCitation>
treated this taxon as a subspecies of
<taxonomicName id="E6C14D7DFFFFFFBAFCE3F903FBB400F6" authority="Becker" authorityName="Becker" box="[823,1090,1735,1760]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="angustifrons">
<emphasis id="13B5EAECFFFFFFBAFCE3F903FC1300F6" box="[823,997,1736,1760]" italics="true" pageId="42" pageNumber="43">U. angustifrons</emphasis>
Becker
</taxonomicName>
and he wrote that he could understand why Becker described it again under the name
<taxonomicName id="E6C14D7DFFFFFFBAFCF1F927FC660113" box="[805,912,1772,1797]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFFFFBAFCF1F927FC660113" box="[805,912,1772,1797]" italics="true" pageId="42" pageNumber="43">U. putilla</emphasis>
</taxonomicName>
. If Engel had the
<typeStatus id="FE7A885CFFFFFFBAFB8CF927FB650113" box="[1112,1171,1773,1797]" pageId="42" pageNumber="43">types</typeStatus>
of
<taxonomicName id="E6C14D7DFFFFFFBAFB6CF927FAD20113" box="[1208,1316,1772,1797]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFFFFBAFB6CF927FAD20113" box="[1208,1316,1772,1797]" italics="true" pageId="42" pageNumber="43">U. putilla</emphasis>
</taxonomicName>
in front of him then his decision is inexplicable, even without dissection these two species are abundantly distinct. Perhaps Engel did not have the
<taxonomicName id="E6C14D7DFFFFFFBAFE74F8FFFDFA015B" box="[416,524,1844,1869]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFFFFBAFE74F8FFFDFA015B" box="[416,524,1844,1869]" italics="true" pageId="42" pageNumber="43">U. putilla</emphasis>
</taxonomicName>
<typeStatus id="FE7A885CFFFFFFBAFDC0F8FFFDB9015B" box="[532,591,1845,1869]" pageId="42" pageNumber="43">types</typeStatus>
available, in which case he might have worked from Becker’s description alone.
<bibRefCitation id="45504B0FFFFFFFBAFF35F89DFE660166" author="Becker, T." box="[225,400,1879,1904]" firstAuthor="Becker" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="42" pageNumber="43" pagination="193 - 228" part="50" refId="ref58401" refString="Becker, T. (1906 b) Usia Latr. Berliner Entomologische Zeitschrift, 50 (1905), 193 - 228." title="Usia Latr" type="journal article" year="1906" yearSuffix="b">Becker (1906b)</bibRefCitation>
did list some useless features in his key (e.g., colour of wing membrane and position of vein r-m), but the width of the frons and nature of occiput, accurately described by Becker, should have been sufficient to convince Engel he was dealing with a distinct taxon. In his description
<bibRefCitation id="45504B0FFFFFFFBAFBF7F86AFB4A01AE" author="Engel, E. O." box="[1059,1212,1952,1977]" editor="Lindner, E." firstAuthor="Engel" journalOrPublisher="E. Schweizerbart, Stuttgart" pageId="42" pageNumber="43" pagination="49 - 96" part="4" refId="ref58498" refString="Engel, E. O. (1932) Bombyliidae. In: Lindner, E., (Ed.), Die Fliegen der palaarktischen Region. Vol. 4. pt. 3. E. Schweizerbart, Stuttgart, pp. 49 - 96." title="Bombyliidae" type="book chapter" volumeTitle="Die Fliegen der palaarktischen Region" year="1932">Engel (1932)</bibRefCitation>
mentioned several characters (entirely dusted lower frons and dusted occiput) that do not fit Becker’s description of
<taxonomicName id="E6C14D7DFFFFFFBAFB0DF80FFAB201CB" box="[1241,1348,1988,2013]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFFFFBAFB0DF80FFAB201CB" box="[1241,1348,1988,2013]" italics="true" pageId="42" pageNumber="43">U. putilla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="75BE6676FFFEFFBBFF43F850FC6601D9" ID-DOI="http://doi.org/10.5281/zenodo.6376342" ID-Zenodo-Dep="6376342" httpUri="https://zenodo.org/record/6376342/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,234,1946,1968]" targetBox="[206,1372,234,1903]" targetPageId="43">
<paragraph id="217E36FEFFFEFFBBFF43F850FC6601D9" blockId="43.[151,1435,1946,1999]" pageId="43" pageNumber="44">
<emphasis id="13B5EAECFFFEFFBBFF43F850FEE201A6" bold="true" box="[151,276,1946,1968]" pageId="43" pageNumber="44">PLATE IX.</emphasis>
<taxonomicName id="E6C14D7DFFFEFFBBFECFF851FDD901A6" authority="Becker, 1906" authorityName="Becker" authorityYear="1906" box="[283,559,1946,1968]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="putilla">
<emphasis id="13B5EAECFFFEFFBBFECFF851FE6501A6" box="[283,403,1946,1968]" italics="true" pageId="43" pageNumber="44">Usia putilla</emphasis>
<bibRefCitation id="45504B0FFFFEFFBBFE4EF850FDD901A6" author="Becker, T." box="[410,559,1946,1968]" firstAuthor="Becker" journalOrPublisher="Berliner Entomologische Zeitschrift" pageId="43" pageNumber="44" pagination="193 - 228" part="50" refId="ref58401" refString="Becker, T. (1906 b) Usia Latr. Berliner Entomologische Zeitschrift, 50 (1905), 193 - 228." title="Usia Latr" type="journal article" year="1906" yearSuffix="b">Becker, 1906b</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="45504B0FFFFEFFBBFDEFF850FDBE01A6" author="Becker, T." box="[571,584,1946,1968]" journalOrPublisher="Zeitschrift fur Systematische Hymenopterologie und Dipterologie" pageId="43" pageNumber="44" pagination="1 - 16" part="6" refId="ref58366" refString="Becker, T. (1906 a) Die Ergebnisse meiner dipterologischen Fruhjahrsreise nach Algier und Tunis 1906 [part]. Zeitschrift fur Systematische Hymenopterologie und Dipterologie, 6, 1 - 16." title="Die Ergebnisse meiner dipterologischen Fruhjahrsreise nach Algier und Tunis 1906 [part]" type="journal article" year="1906" yearSuffix="a">
<emphasis id="13B5EAECFFFEFFBBFDEFF850FDBE01A6" bold="true" box="[571,584,1946,1968]" pageId="43" pageNumber="44">a</emphasis>
</bibRefCitation>
epiphallic complex ventral,
<emphasis id="13B5EAECFFFEFFBBFCB8F850FC8C01A6" bold="true" box="[876,890,1946,1968]" pageId="43" pageNumber="44">b</emphasis>
epiphallic complex lateral,
<emphasis id="13B5EAECFFFEFFBBFB4DF850FB5201A6" bold="true" box="[1177,1188,1946,1968]" pageId="43" pageNumber="44">c</emphasis>
gonocoxite &amp; gonostyli ventral,
<emphasis id="13B5EAECFFFEFFBBFF3EF873FF0E01D9" bold="true" box="[234,248,1977,1999]" pageId="43" pageNumber="44">d</emphasis>
epandrium dorsal,
<emphasis id="13B5EAECFFFEFFBBFE6FF873FE3001D9" bold="true" box="[443,454,1977,1999]" pageId="43" pageNumber="44">e</emphasis>
epandrium lateral,
<emphasis id="13B5EAECFFFEFFBBFD5FF873FD6201D9" bold="true" box="[651,660,1977,1999]" pageId="43" pageNumber="44">f</emphasis>
female genitalia ventral.
</paragraph>
</caption>
<subSubSection id="69DB6575FFF9FFBCFF13FF5DFBEF06C3" pageId="44" pageNumber="45" type="distribution">
<paragraph id="217E36FEFFF9FFBCFF13FF5DFBEF06C3" blockId="44.[151,1436,151,213]" pageId="44" pageNumber="45">
<emphasis id="13B5EAECFFF9FFBCFF13FF5DFE9606A6" bold="true" box="[199,352,151,176]" pageId="44" pageNumber="45">Distribution.</emphasis>
A rare and little known species so far only known from the Rhone valley near Lyon, and the Mediterranean coast, southern
<collectingCountry id="59D6766EFFF9FFBCFE24FF77FDCA06C2" box="[496,572,189,212]" name="France" pageId="44" pageNumber="45">France</collectingCountry>
and a single specimen from central
<collectingCountry id="59D6766EFFF9FFBCFC06FF76FBE306C3" box="[978,1045,188,213]" name="Spain" pageId="44" pageNumber="45">Spain</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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