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<document id="C4F5EE4B7E912D5F9419190E77EDF5C0" ID-DOI="10.1649/0010-065x-67.4.539" ID-ISSN="1938-4394" ID-Zenodo-Dep="10104039" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630165750611" checkinUser="carolina" docAuthor="Gómez, R. Antonio" docDate="2013" docId="03BDE56C7407272DBAEDFAEEFB88F90E" docLanguage="en" docName="0010-065X-67.4.539.pdf.imf" docOrigin="The Coleopterists Bulletin 67 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-67.4.539" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Cryptocephalomorpha genieri Baehr 1997" docType="treatment" docVersion="3" lastPageNumber="542" masterDocId="FF849D147406272EBA58FFA3FFE7FFCF" masterDocTitle="The Afrotropical Species Cryptocephalomorpha genieri Baehr (Coleoptera: Carabidae: Pseudomorphinae): Discovery and Description of the Male and Relationships of the Species" masterLastPageNumber="543" masterPageNumber="539" pageNumber="540" updateTime="1699628267248" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="7707D08EC8C99472B1074E02E4A7D688">
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<mods:title id="B7A7AAED001F7BC90870EE557F620821">The Afrotropical Species Cryptocephalomorpha genieri Baehr (Coleoptera: Carabidae: Pseudomorphinae): Discovery and Description of the Male and Relationships of the Species</mods:title>
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<mods:namePart id="355185DC62C3D136E165CB670C3A3D53">Gómez, R. Antonio</mods:namePart>
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<mods:typeOfResource id="527237436E1A7A85AEB7208B403AE676">text</mods:typeOfResource>
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<mods:title id="74A07B5860378732F9C26A65E27198D5">The Coleopterists Bulletin</mods:title>
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<mods:part id="8142B2AFEE9AFF3ED3028C4B5115BBE4">
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<mods:date id="5FC8D72B184980EDACB959C1ECC01A75">2013</mods:date>
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<mods:number id="437ED4A1D4B277C5E38B3962AD2FCC74">2013-12-31</mods:number>
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<mods:number id="3F76E294B749FE444E7E0AEB520EE1BD">67</mods:number>
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<mods:number id="27B3628F509395880B54E609AA14FDEF">4</mods:number>
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<mods:url id="C53793250210CEB4A4C5401FF74F949F">http://dx.doi.org/10.1649/0010-065x-67.4.539</mods:url>
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<mods:classification id="E05A2E27571B56C76F973D235C2BE83C">journal article</mods:classification>
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<mods:identifier id="0FB78ADB162A92316E762EA94EFC3EBC" type="DOI">10.1649/0010-065x-67.4.539</mods:identifier>
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<mods:identifier id="AD38BF565423D6D6B72F0448475981DD" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="A82372738E8E99E2FACE70A641CAFE9B" type="Zenodo-Dep">10104039</mods:identifier>
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</mods:mods>
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<treatment id="03BDE56C7407272DBAEDFAEEFB88F90E" LSID="urn:lsid:plazi:treatment:03BDE56C7407272DBAEDFAEEFB88F90E" httpUri="http://treatment.plazi.org/id/03BDE56C7407272DBAEDFAEEFB88F90E" lastPageId="3" lastPageNumber="542" pageId="1" pageNumber="540">
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<subSubSection id="C30E07F17407272FBAEDFAEEFD87FAAC" box="[181,608,1357,1379]" pageId="1" pageNumber="540" type="nomenclature">
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<paragraph id="8BAB547A7407272FBAEDFAEEFD87FAAC" blockId="1.[181,608,1357,1379]" box="[181,608,1357,1379]" pageId="1" pageNumber="540">
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<heading id="D0E3E3167407272FBAEDFAEEFD87FAAC" box="[181,608,1357,1379]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="540" reason="2">
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<taxonomicName id="4C142FF97407272FBAEDFAEEFD87FAAC" ID-CoL="ZVT5" authority="Baehr, 1997" authorityName="Baehr" authorityYear="1997" box="[181,608,1357,1379]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="genieri">
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<emphasis id="B96088687407272FBAEDFAEEFD87FAAC" bold="true" box="[181,608,1357,1379]" pageId="1" pageNumber="540">
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<emphasis id="B96088687407272FBAEDFAEEFE3BFAAD" bold="true" box="[181,476,1357,1378]" italics="true" pageId="1" pageNumber="540">Cryptocephalomorpha genieri</emphasis>
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<bibRefCitation id="EF85298B7407272FBBBCFAEDFD87FAAC" author="Baehr" box="[484,608,1358,1379]" pageId="1" pageNumber="540" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr, 1997</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E07F17407272FBACFFADBFD99FA42" box="[151,638,1400,1422]" pageId="1" pageNumber="540" type="reference_group">
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<paragraph id="8BAB547A7407272FBACFFADBFD99FA42" blockId="1.[151,638,1400,1422]" box="[151,638,1400,1422]" pageId="1" pageNumber="540">
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<treatmentCitationGroup id="AB0473547407272FBACFFADBFD99FA42" box="[151,638,1400,1422]" pageId="1" pageNumber="540">
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<treatmentCitation id="0AB5726B7407272FBACFFADBFD99FA42" author="Baehr" box="[151,638,1400,1422]" page="382" pageId="1" pageNumber="540" year="1997">
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<taxonomicName id="4C142FF97407272FBACFFADBFD99FA42" ID-CoL="ZVT5" authority="Baehr 1997: 381 - 382" authorityName="Baehr" authorityYear="1997" box="[151,638,1400,1422]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="genieri">
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<emphasis id="B96088687407272FBACFFADBFE4AFA42" box="[151,429,1400,1421]" italics="true" pageId="1" pageNumber="540">Cryptocephalomorpha genieri</emphasis>
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<bibRefCitation id="EF85298B7407272FBBECFADBFD99FA42" author="Baehr" box="[436,638,1400,1422]" pageId="1" pageNumber="540" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr 1997: 381–382</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30E07F17407272DBAF7FA00FB88F90E" lastPageId="3" lastPageNumber="542" pageId="1" pageNumber="540" type="description">
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<paragraph id="8BAB547A7407272FBAF7FA00FE9EF943" blockId="1.[151,638,1442,1729]" pageId="1" pageNumber="540">
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<emphasis id="B96088687407272FBAF7FA00FD9AFA77" bold="true" box="[175,637,1442,1464]" pageId="1" pageNumber="540">
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Supplement to Original Description (
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<figureCitation id="132F48FF7407272FB84AFA00FD97FA77" box="[530,624,1442,1464]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="1.[709,741,887,906]" captionStartId-1="2.[197,229,867,886]" captionTargetBox-0="[739,1171,243,863]" captionTargetBox-1="[208,1162,195,846]" captionTargetId-0="figure-618@1.[685,1172,183,921]" captionTargetId-1="figure-371@2.[172,1193,183,857]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Cryptocephalomorpha genieri, adult male, dorsal habitus. Scale bar = 1 mm." captionText-1="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi-0="http://doi.org/10.5281/zenodo.10104041" figureDoi-1="http://doi.org/10.5281/zenodo.10104043" httpUri-0="https://zenodo.org/record/10104041/files/figure.png" httpUri-1="https://zenodo.org/record/10104043/files/figure.png" pageId="1" pageNumber="540">Figs. 1–2</figureCitation>
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).
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</emphasis>
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The description of the female
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<typeStatus id="54AFEAD87407272FBB9EFA1EFDFCFA1D" box="[454,539,1469,1490]" pageId="1" pageNumber="540" type="holotype">holotype</typeStatus>
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by
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<bibRefCitation id="EF85298B7407272FB81CFA1EFF3FFA22" author="Baehr" pageId="1" pageNumber="540" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
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covers most of the morphological characters of the males examined here, excluding the sexually dimorphic and male genitalic characters, which are described below. These additional specimens, including the
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<specimenCount id="9D129FF37407272FBBCCF9E1FE13F998" box="[404,500,1602,1623]" pageId="1" pageNumber="540" type="adult">four adult</specimenCount>
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females, also provide the opportunity to describe intraspecfic variation of
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<taxonomicName id="4C142FF97407272FBB4AF9D4FE94F943" box="[274,371,1655,1676]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="genieri">
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<emphasis id="B96088687407272FBB4AF9D4FE94F943" box="[274,371,1655,1676]" italics="true" pageId="1" pageNumber="540">C. genieri</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BAB547A7407272FBAF7F931FB83FB2C" blockId="1.[151,638,1442,1729]" lastBlockId="1.[685,1172,991,1729]" pageId="1" pageNumber="540">
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Apart from primary sexually dimorphic characters, males differ from females in possessing a pilose, glandular area on abdominal sternum VII (
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<figureCitation id="132F48FF7407272FB8ECFC59FD10FBC0" box="[692,759,1018,1039]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="1" pageNumber="540">Fig. 2J</figureCitation>
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) similar to males of
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<taxonomicName id="4C142FF97407272FB9E4FC5AFC81FBE6" authority="Ritsema" authorityName="Ritsema" authorityYear="1875" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="540" phylum="Arthropoda" rank="species" species="gaverei">
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<emphasis id="B96088687407272FB9E4FC5AFD1BFBE6" italics="true" pageId="1" pageNumber="540">Cryptocephalomorpha gaverei</emphasis>
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Ritsema
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</taxonomicName>
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, in lacking squamulae on tarsomere III on all tarsi, and in being slightly smaller. Although little variation was observed in antennal shape (
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<figureCitation id="132F48FF7407272FB922FBC7FC2BFBB6" box="[890,972,1124,1145]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="1" pageNumber="540">Fig. 2C</figureCitation>
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) and shape of the terminal palpomeres of the maxillary and labial palpi (
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<figureCitation id="132F48FF7407272FB8B2FB3AFCC5FB61" box="[746,802,1177,1198]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="1" pageNumber="540">Fig. 2</figureCitation>
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A-B) between the sexes, these features in the male were also illustrated in order to follow the taxonomic style of Baehr’ s (1997) plates.
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</paragraph>
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<paragraph id="8BAB547A7407272FB89DFB4AFC67FAA7" blockId="1.[685,1172,991,1729]" pageId="1" pageNumber="540">
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<emphasis id="B96088687407272FB89DFB4AFC27FB31" bold="true" box="[709,960,1257,1278]" pageId="1" pageNumber="540">Measurements of Males.</emphasis>
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TL =
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<quantity id="4CECF99F7407272FBE56FB4AFB77FB31" box="[1038,1168,1256,1278]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.4" metricValueMin="4.3" pageId="1" pageNumber="540" unit="mm" value="4.35" valueMax="4.4" valueMin="4.3">4.3–4.4 mm</quantity>
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; GW = 2.0–
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<quantity id="4CECF99F7407272FB97EFAA0FC9CFAD7" box="[806,891,1283,1304]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="540" unit="mm" value="2.1">2.1 mm</quantity>
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; PL =
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<quantity id="4CECF99F7407272FB99FFAA0FBAEFAD7" box="[967,1097,1283,1305]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="540" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
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; PW = 2.0–
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<quantity id="4CECF99F7407272FB88EFABDFCC7FAFC" box="[726,800,1310,1331]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="540" unit="mm" value="2.1">2.1 mm</quantity>
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; HW =
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<quantity id="4CECF99F7407272FB93CFABDFC30FAFC" box="[868,983,1309,1331]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="1" pageNumber="540" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.3–1.4 mm</quantity>
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; EW = 0.9–1.0 mm; TL/GW = 2.09–2.15; PL/PW = 0.70–0.71; HW/EW = 1.40–1.44.
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</paragraph>
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<paragraph id="8BAB547A7407272FB89DFACDFC69FA1D" blockId="1.[685,1172,991,1729]" pageId="1" pageNumber="540">
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<emphasis id="B96088687407272FB89DFACDFC8CFA4C" bold="true" box="[709,875,1390,1411]" pageId="1" pageNumber="540">Ventral Surface.</emphasis>
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Last visible abdominal sternum (sternum VII) of male with a large, more or less circular, glandular, densely pilose area near basal margin (
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<figureCitation id="132F48FF7407272FB962FA1EFC65FA1D" box="[826,898,1469,1490]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="1" pageNumber="540">Fig. 2J</figureCitation>
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).
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</paragraph>
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<paragraph id="8BAB547A7407272FB89DFA7BFBF5F9BD" blockId="1.[685,1172,991,1729]" pageId="1" pageNumber="540">
|
|||
|
|
<emphasis id="B96088687407272FB89DFA7BFD1BFA22" bold="true" box="[709,764,1496,1517]" pageId="1" pageNumber="540">Legs.</emphasis>
|
|||
|
|
Male protarsus not broadened. Tarsomere IV of all legs ventrally with biseriate squamose setae; pro- and mesotarsomeres II-III each with two ventral spatulate setae, metatarsomeres II-III each with only a single ventral spatulate seta, located proximally on the tarsomere.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7407272CB89DF9D4FE87FA22" blockId="1.[685,1172,991,1729]" lastBlockId="2.[173,659,1071,1729]" lastPageId="2" lastPageNumber="541" pageId="1" pageNumber="540">
|
|||
|
|
<emphasis id="B96088687407272FB89DF9D4FCB8F943" bold="true" box="[709,863,1655,1676]" pageId="1" pageNumber="540">Male Genitalia.</emphasis>
|
|||
|
|
Ring sclerite rather broad, ovoid, slightly asymmetrical, with short, broad apex, with large, asymmetrical, deeply emarginate base (
|
|||
|
|
<figureCitation id="132F48FF7404272CBAECFB8CFF19FB8B" box="[180,254,1071,1092]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2E</figureCitation>
|
|||
|
|
). Sternum VIII symmetrical, not membranous apically, base strongly emarginate, basal angles slightly angulate (
|
|||
|
|
<figureCitation id="132F48FF7404272CBBF5FBC7FDE7FBB6" box="[429,512,1124,1145]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2D</figureCitation>
|
|||
|
|
). Median lobe rather short, symmetrical; in lateral aspect, slen- der and curved medially before broadening subapically, apex slightly narrowed, tip short, bent ventrad (
|
|||
|
|
<figureCitation id="132F48FF7404272CBB5DFB6DFEB1FB2C" box="[261,342,1230,1251]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2G</figureCitation>
|
|||
|
|
). Median lobe in ventral aspect bottle-shaped, very broad basally, margins curved basally, slightly divergent apically, apically narrowed, apex broadly rounded (
|
|||
|
|
<figureCitation id="132F48FF7404272CBBB2FABDFDDEFAFC" box="[490,569,1310,1331]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2F</figureCitation>
|
|||
|
|
). Orifice symmetrical, broad; internal sac rather simply folded (
|
|||
|
|
<figureCitation id="132F48FF7404272CBAA5FAF0FEDBFAA7" box="[253,316,1363,1384]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2</figureCitation>
|
|||
|
|
F-G). Lateral lobes elongate, dissimilar, left broader than right, apices bearing fine setae. Left lateral lobe sinuate, apically curved dorsad, apex broadly rounded (
|
|||
|
|
<figureCitation id="132F48FF7404272CBBAAFA01FDA1FA78" box="[498,582,1442,1463]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2H</figureCitation>
|
|||
|
|
). Right lateral lobe rather straight and slender, apex rounded (
|
|||
|
|
<figureCitation id="132F48FF7404272CBB55FA7BFEB3FA22" box="[269,340,1496,1517]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541">Fig. 2I</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF6B04F27404272CBA9DFCC0FEEFFC19" ID-DOI="http://doi.org/10.5281/zenodo.10104043" ID-Zenodo-Dep="10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="2" pageNumber="541" startId="2.[197,229,867,886]" targetBox="[208,1162,195,846]" targetPageId="2">
|
|||
|
|
<paragraph id="8BAB547A7404272CBA9DFCC0FEEFFC19" blockId="2.[173,1193,867,982]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CBA9DFCC0FEE5FCB9" bold="true" box="[197,258,867,886]" pageId="2" pageNumber="541">Fig. 2.</emphasis>
|
|||
|
|
<taxonomicName id="4C142FF97404272CBB4FFCC0FDC5FCB9" box="[279,546,867,886]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687404272CBB4FFCC0FDC5FCB9" box="[279,546,867,886]" italics="true" pageId="2" pageNumber="541">Cryptocephalomorpha genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BAB547A7404272CBA9DFA51FDAEF90E" blockId="2.[173,659,1071,1729]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CBA9DFA51FECEF9C8" bold="true" box="[197,297,1522,1543]" pageId="2" pageNumber="541">Variation.</emphasis>
|
|||
|
|
Apart from the sexually dimorphic characters, there is slight variation in body size between individual specimens and minor differences in coloration. On all specimens, the area adjacent to the elytral suture is slightly lighter than the disk to a varying degree, and in some specimens the yellowish apical area on the elytra is lighter and more extensive than in others.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7404272CB883FB8CFB7EFB61" blockId="2.[707,1194,1071,1729]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CB883FB8CFCDFFB8B" bold="true" box="[731,824,1071,1092]" pageId="2" pageNumber="541">Vivipary.</emphasis>
|
|||
|
|
Not confirmed in the material examined. Two of the four available adult females were dissected in order to confirm the identification of the material as members of
|
|||
|
|
<taxonomicName id="4C142FF97404272CB98EFBDDFBD0FB5C" box="[982,1079,1150,1171]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687404272CB98EFBDDFBD0FB5C" box="[982,1079,1150,1171]" italics="true" pageId="2" pageNumber="541">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Numerous eggs were observed, but no larvae were present.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7404272CB883FB10FCA1FA78" blockId="2.[707,1194,1071,1729]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CB883FB10FCC2FB07" bold="true" box="[731,805,1203,1224]" pageId="2" pageNumber="541">Habits.</emphasis>
|
|||
|
|
As reported by
|
|||
|
|
<bibRefCitation id="EF85298B7404272CB989FB10FBB1FB07" author="Baehr" box="[977,1110,1203,1224]" pageId="2" pageNumber="541" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
, next to nothing is known of the way of life of
|
|||
|
|
<taxonomicName id="4C142FF97404272CBE1AFB6EFB44FB2D" box="[1090,1187,1229,1250]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687404272CBE1AFB6EFB44FB2D" box="[1090,1187,1229,1250]" italics="true" pageId="2" pageNumber="541">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. All specimens thus far known have been collected using light traps, when collecting methods are specified. This series from
|
|||
|
|
<collectingCountry id="F30314EA7404272CB98BFABDFBFCFAFC" box="[979,1051,1310,1331]" name="Zambia" pageId="2" pageNumber="541">Zambia</collectingCountry>
|
|||
|
|
was collected in late October and November, around the same time of year that the female
|
|||
|
|
<typeStatus id="54AFEAD87404272CB9B0FAF0FBDBFAA7" box="[1000,1084,1363,1384]" pageId="2" pageNumber="541" type="holotype">holotype</typeStatus>
|
|||
|
|
was taken in
|
|||
|
|
<collectingCountry id="F30314EA7404272CB884FACEFCBEFA4D" box="[732,857,1389,1410]" name="South Africa" pageId="2" pageNumber="541">South Africa</collectingCountry>
|
|||
|
|
. The remaining specimens reported in literature were collected in January and December (
|
|||
|
|
<bibRefCitation id="EF85298B7404272CB891FA01FCDDFA78" author="Baehr" box="[713,826,1442,1463]" pageId="2" pageNumber="541" pagination="195 - 199" refId="ref3101" refString="Baehr, M. 2009. A new and a little known species of Cryptocephalomorpha Ritsema. 11 th Supplement to the " Revision of the Pseudomorphinae of the Australian Region. " (Insecta: Coleoptera: Pseudomorphinae). Stuttgarter Beitrage Naturkunde A, Neue Serie 2: 195 - 199." type="journal article" year="2009">Baehr 2009</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7404272CB883FA1EFB4DF943" blockId="2.[707,1194,1071,1729]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CB883FA1EFC16FA1D" bold="true" box="[731,1009,1469,1490]" pageId="2" pageNumber="541">Geographical Distribution.</emphasis>
|
|||
|
|
<taxonomicName id="4C142FF97404272CB9A6FA1EFCB9FA23" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687404272CB9A6FA1EFCB9FA23" italics="true" pageId="2" pageNumber="541">Cryptocephalomorpha genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<materialsCitation id="3B7C5E277404272CB930FA7BFCF3F9F3" country="South Africa" location="south central Africa" pageId="2" pageNumber="541" specimenCount="1">
|
|||
|
|
is known from a few collection sites in tropical
|
|||
|
|
<location id="8ECB02A17404272CB930FA51FBD2F9C8" LSID="urn:lsid:plazi:treatment:03BDE56C7407272DBAEDFAEEFB88F90E:8ECB02A17404272CB930FA51FBD2F9C8" box="[872,1077,1522,1543]" country="South Africa" name="south central Africa" pageId="2" pageNumber="541">south central Africa</location>
|
|||
|
|
: one in the far north of the
|
|||
|
|
<collectingCountry id="F30314EA7404272CB903F9AEFBAAF9ED" box="[859,1101,1549,1570]" name="South Africa" pageId="2" pageNumber="541">Republic of South Africa</collectingCountry>
|
|||
|
|
(the
|
|||
|
|
<typeStatus id="54AFEAD87404272CBED8F9AEFB4EF9ED" box="[1152,1193,1549,1570]" pageId="2" pageNumber="541">type</typeStatus>
|
|||
|
|
locality)
|
|||
|
|
</materialsCitation>
|
|||
|
|
, one in
|
|||
|
|
<collectingCountry id="F30314EA7404272CB93BF984FC24F9F3" box="[867,963,1575,1596]" name="Botswana" pageId="2" pageNumber="541">Botswana</collectingCountry>
|
|||
|
|
, and two nearby localities in
|
|||
|
|
<collectingCountry id="F30314EA7404272CB950F9E1FCB7F998" box="[776,848,1602,1623]" name="Zambia" pageId="2" pageNumber="541">Zambia</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="132F48FF7404272CB938F9E1FC7BF998" box="[864,924,1602,1623]" captionStart="Fig" captionStartId="3.[175,207,836,855]" captionTargetBox="[187,629,199,810]" captionTargetId="figure-602@3.[151,638,183,813]" captionTargetPageId="3" captionText="Fig. 3. Map of southern Africa, showing positions of localities for Cryptocephalomorpha genieri based on examined specimens and published reports." figureDoi="http://doi.org/10.5281/zenodo.10104046" httpUri="https://zenodo.org/record/10104046/files/figure.png" pageId="2" pageNumber="541">Fig. 3</figureCitation>
|
|||
|
|
).
|
|||
|
|
<materialsCitation id="3B7C5E277404272CB9E9F9E1FB42F943" country="Democratic Republic of the Congo" location="The" pageId="2" pageNumber="541" specimenCount="1">
|
|||
|
|
<location id="8ECB02A17404272CB9E9F9E1FC31F998" LSID="urn:lsid:plazi:treatment:03BDE56C7407272DBAEDFAEEFB88F90E:8ECB02A17404272CB9E9F9E1FC31F998" box="[945,982,1602,1623]" country="Democratic Republic of the Congo" name="The" pageId="2" pageNumber="541">The</location>
|
|||
|
|
range of this species also likely includes nearby countries such as
|
|||
|
|
<collectingCountry id="F30314EA7404272CBED3F9FFFBE8F943" name="Democratic Republic of the Congo" pageId="2" pageNumber="541">the Democratic Republic of the Congo</collectingCountry>
|
|||
|
|
and
|
|||
|
|
<collectingCountry id="F30314EA7404272CBE65F9D4FB42F943" box="[1085,1189,1655,1676]" name="Zimbabwe" pageId="2" pageNumber="541">Zimbabwe</collectingCountry>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7404272CB883F931FB4FF90E" blockId="2.[707,1194,1071,1729]" pageId="2" pageNumber="541">
|
|||
|
|
<emphasis id="B96088687404272CB883F931FCBCF968" bold="true" box="[731,859,1682,1703]" pageId="2" pageNumber="541">Recognition.</emphasis>
|
|||
|
|
Presently,
|
|||
|
|
<taxonomicName id="4C142FF97404272CB98FF932FBA7F969" box="[983,1088,1681,1702]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687404272CB98FF932FBA7F969" box="[983,1088,1681,1702]" italics="true" pageId="2" pageNumber="541">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is one of three species of
|
|||
|
|
<taxonomicName id="4C142FF97404272CB93BF90FFBDDF90E" box="[867,1082,1708,1729]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="541" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687404272CB93BF90FFBDDF90E" box="[867,1082,1708,1729]" italics="true" pageId="2" pageNumber="541">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
that lack a
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DB885FF61FE8EFB6E" blockId="3.[733,1173,194,321]" lastBlockId="3.[151,638,951,1238]" pageId="3" pageNumber="542">
|
|||
|
|
tinctly punctate. Lateral margin of mandible excised and with acute spine. Male lateral lobes asetose; male median lobe in ventral aspect simply shaped, without subapical sinuation.
|
|||
|
|
<collectingCountry id="F30314EA7405272DB885FE8FFCD3FE8E" box="[733,820,300,321]" name="Australia" pageId="3" pageNumber="542">Australia</collectingCountry>
|
|||
|
|
, New
|
|||
|
|
<collectingCountry id="F30314EA7405272DB92AFE8FFC5FFE8E" box="[882,952,300,321]" name="Guinea-Bissau" pageId="3" pageNumber="542">Guinea</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F30314EA7405272DB99AFE8FFBB8FE8E" box="[962,1119,300,321]" name="Solomon Islands" pageId="3" pageNumber="542">Solomon Islands</collectingCountry>
|
|||
|
|
...... 5 distinct elytral spot, and because of the discovery of the male of
|
|||
|
|
<taxonomicName id="4C142FF97405272DBB6AFC72FE72FC29" box="[306,405,977,998]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DBB6AFC72FE72FC29" box="[306,405,977,998]" italics="true" pageId="3" pageNumber="542">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the males of all these species are known. Therefore, although the key presented by
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBB15FBA4FE37FBD3" author="Baehr" box="[333,464,1031,1052]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
does not require significant changes in order to identify adults of
|
|||
|
|
<taxonomicName id="4C142FF97405272DBAEFFB98FEFCFB9F" box="[183,283,1083,1104]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DBAEFFB98FEFCFB9F" box="[183,283,1083,1104]" italics="true" pageId="3" pageNumber="542">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the incorporation of male characters and their respective illustrations presented here as well as in
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBB3AFBD2FE0FFB49" author="Baehr" box="[354,488,1137,1158]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
should aid in future identifications.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF6B04F27405272DBAF7FCE7FDE8FC48" ID-DOI="http://doi.org/10.5281/zenodo.10104046" ID-Zenodo-Dep="10104046" httpUri="https://zenodo.org/record/10104046/files/figure.png" pageId="3" pageNumber="542" startId="3.[175,207,836,855]" targetBox="[187,629,199,810]" targetPageId="3">
|
|||
|
|
<paragraph id="8BAB547A7405272DBAF7FCE7FDE8FC48" blockId="3.[151,638,836,903]" pageId="3" pageNumber="542">
|
|||
|
|
<emphasis id="B96088687405272DBAF7FCE7FF0AFC98" bold="true" box="[175,237,836,855]" pageId="3" pageNumber="542">Fig. 3.</emphasis>
|
|||
|
|
Map of southern Africa, showing positions of localities for
|
|||
|
|
<taxonomicName id="4C142FF97405272DBB7DFCFFFDCFFCA0" box="[293,552,860,879]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DBB7DFCFFFDCFFCA0" box="[293,552,860,879]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
based on examined specimens and published reports.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BAB547A7405272DBAF7FB05FEB6FB19" blockId="3.[151,638,951,1238]" pageId="3" pageNumber="542">
|
|||
|
|
Modification of second statement of couplet 1 of
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBACFFB62FEFDFB19" author="Baehr" box="[151,282,1217,1238]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
: 377:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DBACFFB53FD9AFABF" blockId="3.[151,638,1264,1392]" pageId="3" pageNumber="542">
|
|||
|
|
– Elytra without distinct light spot. Apex of aedeagus regularly acute, or broadly rounded in ventral aspect (
|
|||
|
|
<figureCitation id="132F48FF7405272DBB22FA85FE22FAF4" box="[378,453,1318,1339]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="3" pageNumber="542">Fig. 2F</figureCitation>
|
|||
|
|
) and slightly bent ventrad in lateral aspect (
|
|||
|
|
<figureCitation id="132F48FF7405272DBB9BFAE3FDF4FA9A" box="[451,531,1344,1365]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="3" pageNumber="542">Fig. 2G</figureCitation>
|
|||
|
|
). Distribution different..................................................4
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DBAF7FA26FD99FA55" blockId="3.[175,638,1413,1434]" box="[175,638,1413,1434]" pageId="3" pageNumber="542">
|
|||
|
|
Modification of couplet 4 of
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBB9FFA26FDAEFA55" author="Baehr" box="[455,585,1413,1434]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
: 378:
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DBACFFA16FD99F94B" blockId="3.[151,639,1461,1729]" pageId="3" pageNumber="542">
|
|||
|
|
4. Larger, wider species, length>
|
|||
|
|
<quantity id="4CECF99F7405272DBBACFA16FDD7FA05" box="[500,560,1461,1482]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="542" unit="mm" value="4.0">4 mm</quantity>
|
|||
|
|
. Lateral margin of elytra near base with elongate setae. Surface not distinctly punctate. Lateral margin of mandible regularly curved. Male lateral lobes with 3–4 fine setae at apex (
|
|||
|
|
<figureCitation id="132F48FF7405272DBB87F9BCFDFDF9FB" box="[479,538,1567,1588]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="3" pageNumber="542">Fig. 2</figureCitation>
|
|||
|
|
H-I); male median lobe in ventral aspect bottle-shaped (
|
|||
|
|
<figureCitation id="132F48FF7405272DBA97F9F6FEFAF9A5" box="[207,285,1621,1642]" captionStart="Fig" captionStartId="2.[197,229,867,886]" captionTargetBox="[208,1162,195,846]" captionTargetId="figure-371@2.[172,1193,183,857]" captionTargetPageId="2" captionText="Fig. 2. Cryptocephalomorpha genieri, male morphological characters. A) Terminal palpomeres of maxillary palpus, B) Terminal palpomeres of labial palpus, C) Antennomeres V–XI, D) Abdominal sternum VIII, E) Ring sclerite, F) Median lobe, ventral aspect, G) Median lobe, lateral aspect, H) Left lateral lobe, lateral aspect, I) Right lateral lobe, lateral aspect, J) Abdominal sternum VII, ventral aspect, with arrow indicating glandular, pilose area near base." figureDoi="http://doi.org/10.5281/zenodo.10104043" httpUri="https://zenodo.org/record/10104043/files/figure.png" pageId="3" pageNumber="542">Fig. 2F</figureCitation>
|
|||
|
|
).
|
|||
|
|
<collectingCountry id="F30314EA7405272DBB69F9F6FE52F9A5" box="[305,437,1621,1642]" name="South Africa" pageId="3" pageNumber="542">South Africa</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F30314EA7405272DBB9AF9F6FDC1F9A5" box="[450,550,1621,1642]" name="Botswana" pageId="3" pageNumber="542">Botswana</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F30314EA7405272DB86BF9F6FD9AF9A5" box="[563,637,1621,1642]" name="Zambia" pageId="3" pageNumber="542">Zambia</collectingCountry>
|
|||
|
|
.............................................
|
|||
|
|
<taxonomicName id="4C142FF97405272DBBB4F9CCFD99F94B" authority="Baehr" authorityName="Baehr" box="[492,638,1647,1668]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DBBB4F9CCFD99F94B" bold="true" box="[492,638,1647,1668]" pageId="3" pageNumber="542">
|
|||
|
|
<emphasis id="B96088687405272DBBB4F9CCFDD4F94B" bold="true" box="[492,563,1647,1668]" italics="true" pageId="3" pageNumber="542">genieri</emphasis>
|
|||
|
|
Baehr
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DBACFF932FD9AF90E" blockId="3.[151,639,1461,1729]" pageId="3" pageNumber="542">
|
|||
|
|
– Smaller, narrower species, length <
|
|||
|
|
<quantity id="4CECF99F7405272DB872F931FD9DF968" box="[554,634,1682,1703]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="542" unit="mm" value="3.5">3.5 mm</quantity>
|
|||
|
|
. Lateral margin of elytra asetose. Surface dis-
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DB89DFEC2FB8CFC70" blockId="3.[685,1173,353,1729]" pageId="3" pageNumber="542">
|
|||
|
|
<emphasis id="B96088687405272DB89DFEC2FC7EFEB9" bold="true" box="[709,921,353,374]" pageId="3" pageNumber="542">Phylogenetic Aspects.</emphasis>
|
|||
|
|
<bibRefCitation id="EF85298B7405272DB9FEFEC2FBFBFEB9" author="Baehr" box="[934,1052,353,374]" pageId="3" pageNumber="542" pagination="11 - 17" refId="ref2927" refString="Baehr, M. 1994. Phylogenetic relations and biogeography of the genera of Pseudomorphinae (Coleoptera, Carabidae) [pp. 11 - 17]. In: Carabid Beetles: Ecology and Evolution (K. Desender, M. Dufrene, M. Loreau, M. L. Luff, and J. - P. Maelfait, editors). Kluwer Academic Press, Dordrecht, The Netherlands." type="book chapter" year="1994">Baehr (1994</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBE7DFEC2FBB2FEB9" author="Baehr" box="[1061,1109,353,374]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">1997</bibRefCitation>
|
|||
|
|
) listed several reasons for the challenges associated with inferring the evolutionary relationships within the
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8F5FE12FCBBFE09" box="[685,860,433,454]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="subFamily" subFamily="Pseudomorphinae">Pseudomorphinae</taxonomicName>
|
|||
|
|
, including the relative scarcity of specimens (depending on the species), the divergent morphological features of the group that include many putative character reductions, and, perhaps most importantly, the unsettled sister group of the subfamily, which greatly impacts postulation of the polarity of characters. The systematic position of
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8ACFDC8FC20FD4F" box="[756,967,619,640]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB8ACFDC8FC20FD4F" box="[756,967,619,640]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is not fully resolved, but current thinking suggests
|
|||
|
|
<taxonomicName id="4C142FF97405272DB998FD26FB74FD55" box="[960,1171,645,666]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB998FD26FB74FD55" box="[960,1171,645,666]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is either sister to the clade that includes
|
|||
|
|
<taxonomicName id="4C142FF97405272DBE73FD03FD01FD1F" authority="Hope" authorityName="Hope" class="Insecta" family="Carabidae" genus="Adelotopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DBE73FD03FB73FD7A" box="[1067,1172,672,693]" italics="true" pageId="3" pageNumber="542">Adelotopus</emphasis>
|
|||
|
|
Hope
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8ABFD19FC35FD1F" authority="Notman" authorityName="Notman" authorityYear="1925" box="[755,978,698,720]" class="Insecta" family="Carabidae" genus="Cainogenion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB8ABFD19FC92FD00" box="[755,885,698,719]" italics="true" pageId="3" pageNumber="542">Cainogenion</emphasis>
|
|||
|
|
Notman
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="4C142FF97405272DBE54FD19FCF9FD25" authority="Waterhouse" authorityName="Waterhouse" class="Insecta" family="Carabidae" genus="Paussotropus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DBE54FD19FB74FD00" box="[1036,1171,698,719]" italics="true" pageId="3" pageNumber="542">Paussotropus</emphasis>
|
|||
|
|
Waterhouse
|
|||
|
|
</taxonomicName>
|
|||
|
|
or represents an independent lineage within
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8A8FD53FC70FCCA" box="[752,919,752,773]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="subFamily" subFamily="Pseudomorphinae">Pseudomorphinae</taxonomicName>
|
|||
|
|
that convergently evolved several characters states similar to those characteristic of this clade (
|
|||
|
|
<bibRefCitation id="EF85298B7405272DB91DFC86FC53FCF5" author="Baehr" box="[837,948,805,826]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr 1997</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<taxonomicName id="4C142FF97405272DB99CFC86FD08FC9B" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DB99CFC86FD08FC9B" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is an important taxon because it is thought to be the sister to the remaining members of the genus (
|
|||
|
|
<bibRefCitation id="EF85298B7405272DB8ACFCD6FC8FFC45" author="Baehr" box="[756,872,885,906]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr 1997</bibRefCitation>
|
|||
|
|
), and therefore it is significant for inferring the ground plan morphology for the genus and its placement within the subfamily.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BAB547A7405272DB89DFC67FB88F90E" blockId="3.[685,1173,353,1729]" pageId="3" pageNumber="542">
|
|||
|
|
As stated above, members of
|
|||
|
|
<taxonomicName id="4C142FF97405272DB991FC67FB73FC16" box="[969,1172,964,985]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB991FC67FB73FC16" box="[969,1172,964,985]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are very similar to one another, and species diagnoses rely on detailed examination of structural features (
|
|||
|
|
<emphasis id="B96088687405272DB8B0FBB7FCE4FBE6" box="[744,771,1044,1065]" italics="true" pageId="3" pageNumber="542">e.g</emphasis>
|
|||
|
|
., presence or absence of microreticulation, shape of the median lobe). However, within the subfamily, the genus is unusual in possessing greatly reduced mouthparts (
|
|||
|
|
<emphasis id="B96088687405272DB9D4FBC0FC43FBB7" box="[908,932,1123,1144]" italics="true" pageId="3" pageNumber="542">i.e</emphasis>
|
|||
|
|
., small mandibles and potentially non-functional laciniae), a concealed labrum, and a hypognathous head capsule among other autapomorphic character states (
|
|||
|
|
<bibRefCitation id="EF85298B7405272DBE4FFB17FB6DFB06" author="Baehr" box="[1047,1162,1204,1225]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr 1997</bibRefCitation>
|
|||
|
|
). The discovery of the adult male confirms the hypothesis that
|
|||
|
|
<taxonomicName id="4C142FF97405272DB90AFB4BFC5EFB32" box="[850,953,1256,1277]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="genieri">
|
|||
|
|
<emphasis id="B96088687405272DB90AFB4BFC5EFB32" box="[850,953,1256,1277]" italics="true" pageId="3" pageNumber="542">C. genieri</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is sister to all other
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8F5FAA0FC6BFAD7" box="[685,908,1283,1304]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB8F5FAA0FC6BFAD7" box="[685,908,1283,1304]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Plesiomorphic character states discovered here are the presence of apical setae on the male lateral lobes (absent from all other
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8BDFAF1FC5FFAA8" box="[741,952,1362,1383]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B96088687405272DB8BDFAF1FC5FFAA8" box="[741,952,1362,1383]" italics="true" pageId="3" pageNumber="542">Cryptocephalomorpha</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
excluding
|
|||
|
|
<taxonomicName id="4C142FF97405272DBE7DFAF1FB60FAA8" authorityName="Ritsema" authorityYear="1875" box="[1061,1159,1362,1383]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="gaverei">
|
|||
|
|
<emphasis id="B96088687405272DBE7DFAF1FB60FAA8" box="[1061,1159,1362,1383]" italics="true" pageId="3" pageNumber="542">C. gaverei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), the bilaterally symmetrical shape of the male median lobe with a symmetrical apex, and the fairly elongate orifice. The presence of a densely pilose, glandular area on abdominal sternum VII in the male was not coded by
|
|||
|
|
<bibRefCitation id="EF85298B7405272DB9B8FA7BFB86FA22" author="Baehr" box="[992,1121,1496,1517]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
, perhaps because it was only known in the male of
|
|||
|
|
<taxonomicName id="4C142FF97405272DB8F5F9AFFCF5F9EE" authorityName="Ritsema" authorityYear="1875" box="[685,786,1548,1569]" class="Insecta" family="Carabidae" genus="Cryptocephalomorpha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="542" phylum="Arthropoda" rank="species" species="gaverei">
|
|||
|
|
<emphasis id="B96088687405272DB8F5F9AFFCF5F9EE" box="[685,786,1548,1569]" italics="true" pageId="3" pageNumber="542">C. gaverei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but it may be an important character for future reconstructed phylogenies. The confirmation of the systematic position of this taxon provides further evidence that the genus likely evolved in the Southern Hemisphere and attained its current distribution following the breakup of Gondwana, as hypothesized by
|
|||
|
|
<bibRefCitation id="EF85298B7405272DB9B1F90FFB8CF90E" author="Baehr" box="[1001,1131,1708,1729]" pageId="3" pageNumber="542" pagination="1 - 510" refId="ref3002" refString="Baehr, M. 1997. Revision of the Pseudomorphinae of the Australian Region 2. The genera Pseudomorpha Kirby, Adelotopus Hope, Cainogenion Notman, Paussotropus Waterhouse, and Cryptocephalomorpha Ritsema. Taxonomy, phylogeny, zoogeography (Insecta, Coleoptera, Carabidae). Spixiana Supplement 23: 1 - 510." type="journal article" year="1997">Baehr (1997)</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|