treatments-xml/data/03/85/63/03856357CC73FFD4FFD059F9FB1F6781.xml

<document id="A21EBF17C69F2FAD82C4D91896AA5B98" ID-DOI="10.5281/zenodo.11512244" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512244" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1717719192119" checkinUser="felipe" docAuthor="Zug, George R." docDate="2015" docId="03856357CC73FFD4FFD059F9FB1F6781" docLanguage="en" docName="ProcCalifAcadSci.62.5.135-190.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (5)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Kalophrynus orangensis Dutta, Ahmed, and Das 2000" docType="treatment" docVersion="3" lastPageNumber="172" masterDocId="FFBC1B2FCC57FFF1FFAB5C0AFF8C611C" masterDocTitle="Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs" masterLastPageNumber="190" masterPageNumber="135" pageNumber="171" updateTime="1720720661996" updateUser="carolina" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F445C97AA833A1FA640B89D5FCBDF76B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C03E1132CCBE2565761E694024165E7D">
<mods:title id="4F9A7BF48166FD9330A1F28CD6E09E52">Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs</mods:title>
</mods:titleInfo>
<mods:name id="BCFA3B5193ADBD30C16AEF3BB5C30FCE" type="personal">
<mods:role id="20080BD1D7D3967F95013CBBBD6432E2">
<mods:roleTerm id="20303B5EBB0028853D98D6472988091D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="74DCBDEFB4C108F592F6C4EF15EACF69">Zug, George R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1B54DF66AE6B6D3BFB0924107406FD70">text</mods:typeOfResource>
<mods:relatedItem id="33A43215E2577F4C481D3C9ACC74FF70" type="host">
<mods:titleInfo id="8714315948211D1E7EF3D5E4C986357D">
<mods:title id="B41CB03A5E52FBE324303EF3B844616F">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="DCD9941005226A7365DB3F74040FD30D">
<mods:date id="7E788B872936387A92EF13D13069501D">2015</mods:date>
<mods:detail id="C9166A9430879A5BD827D414FBB6EC7C" type="pubDate">
<mods:number id="6900942576E63BE0269729912503C463">2015-04-15</mods:number>
</mods:detail>
<mods:detail id="336D62ACAAB216187DB90381C614D167" type="volume">
<mods:number id="91FCD0C885BE070F8FEC0F005241E0FB">62</mods:number>
</mods:detail>
<mods:detail id="D9A7BBAB31839C38DDA9132B11FF0970" type="issue">
<mods:number id="F2F5F905A115FAABF0D9F6416206D8AD">5</mods:number>
</mods:detail>
<mods:extent id="DA84228A4E7652A657101D02146FDE35" unit="page">
<mods:start id="8A536968143DD5E056A609C0114E202B">135</mods:start>
<mods:end id="CD58FC394FB8BA9D1FE29CC8BB948160">190</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="58418A4B12EE6FA668BB1C25AE401CA2">journal article</mods:classification>
<mods:identifier id="43803742F132C77CF6E134E41D2B1FE2" type="DOI">10.5281/zenodo.11512244</mods:identifier>
<mods:identifier id="ABEFE279DC2C8B6F99047C4D83F8BFDA" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="D0B0FDA59A85998375A593C254F9734C" type="Zenodo-Dep">11512244</mods:identifier>
<mods:identifier id="7E234E909C9E6A3162D7F6410903C30A" type="ZooBank">AD651C54-BC39-4A21-A0CA-3B9B8309A0BB</mods:identifier>
</mods:mods>
<treatment id="03856357CC73FFD4FFD059F9FB1F6781" LSID="urn:lsid:plazi:treatment:03856357CC73FFD4FFD059F9FB1F6781" httpUri="http://treatment.plazi.org/id/03856357CC73FFD4FFD059F9FB1F6781" lastPageId="37" lastPageNumber="172" pageId="36" pageNumber="171">
<subSubSection id="C33681CACC73FFD5FFD059F9FB026717" box="[123,1166,1523,1547]" pageId="36" pageNumber="171" type="nomenclature">
<paragraph id="8B93D241CC73FFD5FFD059F9FB026717" blockId="36.[123,1166,1523,1577]" box="[123,1166,1523,1547]" pageId="36" pageNumber="171">
<heading id="D0DB652DCC73FFD5FFD059F9FB026717" box="[123,1166,1523,1547]" centered="true" fontSize="10" level="2" pageId="36" pageNumber="171" reason="4">
<taxonomicName id="4C2CA9C2CC73FFD5FFD059F9FB026717" ID-CoL="6NJSW" authority="Dutta, Ahmed, and Das, 2000 Indian Striped Sticky Frog" authorityName="Dutta, Ahmed, and Das" authorityYear="2000" box="[123,1166,1523,1547]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="36" pageNumber="171" phylum="Chordata" rank="species" species="orangensis">
<emphasis id="B9580E53CC73FFD5FFD059F9FB026717" bold="true" box="[123,1166,1523,1547]" pageId="36" pageNumber="171">
<emphasis id="B9580E53CC73FFD5FFD059F9FE046717" bold="true" box="[123,392,1523,1547]" italics="true" pageId="36" pageNumber="171">Kalophrynus orangensis</emphasis>
<bibRefCitation id="EFBDAFB0CC73FFD5FE3B59F9FD536717" author="DUTTA, S. K. &amp; AHMED M. F. &amp; I. DAS" box="[400,735,1523,1547]" pageId="36" pageNumber="171" pagination="67 - 74" refId="ref37371" refString="DUTTA, S. K., AHMED M. F., AND I. DAS 2000. Kalophrynus (Anura: Microhylidae), a new genus for India, with the description of a new species, Kalophrynus orangensis, from Assam State. Hamadryad 25 (2): 67 - 74." type="journal article" year="2000">Dutta, Ahmed, and Das, 2000</bibRefCitation>
Indian Striped Sticky Frog
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC73FFD5FFD05A19FF5E6735" box="[123,210,1555,1577]" pageId="36" pageNumber="171" type="description">
<paragraph id="8B93D241CC73FFD5FFD05A19FF5E6735" blockId="36.[123,1166,1523,1577]" box="[123,210,1555,1577]" pageId="36" pageNumber="171">
<figureCitation id="1317CEC4CC73FFD5FFD05A19FF406735" box="[123,204,1555,1577]" captionStart="FIGURE" captionStartId="19.[155,167,219,238]" captionTargetBox="[696,1176,216,1012]" captionTargetId="figure-544@19.[696,1176,216,1012]" captionTargetPageId="19" captionText="FIGURE 6. Burmese distribution of voucher specimens of Kalophrynus anya sp. nov. in northern Myanmar and K. inter- lineatus in southern and peninsular Myanmar. The three report- ed localities of K. orangensis are also shown; all lie within the Brahmaputra river valley. Solid circles denote specimens exam- ined in this study; open circles are literature or unexamined museum records; and stars represent type localities of available names." figureDoi="http://doi.org/10.5281/zenodo.11512252" httpUri="https://zenodo.org/record/11512252/files/figure.png" pageId="36" pageNumber="171">Figure 6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC73FFD5FFD05A33FCD36773" pageId="36" pageNumber="171" type="reference_group">
<paragraph id="8B93D241CC73FFD5FFD05A33FCD36773" blockId="36.[123,1174,1593,1647]" pageId="36" pageNumber="171">
<taxonomicName id="4C2CA9C2CC73FFD5FFD05A33FD156753" ID-CoL="6NJSW" authority="Dutta, Ahmed, and Das, 2000" authorityName="Dutta, Ahmed, and Das" authorityYear="2000" box="[123,665,1593,1615]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="36" pageNumber="171" phylum="Chordata" rank="species" species="orangensis">
<emphasis id="B9580E53CC73FFD5FFD05A33FEE46753" box="[123,360,1593,1615]" italics="true" pageId="36" pageNumber="171">Kalophrynus orangensis</emphasis>
<materialsCitation id="3B44D81CCC73FFD5FEDA5A30FCD56773" country="India" latitude="26.5" location="Orang National Park" longLatPrecision="1241" longitude="92.25" municipality="Darrang District" pageId="36" pageNumber="171" specimenCount="1" stateProvince="Assam">
<bibRefCitation id="EFBDAFB0CC73FFD5FEDA5A30FD156753" author="DUTTA, S. K. &amp; AHMED M. F. &amp; I. DAS" box="[369,665,1593,1615]" pageId="36" pageNumber="171" pagination="67 - 74" refId="ref37371" refString="DUTTA, S. K., AHMED M. F., AND I. DAS 2000. Kalophrynus (Anura: Microhylidae), a new genus for India, with the description of a new species, Kalophrynus orangensis, from Assam State. Hamadryad 25 (2): 67 - 74." type="journal article" year="2000">Dutta, Ahmed, and Das, 2000</bibRefCitation>
, 
<emphasis id="B9580E53CC73FFD5FD0C5A33FC966752" box="[679,794,1593,1614]" italics="true" pageId="36" pageNumber="171">Hamadryad</emphasis>
25:68 [
<typeStatus id="54976CE3CC73FFD5FCC15A30FC1F6753" box="[874,915,1594,1615]" pageId="36" pageNumber="171">type</typeStatus>
locality: “ 
<location id="8EF3849ACC73FFD5FC555A30FF4B6773" LSID="urn:lsid:plazi:treatment:03856357CC73FFD4FFD059F9FB1F6781:8EF3849ACC73FFD5FC555A30FF4B6773" country="India" latitude="26.5" longLatPrecision="1241" longitude="92.25" municipality="Darrang District" name="Orang National Park" pageId="36" pageNumber="171" stateProvince="Assam">Orang National Park</location>
(
<geoCoordinate id="EE18B486CC73FFD5FF7D5A53FEAA6773" box="[214,294,1624,1647]" degrees="26" direction="north" minutes="30" orientation="latitude" pageId="36" pageNumber="171" precision="925" value="26.5">26°30′N</geoCoordinate>
; 
<geoCoordinate id="EE18B486CC73FFD5FE985A50FE086773" box="[307,388,1625,1647]" degrees="92" direction="east" minutes="15" orientation="longitude" pageId="36" pageNumber="171" precision="925" value="92.25">92°15´E</geoCoordinate>
), 
<collectingMunicipality id="6BF7483BCC73FFD5FE335A50FDB96773" box="[408,565,1625,1647]" pageId="36" pageNumber="171">Darrang District</collectingMunicipality>
, 
<collectingRegion id="49E81CA3CC73FFD5FDE95A50FD0F6773" box="[578,643,1626,1647]" country="India" name="Assam" pageId="36" pageNumber="171">Assam</collectingRegion>
, north-eastern 
<collectingCountry id="F33B92D1CC73FFD5FCBE5A53FCCA6773" box="[789,838,1625,1647]" name="India" pageId="36" pageNumber="171">India</collectingCountry>
”]
</materialsCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC73FFD4FF005A8FFF34624B" lastPageId="37" lastPageNumber="172" pageId="36" pageNumber="171" type="materials_examined">
<paragraph id="8B93D241CC73FFD5FF005A8FFB196781" blockId="36.[123,1173,1669,1726]" box="[171,1173,1669,1694]" pageId="36" pageNumber="171">
<materialsCitation id="3B44D81CCC73FFD5FF005A8FFC576781" box="[171,987,1669,1694]" collectionCode="TYPE, MATERIAL" collectorName="Kolkata A" country="India" location="India" pageId="36" pageNumber="171" specimenCount="1" typeStatus="holotype">
<emphasis id="B9580E53CC73FFD5FF005A8FFED66780" bold="true" box="[171,346,1669,1693]" pageId="36" pageNumber="171">
<smallCapsWord id="8D75449DCC73FFD5FF005A8FFF686780" baselines="1687,1687" box="[171,228,1669,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="36" pageNumber="171">
<collectionCode id="ED3D4A84CC73FFD5FF005A8FFF686780" box="[171,228,1669,1693]" pageId="36" pageNumber="171">TYPE</collectionCode>
</smallCapsWord>
<smallCapsWord id="8D75449DCC73FFD5FF465A83FED66780" baselines="1687" box="[237,346,1673,1692]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="36" pageNumber="171">
<collectionCode id="ED3D4A84CC73FFD5FF465A83FED66780" box="[237,346,1673,1692]" pageId="36" pageNumber="171">MATERIAL</collectionCode>
</smallCapsWord>
</emphasis>
.— 
<smallCapsWord id="8D75449DCC73FFD5FE295A8CFE7C6780" baselines="1687,1687" box="[386,496,1670,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Holotype" pageId="36" pageNumber="171">
<typeStatus id="54976CE3CC73FFD5FE295A8CFE7C6780" box="[386,496,1670,1693]" pageId="36" pageNumber="171" type="holotype">HOLOTYPE</typeStatus>
</smallCapsWord>
: Zoological Survey of 
<collectingCountry id="F33B92D1CC73FFD5FD5A5A8FFCA46781" box="[753,808,1669,1693]" name="India" pageId="36" pageNumber="171">India</collectingCountry>
, 
<collectorName id="26D9B797CC73FFD5FC9C5A8FFC2A6782" box="[823,934,1669,1694]" pageId="36" pageNumber="171">Kolkata A</collectorName>
9087
</materialsCitation>
. 
<materialsCitation id="3B44D81CCC73FFD5FC405A8CFB196781" box="[1003,1173,1669,1693]" collectionCode="ZSI" country="India" pageId="36" pageNumber="171" specimenCount="1" typeStatus="paratype">
<smallCapsWord id="8D75449DCC73FFD5FC405A8CFBD16780" baselines="1687,1687" box="[1003,1117,1670,1693]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Paratypes" pageId="36" pageNumber="171">
<typeStatus id="54976CE3CC73FFD5FC405A8CFBD16780" box="[1003,1117,1670,1693]" pageId="36" pageNumber="171" type="paratype">PARATYPES</typeStatus>
</smallCapsWord>
: 
<collectionCode id="ED3D4A84CC73FFD5FBC65A8FFB196781" box="[1133,1173,1669,1693]" pageId="36" pageNumber="171">ZSI</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="8B93D241CC73FFD5FFD05AACFD1A67A1" blockId="36.[123,1173,1669,1726]" box="[123,662,1701,1726]" pageId="36" pageNumber="171">A9088–91. All specimens by original designation.</paragraph>
<paragraph id="8B93D241CC72FFD4FF005CD1FBC360C8" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">
<smallCapsWord id="8D75449DCC72FFD4FF005CD1FEA561ED" baselines="237,236" box="[171,297,219,243]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Definition" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FF005CD1FEA561ED" bold="true" box="[171,297,219,243]" pageId="37" pageNumber="172">DEFINITION</emphasis>
</smallCapsWord>
.— Medium-sized, adult females to 
<quantity id="4CD47FA4CC72FFD4FD135CD1FCA661EE" box="[696,810,219,243]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="37" pageNumber="172" unit="mm" value="36.5" valueMax="38.0" valueMin="35.0">35–38 mm</quantity>
<collectionCode id="ED3D4A84CC72FFD4FC9E5CD1FCEB61EF" box="[821,871,219,243]" pageId="37" pageNumber="172">SVL</collectionCode>
(
<emphasis id="B9580E53CC72FFD4FCD15CD6FC0B61EF" box="[890,903,220,243]" italics="true" pageId="37" pageNumber="172">n</emphasis>
= 3), adult males 
<quantity id="4CD47FA4CC72FFD4FBF85CD1FF28600F" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="37" pageNumber="172" unit="mm" value="37.0" valueMax="38.0" valueMin="36.0">36–38 mm</quantity>
<collectionCode id="ED3D4A84CC72FFD4FF1A5CF1FF6F600F" box="[177,227,251,275]" pageId="37" pageNumber="172">SVL</collectionCode>
(
<emphasis id="B9580E53CC72FFD4FF535CF6FE89600F" box="[248,261,252,275]" italics="true" pageId="37" pageNumber="172">n</emphasis>
= 2); head modest length NA % HeadL/ 
<collectionCode id="ED3D4A84CC72FFD4FD7E5CF1FC8B600F" box="[725,775,251,275]" pageId="37" pageNumber="172">SVL</collectionCode>
; head wider than long 118–133 % HeadW/ 
<collectionCode id="ED3D4A84CC72FFD4FF7F5D16FE8A6028" box="[212,262,284,308]" pageId="37" pageNumber="172">SVL</collectionCode>
; snout moderately broad?38–43 % SnW/HeadL; naris closer to snout than to eye NA?72–80 % NarEye/SnEye; eye moderate?42–44 % EyeD/HeadL; tympanum visible and nearly as large as eye?83–94 % Tymp/ 
<collectionCode id="ED3D4A84CC72FFD4FE6C5D56FE756068" box="[455,505,348,372]" pageId="37" pageNumber="172">SVL</collectionCode>
; moderate length forelimb NA % Forarm/ 
<collectionCode id="ED3D4A84CC72FFD4FC785D56FB896068" box="[979,1029,348,372]" pageId="37" pageNumber="172">SVL</collectionCode>
and forearm equal crus length NA % Forarm/CrusL; hindlimb moderately long NA % HndlL/ 
<collectionCode id="ED3D4A84CC72FFD4FBA65D76FBB36088" box="[1037,1087,380,404]" pageId="37" pageNumber="172">SVL</collectionCode>
, NA % CrusL/ 
<collectionCode id="ED3D4A84CC72FFD4FF6E5D96FF7B60A8" box="[197,247,412,436]" pageId="37" pageNumber="172">SVL</collectionCode>
, and NA % CrusL/ThghL; hindfoot well developed NA % HndfL/CrusL; limb measurements not reported. Question marks preceding data indicate uncertainty of original data.
</paragraph>
<paragraph id="8B93D241CC72FFD4FF005DD7FE196309" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">Vomerine teeth presence or absence not reported; palatal fold morphology not reported; tongue elongate with rounded tip.</paragraph>
<paragraph id="8B93D241CC72FFD4FF005E17FEDB63AA" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">Fingers not webbed; lengths 3&gt;2&gt;1&gt;4; tips rounded and not dilated; subarticular tubercles round, one beneath fingers 1, 2 and 4, two beneath finger 3; fleshy palmar tubercle, size and shape not reported. Toes webbed at base, lengths 4&gt;3&gt;5&gt;2&gt;1; tips rounded, not dilated; large round subarticular tubercles, one of 1 and 2, two on 3 and 5, and 3 on 4; small, round inner and outer metatarsal tubercles.</paragraph>
<paragraph id="8B93D241CC72FFD4FF005EB5FF34624B" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">In life, dorsum reddish brown to cream with dark brown reverse-Y shaped mark with base between eyes, bifurcating on nape and limbs extending diagonally to top of hindlimbs, distinct narrow white stripe from above eyes to shoulder and extending diagonally to mid inguina; black white-edged inguinal ocellus. Venter yellowish cream with black speckled throat and anterior chest.</paragraph>
</subSubSection>
<subSubSection id="C33681CACC72FFD4FF005F6AFBD4626B" box="[171,1112,863,888]" pageId="37" pageNumber="172" type="etymology">
<paragraph id="8B93D241CC72FFD4FF005F6AFBD4626B" blockId="37.[123,1175,219,1693]" box="[171,1112,863,888]" pageId="37" pageNumber="172">
<smallCapsWord id="8D75449DCC72FFD4FF005F6AFEB9626A" baselines="882,881" box="[171,309,864,888]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FF005F6AFEB9626A" bold="true" box="[171,309,864,888]" pageId="37" pageNumber="172">ETYMOLOGY</emphasis>
</smallCapsWord>
.— The specific name derives from the 
<typeStatus id="54976CE3CC72FFD4FD4A5F6AFC836264" box="[737,783,864,888]" pageId="37" pageNumber="172">type</typeStatus>
locality, Orang National Park.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC72FFD4FF005F8AFF6A65A5" pageId="37" pageNumber="172" type="distribution">
<paragraph id="8B93D241CC72FFD4FF005F8AFC8662E4" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">
<smallCapsWord id="8D75449DCC72FFD4FF005F8AFECF628A" baselines="914,913" box="[171,323,896,920]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FF005F8AFECF628A" bold="true" box="[171,323,896,920]" pageId="37" pageNumber="172">DISTRIBUTION</emphasis>
</smallCapsWord>
.— (
<figureCitation id="1317CEC4CC72FFD4FEDF5F8AFE3B6284" box="[372,439,896,920]" captionStart="FIGURE" captionStartId="19.[155,167,219,238]" captionTargetBox="[696,1176,216,1012]" captionTargetId="figure-544@19.[696,1176,216,1012]" captionTargetPageId="19" captionText="FIGURE 6. Burmese distribution of voucher specimens of Kalophrynus anya sp. nov. in northern Myanmar and K. inter- lineatus in southern and peninsular Myanmar. The three report- ed localities of K. orangensis are also shown; all lie within the Brahmaputra river valley. Solid circles denote specimens exam- ined in this study; open circles are literature or unexamined museum records; and stars represent type localities of available names." figureDoi="http://doi.org/10.5281/zenodo.11512252" httpUri="https://zenodo.org/record/11512252/files/figure.png" pageId="37" pageNumber="172">Fig. 6</figureCitation>
) Presently known from the 
<typeStatus id="54976CE3CC72FFD4FD515F8AFCA46284" box="[762,808,896,920]" pageId="37" pageNumber="172">type</typeStatus>
locality in 
<location id="8EF3849ACC72FFD4FC045F8AFB196284" LSID="urn:lsid:plazi:treatment:03856357CC73FFD4FFD059F9FB1F6781:8EF3849ACC72FFD4FC045F8AFB196284" box="[943,1173,896,920]" country="India" county="Brahmaputra River" latitude="26.5" longLatPrecision="1241" longitude="92.25" municipality="All" name="Orang National Park" pageId="37" pageNumber="172" stateProvince="Assam">Orang National Park</location>
(
<geoCoordinate id="EE18B486CC72FFD4FF2F5FAAFF5362AB" box="[132,223,928,952]" degrees="26" direction="north" minutes="30" orientation="latitude" pageId="37" pageNumber="172" precision="925" value="26.5">26°30´N</geoCoordinate>
, 
<geoCoordinate id="EE18B486CC72FFD4FF455FAAFECB62AB" box="[238,327,928,952]" degrees="92" direction="east" minutes="15" orientation="longitude" pageId="37" pageNumber="172" precision="925" value="92.25">92°15´E</geoCoordinate>
) in central 
<collectingRegion id="49E81CA3CC72FFD4FE6E5FAAFD8262A4" box="[453,526,928,952]" country="India" name="Assam" pageId="37" pageNumber="172">Assam</collectingRegion>
, 
<collectingCountry id="F33B92D1CC72FFD4FDB75FAAFDDF62A4" box="[540,595,928,952]" name="India" pageId="37" pageNumber="172">India</collectingCountry>
, and two other localities: 
<collectingRegion id="49E81CA3CC72FFD4FCC65FAAFC7962A4" box="[877,1013,928,952]" country="India" name="West Bengal" pageId="37" pageNumber="172">West Bengal</collectingRegion>
, 
<collectingCountry id="F33B92D1CC72FFD4FBA85FAAFBB662A4" box="[1027,1082,928,952]" name="India" pageId="37" pageNumber="172">India</collectingCountry>
(
<bibRefCitation id="EFBDAFB0CC72FFD4FBE05FAAFF5E62C4" author="PAUL, S. &amp; M. C. BISWAS &amp; K. DEUTI" pageId="37" pageNumber="172" pagination="97 - 98" refId="ref38384" refString="PAUL, S., M. C. BISWAS, AND K. DEUTI. 2007. Kalophrynus orangensis (Orang Sticky Frog). Herpetological Review 38 (1): 97 - 98." type="journal article" year="2007">Paul et al. 2007</bibRefCitation>
); 
<location id="8EF3849ACC72FFD4FF405FCAFEF462C4" LSID="urn:lsid:plazi:treatment:03856357CC73FFD4FFD059F9FB1F6781:8EF3849ACC72FFD4FF405FCAFEF462C4" box="[235,376,960,984]" country="India" county="Brahmaputra River" latitude="26.5" longLatPrecision="1241" longitude="92.25" municipality="All" name="Mymensignh" pageId="37" pageNumber="172" stateProvince="Assam">Mymensignh</location>
, 
<collectingCountry id="F33B92D1CC72FFD4FE2C5FCAFD8F62C4" box="[391,515,960,984]" name="Bangladesh" pageId="37" pageNumber="172">Bangladesh</collectingCountry>
(Mahony and Ali Reza 2007). 
<collectingMunicipality id="6BF7483BCC72FFD4FCFC5FCAFCF562C4" box="[855,889,960,984]" pageId="37" pageNumber="172">All</collectingMunicipality>
three localities are within the lower middle portion of the 
<collectingCounty id="62F2AACDCC72FFD4FE795FEAFD1362E4" box="[466,671,992,1016]" pageId="37" pageNumber="172">Brahmaputra River</collectingCounty>
drainage.
</paragraph>
<paragraph id="8B93D241CC72FFD4FF00580BFF6A65A5" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">
<smallCapsWord id="8D75449DCC72FFD4FF00580BFE9D650B" baselines="1043,1042" box="[171,273,1025,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Natural" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FF00580BFE9D650B" bold="true" box="[171,273,1025,1049]" pageId="37" pageNumber="172">NATURAL</emphasis>
</smallCapsWord>
<smallCapsWord id="8D75449DCC72FFD4FEB3580BFEF5650B" baselines="1043,1042" box="[280,377,1025,1049]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="History" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FEB3580BFEF5650B" bold="true" box="[280,377,1025,1049]" pageId="37" pageNumber="172">HISTORY</emphasis>
</smallCapsWord>
. 
<emphasis id="B9580E53CC72FFD4FED4580BFE166505" bold="true" box="[383,410,1025,1049]" pageId="37" pageNumber="172">—</emphasis>
The individuals from the 
<typeStatus id="54976CE3CC72FFD4FD1E580BFD6F6505" box="[693,739,1025,1049]" pageId="37" pageNumber="172">type</typeStatus>
series were found on the ground [litter] in grassland adjacent to dry deciduous forest [secondary] or among trees at forest edging on grassland. Breeding occurred in this area in June with females depositing eggs in small puddles; males called beneath vegetation overhanging these puddles. The West Bengal individual was found 
<quantity id="4CD47FA4CC72FFD4FBFF586BFB1A6565" box="[1108,1174,1121,1145]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="172" unit="cm" value="60.0">60 cm</quantity>
high in a bush in a primary semi-evergreen forest. No habitat data are available for the 
<collectingCountry id="F33B92D1CC72FFD4FBB25888FB196586" box="[1049,1173,1154,1178]" name="Bangladesh" pageId="37" pageNumber="172">Bangladesh</collectingCountry>
specimen.
</paragraph>
</subSubSection>
<subSubSection id="C33681CACC72FFD4FF0058C8FB1F6781" pageId="37" pageNumber="172" type="discussion">
<paragraph id="8B93D241CC72FFD4FF0058C8FC1F65E6" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">
<smallCapsWord id="8D75449DCC72FFD4FF0058C8FEA465C4" baselines="1236,1235" box="[171,296,1218,1242]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Comments" pageId="37" pageNumber="172">
<emphasis id="B9580E53CC72FFD4FF0058C8FEA465C4" bold="true" box="[171,296,1218,1242]" pageId="37" pageNumber="172">COMMENTS</emphasis>
</smallCapsWord>
.— Preceding information extracted from 
<bibRefCitation id="EFBDAFB0CC72FFD4FD5958C9FC3165C6" author="DUTTA, S. K. &amp; AHMED M. F. &amp; I. DAS" box="[754,957,1218,1242]" pageId="37" pageNumber="172" pagination="67 - 74" refId="ref37371" refString="DUTTA, S. K., AHMED M. F., AND I. DAS 2000. Kalophrynus (Anura: Microhylidae), a new genus for India, with the description of a new species, Kalophrynus orangensis, from Assam State. Hamadryad 25 (2): 67 - 74." type="journal article" year="2000">Dutta et al. (2000)</bibRefCitation>
. Question marks in the Definition indicate my uncertainty on the accuracy of the proportions.
</paragraph>
<paragraph id="8B93D241CC72FFD4FF005909FB1F6781" blockId="37.[123,1175,219,1693]" pageId="37" pageNumber="172">
<taxonomicName id="4C2CA9C2CC72FFD4FF005909FECD6406" authorityName="Dutta, Ahmed, and Das" authorityYear="2000" box="[171,321,1283,1306]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="species" species="orangensis">
<emphasis id="B9580E53CC72FFD4FF005909FECD6406" box="[171,321,1283,1306]" italics="true" pageId="37" pageNumber="172">K. orangensis</emphasis>
</taxonomicName>
is recently described and is based on 
<specimenCount id="9D2A19C8CC72FFD4FD4D5908FCC16406" box="[742,845,1282,1306]" count="5" pageId="37" pageNumber="172" type="adult">five adult</specimenCount>
specimens, 
<specimenCount id="9D2A19C8CC72FFD4FC7E5908FBEA6406" box="[981,1126,1282,1306]" count="3" pageId="37" pageNumber="172" type="female">three females</specimenCount>
and 
<specimenCount id="9D2A19C8CC72FFD4FFD05929FF6B6427" box="[123,231,1315,1339]" count="2" pageId="37" pageNumber="172" type="male">two males</specimenCount>
. Because the variation in head striping and outer metatarsal tubercle size, the two major diagnostic features identified by 
<bibRefCitation id="EFBDAFB0CC72FFD4FE705949FD2E6447" author="DUTTA, S. K. &amp; AHMED M. F. &amp; I. DAS" box="[475,674,1347,1371]" pageId="37" pageNumber="172" pagination="67 - 74" refId="ref37371" refString="DUTTA, S. K., AHMED M. F., AND I. DAS 2000. Kalophrynus (Anura: Microhylidae), a new genus for India, with the description of a new species, Kalophrynus orangensis, from Assam State. Hamadryad 25 (2): 67 - 74." type="journal article" year="2000">Dutta et al. (2000)</bibRefCitation>
, were encompassed in a wider sample of Thai and Vietnamese 
<taxonomicName id="4C2CA9C2CC72FFD4FE81596EFD776467" authority=", Ohler and Grosjean (2005)" authorityName=", Ohler and Grosjean" authorityYear="2005" box="[298,763,1379,1403]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC72FFD4FE81596EFE5C6467" box="[298,464,1379,1403]" italics="true" pageId="37" pageNumber="172">K. interlineatus</emphasis>
, Ohler and Grosjean (2005)
</taxonomicName>
proposed that 
<collectingRegion id="49E81CA3CC72FFD4FC335969FC6D6467" box="[920,993,1379,1403]" country="India" name="Assam" pageId="37" pageNumber="172">Assam</collectingRegion>
and Indochinese specimens were conspecific; they thus considered 
<taxonomicName id="4C2CA9C2CC72FFD4FD37598EFCBE6487" authorityName="Dutta, Ahmed, and Das" authorityYear="2000" box="[668,818,1412,1435]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="species" species="orangensis">
<emphasis id="B9580E53CC72FFD4FD37598EFCBE6487" box="[668,818,1412,1435]" italics="true" pageId="37" pageNumber="172">K. orangensis</emphasis>
</taxonomicName>
a junior synonym of 
<taxonomicName id="4C2CA9C2CC72FFD4FBB6598EFF3F64A7" authorityName=", Ohler and Grosjean" authorityYear="2005" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="species" species="interlineatus">
<emphasis id="B9580E53CC72FFD4FBB6598EFF3F64A7" italics="true" pageId="37" pageNumber="172">K. interlineatus</emphasis>
</taxonomicName>
. Matsui and colleagues (2009, 2011, 2012) continued to recognize 
<taxonomicName id="4C2CA9C2CC72FFD4FC3B59AEFBA964A7" authorityName="Dutta, Ahmed, and Das" authorityYear="2000" box="[912,1061,1444,1467]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="species" species="orangensis">
<emphasis id="B9580E53CC72FFD4FC3B59AEFBA964A7" box="[912,1061,1444,1467]" italics="true" pageId="37" pageNumber="172">K. orangensis</emphasis>
</taxonomicName>
as a valid species. An additional difficulty in evaluating the status of this species is presumed errors in the measurements of the head of the 
<typeStatus id="54976CE3CC72FFD4FE4B59EFFD8264E1" box="[480,526,1509,1533]" pageId="37" pageNumber="172">type</typeStatus>
series (
<bibRefCitation id="EFBDAFB0CC72FFD4FDC959EFFC9964E0" author="DUTTA, S. K. &amp; AHMED M. F. &amp; I. DAS" box="[610,789,1508,1532]" pageId="37" pageNumber="172" pagination="67 - 74" refId="ref37371" refString="DUTTA, S. K., AHMED M. F., AND I. DAS 2000. Kalophrynus (Anura: Microhylidae), a new genus for India, with the description of a new species, Kalophrynus orangensis, from Assam State. Hamadryad 25 (2): 67 - 74." type="journal article" year="2000">Dutta et al. 2000</bibRefCitation>
: table 1; see comments in 
<emphasis id="B9580E53CC72FFD4FB9C59EEFF336700" italics="true" pageId="37" pageNumber="172">Morphometric</emphasis>
subsection of the above 
<emphasis id="B9580E53CC72FFD4FE625A0EFD696700" box="[457,741,1540,1564]" italics="true" pageId="37" pageNumber="172">Morphology and Variation</emphasis>
section). I have not examined any specimens of this population, and even though, this taxon is poorly characterized in its original description, I hesitate to consider it a synonym without a thorough comparison. Additionally, recent molecular analyses have demonstrated considerable genetic diversity in 
<taxonomicName id="4C2CA9C2CC72FFD4FC385A6FFB916761" authorityName="Tschudi" authorityYear="1838" box="[915,1053,1637,1661]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="37" pageNumber="172" phylum="Chordata" rank="genus">
<emphasis id="B9580E53CC72FFD4FC385A6FFB916761" box="[915,1053,1637,1661]" italics="true" pageId="37" pageNumber="172">Kalophrynus</emphasis>
</taxonomicName>
, hence the geographic isolation of the Indian populations likely will be matched by their genetic uniqueness.
</paragraph>
</subSubSection>
</treatment>
</document>