treatments-xml/data/29/01/87/290187FFFFBFFFE1FE76EE3B454B6F32.xml

<document id="D6F5D01F94F83BE76BE6F908FBB34759" ID-CLB-Dataset="8569" ID-DOI="10.5281/zenodo.4468289" ID-GBIF-Dataset="0657afbe-a3ef-4f09-9361-95b1871b860a" ID-ISSN="ISSN 0374-1036" ID-Zenodo-Dep="4468289" ID-ZooBank="879078EE-3058-46B0-A3A1-19E8945F42AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611664279813" checkinUser="felipe" docAuthor="Kvifte, Gunnar M." docDate="2015" docId="290187FFFFBFFFE1FE76EE3B454B6F32" docLanguage="en" docName="ActaEntMusNatPra.55.2.495-503.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Psychomasina paedagogica Kvifte 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="499" masterDocId="D538FF87FFBDFFE5FFCFEC5946426C41" masterDocTitle="Review of the genus Psychomasina (Diptera: Psychodidae: Psychodinae), with descriptions of two new species" masterLastPageNumber="503" masterPageNumber="495" pageNumber="497" updateTime="1698925516382" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4EA04E4D692D53D2E02229879F31B198">Review of the genus Psychomasina (Diptera: Psychodidae: Psychodinae), with descriptions of two new species</mods:title>
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<mods:namePart id="AFB615A21DEA7F7B2CB940ED6CFCB5FC">Kvifte, Gunnar M.</mods:namePart>
<mods:affiliation id="6EF6B2F83F82CAA1DBFA16BF3CADDD32">1,2) &amp;) Department of Limnology, Institute of Biology, University of Kassel, Heinrich-Plett Str. 40, DE- 34132 Kassel-Oberzwehren, Germany; e-mail: Gunnar. Kvifte @ uib. no &amp; 1,2) &amp;) Department of Natural History, University Museum of Bergen, P. O. Box 7800, University of Bergen, NO- 5020 Bergen, Norway</mods:affiliation>
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<paragraph id="A11736E9FFBFFFE7FE76EE3B45076E3B" blockId="2.[441,837,610,634]" box="[441,837,610,634]" pageId="2" pageNumber="497">
<heading id="FA5F8185FFBFFFE7FE76EE3B45076E3B" bold="true" box="[441,837,610,634]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="497" reason="3">
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<taxonomicName id="66A84D6AFFBFFFE7FE76EE3B44AB6E3B" authority="Kvifte, 2015" authorityName="Kvifte" authorityYear="2015" box="[441,745,610,634]" class="Insecta" family="Psychodidae" genus="Psychomasina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="paedagogica" status="sp. nov.">
<emphasis id="93DCEAFBFFBFFFE7FE76EE3B44AB6E3B" bold="true" box="[441,745,610,634]" italics="true" pageId="2" pageNumber="497">Psychomasina paedagogica</emphasis>
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<paragraph id="A11736E9FFBFFFE7FD8CEED844F86ED5" blockId="2.[151,1127,641,729]" box="[579,698,641,660]" pageId="2" pageNumber="497">
(
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, 
<figureCitation id="39932A6CFFBFFFE7FD50EED844F76ED5" box="[671,693,641,660]" captionStart="Figs 6–12. 6–9" captionStartId="5.[113,150,1492,1511]" captionTargetBox="[123,1074,488,1471]" captionTargetId="figure-109@5.[113,1088,482,1480]" captionTargetPageId="5" captionText="Figs 6–12. 6–9 – Psychomasina violina sp. nov., holotype, male. 6 – head; 7 – wing; 8 – aedeagus and gonopods; 9 – epandrium and proctiger. 10–12 – aedeagal joint shape in Psychomasina spp.: 10 – P. paedagogica sp. nov., 11 – P. violina sp. nov., 12 – P. armata Ježek, 2004 (Fig. 12 redrawn from JEŽEK 2004). For abbreviations see Material and methods. Length of scale bars in μm: 100 (6, 8, 9) 250 (7)" figureDoi="http://doi.org/10.5281/zenodo.4468293" httpUri="https://zenodo.org/record/4468293/files/figure.png" pageId="2" pageNumber="497">10</figureCitation>
)
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<subSubSection id="E9B26562FFBFFFE7FF58EEF244726FC9" pageId="2" pageNumber="497" type="materials_examined">
<paragraph id="A11736E9FFBFFFE7FF58EEF244B86E98" blockId="2.[151,1127,641,729]" pageId="2" pageNumber="497">
<emphasis id="93DCEAFBFFBFFFE7FF58EEF2475F6EFF" bold="true" box="[151,285,683,702]" pageId="2" pageNumber="497">Type material.</emphasis>
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: ♁, ‘ 
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: 
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, 
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station (
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, 
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), 
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.a.s.l., 
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, 
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leg.ʼ (coll. 
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).
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<emphasis id="93DCEAFBFFBFFFE7FF58EEA847D56F49" bold="true" box="[151,407,753,776]" pageId="2" pageNumber="497">Diagnostic characters.</emphasis>
<taxonomicName id="66A84D6AFFBFFFE7FE6FEEA8448C6F49" authorityName="Kvifte" authorityYear="2015" box="[416,718,753,776]" class="Insecta" family="Psychodidae" genus="Psychomasina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="paedagogica">
<emphasis id="93DCEAFBFFBFFFE7FE6FEEA8448C6F49" box="[416,718,753,776]" italics="true" pageId="2" pageNumber="497">Psychomasina paedagogica</emphasis>
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can be separated from all other 
<taxonomicName id="66A84D6AFFBFFFE7FBF8EEA847496F69" authorityName="Jezek" authorityYear="2004" class="Insecta" family="Psychodidae" genus="Psychomasina" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="497" phylum="Arthropoda" rank="genus">
<emphasis id="93DCEAFBFFBFFFE7FBF8EEA847496F69" italics="true" pageId="2" pageNumber="497">Psychomasina</emphasis>
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species by the eyebridges separated by one facet diameter, wing veins without expanded areas, gonostylus with a row of five spiniform sensilla subapically on mesal surface and without terminal hook, and distal phallomeres of aedeagus without teeth or spines on mesal or lateral surfaces (
<figureCitation id="39932A6CFFBFFFE7FE67EF2844616FC9" box="[424,547,881,904]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="2" pageNumber="497">Figs 1, 2, 4</figureCitation>
).
</paragraph>
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<subSubSection id="E9B26562FFBFFFE1FF58EFC8426E6DD2" lastPageId="4" lastPageNumber="499" pageId="2" pageNumber="497" type="description">
<paragraph id="A11736E9FFBFFFE7FF58EFC844E468C9" blockId="2.[151,1127,753,1576]" pageId="2" pageNumber="497">
<emphasis id="93DCEAFBFFBFFFE7FF58EFC847226FE9" bold="true" box="[151,352,912,936]" pageId="2" pageNumber="497">
Description. 
<emphasis id="93DCEAFBFFBFFFE7FEE7EFC947226FE9" bold="true" box="[296,352,912,936]" italics="true" pageId="2" pageNumber="497">Male</emphasis>
</emphasis>
(n = 1). Head (
<figureCitation id="39932A6CFFBFFFE7FDCFEFC8447F6FE9" box="[512,573,913,936]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="2" pageNumber="497">Fig. 1</figureCitation>
) about as long as wide, vertex covering about 0.3 of total head length; five pairs of postocular setae present on dorsal side, reaching fourth facet column from middle, only slightly larger than other setae on vertex; eyebridge comprised of four rows of facets, separated by one facet diameter; interocular suture present, broadly U-shaped; frontal scar patch nearly square, posteriorly with line of hairs reaching interocular suture; sides of frons area sharply delimited from rest of face by keels; clypeus with anterior margin straight; labellae bulbous with seven setae; palps with only segments 1–3 present, length of preserved palp segments 80: 155: 150.
</paragraph>
<paragraph id="A11736E9FFBFFFE7FF7AE8C8458968A9" blockId="2.[151,1127,753,1576]" pageId="2" pageNumber="497">Antennae not completely preserved; scapus cylindrical, pedicel round; flagellomeres subsymmetrically nodiform with node onion-shaped; slightly projected on mesal side; length of preserved antennal segments 100: 65: 115: 100: 95; ascoids not preserved.</paragraph>
<paragraph id="A11736E9FFBFFFE7FF7AE8A845E26909" blockId="2.[151,1127,753,1576]" pageId="2" pageNumber="497">
Thorax with dorsum, anepisternum and scutellum densely setose; laterotergite sparsely haired; other thoracic sclerites bare; coxae with dorsal stripe of setae; mid coxae with strong mesal protuberance covered in pores (
<figureCitation id="39932A6CFFBFFFE7FDE2E968442C6909" box="[557,622,1329,1352]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="2" pageNumber="497">Fig. 3</figureCitation>
); legs otherwise not present.
</paragraph>
<paragraph id="A11736E9FFBFFFE7FF7AE908472B69A9" blockId="2.[151,1127,753,1576]" pageId="2" pageNumber="497">
Wing (
<figureCitation id="39932A6CFFBFFFE7FECFE90847036929" box="[256,321,1361,1384]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="2" pageNumber="497">Fig. 2</figureCitation>
) 
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long, 
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wide; costa with single break, thickened basally over S c; S c elongate, weakly curved, almost reaching C, distally narrowed; R 5 terminating nearly in wing apex; radial fork distal to medial fork, apex of CuA 2 between medial and radial fork; jugum broadly rounded triangular; section of R 1 that runs parallel to Sc reduced; crossveins reduced.
</paragraph>
<paragraph id="A11736E9FFBFFFE6FF7AE9A8479B6D8E" blockId="2.[151,1127,753,1576]" lastBlockId="3.[113,1089,216,463]" lastPageId="3" lastPageNumber="498" pageId="2" pageNumber="497">
Genitalia (
<figureCitation id="39932A6CFFBFFFE7FEE7E9A847286A49" box="[296,362,1521,1544]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="2" pageNumber="497">Fig. 4</figureCitation>
) with hypandrium narrow, connected laterally to gonocoxal condyles; gonocoxite about two thirds of gonostylus length; dorsally without mesal seta or projections; gonocoxal condyles broadly fused with paramere, forming a broad ventral bridge; gonostylus bent towards the lateral side; basally bulb-shaped with long tube-shaped main shaft, apical fifth narrowed, blade-like; row of five sensillae present subapically on mesal surface; aedeagus with basiphallus narrow in dorsal and ventral views, distally Y-shaped; distiphallus consisting of two symmetrical phallomeres shaped like the number 3 or the calligraphic letter G, jointed to basiphallus; with distal mesal joint to ventral epandrial plate and basal lateral joint to basiphallus; parameral sclerite elongate V-shaped, weakly sclerotized distally; more than twice as long as distiphallus.
</paragraph>
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<paragraph id="A11736E9FFBEFFE6FFBEE9A947236A79" blockId="3.[113,1090,1520,1592]" pageId="3" pageNumber="498">
Figs 1–5. 
<taxonomicName id="66A84D6AFFBEFFE6FF09E9A947F66A42" authority="Kvifte, 2015" authorityName="Kvifte" authorityYear="2015" box="[198,436,1520,1539]" class="Insecta" family="Psychodidae" genus="Psychomasina" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="paedagogica" status="sp. nov.">
<emphasis id="93DCEAFBFFBEFFE6FF09E9A947F66A42" box="[198,436,1520,1539]" italics="true" pageId="3" pageNumber="498">Psychomasina paedagogica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88EF5780FFBEFFE6FE76E9A947BC6A42" box="[441,510,1520,1539]" pageId="3" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)
</paragraph>
</caption>
<paragraph id="A11736E9FFB9FFE1FF7AEC85426E6DD2" blockId="4.[151,1127,220,883]" pageId="4" pageNumber="499">
Epandrium (
<figureCitation id="39932A6CFFB9FFE1FEF4EC8547396CB2" box="[315,379,220,243]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="4" pageNumber="499">Fig. 5</figureCitation>
) about as wide as long, with long curved anterior projections half the length of basal epandrial body; aperture not evident; ventral epandrial plate large, U-shaped, well-developed; surstyli tapering, strongly curved in the dorsoventral plane, with tenacula along entire dorsal surface; tenacula of varying length, distalmost one almost as long as surstylus; with feathery apices; proctiger with hypoproct diamond-shaped, epiproct densely setose, small and circular; areas of surstylus flanking proctiger with dense microsetae.
</paragraph>
</subSubSection>
<subSubSection id="E9B26562FFB9FFE1FF58EDC5427C6DF2" box="[151,1086,412,435]" pageId="4" pageNumber="499" type="biology_ecology">
<paragraph id="A11736E9FFB9FFE1FF58EDC5427C6DF2" blockId="4.[151,1127,220,883]" box="[151,1086,412,435]" pageId="4" pageNumber="499">
<emphasis id="93DCEAFBFFB9FFE1FF58EDC546B16DF2" bold="true" box="[151,243,412,435]" pageId="4" pageNumber="499">Biology.</emphasis>
The only known specimen was collected in a Malaise trap in a montane forest.
</paragraph>
</subSubSection>
<subSubSection id="E9B26562FFB9FFE1FF58EDE544956E52" pageId="4" pageNumber="499" type="etymology">
<paragraph id="A11736E9FFB9FFE1FF58EDE544956E52" blockId="4.[151,1127,220,883]" pageId="4" pageNumber="499">
<emphasis id="93DCEAFBFFB9FFE1FF58EDE5475A6D92" bold="true" box="[151,280,444,467]" pageId="4" pageNumber="499">Etymology.</emphasis>
The specific epithet is an adjective, derived from Greek 
<emphasis id="93DCEAFBFFB9FFE1FCB2EDE245B86D92" box="[893,1018,443,467]" italics="true" pageId="4" pageNumber="499">παΙΔαγωγός</emphasis>
(= 
<emphasis id="93DCEAFBFFB9FFE1FBEEEDE546956DB2" italics="true" pageId="4" pageNumber="499">paidagogos</emphasis>
), pedagogue; referring to the possible morphological insights the species reveals into the ontogenesis of parameral appendages and surstyli.
</paragraph>
</subSubSection>
<subSubSection id="E9B26562FFB9FFE1FF58EE45454B6F32" pageId="4" pageNumber="499" type="discussion">
<paragraph id="A11736E9FFB9FFE1FF58EE45454B6F32" blockId="4.[151,1127,220,883]" pageId="4" pageNumber="499">
<emphasis id="93DCEAFBFFB9FFE1FF58EE4547446E72" bold="true" box="[151,262,540,563]" pageId="4" pageNumber="499">Remarks.</emphasis>
In 
<taxonomicName id="66A84D6AFFB9FFE1FEE6EE4544166E72" authority="Kvifte, 2015" authorityName="Kvifte" authorityYear="2015" box="[297,596,540,563]" class="Insecta" family="Psychodidae" genus="Psychomasina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="species" species="paedagogica" status="sp. nov.">
<emphasis id="93DCEAFBFFB9FFE1FEE6EE4544166E72" box="[297,596,540,563]" italics="true" pageId="4" pageNumber="499">Psychomasina paedagogica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="88EF5780FFB9FFE1FD95EE4544ED6E72" box="[602,687,540,563]" pageId="4" pageNumber="499" rank="species">sp. nov.</taxonomicNameLabel>
, the surstylus is jointed by one sclerite to the ventral epandrial plate and by another to the lateral sides of the proctiger. This may be interpreted as evidence of a dual ontogenetic origin of the surstyli in 
<taxonomicName id="66A84D6AFFB9FFE1FCA2EE0545B56E32" authorityName="Newman" authorityYear="1834" box="[877,1015,604,627]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
, or at least in some 
<taxonomicName id="66A84D6AFFB9FFE1FF39EE25473F6ED2" authorityName="Newman" authorityYear="1834" box="[246,381,636,659]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
– rather than being an appendage derived only from the epandrium (surstylus, see 
<bibRefCitation id="C5394B18FFB9FFE1FEF8EEC5440E6EF2" author="CURLER G. R. &amp; MOULTON J. K." box="[311,588,668,691]" pageId="4" pageNumber="499" pagination="603 - 616" refId="ref3576" refString="CURLER G. R. &amp; MOULTON J. K. 2012: Phylogeny of psychodid subfamilies (Diptera: Psychodidae) inferred from nuclear DNA sequences with a review of morphological evidence for relationships. Systematic Entomology 37: 603 - 616." type="journal article" year="2012">CURLER &amp; MOULTON 2012</bibRefCitation>
) or an appendage derived only from the proctiger (cercus, e.g. 
<bibRefCitation id="C5394B18FFB9FFE1FED1EEE5447B6E92" author="QUATE L. W. &amp; VOCKEROTH J. R." box="[286,569,700,723]" pageId="4" pageNumber="499" pagination="293 - 300" refId="ref3950" refString="QUATE L. W. &amp; VOCKEROTH J. R. 1981: Psychodidae. Pp. 293 - 300. In: McALPINE J. F., PETERSON B. V., SHEWELL G. E., TESKEY H. J., VOCKEROTH J. R. &amp; WOOD D. M. (eds): Manual of Nearctic Diptera, Vol. 1. Biosystematics Research Institute, Ottawa, 647 pp." type="book chapter" year="1981">QUATE &amp; VOCKEROTH 1981</bibRefCitation>
), it may be a fusion of both. A dual origin could also explain the difference in sclerotization, pigmentation and setation of the basal mesal areas of the surstyli in some 
<taxonomicName id="66A84D6AFFB9FFE1FE47EEA5444B6F52" baseAuthorityName="KVIFTE" baseAuthorityYear="2014" box="[392,521,764,787]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="tribe" tribe="Pericomaini">Pericomaini</taxonomicName>
(e.g. 
<bibRefCitation id="C5394B18FFB9FFE1FD8BEEA545466F52" author="KVIFTE G. M. &amp; IVKOVIC M. &amp; KLARIC A." box="[580,772,764,787]" pageId="4" pageNumber="499" pagination="57 - 67" refId="ref3848" refString="KVIFTE G. M., IVKOVIC M. &amp; KLARIC A. 2013: New records of moth flies (Diptera: Psychodidae) from Croatia, with the description of Berdeniella keroveci sp. nov. Zootaxa 3737: 57 - 67." type="journal article" year="2013">KVIFTE et al. 2013</bibRefCitation>
, 
<figureCitation id="39932A6CFFB9FFE1FCC1EEA545226F52" box="[782,864,764,787]" captionStart="Figs 1–5" captionStartId="3.[113,150,1520,1539]" captionTargetBox="[117,1080,485,1498]" captionTargetId="figure-9@3.[113,1089,479,1498]" captionTargetPageId="3" captionText="Figs 1–5. Psychomasina paedagogica sp. nov., holotype, male. 1 – head; 2 – wing; 3 – mid coxa; 4 – aedeagus and gonopods; 5 – epandrium and proctiger. For abbreviations see Material and methods. Length of scale bars in μm: 100 (1, 4, 5), 250 (2), 50 (3)" figureDoi="http://doi.org/10.5281/zenodo.4468291" httpUri="https://zenodo.org/record/4468291/files/figure.png" pageId="4" pageNumber="499">Fig. 2D</figureCitation>
) and Paramormiini (
<figureCitation id="39932A6CFFB9FFE1FBF4EEA546E46F72" captionStart="Figs 6–12. 6–9" captionStartId="5.[113,150,1492,1511]" captionTargetBox="[123,1074,488,1471]" captionTargetId="figure-109@5.[113,1088,482,1480]" captionTargetPageId="5" captionText="Figs 6–12. 6–9 – Psychomasina violina sp. nov., holotype, male. 6 – head; 7 – wing; 8 – aedeagus and gonopods; 9 – epandrium and proctiger. 10–12 – aedeagal joint shape in Psychomasina spp.: 10 – P. paedagogica sp. nov., 11 – P. violina sp. nov., 12 – P. armata Ježek, 2004 (Fig. 12 redrawn from JEŽEK 2004). For abbreviations see Material and methods. Length of scale bars in μm: 100 (6, 8, 9) 250 (7)" figureDoi="http://doi.org/10.5281/zenodo.4468293" httpUri="https://zenodo.org/record/4468293/files/figure.png" pageId="4" pageNumber="499">Fig. 9</figureCitation>
, cr sst). The occurence of the latter kind of surstylus texture heterogeneity, however, is a variable character in 
<taxonomicName id="66A84D6AFFB9FFE1FEB9EF6547BF6F12" authorityName="Newman" authorityYear="1834" box="[374,509,828,851]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="499" phylum="Arthropoda" rank="subFamily" subFamily="Psychodinae">Psychodinae</taxonomicName>
and must be examined for a greater taxon sample in order to determine whether it has any phylogenetic implications.
</paragraph>
</subSubSection>
</treatment>
</document>